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<document id="6A8B7D142C70012B8CBB0803914B2BA2" ID-DOI="10.5252/zoosystema2022v44a22" ID-GBIF-Dataset="5d8905c9-7890-4f01-b4ee-f8bce829fa93" ID-ISSN="1638-9387" ID-Zenodo-Dep="7428311" ID-ZooBank="urn:lsid:zoobank.org:pub:27DA11CC-1A56-4E2A-BF0E-77F630407E7D" checkinTime="1670848154943" checkinUser="felipe" docAuthor="Webb, Michael D. &amp; Xu, Ye" docDate="2022" docId="03E1879FFF98FFE579DDFE1DFB686141" docLanguage="en" docName="Zoosystema.44.22.549-563.pdf" docOrigin="Zoosystema 44 (22)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.3:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="3" docTitle="Dayus takagii Dworakowska 1971" docType="treatment" docVersion="7" lastPageNumber="555" masterDocId="FFD8FFE7FF90FFED7A72FF8AFF94642C" masterDocTitle="Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific" masterLastPageNumber="563" masterPageNumber="549" pageNumber="555" updateTime="1698310541687" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="184270EE4BBE350520FDB94C4FC63E9F">Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific</mods:title>
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<mods:name id="EB3E85C61FEB6F457D7B26449D3F4200" type="personal">
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<mods:roleTerm id="4B725A005FC237D03DCA528A22457BCB">Author</mods:roleTerm>
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<mods:namePart id="F325329880207722E77130C3F96EBA20">Webb, Michael D.</mods:namePart>
<mods:affiliation id="38A518C34EF25A71D87E0A82F88D781E">Department of Life Sciences (Insects), The Natural History Museum, London, SW 7 5 BD (United Kingdom)</mods:affiliation>
<mods:nameIdentifier id="D9CC82BA531D17D65E610B299EB14F64" type="email">webb@nhm.ac.uk</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="0F742D8A95FE83539E18255E8E275C8B">Xu, Ye</mods:namePart>
<mods:affiliation id="1F7EA846530822B6E0BB8E277F4C1E0A">Institute of Entomology, Jiangxi Agricultural University, Nanchang 330045 (China)</mods:affiliation>
<mods:nameIdentifier id="22CFC8D648D0617CAC3A18259CEEC5C2" type="email">yexu@jxau.edu.cn</mods:nameIdentifier>
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<mods:title id="2B2F8AC438C9FA5E5101AFEBEBE1B3B5">Zoosystema</mods:title>
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<mods:part id="0D20600826953ED5E47C2D207A45172C">
<mods:date id="26183B4BBFCF1D08C14B728F7CD1335A">2022</mods:date>
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<mods:number id="DF6BAECB169D84CAE68FDF15DE9D4C15">2022-12-07</mods:number>
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<mods:number id="2CC30F511C431C7DE05E6558E9EE9490">44</mods:number>
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<mods:identifier id="3D9F2CBFBC616E4174A04C47CEE342FF" type="DOI">10.5252/zoosystema2022v44a22</mods:identifier>
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<treatment id="03E1879FFF98FFE579DDFE1DFB686141" ID-DOI="http://doi.org/10.5281/zenodo.7428354" ID-GBIF-Taxon="206232227" ID-Zenodo-Dep="7428354" LSID="urn:lsid:plazi:treatment:03E1879FFF98FFE579DDFE1DFB686141" httpUri="http://treatment.plazi.org/id/03E1879FFF98FFE579DDFE1DFB686141" lastPageNumber="555" pageId="8" pageNumber="555">
<subSubSection id="C3526502FF98FFE579DDFE1DFAB8659D" box="[943,1324,406,433]" pageId="8" pageNumber="555" type="nomenclature">
<paragraph id="8BF73689FF98FFE579DDFE1DFAB8659D" blockId="8.[943,1324,406,433]" box="[943,1324,406,433]" pageId="8" pageNumber="555">
<heading id="D0BF81E5FF98FFE579DDFE1DFAB8659D" box="[943,1324,406,433]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="555" reason="2">
<taxonomicName id="4C484D0AFF98FFE579DDFE1DFAB8659D" ID-CoL="34CDB" authority="Dworakowska, 1971" authorityName="Dworakowska" authorityYear="1971" box="[943,1324,406,433]" class="Insecta" family="Cicadellidae" genus="Dayus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="takagii">
<emphasis id="B93CEA9BFF98FFE579DDFE1DFBD2659D" bold="true" box="[943,1094,406,433]" italics="true" pageId="8" pageNumber="555">Dayus takagii</emphasis>
<bibRefCitation id="EFD94B78FF98FFE57E3FFE1CFAB8659D" author="Dworakowska" box="[1101,1324,406,433]" firstAuthor="Dworakowska" pageId="8" pageNumber="555" pagination="501 - 509" refId="ref9773" refString="DWORAKOWSKA I. 1971. - Dayus takagii sp. n. and some other Empoascini (Auchenorrhyncha, Cicadellidae, Typhlocybinae). Bulletin de l'Academie polonaise des Sciences. Serie des Sciences biologiques 19: 501 - 509." type="journal article" year="1971">Dworakowska, 1971</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3526502FF98FFE5795FFE5AFC0366AD" pageId="8" pageNumber="555" type="reference_group">
<paragraph id="8BF73689FF98FFE5795FFE5AFC2C662E" blockId="8.[813,1455,463,514]" pageId="8" pageNumber="555">
<treatmentCitationGroup id="AB5811A7FF98FFE5795FFE5AFC2C662E" pageId="8" pageNumber="555">
<taxonomicName id="4C484D0AFF98FFE5795FFE5AFC2E65C4" ID-CoL="8J422" authorityName="Melichar" authorityYear="1903" box="[813,954,464,488]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B93CEA9BFF98FFE5795FFE5AFC2E65C4" box="[813,954,464,488]" italics="true" pageId="8" pageNumber="555">Empoasca rufa</emphasis>
</taxonomicName>
<treatmentCitation id="0AE91098FF98FFE579A5FE5AFB3565CB" author="MATSUMURA S." box="[983,1185,463,488]" page="5" pageId="8" pageNumber="555" year="1934">
<bibRefCitation id="EFD94B78FF98FFE579A5FE5AFB3565CB" author="Matsumura" box="[983,1185,463,488]" firstAuthor="Matsumura" page="5" pageId="8" pageNumber="555" pagination="1 - 15" refId="ref10879" refString="MATSUMURA S. 1934. - Eupterygidae. Catalogue of Japanese Insects 3: 1 - 15." type="journal article" year="1934">Matsumura 1934: 5</bibRefCitation>
</treatmentCitation>
(nec
<bibRefCitation id="EFD94B78FF98FFE57EA9FE45FAE565C4" author="Melichar" box="[1243,1393,463,488]" firstAuthor="Melichar" pageId="8" pageNumber="555" refId="ref10954" refString="MELICHAR L. 1903. - Homopteren-Fauna von Ceylon. Berlin, Felix L. Dames. iv, 248 p., 6 pls." type="book" year="1903">Melichar 1903</bibRefCitation>
, misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF73689FF98FFE5795FFD90FC0366AD" blockId="8.[813,1455,537,641]" pageId="8" pageNumber="555">
<treatmentCitationGroup id="AB5811A7FF98FFE5795FFD90FC0366AD" pageId="8" pageNumber="555">
<treatmentCitation id="0AE91098FF98FFE5795FFD90FB2E661D" author="DWORAKOWSKA I." box="[813,1210,537,562]" page="501" pageId="8" pageNumber="555" year="1971">
<taxonomicName id="4C484D0AFF98FFE5795FFD90FB2E661D" ID-CoL="34CDB" authority="Dworakowska, 1971: 501" authorityName="Dworakowska" authorityPageNumber="501" authorityYear="1971" box="[813,1210,537,562]" class="Insecta" family="Cicadellidae" genus="Dayus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="takagii">
<emphasis id="B93CEA9BFF98FFE5795FFD90FC24661E" box="[813,944,538,562]" italics="true" pageId="8" pageNumber="555">Dayus takagii</emphasis>
<bibRefCitation id="EFD94B78FF98FFE579CAFD90FB2E661D" author="Dworakowska" box="[952,1210,537,562]" firstAuthor="Dworakowska" page="501" pageId="8" pageNumber="555" pagination="501 - 509" refId="ref9773" refString="DWORAKOWSKA I. 1971. - Dayus takagii sp. n. and some other Empoascini (Auchenorrhyncha, Cicadellidae, Typhlocybinae). Bulletin de l'Academie polonaise des Sciences. Serie des Sciences biologiques 19: 501 - 509." type="journal article" year="1971">Dworakowska, 1971: 501</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1-11. —
<treatmentCitation id="0AE91098FF98FFE57F3EFD90FC166660" author="LEE C. E." page="565" pageId="8" pageNumber="555" year="1979">
<bibRefCitation id="EFD94B78FF98FFE57F3EFD90FC166660" author="Lee" firstAuthor="Lee" page="565 - 566" pageId="8" pageNumber="555" refId="ref10665" refString="LEE C. E. 1979. - Illustrated flora and fauna of Korea, Vol. 23 Insecta (VII). Seoul, Samhwa Publishing Co. Ltd. 1070 p. (In Korean)." type="book" year="1979">Lee 1979: 565-566</bibRefCitation>
</treatmentCitation>
, plate 48, fig. 264, figs 528-530 (
<collectingCountry id="F35F7619FF98FFE57EC9FDBEFB626660" box="[1211,1270,564,588]" name="South Korea" pageId="8" pageNumber="555">Korea</collectingCountry>
). —
<treatmentCitation id="0AE91098FF98FFE57F57FDBEFC12664A" author="DWORAKOWSKA I." page="54" pageId="8" pageNumber="555" year="1982">
<bibRefCitation id="EFD94B78FF98FFE57F57FDBEFC12664A" author="Dworakowska" firstAuthor="Dworakowska" page="54" pageId="8" pageNumber="555" pagination="33 - 57" refId="ref9853" refString="DWORAKOWSKA I. 1982. - Empoascini of Japan, Korea and Northeast part of China (Homoptera, Auchenorrhyncha, Cicadellidae, Typhlocybinae). Reichenbachia 20: 33 - 57." type="journal article" year="1982">Dworakowska 1982: 54</bibRefCitation>
</treatmentCitation>
, figs 282-288 (
<collectingCountry id="F35F7619FF98FFE57E61FDC5FBC4664B" box="[1043,1104,591,615]" name="Japan" pageId="8" pageNumber="555">Japan</collectingCountry>
). —
<treatmentCitation id="0AE91098FF98FFE57E0FFDC4FAF4664B" author="QIN D. Z. &amp; ZHANG Y. L." box="[1149,1376,590,615]" httpUri="http://treatment.plazi.org/id/2F5D87EBFF96924FFF13018A34EEF815" page="48" pageId="8" pageNumber="555" year="2007">
<bibRefCitation id="EFD94B78FF98FFE57E0FFDC4FAF4664B" author="Qin &amp; Zhang" box="[1149,1376,590,615]" firstAuthor="Qin" page="48" pageId="8" pageNumber="555" pagination="43 - 51" refId="ref11087" refString="QIN D. Z. &amp; ZHANG Y. L. 2007. - Revision of the Chinese species of the genus Dayus Mahmood (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini), with description of three new species. Zootaxa 1624 (1): 43 - 51. https: // doi. org / 10.11646 / zootaxa. 1624.1.4" type="journal article" year="2007">Qin &amp; Zhang 2007: 48</bibRefCitation>
</treatmentCitation>
, figs 32- 40,
<collectingCountry id="F35F7619FF98FFE57921FDE3FC0766AD" box="[851,915,617,641]" name="China" pageId="8" pageNumber="555">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3526502FF98FFE5795FFD13FC3966E7" pageId="8" pageNumber="555" type="distribution">
<paragraph id="8BF73689FF98FFE5795FFD13FC3966E7" blockId="8.[813,1455,665,715]" pageId="8" pageNumber="555">
DISTRIBUTION. —
<collectingCountry id="F35F7619FF98FFE5799CFD13FBBD669C" box="[1006,1065,665,688]" name="Japan" pageId="8" pageNumber="555">Japan</collectingCountry>
,
<collectingCountry id="F35F7619FF98FFE57E47FD13FBE7669C" box="[1077,1139,665,688]" name="South Korea" pageId="8" pageNumber="555">Korea</collectingCountry>
,
<collectingCountry id="F35F7619FF98FFE57E0DFD13FB54669D" box="[1151,1216,665,689]" name="China" pageId="8" pageNumber="555">China</collectingCountry>
,
<collectingCountry id="F35F7619FF98FFE57EBEFD13FADC669D" box="[1228,1352,665,689]" name="Hong Kong" pageId="8" pageNumber="555">Hong Kong</collectingCountry>
(?) see Remarks below.
</paragraph>
</subSubSection>
<subSubSection id="C3526502FF98FFE5795FFD69FCE56739" pageId="8" pageNumber="555" type="materials_examined">
<paragraph id="8BF73689FF98FFE5795FFD69FCE56739" blockId="8.[811,1455,738,789]" pageId="8" pageNumber="555">
MATERIAL EXAMINED. —
<materialsCitation id="3B203CD4FF98FFE57E6CFD69FCF96739" ID-GBIF-Occurrence="3990056320" collectingDate="1958-01-10" collectionCode="NHM" location="Kyushu" pageId="8" pageNumber="555" specimenCount="1" specimenCount-female="1" stateProvince="Fukuoka">
<emphasis id="B93CEA9BFF98FFE57E6CFD69FBC266D7" bold="true" box="[1054,1110,739,763]" pageId="8" pageNumber="555">
<collectingCountry id="F35F7619FF98FFE57E6CFD69FBC266D7" box="[1054,1110,739,763]" name="Japan" pageId="8" pageNumber="555">Japan</collectingCountry>
</emphasis>
<specimenCount id="9D4EFD00FF98FFE57E1AFD69FB1D66D6" box="[1128,1161,739,763]" pageId="8" pageNumber="555" type="female">1 ♀</specimenCount>
;
<location id="8E976052FF98FFE57EE6FD69FB7566D7" LSID="urn:lsid:plazi:treatment:03E1879FFF98FFE579DDFE1DFB686141:8E976052FF98FFE57EE6FD69FB7566D7" box="[1172,1249,739,763]" name="Kyushu" pageId="8" pageNumber="555" stateProvince="Fukuoka">Kyushu</location>
;
<collectingRegion id="498CF86BFF98FFE57E9BFD69FAD566D7" box="[1257,1345,739,763]" country="Japan" name="Fukuoka" pageId="8" pageNumber="555">Fukuoka</collectingRegion>
;
<date id="FFF61049FF98FFE57F3BFD68FA3F66D6" box="[1353,1451,738,762]" pageId="8" pageNumber="555" value="1958-01-10">
<collectingDate id="EFB2E9A1FF98FFE57F3BFD68FA3F66D6" box="[1353,1451,738,762]" pageId="8" pageNumber="555" value="1958-01-10">10.I.1958</collectingDate>
</date>
;
<collectionCode id="ED59AE4CFF98FFE57959FD74FCF96739" box="[811,877,766,789]" pageId="8" pageNumber="555">NHM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526502FF98FFE5795FFCBCFB686141" pageId="8" pageNumber="555" type="discussion">
<paragraph id="8BF73689FF98FFE5795FFCBCFC056762" blockId="8.[811,1456,822,1389]" box="[813,913,822,848]" pageId="8" pageNumber="555">REMARKS</paragraph>
<paragraph id="8BF73689FF98FFE5795FFCDFFB686141" blockId="8.[811,1456,822,1389]" pageId="8" pageNumber="555">
This species was described from the
<typeStatus id="54F3882BFF98FFE57ED0FCDFFA946743" box="[1186,1280,853,879]" pageId="8" pageNumber="555">holotype</typeStatus>
male and several
<typeStatus id="54F3882BFF98FFE57959FCFFFC0467A3" box="[811,912,885,911]" pageId="8" pageNumber="555">paratypes</typeStatus>
(male and female) from Kyushu (
<collectingCountry id="F35F7619FF98FFE57F76FCFFFAD067A3" box="[1284,1348,885,911]" name="Japan" pageId="8" pageNumber="555">Japan</collectingCountry>
) and
<specimenCount id="9D4EFD00FF98FFE57FF6FCFFFCE06783" pageId="8" pageNumber="555" type="female">two female</specimenCount>
<typeStatus id="54F3882BFF98FFE57908FC1FFC4B6783" box="[890,991,917,943]" pageId="8" pageNumber="555">paratypes</typeStatus>
from
<collectingCountry id="F35F7619FF98FFE57E52FC1FFB316783" box="[1056,1189,917,943]" name="Hong Kong" pageId="8" pageNumber="555">Hong Kong</collectingCountry>
. Although no depository for the types is indicated in the original description, it states in the acknowledgements to the paper that the type series was borrowed from EIHU, and this is presumed to be Matsumuras collection. Therefore, Matsumuras misidentification given in the original description (see above) refers to
<bibRefCitation id="EFD94B78FF98FFE57F41FBBFFCE26042" author="Matsumura" firstAuthor="Matsumura" pageId="8" pageNumber="555" pagination="1 - 15" refId="ref10879" refString="MATSUMURA S. 1934. - Eupterygidae. Catalogue of Japanese Insects 3: 1 - 15." type="journal article" year="1934">Matsumura (1934)</bibRefCitation>
where
<taxonomicName id="4C484D0AFF98FFE579CDFBDFFB226042" authority="Melichar" authorityName="Melichar" authorityYear="1903" box="[959,1206,1108,1134]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B93CEA9BFF98FFE579CDFBDFFBC76042" box="[959,1107,1108,1134]" italics="true" pageId="8" pageNumber="555">Empoasca rufa</emphasis>
Melichar
</taxonomicName>
is listed from
<collectingCountry id="F35F7619FF98FFE57F34FBDFFA166043" box="[1350,1410,1109,1135]" name="Japan" pageId="8" pageNumber="555">Japan</collectingCountry>
and not (as stated)
<bibRefCitation id="EFD94B78FF98FFE579B7FBFFFB0560A2" author="Matsumura" box="[965,1169,1140,1167]" firstAuthor="Matsumura" pageId="8" pageNumber="555" pagination="55 - 91" refId="ref10793" refString="MATSUMURA S. 1931. - A revision of the Palaearctic and Oriental Typhlocybid genera, with descriptions of new species and new genera. Insecta Matsumurana 6: 55 - 91, pls ii-iii, figs 1 - 6. http: // hdl. handle. net / 2115 / 9233" type="journal article" year="1931">Matsumura (1931)</bibRefCitation>
where it is listed from
<collectingCountry id="F35F7619FF98FFE57F0DFBFFFCFD6082" name="Taiwan" pageId="8" pageNumber="555">Formosa</collectingCountry>
. The female
<typeStatus id="54F3882BFF98FFE57982FB1EFBC16082" box="[1008,1109,1172,1198]" pageId="8" pageNumber="555">paratypes</typeStatus>
from
<collectingCountry id="F35F7619FF98FFE57EE5FB1EFA8C6082" box="[1175,1304,1172,1198]" name="Hong Kong" pageId="8" pageNumber="555">Hong Kong</collectingCountry>
could equally be
<taxonomicName id="4C484D0AFF98FFE5793EFB3FFC5360E2" authorityName="Dworakowska &amp; Viraktamath" authorityYear="1978" box="[844,967,1204,1230]" class="Insecta" family="Cicadellidae" genus="Dayus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B93CEA9BFF98FFE5793EFB3FFCF560E2" box="[844,865,1205,1230]" italics="true" pageId="8" pageNumber="555">D</emphasis>
.
<emphasis id="B93CEA9BFF98FFE5791CFB3EFC5360E2" box="[878,967,1204,1230]" italics="true" pageId="8" pageNumber="555">formosus</emphasis>
</taxonomicName>
examined here from a male from
<collectingCountry id="F35F7619FF98FFE57F5DFB3EFA3B60E2" box="[1327,1455,1204,1230]" name="Hong Kong" pageId="8" pageNumber="555">Hong Kong</collectingCountry>
and hence the question mark under Distribution. The colour of
<taxonomicName id="4C484D0AFF98FFE5793AFB7EFC256121" authorityName="Dworakowska" authorityYear="1971" box="[840,945,1267,1294]" class="Insecta" family="Cicadellidae" genus="Dayus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="takagii">
<emphasis id="B93CEA9BFF98FFE5793AFB7EFCC96121" box="[840,861,1268,1293]" italics="true" pageId="8" pageNumber="555">D</emphasis>
.
<emphasis id="B93CEA9BFF98FFE5791BFB79FC256121" box="[873,945,1267,1293]" italics="true" pageId="8" pageNumber="555">takagii</emphasis>
</taxonomicName>
was not originally described except that the species resembled
<taxonomicName id="4C484D0AFF98FFE579B9FA9EFBF86101" authorityName="d'Australie" baseAuthorityName="Kirkaldy" baseAuthorityYear="1907" box="[971,1132,1299,1326]" class="Insecta" family="Cicadellidae" genus="Dayus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="555" phylum="Arthropoda" rank="species" species="euryphaessus">
<emphasis id="B93CEA9BFF98FFE579B9FA9EFC746101" box="[971,992,1300,1325]" italics="true" pageId="8" pageNumber="555">D</emphasis>
.
<emphasis id="B93CEA9BFF98FFE5799EFA99FBF86101" box="[1004,1132,1299,1325]" italics="true" pageId="8" pageNumber="555">euryphaessus</emphasis>
</taxonomicName>
but with paler coloration. The female tentatively identified here as this species (based on its locality
<collectingCountry id="F35F7619FF98FFE579F0FAD9FC576141" box="[898,963,1363,1389]" name="Japan" pageId="8" pageNumber="555">Japan</collectingCountry>
), is mainly reddish in colour.
</paragraph>
</subSubSection>
</treatment>
</document>