treatments-xml/data/03/DF/BE/03DFBE05E00EFFE0FF64D70DFAE4FE78.xml
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<document id="DA2B43472F52B88DE78C70D4E3131CD6" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. &amp; Colville, Jonathan F." docDate="2020" docId="03DFBE05E00EFFE0FF64D70DFAE4FE78" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia pallidibrunnea Dombrow 2020, new species" docType="treatment" docUuid="725E4997-D1E7-4D08-A3FF-95E8A92913D0" docUuidSource="ZooBank" docVersion="9" lastPageNumber="22" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="19" updateTime="1698849823920" updateUser="ExternalLinkService">
<mods:mods id="3C7593DA1E0754B0419BF07823798047" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C064076284B60105DA0C2E16C29B63A8">
<mods:title id="E4080C709AA6D1BC5791A7D398F7FA54">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
</mods:titleInfo>
<mods:name id="F0A04E3DBF1E509979D122F48EBBB145" type="personal">
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</mods:role>
<mods:namePart id="362AE388BB13471AA48FD7255CAF0689">Dombrow, Holger E.</mods:namePart>
<mods:nameIdentifier id="AD7ACAAA3BDEB9B77606173CEE45ED60" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="0CA54FD1B31EA24FFCD8B61C8D8A602C">Colville, Jonathan F.</mods:namePart>
<mods:nameIdentifier id="4A79EA773C0737AB10740E47CB045818" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
<mods:nameIdentifier id="44DF6C7F51B09158CFAFFB6336A74412" type="email">jonathan.colville@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo id="D7B718953872D5FE2FA9512C647A3614">
<mods:title id="859DBA041B0E2F3CB6C61560E3EE26C0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C51F925BE6FEC9B61F4B4933A9D9DFCC">
<mods:date id="8FEA5664DAD58314B09293C6DB4A5A86">2020</mods:date>
<mods:detail id="52F6223979EAE850B6B4772B73C7B016" type="pubDate">
<mods:number id="3B3F262CF8A42D28669A019E02454962">2020-08-10</mods:number>
</mods:detail>
<mods:detail id="E221ACD0D0704AD279ED37E620CE4D4B" type="volume">
<mods:number id="30C2A0ADACB2254A44E0057153C78FCB">4823</mods:number>
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<mods:start id="094DF64D24B1A12293820FBA8CA2FC23">1</mods:start>
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<mods:classification id="C57FFDEDA411313277EC843A5775AB95">journal article</mods:classification>
<mods:identifier id="D292B3AC4E727C60820DE4FD3F135AE2" type="CLB-Dataset">8721</mods:identifier>
<mods:identifier id="B8097A6B839E6CA5D77419AC79225DFB" type="DOI">10.11646/zootaxa.4823.1.1</mods:identifier>
<mods:identifier id="2659DE7DB852F35762D98064AEDB1E73" type="GBIF-Dataset">a43d0118-9df5-4970-a316-0dae5c9caebf</mods:identifier>
<mods:identifier id="D72E094B8D6B64D9606DD0DD4ADBC9C4" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03DFBE05E00EFFE0FF64D70DFAE4FE78" ID-DOI="http://doi.org/10.5281/zenodo.4455275" ID-GBIF-Taxon="177013529" ID-Zenodo-Dep="4455275" LSID="urn:lsid:zoobank.org:act:725E4997-D1E7-4D08-A3FF-95E8A92913D0" httpUri="http://treatment.plazi.org/id/03DFBE05E00EFFE0FF64D70DFAE4FE78" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19">
<subSubSection id="C36C5C98E00EFFE7FF64D70DFDB2FCDE" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BC90F13E00EFFE7FF64D70DFD41FCBD" blockId="18.[151,858,790,886]" box="[151,764,790,817]" pageId="18" pageNumber="19">
<heading id="D081B87FE00EFFE7FF64D70DFD41FCBD" bold="true" box="[151,764,790,817]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="B902D301E00EFFE7FF64D70DFD41FCBD" bold="true" box="[151,764,790,817]" pageId="18" pageNumber="19">
<taxonomicName id="4C767490E00EFFE7FF64D70DFDE2FCBC" authority="Dombrow" authorityName="Dombrow" authorityYear="2020" box="[151,607,790,817]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea" status="sp. nov.">
<emphasis id="B902D301E00EFFE7FF64D70DFE67FCBC" bold="true" box="[151,474,790,817]" italics="true" pageId="18" pageNumber="19">Beckhoplia pallidibrunnea</emphasis>
Dombrow
</taxonomicName>
,
<taxonomicNameLabel id="A2316E7AE00EFFE7FD9FD70CFD41FCBD" box="[620,764,790,817]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC90F13E00EFFE7FF64D722FDB2FCDE" blockId="18.[151,858,790,886]" box="[151,527,825,852]" pageId="18" pageNumber="19">
(
<figureCitation id="134D1396E00EFFE7FF6CD722FF40FCDE" box="[159,253,825,851]" captionStart="FIGURES 3638" captionStartId="23.[151,275,402,428]" captionTargetBox="[304,1284,185,355]" captionTargetId="figure-17@23.[301,1287,181,358]" captionText="FIGURES 3638. General drawings of the pygidium of Beckhoplia Dombrow, 2005 species, in posterior view ♂♂. 36, upper margin moderately convex, linear medially, lateral margin subparallel in upper fifth, linearly attenuate apically; 37, upper margin slightly convex, lateral margin linearly attenuate apically; 38, upper margin moderately convex, linear medially, lateral margin slightly convex attenuate apically. Scale bar 1 mm" figureDoi="http://doi.org/10.5281/zenodo.4455243" httpUri="https://zenodo.org/record/4455243/files/figure.png" pageId="18" pageNumber="19">Figs. 38</figureCitation>
, 52, 57, 61, 6465, 101)
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E00EFFE7FF64D745FCE7FCFB" box="[151,858,862,886]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BC90F13E00EFFE7FF64D745FCE7FCFB" blockId="18.[151,858,790,886]" box="[151,858,862,886]" pageId="18" pageNumber="19">
http://zoobank.org/
<uuid id="FFD035C6E00EFFE7FE9BD745FCE7FCFB" box="[360,858,862,886]" pageId="18" pageNumber="19">725E4997-D1E7-4D08-A3FF-95E8A92913D0</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E00EFFE7FF64D7B8FA31FA0D" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BC90F13E00EFFE7FF64D7B8FBDAFC30" blockId="18.[151,1127,930,958]" box="[151,1127,930,958]" pageId="18" pageNumber="19">
<materialsCitation id="3B1E054EE00EFFE7FF64D7B8FBDEFC30" ID-GBIF-Occurrence="3022772301" box="[151,1123,930,958]" collectionCode="NCP" country="South Africa" location="RSA" pageId="18" pageNumber="19" specimenCount="1" typeStatus="holotype">
<emphasis id="B902D301E00EFFE7FF64D7B8FE8BFC30" bold="true" box="[151,310,931,957]" pageId="18" pageNumber="19">
<typeStatus id="54CDB1B1E00EFFE7FF64D7B8FF6FFC30" box="[151,210,931,957]" pageId="18" pageNumber="19">Type</typeStatus>
locality.
</emphasis>
[
<collectingCountry id="F3614F83E00EFFE7FEB0D7B8FEC2FC30" box="[323,383,931,957]" name="South Africa" pageId="18" pageNumber="19">RSA</collectingCountry>
,
<collectionCode id="ED6797D6E00EFFE7FE79D7B9FE7AFC30" box="[394,455,930,957]" country="United Kingdom" name="National Collection of Passiflora" pageId="18" pageNumber="19" type="Herbarium">NCP</collectionCode>
]: Farm Glenlyon [now Hantam National Botanical Garden]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC90F13E00EFFE7FF64D7F3FCABFAB7" blockId="18.[151,1437,1000,1339]" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF64D7F3FEFAFB8F" bold="true" box="[151,327,1000,1026]" pageId="18" pageNumber="19">Type material.</emphasis>
[20]
<emphasis id="B902D301E00EFFE7FE75D7F3FE4EFB8F" bold="true" box="[390,499,1000,1026]" pageId="18" pageNumber="19">
<typeStatus id="54CDB1B1E00EFFE7FE75D7F3FE4EFB8F" box="[390,499,1000,1026]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE00EFFE7FE0FD7F2FDA9FB8E" box="[508,532,1001,1027]" pageId="18" pageNumber="19" type="male"></specimenCount>
: yellow label hw «30» pr «.276 | coll. Dombrow» ||white label pr «
<collectingCountry id="F3614F83E00EFFE7FAE6D7F3FE9BFBA8" name="South Africa" pageId="18" pageNumber="19">Republic of South Africa</collectingCountry>
|
<collectionCode id="ED6797D6E00EFFE7FEC9D010FED9FBA8" box="[314,356,1035,1061]" pageId="18" pageNumber="19">NC</collectionCode>
, Bokkeveldberge | Farm Glenlyon | 31.23.841S19.08.397E» bs pr «altitude
<quantity id="4C8EA2F6E00EFFE7FB30D010FAB7FBAB" box="[1219,1290,1035,1062]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="18" pageNumber="19" unit="m" value="732.0">732 m</quantity>
|
<date id="FFC829D3E00EFFE7FAEDD010FA21FBA8" box="[1310,1436,1035,1061]" pageId="18" pageNumber="19" value="2008-04-09">04.09.2008</date>
| #522 | leg.: H. &amp; E. Dombrow» || green label pr «Host plant: |
<taxonomicName id="4C767490E00EFFE7FC81D035FB89FBCA" authorityName="Sweet" authorityYear="1826" baseAuthorityName="Andrews" box="[882,1076,1070,1095]" class="Liliopsida" family="Iridaceae" genus="Homeria" higherTaxonomySource="CoL" kingdom="Plantae" order="Asparagales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="miniata">
<emphasis id="B902D301E00EFFE7FC81D035FB89FBCA" box="[882,1076,1070,1095]" italics="true" pageId="18" pageNumber="19">Homeria miniata</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E00EFFE7FBB9D035FB46FBC5" box="[1098,1275,1070,1096]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="acaulis">
<emphasis id="B902D301E00EFFE7FBB9D035FB46FBC5" box="[1098,1275,1070,1096]" italics="true" pageId="18" pageNumber="19">Arctotis acaulis</emphasis>
</taxonomicName>
» || blue label pr «Bioregion: | Shale Renosterveld | Vegetation Type: FRs2 | Nieuwoudtville Shale Renosterveld» || yellow label pr «SAMCOL | A071227» || red label pr «
<typeStatus id="54CDB1B1E00EFFE7FD32D068FC96FB00" box="[705,811,1139,1165]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
|
<taxonomicName id="4C767490E00EFFE7FCB2D068FC02FB00" authorityName="Dombrow" authorityYear="2005" box="[833,959,1139,1165]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E00EFFE7FCB2D068FC02FB00" box="[833,959,1139,1165]" italics="true" pageId="18" pageNumber="19">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E00EFFE7FC26D068FBC6FB00" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[981,1147,1139,1165]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea" status="sp. nov.">
<emphasis id="B902D301E00EFFE7FC26D068FBC6FB00" box="[981,1147,1139,1165]" italics="true" pageId="18" pageNumber="19">pallidibrunnea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE00EFFE7FB70D06FFB05FB03" box="[1155,1208,1140,1166]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E00EFFE7FF1ED08DFEFEFB3D" box="[237,323,1174,1200]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="18" pageNumber="19" type="Museum">SAMC</collectionCode>
).
<emphasis id="B902D301E00EFFE7FE98D08DFE5EFB3D" bold="true" box="[363,483,1174,1200]" pageId="18" pageNumber="19">
<typeStatus id="54CDB1B1E00EFFE7FE98D08DFE5EFB3D" box="[363,483,1174,1200]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE00EFFE7FE19D08DFDAAFB3D" box="[490,535,1174,1200]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D70C49AE00EFFE7FDD7D08DFDD8FB3D" box="[548,613,1174,1200]" pageId="18" pageNumber="19" type="female">2 ♀♀</specimenCount>
: same labels as
<typeStatus id="54CDB1B1E00EFFE7FCEFD08DFCC3FB3D" box="[796,894,1174,1200]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
but || yellow label hw «30» pr «.266,.280,.287 | coll. Dombrow» || yellow label pr «SAMCOL | A071227» || red label pr «
<typeStatus id="54CDB1B1E00EFFE7FC18D0A2FBF2FB5E" box="[1003,1103,1209,1235]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
|
<taxonomicName id="4C767490E00EFFE7FB97D0A3FB5FFB5F" authorityName="Dombrow" authorityYear="2005" box="[1124,1250,1208,1234]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E00EFFE7FB97D0A3FB5FFB5F" box="[1124,1250,1208,1234]" italics="true" pageId="18" pageNumber="19">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E00EFFE7FB05D0A3FA21FB5F" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1270,1436,1208,1234]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea" status="sp. nov.">
<emphasis id="B902D301E00EFFE7FB05D0A3FA21FB5F" box="[1270,1436,1208,1234]" italics="true" pageId="18" pageNumber="19">pallidibrunnea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE00EFFE7FF64D0C7FF71FB7B" box="[151,204,1244,1270]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E00EFFE7FDE1D0C0FDDAFB78" box="[530,615,1243,1269]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="18" pageNumber="19" type="Museum">SAMC</collectionCode>
);
<specimenCount id="9D70C49AE00EFFE7FD8ED0C0FD73FB7B" box="[637,718,1243,1270]" pageId="18" pageNumber="19" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D70C49AE00EFFE7FD28D0C0FCA3FB7B" box="[731,798,1243,1270]" pageId="18" pageNumber="19" type="female">6 ♀♀</specimenCount>
: same labels as
<typeStatus id="54CDB1B1E00EFFE7FC2AD0C0FB86FB78" box="[985,1083,1243,1269]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
but || yellow label hw «30» pr «.267.268;.270.271;.273.275,.277.279,.281.286 | coll. Dombrow» || red label pr «
<typeStatus id="54CDB1B1E00EFFE7FB65D0E5FB47FA95" box="[1174,1274,1278,1304]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
|
<taxonomicName id="4C767490E00EFFE7FAE3D0E5FA33FA95" authorityName="Dombrow" authorityYear="2005" box="[1296,1422,1278,1304]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E00EFFE7FAE3D0E5FA33FA95" box="[1296,1422,1278,1304]" italics="true" pageId="18" pageNumber="19">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E00EFFE7FF64D13BFE80FAB7" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[151,317,1312,1338]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea" status="sp. nov.">
<emphasis id="B902D301E00EFFE7FF64D13BFE80FAB7" box="[151,317,1312,1338]" italics="true" pageId="18" pageNumber="19">pallidibrunnea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE00EFFE7FEB6D13AFEC7FAB6" box="[325,378,1313,1339]" pageId="18" pageNumber="19" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E00EFFE7FD46D13BFCB6FAB7" box="[693,779,1312,1338]" pageId="18" pageNumber="19">ESDW</collectionCode>
).
</paragraph>
<paragraph id="8BC90F13E00EFFE7FF64D17EFA31FA0D" blockId="18.[151,1420,1381,1408]" box="[151,1420,1381,1408]" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF64D17EFE20FA0D" bold="true" box="[151,413,1381,1408]" pageId="18" pageNumber="19">
Condition of
<typeStatus id="54CDB1B1E00EFFE7FEC1D17DFE24FA0D" box="[306,409,1382,1408]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
.
</emphasis>
Very good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E00EFFE7FF64D1B0FC72F9FF" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BC90F13E00EFFE7FF64D1B0FC72F9FF" blockId="18.[151,1437,1451,1651]" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF64D1B0FE20FA48" bold="true" box="[151,413,1451,1477]" pageId="18" pageNumber="19">Differential diagnosis.</emphasis>
Differs from
<taxonomicName id="4C767490E00EFFE7FDC6D1B7FCB2FA48" authority=": Scutellum" authorityName="Scutellum" box="[565,783,1451,1478]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B902D301E00EFFE7FDC6D1B7FD32FA48" box="[565,655,1451,1477]" italics="true" pageId="18" pageNumber="19">B. fusca</emphasis>
: Scutellum
</taxonomicName>
of
<taxonomicName id="4C767490E00EFFE7FCC1D1B7FC4AFA48" authorityName="Dombrow" authorityYear="2020" box="[818,1015,1451,1477]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea">
<emphasis id="B902D301E00EFFE7FCC1D1B7FC4AFA48" box="[818,1015,1451,1477]" italics="true" pageId="18" pageNumber="19">B. pallidibrunnea</emphasis>
</taxonomicName>
as broad as long.
<taxonomicName id="4C767490E00EFFE7FB4DD1B0FA8DFA48" box="[1214,1328,1451,1477]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="18" pageNumber="70" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
of
<taxonomicName id="4C767490E00EFFE7FAA7D1B7FF0CFA6A" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B902D301E00EFFE7FAA7D1B7FF0CFA6A" italics="true" pageId="18" pageNumber="19">B. fusca</emphasis>
</taxonomicName>
longer than broad. Pretarsomere of metathoracic tarsus of
<taxonomicName id="4C767490E00EFFE7FCB5D1D5FBB1FA65" authorityName="Dombrow" authorityYear="2020" box="[838,1036,1486,1512]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea">
<emphasis id="B902D301E00EFFE7FCB5D1D5FBB1FA65" box="[838,1036,1486,1512]" italics="true" pageId="18" pageNumber="19">B. pallidibrunnea</emphasis>
</taxonomicName>
slightly thickened. Pretarsomere of metathoracic tarsus of
<taxonomicName id="4C767490E00EFFE7FE61D1EAFE51F987" box="[402,492,1520,1546]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B902D301E00EFFE7FE61D1EAFE51F987" box="[402,492,1520,1546]" italics="true" pageId="18" pageNumber="19">B. fusca</emphasis>
</taxonomicName>
slender. Interno-posterior margin of parameres of
<taxonomicName id="4C767490E00EFFE7FBE9D1EAFB63F987" authorityName="Dombrow" authorityYear="2020" box="[1050,1246,1520,1546]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea">
<emphasis id="B902D301E00EFFE7FBE9D1EAFB63F987" box="[1050,1246,1520,1546]" italics="true" pageId="18" pageNumber="19">B. pallidibrunnea</emphasis>
</taxonomicName>
parabola-shaped (
<figureCitation id="134D1396E00EFFE7FF6CD208FF43F9A0" box="[159,254,1555,1581]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="18" pageNumber="19">Figs. 64</figureCitation>
,
<figureCitation id="134D1396E00EFFE7FEF8D208FE85F9A0" box="[267,312,1555,1581]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="18" pageNumber="19">101</figureCitation>
). Interno-posterior margin of parameres of
<taxonomicName id="4C767490E00EFFE7FCD6D20FFC3DF9A0" box="[805,896,1555,1581]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B902D301E00EFFE7FCD6D20FFC3DF9A0" box="[805,896,1555,1581]" italics="true" pageId="18" pageNumber="19">B. fusca</emphasis>
</taxonomicName>
narrowed curly bracket-shaped (
<figureCitation id="134D1396E00EFFE7FB05D208FAE8F9A0" box="[1270,1365,1555,1581]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="18" pageNumber="19">Figs. 66</figureCitation>
,
<figureCitation id="134D1396E00EFFE7FA91D208FA2DF9A0" box="[1378,1424,1555,1581]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="18" pageNumber="19">100</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E00EFFE7FD8FD22DFCFEF9DD" authorityName="Dombrow" authorityYear="2020" box="[636,835,1590,1616]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pallidibrunnea">
<emphasis id="B902D301E00EFFE7FD8FD22DFCFEF9DD" box="[636,835,1590,1616]" italics="true" pageId="18" pageNumber="19">B. pallidibrunnea</emphasis>
</taxonomicName>
concave medially (
<figureCitation id="134D1396E00EFFE7FBD6D22DFB38F9DD" box="[1061,1157,1590,1616]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="18" pageNumber="19">Figs. 64</figureCitation>
,
<figureCitation id="134D1396E00EFFE7FB60D22DFB7DF9DD" box="[1171,1216,1590,1616]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="18" pageNumber="19">101</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E00EFFE7FE57D242FE43F9FF" box="[420,510,1624,1650]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="B902D301E00EFFE7FE57D242FE43F9FF" box="[420,510,1624,1650]" italics="true" pageId="18" pageNumber="19">B. fusca</emphasis>
</taxonomicName>
convex, incised medially (
<figureCitation id="134D1396E00EFFE7FCD8D243FC34F9FF" box="[811,905,1624,1650]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="18" pageNumber="19">Figs. 66</figureCitation>
,
<figureCitation id="134D1396E00EFFE7FC66D243FC7EF9FF" box="[917,963,1624,1650]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="18" pageNumber="19">100</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E00EFFE0FF64D286FCECFF58" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19" type="description">
<paragraph id="8BC90F13E00EFFE7FF64D286FE9CF93A" blockId="18.[151,1437,1693,2032]" box="[151,289,1693,1719]" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF64D286FE9CF93A" bold="true" box="[151,289,1693,1719]" pageId="18" pageNumber="19">Description</emphasis>
</paragraph>
<paragraph id="8BC90F13E00EFFE7FF34D2DBFC83F956" blockId="18.[151,1437,1693,2032]" box="[199,830,1728,1755]" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF34D2DBFE3DF957" bold="true" box="[199,384,1728,1754]" pageId="18" pageNumber="19">Measurements:</emphasis>
length 8.59.0 mm; width 4.35.0 mm.
</paragraph>
<paragraph id="8BC90F13E00EFFE7FF34D2F8FF69F8AD" blockId="18.[151,1437,1693,2032]" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF34D2F8FEE6F970" bold="true" box="[199,347,1763,1789]" pageId="18" pageNumber="19">Integument:</emphasis>
Black; funiculum brown annulated; elytra transparently pale-brown, margins sometimes blackened.
</paragraph>
<paragraph id="8BC90F13E00EFFE6FF34D333FB14FC26" blockId="18.[151,1437,1693,2032]" lastBlockId="19.[151,1437,151,2026]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B902D301E00EFFE7FF34D333FEFAF8CF" bold="true" box="[199,327,1832,1858]" pageId="18" pageNumber="19">Tegument:</emphasis>
<emphasis id="B902D301E00EFFE7FEBDD333FE15F8CF" box="[334,424,1832,1858]" italics="true" pageId="18" pageNumber="19">Clypeus</emphasis>
bare; sparse setose with very short, erect, black setae on the inner side of the vertically reflexed anterior and lateral margin.
<emphasis id="B902D301E00EFFE7FDE5D350FDEEF8E8" box="[534,595,1867,1893]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
loose setose with medium-long, erect, black setae.
<emphasis id="B902D301E00EFFE7FB79D357FAACF8E8" box="[1162,1297,1867,1893]" italics="true" pageId="18" pageNumber="19">Eye canthus</emphasis>
with a loose setose ciliary of long black setae.
<emphasis id="B902D301E00EFFE7FDE5D375FDD8F805" box="[534,613,1902,1928]" italics="true" pageId="18" pageNumber="19">Scapus</emphasis>
loose setose with medium-long, black setae, brown ventrally, sparse and short on
<emphasis id="B902D301E00EFFE7FF05D38BFED5F827" box="[246,360,1936,1962]" italics="true" pageId="18" pageNumber="19">pedicellus</emphasis>
;
<emphasis id="B902D301E00EFFE7FE86D38BFE58F827" box="[373,485,1936,1962]" italics="true" pageId="18" pageNumber="19">funiculum</emphasis>
with few very short, semi-erect, brown setae, sparse on
<emphasis id="B902D301E00EFFE7FBBDD38BFB2BF827" box="[1102,1174,1936,1962]" italics="true" pageId="18" pageNumber="19">clavus</emphasis>
.
<emphasis id="B902D301E00EFFE7FB51D38AFAACF827" box="[1186,1297,1937,1962]" italics="true" pageId="18" pageNumber="19">Pronotum</emphasis>
loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long, black setae on lateral margin.
<emphasis id="B902D301E00EFFE7FF64D3CDFEB8F87D" box="[151,261,2006,2032]" italics="true" pageId="18" pageNumber="19">Scutellum</emphasis>
loose setose with medium-long, erect, black setae.
<emphasis id="B902D301E00EFFE7FCB7D3CDFC37F87D" box="[836,906,2006,2032]" italics="true" pageId="18" pageNumber="19">Elytra</emphasis>
without pattern. Loose setose with short, semi-erect, brownish setae; medium-long, erect, black setae interspersed, long towards humerus; with a sparse juxtasutural row of robust, long, erect, black setae, shorter in posterior half, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, erect, black setae.
<emphasis id="B902D301E00FFFE6FB42D4C7FABEFF7B" box="[1201,1283,220,246]" italics="true" pageId="19" pageNumber="20">Thorax</emphasis>
and
<emphasis id="B902D301E00FFFE6FACBD4C7FA21FF7B" box="[1336,1436,220,246]" italics="true" pageId="19" pageNumber="20">pleurites</emphasis>
loose pilose with long, black hairs.
<emphasis id="B902D301E00FFFE6FDC1D4E4FD01FE95" box="[562,700,255,280]" italics="true" pageId="19" pageNumber="20">Ventrite IV</emphasis>
loose setose with short, reclinate, black setae; long, semi-erect, black setae interspersed.
<emphasis id="B902D301E00FFFE6FE40D53AFDFAFEB6" box="[435,583,289,315]" italics="true" pageId="19" pageNumber="20">Propygidium</emphasis>
loose squamose with reclinate, lanceolate, whitish scales; long, semi-erect, black setae interspersed.
<emphasis id="B902D301E00FFFE6FE40D55FFD9DFED3" box="[435,544,324,350]" italics="true" pageId="19" pageNumber="20">Pygidium</emphasis>
sparse setose with long, erect, black setae; tip of pygidium with a loose ciliary of medium-long, black setae.
<emphasis id="B902D301E00FFFE6FE1FD57CFD02FE0C" box="[492,703,359,385]" italics="true" pageId="19" pageNumber="20">Prothoracic femur</emphasis>
loose pilose with long, black hairs; bare internally.
<emphasis id="B902D301E00FFFE6FAE0D57CFF76FE2E" italics="true" pageId="19" pageNumber="20">Prothoracic tibia</emphasis>
loose setose with short, reclinate, black setae, medium-long and long ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a longer, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth.
<emphasis id="B902D301E00FFFE6FB0CD5EAFF64FDA3" italics="true" pageId="19" pageNumber="20">Mesothoracic femur</emphasis>
loose pilose with long, black hairs; bare internally.
<emphasis id="B902D301E00FFFE6FCD8D60FFBBEFDA3" box="[811,1027,532,558]" italics="true" pageId="19" pageNumber="20">Mesothoracic tibia</emphasis>
loose setose with short, semi-erect, whitish setae; medium-long, semi-erect, black, robust setae interspersed, sparser, finer, medium-long and long internally. Posterior margin with a bristled ciliary of very short and short, black bristles, lengthening dorsally and ventrally, missing internally.
<emphasis id="B902D301E00FFFE6FE14D667FD71FD1B" box="[487,716,636,662]" italics="true" pageId="19" pageNumber="20">Metathoracic femur</emphasis>
loose setose with medium-long to distinctly long, erect, black setae; whitish ventrally; bare internally.
<emphasis id="B902D301E00FFFE6FD9BD684FC83FD34" box="[616,830,671,697]" italics="true" pageId="19" pageNumber="20">Metathoracic tibia</emphasis>
loose setose with very short, reclinate, brown setae; medium-long and long, robust, semi-erect, black setae interspersed, denser and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally.
<emphasis id="B902D301E00FFFE6FEFFD71CFE28FCAC" box="[268,405,775,801]" italics="true" pageId="19" pageNumber="20">Prothoracic</emphasis>
and
<emphasis id="B902D301E00FFFE6FE22D71CFC9EFCAD" box="[465,803,775,801]" italics="true" pageId="19" pageNumber="20">mesothoracic tarsomere IIV</emphasis>
bare.
<emphasis id="B902D301E00FFFE6FC9DD71CFB7CFCAD" box="[878,1217,775,801]" italics="true" pageId="19" pageNumber="20">Metathoracic tarsomere IIV</emphasis>
sparse setose with very short, reclinate, brown setae.
<emphasis id="B902D301E00FFFE6FDEDD731FD6DFCCE" box="[542,720,810,835]" italics="true" pageId="19" pageNumber="20">Tarsomere IIV</emphasis>
of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards
<emphasis id="B902D301E00FFFE6FDAAD757FCC8FCEB" box="[601,885,844,870]" italics="true" pageId="19" pageNumber="20">metathoracic tarsomeres</emphasis>
, fine dorsally and more robust ventrally.
<emphasis id="B902D301E00FFFE6FAB9D756FF55FC05" italics="true" pageId="19" pageNumber="20">Pretarsomere</emphasis>
of all thoracic legs sparse setose with extremely short to very short, semi-erect and reclinate, brown setae; short and semi-erect ventrally; few, medium-long, semi-erect, black setae at apical margin.
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D7AEFB90FB9A" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D7AEFEACFC42" bold="true" box="[199,273,949,975]" pageId="19" pageNumber="20">Head:</emphasis>
Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle III cylindrical; funicle III asymmetric frustumshaped; clavus elliptic, slightly longer than pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D03AFA2AFB0E" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D03AFEF1FBB6" bold="true" box="[199,332,1057,1083]" pageId="19" pageNumber="20">Pronotum:</emphasis>
Widest at anterior two fifth; loose punctate with very small and small, shallow, roundish punctures, dense apically, loose within medial furrow. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow medial furrow. Posterior corner angled.
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D096FE96FB62" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D096FEF6FB2A" bold="true" box="[199,331,1165,1191]" pageId="19" pageNumber="20">Scutellum:</emphasis>
Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures, micropunctate within; with a complete, median longitudinal, narrow, smooth stripe. Lateral margin slightly convex. Tip rounded.
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D0E2FF52FA45" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D0E2FEA3FA9E" bold="true" box="[199,286,1273,1299]" pageId="19" pageNumber="20">Elytra:</emphasis>
Ovate; longer than broad; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus distinct; with a roundish torus beside scutellum; lateral trench missing; broadly, slightly, longitudinally depressed along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, roundish punctures from anterior margin, crossing scutellar torus towards apical torus, terminating in apical quarter.
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D1CAFF40F9DA" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D1CAFEFAFA66" bold="true" box="[199,327,1489,1515]" pageId="19" pageNumber="20">Abdomen:</emphasis>
Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin moderately convex, linear medially; lateral margin slightly convex attenuate apically; tip rounded.
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D27AFF15F8DE" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D27AFE30F9F6" bold="true" box="[199,397,1633,1659]" pageId="19" pageNumber="20">Prothoracic leg:</emphasis>
Femur and tibia robust. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth concave. Tarsomere IIV combined extremely short; lengthening from tarsomere IIV; slightly less than
<superScript id="7C03A25BE00FFFE6FB1FD2D6FB49F956" attach="right" box="[1260,1268,1741,1755]" fontSize="6" pageId="19" pageNumber="20">7</superScript>
/
<quantity id="4C8EA2F6E00FFFE6FB0ED2FBFA91F96A" box="[1277,1324,1741,1774]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="20" unit="in" value="10.0">
<subScript id="17F20D56E00FFFE6FB0ED2FBFAB0F963" attach="left" box="[1277,1293,1760,1774]" fontSize="6" pageId="19" pageNumber="20">10</subScript>
in
</quantity>
length of tibia. Pretarsomere distinctly thickened; with a small, triangular-shaped, median tooth ventrally; slightly longer than tarsomere IIV combined. Internal claw slightly shorter than pretarsomere. External claw slightly more than
<superScript id="7C03A25BE00FFFE6FF64D322FF22F8CA" attach="none" box="[151,159,1849,1863]" fontSize="6" pageId="19" pageNumber="20">7</superScript>
/
</paragraph>
<paragraph id="8BC90F13E00FFFE6FF5BD357FE5BF8DE" blockId="19.[151,1437,151,2026]" box="[168,486,1849,1882]" pageId="19" pageNumber="20">10 in length of internal claw.</paragraph>
<paragraph id="8BC90F13E00FFFE6FF34D346FD7EF86E" blockId="19.[151,1437,151,2026]" pageId="19" pageNumber="20">
<emphasis id="B902D301E00FFFE6FF34D346FE1DF8FA" bold="true" box="[199,416,1885,1911]" pageId="19" pageNumber="20">Mesothoracic leg:</emphasis>
Tibia slightly less than
<superScript id="7C03A25BE00FFFE6FD5DD346FD0BF8E6" attach="right" box="[686,694,1885,1899]" fontSize="6" pageId="19" pageNumber="20">9</superScript>
/
<quantity id="4C8EA2F6E00FFFE6FD4CD36BFD51F8FA" box="[703,748,1885,1918]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="20" unit="in" value="10.0">
<subScript id="17F20D56E00FFFE6FD4CD36BFD72F8F3" attach="left" box="[703,719,1904,1918]" fontSize="6" pageId="19" pageNumber="20">10</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined extremely short; slightly less than
<superScript id="7C03A25BE00FFFE6FE91D39AFED7F802" attach="right" box="[354,362,1921,1935]" fontSize="6" pageId="19" pageNumber="20">2</superScript>
/
<quantity id="4C8EA2F6E00FFFE6FE81D38FFE25F816" box="[370,408,1921,1954]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="19" pageNumber="20" unit="in" value="5.0">
<subScript id="17F20D56E00FFFE6FE81D38FFEC7F82F" attach="left" box="[370,378,1940,1954]" fontSize="6" pageId="19" pageNumber="20">5</subScript>
in
</quantity>
length of tibia. Pretarsomere with a shortened granulate ridge ventrally, roundish cutting missing; as long as tarsomere IIV combined. External claw slightly shorter than pretarsomere. Internal claw slightly more than
<superScript id="7C03A25BE00FFFE6FE9CD3D2FECAF85A" attach="right" box="[367,375,1993,2007]" fontSize="6" pageId="19" pageNumber="20">9</superScript>
/
<quantity id="4C8EA2F6E00FFFE6FE73D3C7FE13F86E" box="[384,430,1993,2026]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="20" unit="in" value="10.0">
<subScript id="17F20D56E00FFFE6FE73D3C7FE2DF867" attach="left" box="[384,400,2012,2026]" fontSize="6" pageId="19" pageNumber="20">10</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D48DFC41FE90" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D48DFE23FF3C" bold="true" box="[199,414,150,177]" pageId="20" pageNumber="21">Metathoracic leg:</emphasis>
Tibia slightly more than
<superScript id="7C03A25BE008FFE1FD4FD48DFD79FF29" attach="right" box="[700,708,150,164]" fontSize="6" pageId="20" pageNumber="21">1</superScript>
/
<subScript id="17F20D56E008FFE1FD3ED4B2FD60FF3A" attach="left" box="[717,733,169,183]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
more in length of femur. Tarsomere IIV combined distinctly short; slightly less than
<superScript id="7C03A25BE008FFE1FE5FD4A0FE09FF44" attach="right" box="[428,436,187,201]" fontSize="6" pageId="20" pageNumber="21">7</superScript>
/
<quantity id="4C8EA2F6E008FFE1FE4FD4D6FE56FF58" box="[444,491,187,219]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="10.0">
<subScript id="17F20D56E008FFE1FE4FD4D6FE71FF56" attach="left" box="[444,460,205,219]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than
<superScript id="7C03A25BE008FFE1FE9ED4C5FEC8FF61" attach="right" box="[365,373,222,236]" fontSize="6" pageId="20" pageNumber="21">1</superScript>
/
<quantity id="4C8EA2F6E008FFE1FE8ED4EAFE1FFF74" box="[381,418,223,255]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="20" pageNumber="21" unit="in" value="2.0">
<subScript id="17F20D56E008FFE1FE8ED4EAFE38FF72" attach="left" box="[381,389,241,255]" fontSize="6" pageId="20" pageNumber="21">2</subScript>
in
</quantity>
length of tarsomere IIV combined. External claw slightly less than
<superScript id="7C03A25BE008FFE1FB42D4C5FB04FF61" attach="right" box="[1201,1209,222,236]" fontSize="6" pageId="20" pageNumber="21">1</superScript>
/
<subScript id="17F20D56E008FFE1FB32D4EAFB6CFF72" attach="left" box="[1217,1233,241,255]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
more in length of pretarsomere. Internal claw slightly more than
<superScript id="7C03A25BE008FFE1FD42D519FD04FE9D" attach="right" box="[689,697,258,272]" fontSize="6" pageId="20" pageNumber="21">4</superScript>
/
<quantity id="4C8EA2F6E008FFE1FD31D50EFD5AFE90" box="[706,743,259,291]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FD31D50EFD77FEAE" attach="left" box="[706,714,277,291]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D53CFD8BFECC" blockId="20.[151,1437,150,2019]" box="[199,566,294,321]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D53CFEE9FECC" bold="true" box="[199,340,295,321]" pageId="20" pageNumber="21">Parameres:</emphasis>
(
<figureCitation id="134D1396E008FFE1FE97D53CFE52FECC" box="[356,495,294,321]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="20" pageNumber="21">Figs. 6465</figureCitation>
,
<figureCitation id="134D1396E008FFE1FE08D53CFD94FECC" box="[507,553,295,321]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="20" pageNumber="21">101</figureCitation>
).
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D557FCC3FEE8" blockId="20.[151,1437,150,2019]" box="[199,894,330,359]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D557FE01FEE8" bold="true" box="[199,444,331,359]" pageId="20" pageNumber="21">
<specimenCount id="9D70C49AE008FFE1FF34D557FF4FFEEA" box="[199,242,332,359]" pageId="20" pageNumber="21" type="female">♀♀</specimenCount>
. Measurements:
</emphasis>
length
<quantity id="4C8EA2F6E008FFE1FDE0D550FD25FEE8" box="[531,664,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="9.7" metricValueMin="9.3" pageId="20" pageNumber="21" unit="mm" value="9.5" valueMax="9.7" valueMin="9.3">9.39.7mm</quantity>
; width
<quantity id="4C8EA2F6E008FFE1FD1ED551FCC7FEE8" box="[749,890,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.7" metricValueMin="4.9" pageId="20" pageNumber="21" unit="mm" value="5.3" valueMax="5.7" valueMin="4.9">4.95.7 mm</quantity>
.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D574FEABFBF0" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D574FEF7FE04" bold="true" box="[199,330,367,393]" pageId="20" pageNumber="21">Tegument:</emphasis>
<emphasis id="B902D301E008FFE1FEA6D574FE0CFE04" box="[341,433,367,393]" italics="true" pageId="20" pageNumber="21">Clypeus</emphasis>
setose with loose, very short, semi-erect brown setae, medium-long on
<emphasis id="B902D301E008FFE1FAE7D574FAF6FE04" box="[1300,1355,367,393]" italics="true" pageId="20" pageNumber="21">head</emphasis>
. Setae on
<emphasis id="B902D301E008FFE1FF4FD588FE93FE20" box="[188,302,403,429]" italics="true" pageId="20" pageNumber="21">funiculum</emphasis>
and
<emphasis id="B902D301E008FFE1FE9BD588FE0CFE20" box="[360,433,403,429]" italics="true" pageId="20" pageNumber="21">clavus</emphasis>
pale-brown.
<emphasis id="B902D301E008FFE1FDBFD588FD65FE20" box="[588,728,403,429]" italics="true" pageId="20" pageNumber="21">Eye canthus</emphasis>
with a loose ciliary of medium-long, brown setae.
<emphasis id="B902D301E008FFE1FAD8D588FA21FE21" box="[1323,1436,403,428]" italics="true" pageId="20" pageNumber="21">Pronotum</emphasis>
without pattern. Loose setose with medium-long and long, erect, brown setae; very short and short, semi-erect, yellowish setae interspersed in medial furrow. With a loose setose ciliary of medium-long and long brown setae on lateral margin.
<emphasis id="B902D301E008FFE1FE95D5E4FE6AFD94" box="[358,471,511,537]" italics="true" pageId="20" pageNumber="21">Scutellum</emphasis>
loose setose with medium-long, semi-erect, brown setae; short semi-erect, yellowish setae interspersed.
<emphasis id="B902D301E008FFE1FE81D638FE04FDB0" box="[370,441,547,573]" italics="true" pageId="20" pageNumber="21">Elytra</emphasis>
without pattern. Loose setose with short, reclinate, pale-brown setae; long, semi-erect and erect, brown setae interspersed, denser towards humerus.
<emphasis id="B902D301E008FFE1FCA4D65CFC14FDEC" box="[855,937,583,609]" italics="true" pageId="20" pageNumber="21">Thorax</emphasis>
and
<emphasis id="B902D301E008FFE1FC13D65CFBF9FDEC" box="[992,1092,583,609]" italics="true" pageId="20" pageNumber="21">pleurites</emphasis>
loose pilose with long, brown hairs.
<emphasis id="B902D301E008FFE1FF2CD670FED4FD09" box="[223,361,619,644]" italics="true" pageId="20" pageNumber="21">Ventrite IV</emphasis>
loose setose with short and medium-long, reclinate, yellowish-white setae; long, semi-erect, brown setae interspersed.
<emphasis id="B902D301E008FFE1FE34D694FDE6FD24" box="[455,603,655,681]" italics="true" pageId="20" pageNumber="21">Propygidium</emphasis>
and
<emphasis id="B902D301E008FFE1FD64D694FCBCFD24" box="[663,769,655,681]" italics="true" pageId="20" pageNumber="21">pygidium</emphasis>
loose setose with short, reclinate, yellowish-white setae; long, semi-erect, brown setae interspersed. Tip of pygidium with a loose setose ciliary of medium-long and long black-brown setae.
<emphasis id="B902D301E008FFE1FE85D6CCFDF5FD7C" box="[374,584,727,753]" italics="true" pageId="20" pageNumber="21">Prothoracic femur</emphasis>
loose pilose with medium-long and long brown hairs; bare internally.
<emphasis id="B902D301E008FFE1FA9AD6CCFE89FC98" italics="true" pageId="20" pageNumber="21">Prothoracic tibia</emphasis>
loose setose with very short and short semi-erect brown setae; erect and medium-long ventrally. With a loose setose ciliary of medium-long brown setae on internal ridge; anterior margin with a loose setaceous ciliary of short pale-brown setae and a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth.
<emphasis id="B902D301E008FFE1FDF5D77CFD50FC0C" box="[518,749,871,897]" italics="true" pageId="20" pageNumber="21">Mesothoracic femur</emphasis>
loose pilose with long black hairs; long and distinctly-long pale-brown ventrally; bare internally.
<emphasis id="B902D301E008FFE1FDA6D790FC8CFC28" box="[597,817,907,933]" italics="true" pageId="20" pageNumber="21">Mesothoracic tibia</emphasis>
loose setose with short, semi-erect, yellowish-white setae; sparser internally; medium-long, semi-erect, pale-brown, robust setae interspersed; medium-long and long, finer and darker internally. Fringe of posterior margin as in male but red-brown.
<emphasis id="B902D301E008FFE1FBD9D7C8FAB1FC60" box="[1066,1292,979,1005]" italics="true" pageId="20" pageNumber="21">Metathoracic femur</emphasis>
loose setose with medium-long and long brown setae; paler ventrally; bare internally.
<emphasis id="B902D301E008FFE1FC00D7ECFB76FB9C" box="[1011,1227,1015,1041]" italics="true" pageId="20" pageNumber="21">Metathoracic tibia</emphasis>
loose setose with short, semi-erect, pale-brown setae; medium-long and robust, semi-erect, red-brown setae interspersed; finer, medium-long and long internally. Ciliary of posterior margin as in male but red-brown. Setae of
<emphasis id="B902D301E008FFE1FAF3D024FA21FBD5" box="[1280,1436,1087,1113]" italics="true" pageId="20" pageNumber="21">thoracic tarsi</emphasis>
red-brown.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D09CFBBDFA80" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D09CFEACFB2C" bold="true" box="[199,273,1159,1185]" pageId="20" pageNumber="21">Head:</emphasis>
Clypeus very slightly depressed laterally; dense punctate with large, shallow, roundish punctures. Anterior margin slightly vertically reflexed. Lateral margin slightly reflexed. Head dense punctate with small to large, shallow, roundish punctures, not merging, not micropunctate. Funicle IIIII short-asymmetric frustumshaped; clavus short elliptic, as long as pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D10CFB47FAD8" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D10CFEF1FABC" bold="true" box="[199,332,1303,1329]" pageId="20" pageNumber="21">Pronotum:</emphasis>
Moderate pulvinate laterally. Dense punctate with small to large, shallow, roundish punctures, denser apically. Lateral margin convex attenuate in posterior three fifth. Posterior corner rounded.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D145FEEEFA10" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D145FEF6FAF5" bold="true" box="[199,331,1374,1400]" pageId="20" pageNumber="21">Scutellum:</emphasis>
With a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior third.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D1BCFF76FA68" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D1BCFEA3FA4C" bold="true" box="[199,286,1447,1473]" pageId="20" pageNumber="21">Elytra:</emphasis>
Widest in anterior third; lateral trench indistinct; scutellar torus missing; not depressed along suture.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D1F5FCBCF9DC" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D1F5FEFAF985" bold="true" box="[199,327,1518,1544]" pageId="20" pageNumber="21">Abdomen:</emphasis>
Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite IIV broadened medially. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper quarter, linearly attenuate apically; tip linear.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D241FCE9F888" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D241FE33F9F8" bold="true" box="[199,398,1626,1653]" pageId="20" pageNumber="21">Prothoracic leg:</emphasis>
Median tooth broad, triangular-shaped. Basal tooth very small. Shape of interspace between teeth concave. Tarsomere IIV combined distinctly short; equal in length; slightly more than
<superScript id="7C03A25BE008FFE1FB0FD265FAB9F901" attach="right" box="[1276,1284,1662,1676]" fontSize="6" pageId="20" pageNumber="21">3</superScript>
/
<quantity id="4C8EA2F6E008FFE1FAFFD28AFA8CF914" box="[1292,1329,1663,1695]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FAFFD28AFAA9F912" attach="left" box="[1292,1300,1681,1695]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; with a smooth ridge ventrally; slightly more than
<superScript id="7C03A25BE008FFE1FB70D2B9FB36F93D" attach="right" box="[1155,1163,1698,1712]" fontSize="6" pageId="20" pageNumber="21">3</superScript>
/
<quantity id="4C8EA2F6E008FFE1FB60D2AEFB05F930" box="[1171,1208,1699,1731]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FB60D2AEFB26F94E" attach="left" box="[1171,1179,1717,1731]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of tarsomere IIV combined. Internal claw slender; not broadened medially; slightly less than
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/
<quantity id="4C8EA2F6E008FFE1FBA4D2C2FB3BF96C" box="[1111,1158,1735,1767]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="10.0">
<subScript id="17F20D56E008FFE1FBA4D2C2FBDAF96A" attach="left" box="[1111,1127,1753,1767]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
in
</quantity>
length of pretarsomere. External claw slightly less than
<superScript id="7C03A25BE008FFE1FDF6D2F1FDB0F975" attach="right" box="[517,525,1770,1784]" fontSize="6" pageId="20" pageNumber="21">9</superScript>
/
<quantity id="4C8EA2F6E008FFE1FDE5D2E6FDF9F888" box="[534,580,1771,1803]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="10.0">
<subScript id="17F20D56E008FFE1FDE5D2E6FD9BF886" attach="left" box="[534,550,1789,1803]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
in
</quantity>
length of internal claw.
</paragraph>
<paragraph id="8BC90F13E008FFE1FF34D315FDA6F818" blockId="20.[151,1437,150,2019]" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D315FE1CF8A4" bold="true" box="[199,417,1806,1833]" pageId="20" pageNumber="21">Mesothoracic leg:</emphasis>
Tibia
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/
<quantity id="4C8EA2F6E008FFE1FE08D33AFD97F8A4" box="[507,554,1807,1839]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="10.0">
<subScript id="17F20D56E008FFE1FE08D33AFDB6F8A2" attach="left" box="[507,523,1825,1839]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined very short; slightly more than
<superScript id="7C03A25BE008FFE1FA77D315FA31F891" attach="right" box="[1412,1420,1806,1820]" fontSize="6" pageId="20" pageNumber="21">2</superScript>
/
<quantity id="4C8EA2F6E008FFE1FA67D33AFF10F8C0" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FA67D33AFA21F8A2" attach="none" box="[1428,1436,1825,1839]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of tibia. Pretarsomere with an undulate ridge ventrally, roundish cutting missing; slightly less than
<superScript id="7C03A25BE008FFE1FA77D329FA31F8CD" attach="right" box="[1412,1420,1842,1856]" fontSize="6" pageId="20" pageNumber="21">4</superScript>
/
<quantity id="4C8EA2F6E008FFE1FA67D35EFF10F8FC" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FA67D35EFA21F8DE" attach="none" box="[1428,1436,1861,1875]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of tarsomere IIV combined. External claw slender; not broadened medially; Internal claw slightly less than
<superScript id="7C03A25BE008FFE1FF23D361FF65F805" attach="right" box="[208,216,1914,1928]" fontSize="6" pageId="20" pageNumber="21">4</superScript>
/
<quantity id="4C8EA2F6E008FFE1FF13D396FEBBF818" box="[224,262,1915,1947]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FF13D396FF55F816" attach="left" box="[224,232,1933,1947]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E008FFE0FF34D385FCECFF58" blockId="20.[151,1437,150,2019]" lastBlockId="21.[151,1437,150,219]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B902D301E008FFE1FF34D385FE1DF834" bold="true" box="[199,416,1950,1977]" pageId="20" pageNumber="21">Metathoracic leg:</emphasis>
Femur slightly thickened. Tibia distinctly extended dorso-apically; slightly less than
<superScript id="7C03A25BE008FFE1FA88D385FA3EF821" attach="right" box="[1403,1411,1950,1964]" fontSize="6" pageId="20" pageNumber="21">9</superScript>
/
<quantity id="4C8EA2F6E008FFE1FA7FD3AAFF10F850" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="10.0">
<subScript id="17F20D56E008FFE1FA7FD3AAFA21F832" attach="none" box="[1420,1436,1969,1983]" fontSize="6" pageId="20" pageNumber="21">10</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined short; slightly more than
<superScript id="7C03A25BE008FFE1FC4ED3D9FC78F85D" attach="right" box="[957,965,1986,2000]" fontSize="6" pageId="20" pageNumber="21">4</superScript>
/
<quantity id="4C8EA2F6E008FFE1FC3ED3CEFC4CF850" box="[973,1009,1987,2019]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="21" unit="in" value="5.0">
<subScript id="17F20D56E008FFE1FC3ED3CEFC68F86E" attach="left" box="[973,981,2005,2019]" fontSize="6" pageId="20" pageNumber="21">5</subScript>
in
</quantity>
length of tibia. Pretarsomere slender; slightly more than
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<quantity id="4C8EA2F6E009FFE0FE75D4B2FE10FF3C" box="[390,429,151,183]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="22" unit="in" value="5.0">
<subScript id="17F20D56E009FFE0FE75D4B2FE33FF3A" attach="left" box="[390,398,169,183]" fontSize="6" pageId="21" pageNumber="22">5</subScript>
in
</quantity>
length of tarsomere IIV combined. External claw slightly shorter than pretarsomere. Internal claw slightly less than
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<quantity id="4C8EA2F6E009FFE0FDFDD4D6FD81FF58" box="[526,572,187,219]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="22" unit="in" value="10.0">
<subScript id="17F20D56E009FFE0FDFDD4D6FDA3FF56" attach="left" box="[526,542,205,219]" fontSize="6" pageId="21" pageNumber="22">10</subScript>
in
</quantity>
length of external claw.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E009FFE0FF64D519FCDBFEE8" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BC90F13E009FFE0FF64D519FBDAFE90" blockId="21.[151,1127,258,285]" box="[151,1127,258,285]" pageId="21" pageNumber="22">
<emphasis id="B902D301E009FFE0FF64D519FE89FE91" bold="true" box="[151,308,258,284]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingRegion id="49B2C1F1E009FFE0FEC8D518FDEFFE90" box="[315,594,259,285]" country="South Africa" name="Northern Cape" pageId="21" pageNumber="22">Northern Cape Province</collectingRegion>
(
<figureCitation id="134D1396E009FFE0FD92D518FD78FE90" box="[609,709,259,285]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="21" pageNumber="22">Fig. 140</figureCitation>
), only known from the
<typeStatus id="54CDB1B1E009FFE0FC20D518FBB9FE90" box="[979,1028,259,285]" pageId="21" pageNumber="22">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BC90F13E009FFE0FF64D551FCDBFEE8" blockId="21.[151,870,330,357]" box="[151,870,330,357]" pageId="21" pageNumber="22">
<emphasis id="B902D301E009FFE0FF64D551FEDFFEE8" bold="true" box="[151,354,330,357]" pageId="21" pageNumber="22">
Vegetation
<typeStatus id="54CDB1B1E009FFE0FEECD550FEE0FEE8" box="[287,349,331,357]" pageId="21" pageNumber="22">Type</typeStatus>
.
</emphasis>
Nieuwoudtville Shale Renosterveld [FRs 2].
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E009FFE0FF64D588FB82FE20" box="[151,1087,403,429]" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="8BC90F13E009FFE0FF64D588FB82FE20" blockId="21.[151,1087,403,429]" box="[151,1087,403,429]" pageId="21" pageNumber="22">
<emphasis id="B902D301E009FFE0FF64D588FE96FE20" bold="true" box="[151,299,403,429]" pageId="21" pageNumber="22">Host Plants.</emphasis>
<taxonomicName id="4C767490E009FFE0FEC1D588FE1DFE20" box="[306,416,403,429]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="family">Iridaceae</taxonomicName>
:
<taxonomicName id="4C767490E009FFE0FE58D588FD68FE20" authority="Andrews" authorityName="Andrews" box="[427,725,403,429]" class="Liliopsida" family="Iridaceae" genus="Moraea" kingdom="Plantae" order="Asparagales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="miniata">
<emphasis id="B902D301E009FFE0FE58D588FDD9FE21" box="[427,612,403,428]" italics="true" pageId="21" pageNumber="22">Moraea miniata</emphasis>
Andrews
</taxonomicName>
;
<taxonomicName id="4C767490E009FFE0FD2CD588FCDCFE20" box="[735,865,403,429]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
:
<taxonomicName id="4C767490E009FFE0FC9FD588FB9DFE20" box="[876,1056,403,429]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="acaulis">
<emphasis id="B902D301E009FFE0FC9FD588FB9DFE20" box="[876,1056,403,429]" italics="true" pageId="21" pageNumber="22">Arctotis acaulis</emphasis>
</taxonomicName>
L.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E009FFE0FF64D5C0FAE4FE78" box="[151,1369,474,501]" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8BC90F13E009FFE0FF64D5C0FAE4FE78" blockId="21.[151,1369,474,501]" box="[151,1369,474,501]" pageId="21" pageNumber="22">
<emphasis id="B902D301E009FFE0FF64D5C0FE9DFE78" bold="true" box="[151,288,475,501]" pageId="21" pageNumber="22">Etymology.</emphasis>
The name derives from the pale-brown elytra [Latin: pallidi- = pale; Latin: brunne- = brown].
</paragraph>
</subSubSection>
</treatment>
</document>