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<mods:title id="3AD7289E4D98170C2E9FF15A069F247F">New orchids in the flora of Vietnam VIII (Orchidaceae: Epidendroideae, tribes Malaxideae, Neottieae and Podochileae)</mods:title>
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<mods:namePart id="A9C03A00149FD821AFEA3D4D153E9FBD">Yudova, Daria A.</mods:namePart>
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Aver. &amp; Vuong
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(
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&amp; 
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= 
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(J.Koenig in 
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) 
<treatmentCitation id="0AD710F6FFEBDB21FDF3F9E4FD5BF9E4" author="Smith, J. J." box="[578,753,1602,1624]" page="119" pageId="31" pageNumber="32" year="1932">
<bibRefCitation id="EFE74B16FFEBDB21FDF3F9E4FD5BF9E4" author="Smith, J. J." box="[578,753,1602,1624]" pageId="31" pageNumber="32" pagination="105 - 150" refId="ref34915" refString="Smith, J. J. (1932) Orchidaceae novae Malayenses XIV. Bulletin du Jardin Botanique de Buitenzorg, ser. 3, 12: 105 - 150." type="journal article" year="1932">Smith (1932: 119)</bibRefCitation>
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pro parte: 
<treatmentCitation id="0AD710F6FFEBDB21FCE8F9E4FC4CF9E4" author="Ho, P. - H." box="[857,998,1602,1623]" page="979" pageId="31" pageNumber="32" year="1993">
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, fig. 9874, 2000: 799, fig. 10954), 
<treatmentCitation id="0AD710F6FFEBDB21FA84F9E4FEB1F9C8" author="Seidenfaden, G." page="9" pageId="31" pageNumber="32" year="1975">
<bibRefCitation id="EFE74B16FFEBDB21FA84F9E4FEB1F9C8" author="Seidenfaden, G." pageId="31" pageNumber="32" refId="ref34311" refString="Seidenfaden, G. (1975 a) Contributions to a revision of the orchid flora of Cambodia, Laos and Vietnam. Kai Olsen, Fredensborg, 117 pp." type="book" year="1975">Seidenfaden (1975a: 9</bibRefCitation>
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, 1992: 200), 
<treatmentCitation id="0AD710F6FFEBDB21FE28F9C0FDCDF9C8" author="Averyanov, L. V." box="[409,615,1638,1659]" page="22" pageId="31" pageNumber="32" year="1988">
<bibRefCitation id="EFE74B16FFEBDB21FE28F9C0FDCDF9C8" author="Averyanov, L. V." box="[409,615,1638,1659]" pageId="31" pageNumber="32" refId="ref27790" refString="Averyanov, L. V. (1988) Preliminary list of the Vietnamese orchids. Part 1 (genera A - L). Botanical Institute of the Academy of Sciences of the USSR, Leningrad, 247 pp. [in Russian]" type="book" year="1988">Averyanov (1988: 22</bibRefCitation>
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, 
<treatmentCitation id="0AD710F6FFEBDB21FDC0F9C0FD62F9C8" author="Averyanov, L. V." box="[625,712,1638,1659]" page="27" pageId="31" pageNumber="32" year="1990">
<bibRefCitation id="EFE74B16FFEBDB21FDC0F9C0FD62F9C8" author="Averyanov, L. V." box="[625,712,1638,1659]" pageId="31" pageNumber="32" pagination="20 - 185" refId="ref27835" refString="Averyanov, L. V. (1990) Orchidaceae. In: Tzvelev, N. N. (Ed.) Vascular plants synopsis of Vietnamese flora. Vol. 1. Nauka, Leningrad, 20 pp. 20 - 185." type="book chapter" year="1990">1990: 27</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="0AD710F6FFEBDB21FD63F9C0FC9FF9CF" author="Averyanov, L. V." box="[722,821,1638,1660]" page="164" pageId="31" pageNumber="32" year="1994">
<bibRefCitation id="EFE74B16FFEBDB21FD63F9C0FC9FF9CF" author="Averyanov, L. V." box="[722,821,1638,1660]" pageId="31" pageNumber="32" refId="ref27880" refString="Averyanov, L. V. (1994) Identification guide to Vietnamese orchids (Orchidaceae Juss.). World and Family, St. Petersburg, 432 pp. [in Russian]" type="book" year="1994">1994: 164</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="0AD710F6FFEBDB21FCF1F9C0FC3CF9CF" author="Averyanov, L. V." box="[832,918,1638,1660]" page="51" pageId="31" pageNumber="32" year="2006">
<bibRefCitation id="EFE74B16FFEBDB21FCF1F9C0FC3CF9CF" author="Averyanov, L. V." box="[832,918,1638,1660]" pageId="31" pageNumber="32" pagination="48 - 89" refId="ref27941" refString="Averyanov, L. V. (2006) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 9 (3): 48 - 89." type="journal article" year="2006">2006: 51</bibRefCitation>
</treatmentCitation>
), 
<treatmentCitation id="0AD710F6FFEBDB21FC18F9C0FB95F9CF" author="Hop, T." box="[937,1087,1638,1660]" page="64" pageId="31" pageNumber="32" year="1998">
<bibRefCitation id="EFE74B16FFEBDB21FC18F9C0FB95F9CF" author="Hop, T." box="[937,1087,1638,1660]" pageId="31" pageNumber="32" refId="ref31178" refString="Hop, T. (1998) The orchids of Vietnam. Nha Xuat Ban Nong Nghiep, Ho Chi Minh City, 703 pp." type="book" year="1998">Hop (1998: 64)</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="0AD710F6FFEBDB21FBF9F9C0FA0DF9CF" author="Averyanov, L. V. &amp; Averyanova, A. L." box="[1096,1447,1638,1660]" page="53" pageId="31" pageNumber="32" year="2002">
<bibRefCitation id="EFE74B16FFEBDB21FBF9F9C0FA0DF9CF" author="Averyanov, L. V. &amp; Averyanova, A. L." box="[1096,1447,1638,1660]" pageId="31" pageNumber="32" pagination="49 - 108" refId="ref28069" refString="Averyanov, L. V. &amp; Averyanova, A. L. (2002) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 5 (4): 49 - 108." type="journal article" year="2002">Averyanov &amp; Averyanova (2002: 53</bibRefCitation>
</treatmentCitation>
, 2003: 12).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC936E7FFEBDB21FF39F908FF56F95B" blockId="31.[136,1452,1602,2020]" pageId="31" pageNumber="32">
<treatmentCitationGroup id="AB6611C9FFEBDB21FF39F908FF56F95B" pageId="31" pageNumber="32">
= 
<taxonomicName id="4C764D64FFEBDB21FF2FF908FE63F970" ID-CoL="7RPTQ" authority="Hooker (1890: 84)" authorityName="Hooker" authorityPageNumber="84" authorityYear="1890" box="[158,457,1710,1732]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="koenigii">
<emphasis id="B902EAF5FFEBDB21FF2FF908FEA2F970" box="[158,264,1710,1731]" italics="true" pageId="31" pageNumber="32">A. koenigii</emphasis>
Hooker (1890: 84)
</taxonomicName>
, 
<emphasis id="B902EAF5FFEBDB21FE67F908FD95F970" box="[470,575,1710,1731]" italics="true" pageId="31" pageNumber="32">nom. illeg.</emphasis>
(≡ 
<taxonomicName id="4C764D64FFEBDB21FDD4F908FD77F970" ID-CoL="682R6" box="[613,733,1710,1731]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="hexandra">
<emphasis id="B902EAF5FFEBDB21FDD4F908FD77F970" box="[613,733,1710,1731]" italics="true" pageId="31" pageNumber="32">A. hexandra</emphasis>
</taxonomicName>
; see 
<bibRefCitation id="EFE74B16FFEBDB21FCA3F908FC7AF977" author="Seidenfaden, G." box="[786,976,1710,1732]" pageId="31" pageNumber="32" pagination="41 - 95" refId="ref34283" refString="Seidenfaden, G. (1973 b) Contributions to the orchid flora of Thailand V. Botanisk Tidsskrift 68 (1): 41 - 95." type="journal article" year="1973">Seidenfaden 1973b</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFEBDB21FC6DF908FBB3F977" author="Seidenfaden, G." box="[988,1049,1710,1732]" pageId="31" pageNumber="32" pagination="1 - 216" refId="ref34611" refString="Seidenfaden, G. (1986 b) Orchid genera in Thailand XIII. Thirty-three epidendroid genera. Opera Botanica 89: 1 - 216." type="journal article" year="1986">1986b</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFEBDB21FB97F908FBCFF977" author="Seidenfaden, G." box="[1062,1125,1710,1732]" pageId="31" pageNumber="32" refId="ref34681" refString="Seidenfaden, G. (1995 b) The Descriptiones Epidendrorum of J. G. Konig 1791. Olsen &amp; Olsen, Fredensborg, 61 pp." type="book" year="1995">1995b</bibRefCitation>
), pro parte: 
<treatmentCitation id="0AD710F6FFEBDB21FB51F908FAD4F970" author="Ho, P. - H." box="[1248,1406,1710,1731]" page="1080" pageId="31" pageNumber="32" year="1972">
<bibRefCitation id="EFE74B16FFEBDB21FB51F908FAD4F970" author="Ho, P. - H." box="[1248,1406,1710,1731]" pageId="31" pageNumber="32" refId="ref30993" refString="Ho, P. - H. (1972) Cay-Co Mien-Nam Viet-Nam. 2. ed., 2, Nam Quoc-Te Phat-Trien Sach, Saigon, 1139 pp." type="book" year="1972">Ho (1972: 1080</bibRefCitation>
</treatmentCitation>
, fig. 5208).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFEBDB21FF39F950FB06F8E7" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BC936E7FFEBDB21FF39F950FB06F8E7" blockId="31.[136,1452,1602,2020]" pageId="31" pageNumber="32">
<emphasis id="B902EAF5FFEBDB21FF39F950FF59F8BF" bold="true" box="[136,243,1782,1804]" pageId="31" pageNumber="32">Diagnosis:</emphasis>
—The new species differs from the morphologically most similar 
<taxonomicName id="4C764D64FFEBDB21FCC7F950FC47F8B8" box="[886,1005,1782,1803]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="hexandra">
<emphasis id="B902EAF5FFEBDB21FCC7F950FC47F8B8" box="[886,1005,1782,1803]" italics="true" pageId="31" pageNumber="32">A. hexandra</emphasis>
</taxonomicName>
in inflorescence up to 
<quantity id="4C8E9B02FFEBDB21FB7DF950FA92F8B8" box="[1228,1336,1782,1804]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="31" pageNumber="32" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 cm</quantity>
long, twice as long as leaves or longer, in continuously elongating rachis usually bearing numerous (more than 30) flowers, lip and column with purple spots on sides, and in almost circular to broadly obovate lip callus with small lateral folds at base.
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFEBDB21FF39F8C4FE11F850" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BC936E7FFEBDB21FF39F8C4FE11F850" blockId="31.[136,1452,1602,2020]" pageId="31" pageNumber="32">
<emphasis id="B902EAF5FFEBDB21FF39F8C4FF68F8C4" bold="true" box="[136,194,1890,1911]" pageId="31" pageNumber="32">Type:</emphasis>
— 
<collectingCountry id="F3617677FFEBDB21FF6AF8C4FEE6F8C4" box="[219,332,1890,1911]" name="Vietnam" pageId="31" pageNumber="32">VIETNAM</collectingCountry>
. 
<collectingRegion id="49B2F805FFEBDB21FEE5F8C4FDB4F8CB" box="[340,542,1890,1912]" country="Vietnam" name="Khanh Hoa" pageId="31" pageNumber="32">Khanh Hoa Province</collectingRegion>
: Cam Lam District, Hon Ba Nature Reserve, around point 
<geoCoordinate id="EE425020FFEBDB21FBF7F8C4FB69F8CB" box="[1094,1219,1890,1912]" degrees="12" direction="north" minutes="06.834" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="12.1139">12°06.834’N</geoCoordinate>
<geoCoordinate id="EE425020FFEBDB21FB79F8C4FAFAF8CB" box="[1224,1360,1890,1912]" degrees="108" direction="east" minutes="59.565" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="108.99275">108°59.565’E</geoCoordinate>
, epiphyte in shady riparian forest on granite at elevation of about 
<quantity id="4C8E9B02FFEBDB21FD45F820FC9FF828" box="[756,821,1926,1948]" metricMagnitude="2" metricUnit="m" metricValue="3.61" pageId="31" pageNumber="32" unit="m" value="361.0">361 m</quantity>
a.s.l., along small stream, 
<date id="FFC81027FFEBDB21FBF4F820FB68F828" box="[1093,1218,1926,1947]" pageId="31" pageNumber="32" value="2017-06-09">9 June 2017</date>
, 
<emphasis id="B902EAF5FFEBDB21FB7EF820FF4BF80C" italics="true" pageId="31" pageNumber="32">Truong Ba Vuong, BV 285</emphasis>
(
<typeStatus id="54CD8845FFEBDB21FF40F80CFEEDF873" box="[241,327,1962,1984]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
<collectionCode id="ED67AE22FFEBDB21FEE1F80CFE3EF873" box="[336,404,1962,1984]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="31" pageNumber="32">LE LE</collectionCode>
01054912! https://en.herbariumle.ru/?t=occ&amp;id=5788, 
<typeStatus id="54CD8845FFEBDB21FC1AF80CFC50F873" box="[939,1018,1962,1984]" pageId="31" pageNumber="32" type="isotype">isotypes</typeStatus>
<collectionCode id="ED67AE22FFEBDB21FBB3F80CFBEDF873" box="[1026,1095,1962,1984]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="31" pageNumber="32">LE LE</collectionCode>
01054910! https://en.herbariumle.ru/?t=occ&amp;id=5786, 
<collectionCode id="ED67AE22FFEBDB21FEDDF868FE07F850" box="[364,429,1998,2019]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="31" pageNumber="32" type="Herbarium">VNM</collectionCode>
!).
</paragraph>
</subSubSection>
<caption id="DF09666FFFD4DB1EFF39F8BEFD38F799" ID-DOI="http://doi.org/10.5281/zenodo.13217447" ID-Zenodo-Dep="13217447" httpUri="https://zenodo.org/record/13217447/files/figure.png" pageId="32" pageNumber="33" startId="32.[136,229,1816,1838]" targetBox="[260,1327,190,1791]" targetPageId="32" targetType="figure">
<paragraph id="8BC936E7FFD4DB1EFF39F8BEFD38F799" blockId="32.[136,1453,1816,2090]" pageId="32" pageNumber="33">
<emphasis id="B902EAF5FFD4DB1EFF39F8BEFE7DF89E" bold="true" box="[136,471,1816,1838]" pageId="32" pageNumber="33">
FIGURE 15. 
<taxonomicName id="4C764D64FFD4DB1EFEBDF8BEFE1FF89E" authorityName="Seidenfaden" authorityYear="1980" baseAuthorityName="Rchb. f." box="[268,437,1816,1837]" class="Liliopsida" family="Orchidaceae" genus="Sunipia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="rimannii">
<emphasis id="B902EAF5FFD4DB1EFEBDF8BEFE1FF89E" bold="true" box="[268,437,1816,1837]" italics="true" pageId="32" pageNumber="33">Sunipia rimannii</emphasis>
</taxonomicName>
. A.
</emphasis>
Flowering plant. 
<emphasis id="B902EAF5FFD4DB1EFDCDF8BEFD38F89E" bold="true" box="[636,658,1816,1837]" pageId="32" pageNumber="33">B.</emphasis>
Pseudobulb and basal part of peduncle. 
<emphasis id="B902EAF5FFD4DB1EFBBBF8BEFB88F89D" bold="true" box="[1034,1058,1816,1838]" pageId="32" pageNumber="33">C.</emphasis>
Leaf apex, adaxial side. 
<emphasis id="B902EAF5FFD4DB1EFAB9F8BEFA8AF89E" bold="true" box="[1288,1312,1816,1837]" pageId="32" pageNumber="33">D.</emphasis>
Inflorescence. 
<emphasis id="B902EAF5FFD4DB1EFF39F89AFF34F8E2" bold="true" box="[136,158,1852,1873]" pageId="32" pageNumber="33">E.</emphasis>
Floral bract, adaxial and abaxial side. 
<emphasis id="B902EAF5FFD4DB1EFDA6F89AFDE3F8E1" bold="true" box="[535,585,1852,1874]" pageId="32" pageNumber="33">F, G.</emphasis>
Flower, side view. 
<emphasis id="B902EAF5FFD4DB1EFCB6F89AFC8AF8E2" bold="true" box="[775,800,1852,1873]" pageId="32" pageNumber="33">H.</emphasis>
Flower, view from below. 
<emphasis id="B902EAF5FFD4DB1EFB96F89AFB9CF8E2" bold="true" box="[1063,1078,1852,1873]" pageId="32" pageNumber="33">I.</emphasis>
Flower, view from below, with lateral sepals (synsepal) removed. 
<emphasis id="B902EAF5FFD4DB1EFE20F8C6FE09F8C6" bold="true" box="[401,419,1888,1909]" pageId="32" pageNumber="33">J.</emphasis>
Ovary, lip and column, view from above. 
<emphasis id="B902EAF5FFD4DB1EFC88F8C6FCF8F8C6" bold="true" box="[825,850,1888,1909]" pageId="32" pageNumber="33">K.</emphasis>
Ovary and lip, view from below. 
<emphasis id="B902EAF5FFD4DB1EFB24F8C6FB06F8C6" bold="true" box="[1173,1196,1888,1909]" pageId="32" pageNumber="33">L.</emphasis>
Median sepal, adaxial and abaxial side. 
<emphasis id="B902EAF5FFD4DB1EFEB7F822FE88F82A" bold="true" box="[262,290,1924,1945]" pageId="32" pageNumber="33">M.</emphasis>
Lateral sepals (synsepal), adaxial and abaxial side. 
<emphasis id="B902EAF5FFD4DB1EFCA4F822FC87F82A" bold="true" box="[789,813,1924,1945]" pageId="32" pageNumber="33">N.</emphasis>
Petals, adaxial and abaxial side. 
<emphasis id="B902EAF5FFD4DB1EFBDBF822FB29F829" bold="true" box="[1130,1155,1924,1946]" pageId="32" pageNumber="33">O.</emphasis>
Lip, adaxial side and half-side view. 
<emphasis id="B902EAF5FFD4DB1EFF73F80EFF7FF80E" bold="true" box="[194,213,1960,1981]" pageId="32" pageNumber="33">P.</emphasis>
Lip, abaxial side. 
<emphasis id="B902EAF5FFD4DB1EFE3AF80EFE0EF80D" bold="true" box="[395,420,1960,1982]" pageId="32" pageNumber="33">Q.</emphasis>
Column and lip, side view. 
<emphasis id="B902EAF5FFD4DB1EFD08F80EFD7AF80E" bold="true" box="[697,720,1960,1981]" pageId="32" pageNumber="33">R.</emphasis>
Lip, side view. 
<emphasis id="B902EAF5FFD4DB1EFCDFF80EFC28F80D" bold="true" box="[878,898,1960,1982]" pageId="32" pageNumber="33">S.</emphasis>
Callus at base of lip, adaxial surface and half-side view. 
<emphasis id="B902EAF5FFD4DB1EFF39F86AFF36F852" bold="true" box="[136,156,1996,2017]" pageId="32" pageNumber="33">T.</emphasis>
Pedicel, ovary, column and column foot, side view. 
<emphasis id="B902EAF5FFD4DB1EFD2EF86AFD1DF852" bold="true" box="[671,695,1996,2017]" pageId="32" pageNumber="33">U.</emphasis>
Column, front, half-side and rear view. 
<emphasis id="B902EAF5FFD4DB1EFBF0F86AFBFFF852" bold="true" box="[1089,1109,1996,2017]" pageId="32" pageNumber="33">V.</emphasis>
Pollinarium and pollinium. Photos by Truong Ba Vuong from living plant prior to the preparation of the voucher herbarium specimen 
<emphasis id="B902EAF5FFD4DB1EFBECF857FACCF7B6" box="[1117,1382,2032,2054]" italics="true" pageId="32" pageNumber="33">Truong Ba Vuong, BV1493</emphasis>
. Photo correction and design by L. Averyanov and T. Maisak.
</paragraph>
</caption>
<caption id="DF09666FFFD5DB1FFF39FB1FFF5DFAA5" ID-DOI="http://doi.org/10.5281/zenodo.13217449" ID-Zenodo-Dep="13217449" httpUri="https://zenodo.org/record/13217449/files/figure.png" pageId="33" pageNumber="34" startId="33.[136,229,1209,1231]" targetBox="[165,1412,196,1190]" targetPageId="33" targetType="figure">
<paragraph id="8BC936E7FFD5DB1FFF39FB1FFF5DFAA5" blockId="33.[136,1453,1209,1302]" pageId="33" pageNumber="34">
<emphasis id="B902EAF5FFD5DB1FFF39FB1FFE50FB7D" bold="true" box="[136,506,1209,1231]" pageId="33" pageNumber="34">
FIGURE 16. 
<emphasis id="B902EAF5FFD5DB1FFEBEFB1FFE77FB7D" bold="true" box="[271,477,1209,1231]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C764D64FFD5DB1FFEBEFB1FFE73FB7D" box="[271,473,1209,1231]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="spicata">Appendicula spicata</taxonomicName>
.
</emphasis>
A.
</emphasis>
Flowering shoot. 
<emphasis id="B902EAF5FFD5DB1FFD16FB1FFD17FB7D" bold="true" box="[679,701,1209,1230]" pageId="33" pageNumber="34">B.</emphasis>
Flower, front view. 
<emphasis id="B902EAF5FFD5DB1FFCCFFB1FFC3CFB7C" bold="true" box="[894,918,1209,1231]" pageId="33" pageNumber="34">C.</emphasis>
Flower, side view. 
<emphasis id="B902EAF5FFD5DB1FFBFEFB1FFBCCFB7D" bold="true" box="[1103,1126,1209,1230]" pageId="33" pageNumber="34">D.</emphasis>
Flower, front view (lip removed). 
<emphasis id="B902EAF5FFD5DB1FFF39FB7BFF34FB41" bold="true" box="[136,158,1245,1266]" pageId="33" pageNumber="34">E.</emphasis>
Lip, view from above and from below. 
<emphasis id="B902EAF5FFD5DB1FFD8AFB7BFDE7FB41" bold="true" box="[571,589,1245,1266]" pageId="33" pageNumber="34">F.</emphasis>
Lip, sagittal section. Drawn from the specimen 
<emphasis id="B902EAF5FFD5DB1FFB8BFB7BFAC1FB41" box="[1082,1387,1245,1267]" italics="true" pageId="33" pageNumber="34">Averyanov, T. Maisak, AL1280</emphasis>
by L. Averyanov.
</paragraph>
</caption>
<subSubSection id="C36C656CFFD5DB1FFF39FA99FC06FAEA" box="[136,940,1343,1369]" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BC936E7FFD5DB1FFF39FA99FC06FAEA" blockId="33.[136,1452,1343,2089]" box="[136,940,1343,1369]" pageId="33" pageNumber="34">
<emphasis id="B902EAF5FFD5DB1FFF39FA99FEB9FAEA" bold="true" box="[136,275,1343,1369]" pageId="33" pageNumber="34">Etymology:</emphasis>
—The species name refers to its long spicate inflorescence.
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFD5DB1FFF0CFAC4FCEAF79A" pageId="33" pageNumber="34" type="description">
<paragraph id="8BC936E7FFD5DB1FFF0CFAC4FCEAF79A" blockId="33.[136,1452,1343,2089]" pageId="33" pageNumber="34">
<emphasis id="B902EAF5FFD5DB1FFF0CFAC4FE01FACE" bold="true" box="[189,427,1378,1405]" pageId="33" pageNumber="34">Description:—Herb</emphasis>
clustering, epiphytic or lithophytic. 
<emphasis id="B902EAF5FFD5DB1FFCFCFAC4FC20FACF" bold="true" box="[845,906,1378,1404]" pageId="33" pageNumber="34">Stem</emphasis>
suberect to pendulous, (16)18–26(30) cm long, (1)1.2–1.4(1.6) mm in diameter,glabrous, leafy throughout,densely covered by imbricate leaf sheaths. 
<emphasis id="B902EAF5FFD5DB1FFB6CFA21FA85FA12" bold="true" box="[1245,1327,1415,1441]" pageId="33" pageNumber="34">Leaves</emphasis>
distichous, spreading, sessile, glabrous, with papyraceous tubular sheaths enveloping stem; leaf blade pale green, narrowly ovate to narrowly triangular oblong, (0.9)1–1.6(1.7) cm long, (4)4.5–7(8) mm wide, shallowly emarginate and minutely apiculate. 
<emphasis id="B902EAF5FFD5DB1FFF4DFA54FE30F9BF" bold="true" box="[252,410,1522,1548]" pageId="33" pageNumber="34">Inflorescence</emphasis>
an axillary spike (often pseudoterminal), 
<quantity id="4C8E9B02FFD5DB1FFCC6FA55FC5FF9BE" box="[887,1013,1523,1550]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="33" pageNumber="34" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 cm</quantity>
long (when fully developed), twice as long as leaves or longer, many-flowered (usually with more than 30 flowers), with flowers developing successively and 2–3(4) of them open at once; peduncle 
<quantity id="4C8E9B02FFD5DB1FFDDAF99DFD62F9E6" box="[619,712,1595,1621]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="33" pageNumber="34" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long, glabrous, covered by 2–4 tubular papyraceous sterile bracts; rachis commonly bent at the base, straight or slightly flexuose, spreading or pendulous, glabrous, densiflorous. 
<emphasis id="B902EAF5FFD5DB1FFAD3F9F9FF79F92E" bold="true" pageId="33" pageNumber="34">Floral bracts</emphasis>
triangular or triangular narrowly ovate, acute, deflexed, (1)1.2–1.6(1.8) mm long, (0.4)0.5–0.7(0.8) mm wide, glabrous, pure white, soon becoming light brownish, papyraceous. 
<emphasis id="B902EAF5FFD5DB1FFCC4F901FC7EF972" bold="true" box="[885,980,1703,1729]" pageId="33" pageNumber="34">Flowers</emphasis>
sessile, white, lip and column with purple spots on sides. 
<emphasis id="B902EAF5FFD5DB1FFE85F96CFE2AF957" bold="true" box="[308,384,1738,1764]" pageId="33" pageNumber="34">Ovary</emphasis>
not resupinate, white to white with greenish tint, terete, distally slightly broadening, (1.5)1.6– 1.9(2) mm long, 
<quantity id="4C8E9B02FFD5DB1FFEE1F949FE70F8BA" box="[336,474,1775,1801]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="33" pageNumber="34" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
in diameter, glabrous. 
<emphasis id="B902EAF5FFD5DB1FFD58F948FC9FF8BB" bold="true" box="[745,821,1774,1800]" pageId="33" pageNumber="34">Sepals</emphasis>
ovate, somewhat cymbiform, blunt to obtuse at apex, (2)2.2–2.7(3) mm long, (1.4)1.6–2(2.4) mm wide, outside sometimes with scattered small brownish scurfy hairs, inside glabrous; lateral sepals slightly oblique, distinctly wider than median sepal, forming almost globular or emarginate mentum 
<quantity id="4C8E9B02FFD5DB1FFF5DF8FDFEF5F8C6" box="[236,351,1883,1909]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="33" pageNumber="34" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
long and wide. 
<emphasis id="B902EAF5FFD5DB1FFDA6F8FDFDF5F8C6" bold="true" box="[535,607,1883,1909]" pageId="33" pageNumber="34">Petals</emphasis>
oblong narrowly obovate, as long as sepals but slightly narrower, blunt or roundish at apex, glabrous. 
<emphasis id="B902EAF5FFD5DB1FFE76F8D8FE5BF82B" bold="true" box="[455,497,1918,1944]" pageId="33" pageNumber="34">Lip</emphasis>
broadly oblong, cymbiform, (2.2)2.4–2.6(2.8) mm long, (1.4)1.6–1.8(2) mm wide, subglobose at base, broadly triangular to blunt or obtuse at apex; basal callus retrorse, flat, almost circular to broadly obovate, with small lateral folds at base. 
<emphasis id="B902EAF5FFD5DB1FFDFBF860FD00F853" bold="true" box="[586,682,1990,2016]" pageId="33" pageNumber="34">Column</emphasis>
stout, 
<quantity id="4C8E9B02FFD5DB1FFD42F861FCD3F852" box="[755,889,1991,2018]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="33" pageNumber="34" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
tall; rostellum broadly conoid, about 
<quantity id="4C8E9B02FFD5DB1FFAA9F861FAC6F852" box="[1304,1388,1991,2017]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="34" unit="mm" value="0.8">0.8 mm</quantity>
long; anther cap bail-shaped, placed behind; column foot 
<quantity id="4C8E9B02FFD5DB1FFD65F84DFCF6F7B6" box="[724,860,2027,2053]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="33" pageNumber="34" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
long, forward directed. 
<emphasis id="B902EAF5FFD5DB1FFBDEF84CFB07F7B7" bold="true" box="[1135,1197,2026,2052]" pageId="33" pageNumber="34">Fruit</emphasis>
a capsule, fusiform to narrowly obovoid, 
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long, with 3 low longitudinal ribs.
</paragraph>
</subSubSection>
<caption id="DF09666FFFD6DB1CFF39F8C8FE24F843" ID-DOI="http://doi.org/10.5281/zenodo.13217451" ID-Zenodo-Dep="13217451" httpUri="https://zenodo.org/record/13217451/files/figure.png" pageId="34" pageNumber="35" startId="34.[136,229,1902,1924]" targetBox="[151,1435,190,1878]" targetPageId="34" targetType="figure">
<paragraph id="8BC936E7FFD6DB1CFF39F8C8FE24F843" blockId="34.[136,1451,1902,2032]" pageId="34" pageNumber="35">
<emphasis id="B902EAF5FFD6DB1CFF39F8C8FD82F837" bold="true" box="[136,552,1902,1924]" pageId="34" pageNumber="35">
FIGURE 17. 
<emphasis id="B902EAF5FFD6DB1CFEA7F8C8FE42F830" bold="true" box="[278,488,1902,1924]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="4C764D64FFD6DB1CFEA7F8C8FE4FF830" box="[278,485,1902,1924]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="spicata">Appendicula spicata</taxonomicName>
.
</emphasis>
A, B.
</emphasis>
Flowering shoots. 
<emphasis id="B902EAF5FFD6DB1CFD5BF8C8FCABF837" bold="true" box="[746,769,1902,1924]" pageId="34" pageNumber="35">C.</emphasis>
Inflorescences. 
<emphasis id="B902EAF5FFD6DB1CFC15F8C9FC16F837" bold="true" box="[932,956,1903,1924]" pageId="34" pageNumber="35">D.</emphasis>
Apical portion of shoot with inflorescence. 
<emphasis id="B902EAF5FFD6DB1CFACBF8C9FA01F837" bold="true" box="[1402,1451,1903,1924]" pageId="34" pageNumber="35">E, F.</emphasis>
Inflorescences, side view. 
<emphasis id="B902EAF5FFD6DB1CFE33F834FE31F81B" bold="true" box="[386,411,1938,1960]" pageId="34" pageNumber="35">G.</emphasis>
Inflorescence, view from apex. 
<emphasis id="B902EAF5FFD6DB1CFD7DF835FD57F81B" bold="true" box="[716,765,1939,1960]" pageId="34" pageNumber="35">H–J.</emphasis>
Flower, half-side, front and side view. 
<emphasis id="B902EAF5FFD6DB1CFBC1F835FB23F81B" bold="true" box="[1136,1161,1939,1960]" pageId="34" pageNumber="35">K.</emphasis>
Column, front view. 
<emphasis id="B902EAF5FFD6DB1CFAE3F835FAC2F81B" bold="true" box="[1362,1384,1939,1960]" pageId="34" pageNumber="35">L.</emphasis>
Young fruit. A–C, G: 
<emphasis id="B902EAF5FFD6DB1CFEA5F811FD88F87F" box="[276,546,1974,1996]" italics="true" pageId="34" pageNumber="35">Averyanov, Maisak, AL1280</emphasis>
, photos by L. Averyanov; D–F, H–K: 
<emphasis id="B902EAF5FFD6DB1CFC20F810FBB6F87F" box="[913,1052,1974,1996]" italics="true" pageId="34" pageNumber="35">Nuraliev 2539</emphasis>
, photos by M. Nuraliev. Photo correction and design by M. Nuraliev.
</paragraph>
</caption>
<subSubSection id="C36C656CFFD7DB1DFF0CFF38FCA6FEB3" pageId="35" pageNumber="36" type="biology_ecology">
<paragraph id="8BC936E7FFD7DB1DFF0CFF38FCA6FEB3" blockId="35.[136,1452,158,2093]" pageId="35" pageNumber="36">
<emphasis id="B902EAF5FFD7DB1DFF0CFF38FE73FF0B" bold="true" box="[189,473,158,184]" pageId="35" pageNumber="36">Ecology and phenology:</emphasis>
—Clustering epiphyte an lithophyte. Primary and old secondary humid evergreen broad-leaved lowland and submontane forests on granite, shale and sandstone at elevations of 
<quantity id="4C8E9B02FFD7DB1DFB31FF64FAA5FF6E" box="[1152,1295,194,221]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="11.5" metricValueMin="2.0" pageId="35" pageNumber="36" unit="m" value="675.0" valueMax="1150.0" valueMin="200.0">200–1150 m</quantity>
a.s.l. Locally common. Flowers in March–June, November–December.
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFD7DB1DFF0CFEACFEAFFBD3" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BC936E7FFD7DB1DFF0CFEACFEB6FE23" blockId="35.[136,1452,158,2093]" pageId="35" pageNumber="36">
<emphasis id="B902EAF5FFD7DB1DFF0CFEACFEF3FE97" bold="true" box="[189,345,266,292]" pageId="35" pageNumber="36">Distribution:</emphasis>
—Endemic to 
<collectingCountry id="F3617677FFD7DB1DFE4AFEACFDF4FE97" box="[507,606,266,292]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
: provinces 
<collectingRegion id="49B2F805FFD7DB1DFD6DFEADFC9DFE97" box="[732,823,266,293]" country="Vietnam" name="Ha Tinh" pageId="35" pageNumber="36">Ha Tinh</collectingRegion>
(Huong Son District), 
<collectingRegion id="49B2F805FFD7DB1DFB86FEACFB14FE97" box="[1079,1214,266,292]" country="Vietnam" name="Quang Binh" pageId="35" pageNumber="36">Quang Binh</collectingRegion>
(Minh Hoa District), Thua Thien Hue (districts 
<taxonomicName id="4C764D64FFD7DB1DFE1AFE88FD7FFEFA" authority=", Huong Thuy, Nam" authorityName="Huong Thuy, Nam" box="[427,725,302,329]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="luoi">A Luoi, Huong Thuy, Nam</taxonomicName>
Dong, Phu Loc), 
<collectingRegion id="49B2F805FFD7DB1DFC2AFE88FB88FEFA" box="[923,1058,302,329]" country="Vietnam" name="Quang Nam" pageId="35" pageNumber="36">Quang Nam</collectingRegion>
(Song Thanh Nature Reserve), 
<collectingRegion id="49B2F805FFD7DB1DFA32FE88FF07FEDF" country="Vietnam" name="Gia Lai" pageId="35" pageNumber="36">Gia Lai</collectingRegion>
(K’Bang District), 
<collectingRegion id="49B2F805FFD7DB1DFE34FEF4FE4FFEDF" box="[389,485,338,364]" country="Vietnam" name="Dak Lak" pageId="35" pageNumber="36">Dak Lak</collectingRegion>
(Chu Yang Sinh Mountains), 
<collectingRegion id="49B2F805FFD7DB1DFC9FFEF4FC04FEDE" box="[814,942,338,365]" country="Vietnam" name="Khanh Hoa" pageId="35" pageNumber="36">Khanh Hoa</collectingRegion>
(Hon Ba Nature Reserve), 
<collectingRegion id="49B2F805FFD7DB1DFB6BFEF4FAF5FEDF" box="[1242,1375,338,364]" country="Vietnam" name="Ninh Thuan" pageId="35" pageNumber="36">Ninh Thuan</collectingRegion>
(Ninh Hai District).
</paragraph>
<paragraph id="8BC936E7FFD7DB1DFF0CFE3CFF6CFE4F" blockId="35.[136,1452,158,2093]" pageId="35" pageNumber="36">
<emphasis id="B902EAF5FFD7DB1DFF0CFE3CFE18FE07" bold="true" box="[189,434,410,436]" pageId="35" pageNumber="36">Conservation status:</emphasis>
—In almost all its known locations, 
<taxonomicName id="4C764D64FFD7DB1DFC8DFE3CFBB5FE07" box="[828,1055,410,436]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="spicata">
<emphasis id="B902EAF5FFD7DB1DFC8DFE3CFBB5FE07" box="[828,1055,410,436]" italics="true" pageId="35" pageNumber="36">Appendicula spicata</emphasis>
</taxonomicName>
was observed to be locally common and even abundant. Accordingly, we preliminarily estimate the conservation status of this species as Least Concern (LC).
</paragraph>
<paragraph id="8BC936E7FFD7DB1DFF0CFDA0FEAFFBD3" blockId="35.[136,1452,158,2093]" pageId="35" pageNumber="36">
<emphasis id="B902EAF5FFD7DB1DFF0CFDA0FEA3FD93" bold="true" box="[189,265,518,544]" pageId="35" pageNumber="36">Notes:</emphasis>
—For a long time, 
<taxonomicName id="4C764D64FFD7DB1DFE41FDA0FD73FD93" box="[496,729,518,544]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="spicata">
<emphasis id="B902EAF5FFD7DB1DFE41FDA0FD73FD93" box="[496,729,518,544]" italics="true" pageId="35" pageNumber="36">Appendicula spicata</emphasis>
</taxonomicName>
has been misidentified in botanical surveys in 
<collectingCountry id="F3617677FFD7DB1DFAB6FDA0FACCFD93" box="[1287,1382,518,544]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
as 
<taxonomicName id="4C764D64FFD7DB1DFA22FDA1FF58FDF7" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="hexandra">
<emphasis id="B902EAF5FFD7DB1DFA22FDA1FF58FDF7" italics="true" pageId="35" pageNumber="36">A. hexandra</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE74B16FFD7DB1DFEB3FD8CFE75FDF6" author="Seidenfaden, G." box="[258,479,554,581]" pageId="35" pageNumber="36" refId="ref34311" refString="Seidenfaden, G. (1975 a) Contributions to a revision of the orchid flora of Cambodia, Laos and Vietnam. Kai Olsen, Fredensborg, 117 pp." type="book" year="1975">
Seidenfaden 1975 
<bibRefCitation id="EFE74B16FFD7DB1DFE63FD8CFE75FDF6" author="Garay, L. A. &amp; Hamer, F. &amp; Siegerist, E. S." box="[466,479,554,581]" pageId="35" pageNumber="36" pagination="139 - 141" refId="ref30664" refString="Garay, L. A., Hamer, F. &amp; Siegerist, E. S. (1992) Beitrage zu kultivierten Bulbophyllum - Arten. Die Orchidee 43: 139 - 141." type="journal article" year="1992">a</bibRefCitation>
</bibRefCitation>
, 1992, 
<bibRefCitation id="EFE74B16FFD7DB1DFD82FD8CFD5BFDF7" author="Averyanov, L. V." box="[563,753,554,580]" pageId="35" pageNumber="36" refId="ref27790" refString="Averyanov, L. V. (1988) Preliminary list of the Vietnamese orchids. Part 1 (genera A - L). Botanical Institute of the Academy of Sciences of the USSR, Leningrad, 247 pp. [in Russian]" type="book" year="1988">Averyanov 1988</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFD4FFD8CFC93FDF7" author="Averyanov, L. V." box="[766,825,554,580]" pageId="35" pageNumber="36" pagination="20 - 185" refId="ref27835" refString="Averyanov, L. V. (1990) Orchidaceae. In: Tzvelev, N. N. (Ed.) Vascular plants synopsis of Vietnamese flora. Vol. 1. Nauka, Leningrad, 20 pp. 20 - 185." type="book chapter" year="1990">1990</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFCF7FD8CFC2BFDF7" author="Averyanov, L. V." box="[838,897,554,580]" pageId="35" pageNumber="36" refId="ref27880" refString="Averyanov, L. V. (1994) Identification guide to Vietnamese orchids (Orchidaceae Juss.). World and Family, St. Petersburg, 432 pp. [in Russian]" type="book" year="1994">1994</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFC3EFD8CFC60FDF7" author="Averyanov, L. V." box="[911,970,554,580]" pageId="35" pageNumber="36" pagination="48 - 89" refId="ref27941" refString="Averyanov, L. V. (2006) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 9 (3): 48 - 89." type="journal article" year="2006">2006</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFC66FD8DFB97FDF7" author="Ho, P. - H." box="[983,1085,554,581]" pageId="35" pageNumber="36" refId="ref31024" refString="Ho, P. - H. (1993) An illustrated flora of Vietnam. 3 (2). Mekong Printing, Montreal, 1176 pp." type="book" year="1993">Ho 1993</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFBFBFD8CFB2FFDF7" author="Ho, P. - H." box="[1098,1157,554,580]" pageId="35" pageNumber="36" refId="ref31053" refString="Ho, P. - H. (2000) An illustrated flora of Vietnam. 3. Nha Xuat Ban Tre, Ho Chi Minh City, 1020 pp." type="book" year="2000">2000</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFB23FD8DFAACFDF7" author="Hop, T." box="[1170,1286,554,581]" pageId="35" pageNumber="36" refId="ref31178" refString="Hop, T. (1998) The orchids of Vietnam. Nha Xuat Ban Nong Nghiep, Ho Chi Minh City, 703 pp." type="book" year="1998">Hop 1998</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFAA3FD8CFEFAFDDB" author="Averyanov, L. V. &amp; Averyanova, A. L." pageId="35" pageNumber="36" pagination="49 - 108" refId="ref28069" refString="Averyanov, L. V. &amp; Averyanova, A. L. (2002) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 5 (4): 49 - 108." type="journal article" year="2002">Averyanov &amp; Averyanova 2002</bibRefCitation>
, 2003). The latter species is known from 
<collectingCountry id="F3617677FFD7DB1DFCABFDE8FC2AFDDB" box="[794,896,590,616]" name="Thailand" pageId="35" pageNumber="36">Thailand</collectingCountry>
, 
<collectingCountry id="F3617677FFD7DB1DFC3AFDE8FBABFDDB" box="[907,1025,590,616]" name="Cambodia" pageId="35" pageNumber="36">Cambodia</collectingCountry>
, 
<collectingCountry id="F3617677FFD7DB1DFBBAFDE8FBC0FDDB" box="[1035,1130,590,616]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
and Sumatra (
<bibRefCitation id="EFE74B16FFD7DB1DFABBFDE8FA0DFDDB" author="Comber, J. B." box="[1290,1447,590,616]" pageId="35" pageNumber="36" refId="ref30035" refString="Comber, J. B. (2001) Orchids of Sumatra. The Royal Botanic Gardens, Kew, 1026 pp." type="book" year="2001">Comber 2001</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFF39FDD4FEE9FD3F" author="Averyanov, L. V." box="[136,323,626,652]" pageId="35" pageNumber="36" pagination="26 - 46" refId="ref28023" refString="Averyanov, L. V. (2013) New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos. Turczaninowia 16 (4): 26 - 46. https: // doi. org / 10.14258 / turczaninowia. 16.4.7" type="journal article" year="2013">Averyanov 2013</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFEFDFDD4FDEEFD3F" author="Averyanov, L. V. &amp; Nguyen, K. S. &amp; Maisak, T. V. &amp; Konstantinov, E. L. &amp; Nguyen, T. H. &amp; Bounphanmy, S." box="[332,580,626,652]" pageId="35" pageNumber="36" pagination="5 - 58" refId="ref28231" refString="Averyanov, L. V., Nguyen, K. S., Maisak, T. V., Konstantinov, E. L., Nguyen, T. H. &amp; Bounphanmy, S. (2016) New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos. Turczaninowia 19 (3): 5 - 58. https: // doi. org / 10.14258 / turczaninowia. 19.3.1" type="journal article" year="2016">
Averyanov 
<emphasis id="B902EAF5FFD7DB1DFE7DFDD5FDA9FD3F" box="[460,515,626,652]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2016
</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFDFCFDD4FCF9FD3F" author="Chantanaorrapint, S." box="[589,851,626,652]" pageId="35" pageNumber="36" pagination="676 - 689" refId="ref29613" refString="Chantanaorrapint, S. (2022) 89. Appendicula. In: Chayamarit, K. &amp; Balslev, H. (Eds.) Flora of Thailand 12 (3). The Forest Herbarium, Royal Forest Department, Bangkok, pp. 676 - 689." type="book chapter" year="2022">Chantanaorrapint 2022</bibRefCitation>
). The two species can therefore occur sympatrically in 
<collectingCountry id="F3617677FFD7DB1DFF39FD30FF41FD03" box="[136,235,662,688]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
. These species share superficially fairly similar flowers and are probably connected by natural hybridization, as soon as the plants with intermediate morphology occur; such plants are difficult for identification, particularly in herbarium collections. Meanwhile, living flowering specimens of the new species readily differ from 
<taxonomicName id="4C764D64FFD7DB1DFAAEFD79FA06FD4B" box="[1311,1452,734,761]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="hexandra">
<emphasis id="B902EAF5FFD7DB1DFAAEFD79FA06FD4B" box="[1311,1452,734,761]" italics="true" pageId="35" pageNumber="36">A. hexandra</emphasis>
</taxonomicName>
(comprehensive descriptions and drawings of which are presented by Seidenfaden 1992 and 
<bibRefCitation id="EFE74B16FFD7DB1DFB11FCA4FA0EFCAF" author="Chantanaorrapint, S." box="[1184,1444,770,796]" pageId="35" pageNumber="36" pagination="676 - 689" refId="ref29613" refString="Chantanaorrapint, S. (2022) 89. Appendicula. In: Chayamarit, K. &amp; Balslev, H. (Eds.) Flora of Thailand 12 (3). The Forest Herbarium, Royal Forest Department, Bangkok, pp. 676 - 689." type="book chapter" year="2022">Chantanaorrapint 2022</bibRefCitation>
) in inflorescence 
<quantity id="4C8E9B02FFD7DB1DFEF0FC80FE14FCF2" box="[321,446,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="35" pageNumber="36" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 cm</quantity>
long, twice as long as the leaves or longer (vs. inflorescence commonly 
<quantity id="4C8E9B02FFD7DB1DFB5FFC80FAC1FCF2" box="[1262,1387,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="35" pageNumber="36" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 cm</quantity>
long, two times shorter than the leaves or shorter), rachis continuously elongating, many-flowered, usually with more than 30 flowers (vs. rachis not continuously elongating, few-flowered, usually with 2–10 flowers), flowers pure white, lip and column with purple spots on sides (vs. flowers uniformly white with a green tint to light greenish), basal callus of lip almost circular to broadly obovate, with small lateral folds at the base (vs. basal callus of lip oblong ovate, without lateral folds at the base). In addition, flowering shoots of 
<taxonomicName id="4C764D64FFD7DB1DFCAFFC7DFC3AFC47" box="[798,912,987,1013]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="spicata">
<emphasis id="B902EAF5FFD7DB1DFCAFFC7DFC3AFC47" box="[798,912,987,1013]" italics="true" pageId="35" pageNumber="36">A. spicata</emphasis>
</taxonomicName>
curiously resemble those of the predominantly Malesian species 
<taxonomicName id="4C764D64FFD7DB1DFEE7FC59FD14FBAB" authority="Blume (1825: 298)" authorityName="Blume" authorityPageNumber="298" authorityYear="1825" box="[342,702,1022,1049]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="pendula">
<emphasis id="B902EAF5FFD7DB1DFEE7FC59FE79FBAB" box="[342,467,1022,1049]" italics="true" pageId="35" pageNumber="36">A. pendula</emphasis>
<bibRefCitation id="EFE74B16FFD7DB1DFE6FFC58FD14FBAB" author="Blume, C. L." box="[478,702,1022,1049]" pageId="35" pageNumber="36" refId="ref29354" refString="Blume, C. L. (1825) Bijdragen tot de flora van Nederlandsch Indie ¨. Ter Lands Drukkerij, Batavia, 1169 pp. https: // doi. org / 10.5962 / bhl. title. 395" type="book" year="1825">Blume (1825: 298)</bibRefCitation>
</taxonomicName>
, which has long pendulous inflorescences (
<bibRefCitation id="EFE74B16FFD7DB1DFB0AFC58FAF7FBAB" author="Comber, J. B." box="[1211,1373,1022,1048]" pageId="35" pageNumber="36" refId="ref30010" refString="Comber, J. B. (1990) Orchids of Java. Bentham-Moxon Trust, Royal Botanic Gardens, Kew, 407 pp." type="book" year="1990">Comber 1990</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFADDFC58FA0DFBAB" author="Comber, J. B." box="[1388,1447,1022,1048]" pageId="35" pageNumber="36" refId="ref30035" refString="Comber, J. B. (2001) Orchids of Sumatra. The Royal Botanic Gardens, Kew, 1026 pp." type="book" year="2001">2001</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFF39FB84FE25FB8F" author="Chantanaorrapint, S." box="[136,399,1058,1084]" pageId="35" pageNumber="36" pagination="676 - 689" refId="ref29613" refString="Chantanaorrapint, S. (2022) 89. Appendicula. In: Chayamarit, K. &amp; Balslev, H. (Eds.) Flora of Thailand 12 (3). The Forest Herbarium, Royal Forest Department, Bangkok, pp. 676 - 689." type="book chapter" year="2022">Chantanaorrapint 2022</bibRefCitation>
). 
<taxonomicName id="4C764D64FFD7DB1DFE12FB84FD23FB8F" box="[419,649,1058,1084]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="spicata">
<emphasis id="B902EAF5FFD7DB1DFE12FB84FD23FB8F" box="[419,649,1058,1084]" italics="true" pageId="35" pageNumber="36">Appendicula spicata</emphasis>
</taxonomicName>
differs, however, from the latter species in having much smaller stems and leaves.
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFD7DB1DFF0CFBCCFC24FAEF" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8BC936E7FFD7DB1DFF0CFBCCFC24FAEF" blockId="35.[136,1452,158,2093]" pageId="35" pageNumber="36">
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<treatmentCitation id="0AD710F6FFD7DB1DFF0CFBCCFD75FB37" author="Guillaumin, A." box="[189,735,1130,1156]" page=")" pageId="35" pageNumber="36" year="1930">
<taxonomicName id="4C764D64FFD7DB1DFF0CFBCCFD75FB37" ID-CoL="FW8X" authority="Guillaumin (1930: 340)" authorityName="Guillaumin" authorityPageNumber="340" authorityYear="1930" box="[189,735,1130,1156]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="annamensis">
<emphasis id="B902EAF5FFD7DB1DFF0CFBCCFE7EFB37" box="[189,468,1130,1156]" italics="true" pageId="35" pageNumber="36">Appendicula annamensis</emphasis>
<bibRefCitation id="EFE74B16FFD7DB1DFE68FBCCFD75FB37" author="Guillaumin, A." box="[473,735,1130,1156]" pageId="35" pageNumber="36" pagination="326 - 340" refId="ref30768" refString="Guillaumin, A. (1930) Especes et localites nouvelles d'Orchidees-Vandees d'Indo-Chine. Bulletin de la Societe Botanique de France 77 (3): 326 - 340. https: // doi. org / 10.1080 / 00378941.1930.10833732" type="journal article" year="1930">Guillaumin (1930: 340)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
and 
<treatmentCitation id="0AD710F6FFD7DB1DFCA3FBCDFB30FB36" author="Guillaumin, A." box="[786,1178,1130,1157]" page=")" pageId="35" pageNumber="36" year="1961">
<taxonomicName id="4C764D64FFD7DB1DFCA3FBCDFB30FB36" ID-CoL="7RPTN" authority="Guillaumin (1961: 562)" authorityName="Guillaumin" authorityPageNumber="562" authorityYear="1961" box="[786,1178,1130,1157]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="inornata">
<emphasis id="B902EAF5FFD7DB1DFCA3FBCDFC25FB37" box="[786,911,1131,1157]" italics="true" pageId="35" pageNumber="36">A. inornata</emphasis>
<bibRefCitation id="EFE74B16FFD7DB1DFC25FBCCFB30FB36" author="Guillaumin, A." box="[916,1178,1130,1157]" pageId="35" pageNumber="36" pagination="562 - 565" refId="ref30857" refString="Guillaumin, A. (1961) Plantes nouvelles, rares ou critiques des serres du Museum (Notules sur quelques Orchidees d'lndochine, XXV). Bulletin du Museum National d'Histoire Naturelle, ser. 2, 32 (6): 562 - 565." type="journal article" year="1961">Guillaumin (1961: 562)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
described from southern 
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were treated by 
<bibRefCitation id="EFE74B16FFD7DB1DFE12FB28FD2FFB1A" author="Seidenfaden, G." box="[419,645,1166,1193]" pageId="35" pageNumber="36" refId="ref34311" refString="Seidenfaden, G. (1975 a) Contributions to a revision of the orchid flora of Cambodia, Laos and Vietnam. Kai Olsen, Fredensborg, 117 pp." type="book" year="1975">Seidenfaden (1975a</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFD20FB28FD70FB1B" author="Seidenfaden, G." box="[657,730,1166,1192]" pageId="35" pageNumber="36" pagination="1 - 216" refId="ref34611" refString="Seidenfaden, G. (1986 b) Orchid genera in Thailand XIII. Thirty-three epidendroid genera. Opera Botanica 89: 1 - 216." type="journal article" year="1986">1986b</bibRefCitation>
, 1992) as synonyms of 
<taxonomicName id="4C764D64FFD7DB1DFC54FB29FBC4FB1B" ID-CoL="682R6" box="[997,1134,1166,1193]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="hexandra">
<emphasis id="B902EAF5FFD7DB1DFC54FB29FBC4FB1B" box="[997,1134,1166,1193]" italics="true" pageId="35" pageNumber="36">A. hexandra</emphasis>
</taxonomicName>
. Based on the studies of the 
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collections and protologues, we agree with his opinion. At the same time, the plants recorded for 
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(S 
<collectingCountry id="F3617677FFD7DB1DFAEEFB14FA0EFB7F" box="[1375,1444,1202,1228]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
) under the name 
<taxonomicName id="4C764D64FFD7DB1DFEF1FB71FE4EFB43" ID-CoL="FW8X" authorityName="Guillaumin" authorityYear="1930" box="[320,484,1239,1265]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="annamensis">
<emphasis id="B902EAF5FFD7DB1DFEF1FB71FE4EFB43" box="[320,484,1239,1265]" italics="true" pageId="35" pageNumber="36">A. annamensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE74B16FFD7DB1DFE45FB70FD35FB43" author="Ormerod, P." box="[500,671,1238,1264]" pageId="35" pageNumber="36" pagination="95 - 101" refId="ref33093" refString="Ormerod, P. (2004) Orchidaceous additions to the floras of China and Taiwan (II). Taiwania 49 (2): 95 - 101. https: // doi. org / 10.6165 / tai. 2004.49 (2). 95" type="journal article" year="2004">Ormerod 2004</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFD1AFB70FCC6FB43" author="Chen, S. C. &amp; Liu, Z. J. &amp; Zhu, G. H. &amp; Lang, K. Y. &amp; Ji, Z. H. &amp; Luo, Y. B. &amp; Jin, X. H. &amp; Cribb, P. J. &amp; Wood, J. J. &amp; Gale, S. W. &amp; Ormerod, P. &amp; Vermeulen, J. J. &amp; Wood, H. P. &amp; Clayton, D. &amp; Bell, A." box="[683,876,1238,1264]" pageId="35" pageNumber="36" pagination="211 - 235" refId="ref29820" refString="Chen, S. C., Liu, Z. J., Zhu, G. H., Lang, K. Y., Ji, Z. H., Luo, Y. B., Jin, X. H., Cribb, P. J., Wood, J. J., Gale, S. W., Ormerod, P., Vermeulen, J. J., Wood, H. P., Clayton, D. &amp; Bell, A. (2009) Orchidaceae. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. (Eds.) Flora of China, vol. 25. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 211 - 235." type="book chapter" year="2009">
Chen 
<emphasis id="B902EAF5FFD7DB1DFD5EFB71FC82FB43" box="[751,808,1238,1264]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2009
</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFCC9FB70FB92FB43" author="Zhou, X. &amp; Cheng, Z. &amp; Liu, Q. &amp; Zhang, J. &amp; Hu, A. &amp; Huang, M. &amp; Hu, C. &amp; Tian, H." box="[888,1080,1238,1264]" pageId="35" pageNumber="36" pagination="1 - 148" refId="ref36068" refString="Zhou, X., Cheng, Z., Liu, Q., Zhang, J., Hu, A., Huang, M., Hu, C. &amp; Tian, H. (2016) An updated checklist of Orchidaceae for China, with two new national records. Phytotaxa 276 (1): 1 - 148. https: // doi. org / 10.11646 / phytotaxa. 276.1.1" type="journal article" year="2016">
Zhou 
<emphasis id="B902EAF5FFD7DB1DFC0DFB71FC5FFB43" box="[956,1013,1238,1264]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2016
</bibRefCitation>
, 
<bibRefCitation id="EFE74B16FFD7DB1DFBF4FB70FB40FB43" author="Jin, X. &amp; Li, J. &amp; Ye, D." box="[1093,1258,1238,1264]" pageId="35" pageNumber="36" refId="ref31426" refString="Jin, X., Li, J. &amp; Ye, D. (2023) Atlas of Chinese native orchids. Springer, Singapore, 787 pp. https: // doi. org / 10.1007 / 978 - 981 - 99 - 4853 - 6" type="book" year="2023">
Jin 
<emphasis id="B902EAF5FFD7DB1DFBDFFB71FB02FB43" box="[1134,1192,1238,1264]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2023
</bibRefCitation>
) possibly belong to 
<taxonomicName id="4C764D64FFD7DB1DFF17FB5DFEB2FAA7" box="[166,280,1275,1301]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="spicata">
<emphasis id="B902EAF5FFD7DB1DFF17FB5DFEB2FAA7" box="[166,280,1275,1301]" italics="true" pageId="35" pageNumber="36">A. spicata</emphasis>
</taxonomicName>
according to their reported characters. The blackish leaves of the dried specimens that are indicated as a diagnostic character of 
<taxonomicName id="4C764D64FFD7DB1DFE15FAB9FDE2FA8B" ID-CoL="FW8X" authorityName="Guillaumin" authorityYear="1930" box="[420,584,1311,1337]" class="Liliopsida" family="Orchidaceae" genus="Appendicula" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="annamensis">
<emphasis id="B902EAF5FFD7DB1DFE15FAB9FDE2FA8B" box="[420,584,1311,1337]" italics="true" pageId="35" pageNumber="36">A. annamensis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFE74B16FFD7DB1DFDC5FAB8FCE8FA8B" author="Chen, S. C. &amp; Liu, Z. J. &amp; Zhu, G. H. &amp; Lang, K. Y. &amp; Ji, Z. H. &amp; Luo, Y. B. &amp; Jin, X. H. &amp; Cribb, P. J. &amp; Wood, J. J. &amp; Gale, S. W. &amp; Ormerod, P. &amp; Vermeulen, J. J. &amp; Wood, H. P. &amp; Clayton, D. &amp; Bell, A." box="[628,834,1310,1336]" pageId="35" pageNumber="36" pagination="211 - 235" refId="ref29820" refString="Chen, S. C., Liu, Z. J., Zhu, G. H., Lang, K. Y., Ji, Z. H., Luo, Y. B., Jin, X. H., Cribb, P. J., Wood, J. J., Gale, S. W., Ormerod, P., Vermeulen, J. J., Wood, H. P., Clayton, D. &amp; Bell, A. (2009) Orchidaceae. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. (Eds.) Flora of China, vol. 25. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 211 - 235." type="book chapter" year="2009">
Chen 
<emphasis id="B902EAF5FFD7DB1DFD07FAB9FD45FA8B" box="[694,751,1310,1336]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
(2009)
</bibRefCitation>
are likely to be a consequence of certain conditions of herbarium preparation, and seem to lack any taxonomic significance.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFD7DB1AFF0CFAC0FE2EF82F" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BC936E7FFD7DB1AFF0CFAC0FE2EF82F" blockId="35.[136,1452,158,2093]" lastBlockId="36.[136,1452,158,1948]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B902EAF5FFD7DB1DFF0CFAC0FCC3FA33" bold="true" box="[189,873,1382,1409]" pageId="35" pageNumber="36">
Additional specimens studied (
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):— 
<collectingCountry id="F3617677FFD7DB1DFD6AFAC0FCCEFA33" box="[731,868,1382,1408]" name="Vietnam" pageId="35" pageNumber="36">VIETNAM</collectingCountry>
.
</emphasis>
<collectingRegion id="49B2F805FFD7DB1DFCC2FAC1FBEEFA33" box="[883,1092,1382,1409]" country="Vietnam" name="Ha Tinh" pageId="35" pageNumber="36">Ha Tinh Province</collectingRegion>
: Huong Son District, Son Kim Municipality, Rao Bun stream, around point 
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<geoCoordinate id="EE425020FFD7DB1DFCABFA2CFC18FA16" box="[794,946,1418,1445]" degrees="105" direction="east" minutes="15" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="12" value="105.253334">105º15’12’’E</geoCoordinate>
, primary broad-leaved evergreen closed wet forest on very steep shale slopes of river canyon at elevation about 
<quantity id="4C8E9B02FFD7DB1DFC14FA08FC45FA7A" box="[933,1007,1454,1481]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="300.0">300 m</quantity>
a.s.l., epiphyte on trees along stream, flowers white, light greenish, lip white-yellowish, with purple side lobes, common, 
<date id="FFC81027FFD7DB1DFBC5FA74FAA1FA5F" box="[1140,1291,1490,1517]" pageId="35" pageNumber="36" value="2004-05-02">2 May 2004</date>
, 
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. 
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. Loc, 
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. Averyanov, 
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. 
<collectionCode id="ED67AE22FFD7DB1DFEAEFA51FE9AF9A3" box="[287,304,1527,1552]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="35" pageNumber="36" type="Museum">V</collectionCode>
. The, 
<collectionCode id="ED67AE22FFD7DB1DFEC2FA51FE23F9A3" box="[371,393,1527,1552]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="35" pageNumber="36" type="Herbarium">N</collectionCode>
. 
<collectionCode id="ED67AE22FFD7DB1DFE3DFA51FE37F9A3" box="[396,413,1527,1552]" name="Tavera, Department of Geology and Geophysics" pageId="35" pageNumber="36">T</collectionCode>
. Vinh, 
<collectionCode id="ED67AE22FFD7DB1DFE58FA51FDB7F9A2" box="[489,541,1527,1553]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15866" lsid="urn:lsid:biocol.org:col:15866" name="Martin-Luther-Universität" pageId="35" pageNumber="36" type="Herbarium">HAL</collectionCode>
5016
</emphasis>
(
<collectionCode id="ED67AE22FFD7DB1DFDDDFA50FD13F9A2" box="[620,697,1526,1553]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="35" pageNumber="36">LE LE</collectionCode>
01054933 https://en.herbariumle.ru/?t=occ&amp;id=5808, 
<collectionCode id="ED67AE22FFD7DB1DFAA8FA50FA91F9A2" box="[1305,1339,1526,1553]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="35" pageNumber="36">LE</collectionCode>
01054945 https://en.herbariumle.ru/?t=occ&amp;id=5820, photos 
<collectionCode id="ED67AE22FFD7DB1DFD68F9BDFC9AF987" box="[729,816,1562,1589]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="35" pageNumber="36">LE LE</collectionCode>
01073293 https://en.herbariumle.ru/?t=occ&amp;id=15903). 
<emphasis id="B902EAF5FFD7DB1DFF39F998FEBDF9EB" bold="true" box="[136,279,1598,1624]" pageId="35" pageNumber="36">
<collectingCountry id="F3617677FFD7DB1DFF39F998FEBBF9EB" box="[136,273,1598,1624]" name="Vietnam" pageId="35" pageNumber="36">VIETNAM</collectingCountry>
.
</emphasis>
<collectingRegion id="49B2F805FFD7DB1DFE92F998FD8FF9EB" box="[291,549,1598,1624]" country="Vietnam" name="Quang Binh" pageId="35" pageNumber="36">Quang Binh Province</collectingRegion>
: Minh Hoa District, Dan Hoa Municipality, Vietnam-Laotian border, around Cha Lo border gate, point 
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<geoCoordinate id="EE425020FFD7DB1DFDE0F9C4FD4DF9CE" box="[593,743,1634,1661]" degrees="105" direction="east" minutes="45" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="54" value="105.765">105°45’54”E</geoCoordinate>
, primary broad-leaved forest on steep rocky mountain slopes composed with ochre-brown shale at elevation 
<quantity id="4C8E9B02FFD7DB1DFD09F920FC97F912" box="[696,829,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="35" pageNumber="36" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300–450 m</quantity>
a.s.l., lithophyte on shady mossy boulders, white to yellowish, very common, 
<date id="FFC81027FFD7DB1DFE1AF90CFDE3F977" box="[427,585,1706,1732]" pageId="35" pageNumber="36" value="2008-04-18">18 April 2008</date>
, 
<emphasis id="B902EAF5FFD7DB1DFDE2F90DFB4AF977" box="[595,1248,1706,1733]" italics="true" pageId="35" pageNumber="36">
<collectionCode id="ED67AE22FFD7DB1DFDE2F90DFDCCF977" box="[595,614,1707,1732]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="35" pageNumber="36" type="Herbarium">L</collectionCode>
. Averyanov, 
<collectionCode id="ED67AE22FFD7DB1DFD43F90DFCAEF977" box="[754,772,1707,1732]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="35" pageNumber="36" type="Herbarium">P</collectionCode>
. 
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. Loc, 
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. Vinh, 
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. Khang, 
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(
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01054883 https://en.herbariumle.ru/?t=occ&amp;id=5759, photos 
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: 
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District, 
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Roang Municipality, Tra Lenh Forestry Department station, around point 
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, primary closed evergreen broad-leaved forest on very steep mountain slopes composed with shale, sandstone and granite at elevation 
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a.s.l., epiphyte on mossy trees along rocky stream valley, flowers white, lip with reddish hypochile, very common, 
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. Averyanov, 
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. 
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. Loc, 
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. Regalado, 
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. 
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. Thao, 
<collectionCode id="ED67AE22FFD7DB1DFE2BF825FE1AF82F" box="[410,432,1923,1948]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="35" pageNumber="36" type="Herbarium">N</collectionCode>
. 
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. Vinh, 
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7270
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(photos 
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01073299 https://en.herbariumle.ru/?t=occ&amp;id=15909). 
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: Huong Thuy District, Duong Hoa Municipality, Huong Thuy forest enterprise territory, around point 
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, mountain and hill slopes composed with shale at elevation of about 
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a.s.l., fractionally logged primary and secondary closed evergreen broad-leaved forest on steep stream slopes, epiphyte and lithophyte along rocky stream valley, flowers white, locally abundant, 
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, 
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. Averyanov, 
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. 
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. Loc, 
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. Thao, 
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. 
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. Vinh, 
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8252
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(
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01054939 https://en.herbariumle.ru/?t=occ&amp;id=5814). 
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<collectingRegion id="49B2F805FFD0DB1AFE90FF64FDF9FF6F" box="[289,595,194,221]" inferredBySuffix="true" pageId="36" pageNumber="37">Thua Thien Hue Province</collectingRegion>
: Nam Dong District, Huong Son Municipality, around point 
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(around plot 
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, 
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), heavily logged primary closed broad-leaved evergreen lowland forest along La Ma River on very steep hill slopes composed with clayey shale, at elevation about 
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a.s.l., epiphyte on mossy trees along stream, very common, 
<date id="FFC81027FFD0DB1AFCF2FE88FC5BFEFA" box="[835,1009,302,329]" pageId="36" pageNumber="37" value="2005-03-26">26 March 2005</date>
, 
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. Averyanov, 
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. 
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. Hiep, 
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. 
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. Loc, 
<collectionCode id="ED67AE22FFD0DB1AFA33FE89FA39FEFB" box="[1410,1427,303,328]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="36" pageNumber="37" type="Herbarium">P</collectionCode>
. 
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. The, 
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. Averyanova, 
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. 
<collectionCode id="ED67AE22FFD0DB1AFE22FEF5FE0EFEDF" box="[403,420,339,364]" name="Tavera, Department of Geology and Geophysics" pageId="36" pageNumber="37">T</collectionCode>
. Vinh, 
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6914
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(
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01054940 https://en.herbariumle.ru/?t=occ&amp;id=5815, photos 
<collectionCode id="ED67AE22FFD0DB1AFA38FEF5FF00FE23" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="36" pageNumber="37">LE LE</collectionCode>
01073300 https://en.herbariumle.ru/?t=occ&amp;id=15910, 
<collectionCode id="ED67AE22FFD0DB1AFCA8FED0FC91FE23" box="[793,827,374,400]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="36" pageNumber="37">LE</collectionCode>
01073302 https://en.herbariumle.ru/?t=occ&amp;id=15912). 
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<collectingRegion id="49B2F805FFD0DB1AFEAFFE3CFDEFFE07" box="[286,581,410,437]" inferredBySuffix="true" pageId="36" pageNumber="37">Thua Thien Hue Province</collectingRegion>
: Nam Dong District, Thuong Lo Municipality, Cha Mang river basin, around point 
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, primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale at elev. 
<quantity id="4C8E9B02FFD0DB1AFE75FE44FDEEFE4E" box="[452,580,482,509]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="36" pageNumber="37" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
a.s.l., epiphyte on mossy trees on very steep rocky shady river slope, flowers white, locally very common, 
<date id="FFC81027FFD0DB1AFE61FDA0FDC4FD93" box="[464,622,518,544]" pageId="36" pageNumber="37" value="2007-04-11">11 April 2007</date>
, 
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. Averyanov, 
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. 
<collectionCode id="ED67AE22FFD0DB1AFC85FDA1FCEEFD92" box="[820,836,519,545]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="36" pageNumber="37" type="Herbarium">S</collectionCode>
. Khang, 
<collectionCode id="ED67AE22FFD0DB1AFC17FDA1FC10FD92" box="[934,954,519,545]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="36" pageNumber="37" type="Herbarium">A</collectionCode>
. Averyanova, 
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.D. Phuong, 
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. 
<collectionCode id="ED67AE22FFD0DB1AFABCFDA1FA8AFD93" box="[1293,1312,519,544]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="37" type="Museum">V</collectionCode>
. Hung, 
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11101
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(
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slope of Bach Ma mt, below Hai Vong Dai peak, around point 16°11’65’’N 
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, at elev. about 
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a.s.l., mixed and broad-leaved evergreen secondary closed mountain forest on steep shale/granite slopes and cliffs, epiphyte and lithophyte on shady waterfall cliff, flowers white with green tint, common, 
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. Averyanov, 
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. 
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: Phu Loc District, Bach Ma National Park, 
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slope of Bach Ma mt system, to the 
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of point 
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, at elev. 
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&amp; 
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along narrow ridge and on very steep sandstone/shale slopes, epiphyte and lithophyte in shady place on waterfall cliffs, flowers light yellowish-green, locally abundant, 
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. Hiep, 
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. Averyanov, 
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. 
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. Vinh et al., HLF 1382
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01054941 https://en.herbariumle.ru/?t=occ&amp;id=5816, photos 
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, elev. 
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. 
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NW of K’Bang Town, forest, river bank, epiphytic, 
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, elev. 
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a.s.l., 
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, 
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(
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01087666 https://en.herbariumle.ru/?t=occ&amp;id=41969; 
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0754803). 
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: Chu Yang Sinh Mountains, 
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, 
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(photos 
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: Chu Yang Sinh Mountains, 
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Nguyen Van Canh, 
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(
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: Krong Bong District, Cu Pui Municipality, elevation from Dak Tour Village to main peak of Chu Yang Sinh mountain system along Dak Tour River (
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slope), primary closed mountain broad-leaved evergreen forest on granite slopes along river canyon at elevation of about 
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a.s.l., epiphyte on mossy trees along river in shady places, flowers white, light yellowish, common, 
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. Hoang, D.X. Du, 
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(
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: Ninh Hai District, Vinh Hai Municipality, NEE slopes on Nui Chua mountains to SW of point 
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, at elev. about 
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. Doan, 
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Su, 
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. Trang, 
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Southern 
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, sine coll., herbarium prepared from wild collected cultivated plants on 
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, 
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. Averyanov, 
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. Maisak, 
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(
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</paragraph>
</subSubSection>
</treatment>
</document>