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<document id="667A3B5CF221DBC46F3DD8F0D7F81F36" ID-DOI="10.5281/zenodo.280870" ID-GBIF-Dataset="59423779-337f-41b0-8993-702ba050a79a" ID-ISSN="1175-5326" ID-Zenodo-Dep="280870" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460476674350" checkinUser="plazi" docAuthor="Matallanas, Jesús &amp; Corbella, Cecília" docDate="2012" docId="03DF87D68D1ED64FFF48E2C1FD90FE60" docLanguage="en" docName="zt03296p018.pdf" docOrigin="Zootaxa 3296" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Patagolycus melastomus Matallanas &amp; Corbella, 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="15" masterDocId="FFE6FFAE8D19D641FFDFE453FF94FFAF" masterDocTitle="Redescription of Iluocoetes Jenyns, 1842; proposal of a new genus, Argentinolycus, for Iluocoetes elongatus (Smitt, 1898), and description of Patagolycus melastomus gen. et sp. nov. (Teleostei, Zoarcidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="8" updateTime="1698306862796" updateUser="plazi">
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<mods:title id="C29A5FB8E63B7A9BE8CD083BA0B464A3">Redescription of Iluocoetes Jenyns, 1842; proposal of a new genus, Argentinolycus, for Iluocoetes elongatus (Smitt, 1898), and description of Patagolycus melastomus gen. et sp. nov. (Teleostei, Zoarcidae)</mods:title>
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<mods:namePart id="867B3F228F3FB6748E52EDFD34947A11">Matallanas, Jesús</mods:namePart>
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<mods:name id="62154876B023BEBB385EE8A4D03EA4D4" type="personal">
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<mods:namePart id="633D1C26468CD77F5E00233BCF9CCE47">Corbella, Cecília</mods:namePart>
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<mods:typeOfResource id="40D392DB126DF325FB996287E516AF8A">text</mods:typeOfResource>
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<mods:date id="39AF95B4D184F81458809A48C685D3F6">2012</mods:date>
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<treatment id="03DF87D68D1ED64FFF48E2C1FD90FE60" ID-DOI="http://doi.org/10.5281/zenodo.6179158" ID-GBIF-Taxon="119516490" ID-Zenodo-Dep="6179158" LSID="urn:lsid:plazi:treatment:03DF87D68D1ED64FFF48E2C1FD90FE60" httpUri="http://treatment.plazi.org/id/03DF87D68D1ED64FFF48E2C1FD90FE60" lastPageId="14" lastPageNumber="15" pageId="7" pageNumber="8">
<subSubSection id="C36C654B8D1ED646FF48E2C1FE4FF960" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BC936C08D1ED646FF48E2C1FD87F904" blockId="7.[151,531,1682,1743]" box="[151,531,1682,1707]" pageId="7" pageNumber="8">
<heading id="D08181AC8D1ED646FF48E2C1FD87F904" bold="true" box="[151,531,1682,1707]" fontSize="10" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="B902EAD28D1ED646FF48E2C1FD87F904" bold="true" box="[151,531,1682,1707]" pageId="7" pageNumber="8">
<taxonomicName id="4C764D438D1ED646FF48E2C1FE26F904" ID-CoL="75SGY" box="[151,434,1682,1707]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="melastomus" status="sp. nov.">
<emphasis id="B902EAD28D1ED646FF48E2C1FE26F904" bold="true" box="[151,434,1682,1707]" italics="true" pageId="7" pageNumber="8">Patagolycus melastomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157A98D1ED646FE66E2C0FD87F904" box="[441,531,1683,1707]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC936C08D1ED646FF48E2E4FE4FF960" blockId="7.[151,531,1682,1743]" box="[151,475,1719,1743]" pageId="7" pageNumber="8">
(
<figureCitation id="134D2A458D1ED646FF40E2E4FE8FF960" box="[159,283,1719,1743]" captionStart-0="FIGURE 3 A" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStart-6="FIGURE 9" captionStart-7="FIGURE 10" captionStartId-0="7.[151,250,964,987]" captionStartId-1="9.[151,250,1476,1499]" captionStartId-2="10.[151,250,855,878]" captionStartId-3="10.[151,250,1719,1742]" captionStartId-4="12.[151,250,922,945]" captionStartId-5="12.[151,250,1541,1564]" captionStartId-6="13.[151,250,1595,1618]" captionStartId-7="14.[151,250,1539,1562]" captionTargetBox-0="[362,1226,704,935]" captionTargetBox-1="[162,1421,211,1450]" captionTargetBox-2="[165,1416,262,819]" captionTargetBox-3="[153,1431,965,1689]" captionTargetBox-4="[151,1419,193,894]" captionTargetBox-5="[164,1414,1070,1518]" captionTargetBox-6="[168,1429,324,1565]" captionTargetBox-7="[427,1167,506,1511]" captionTargetId-0="figure@7.[354,1233,697,943]" captionTargetId-1="figure@9.[151,1436,201,1463]" captionTargetId-2="figure@10.[151,1436,247,835]" captionTargetId-3="figure@10.[151,1436,961,1698]" captionTargetId-4="figure@12.[151,1436,193,901]" captionTargetId-5="figure@12.[151,1436,1056,1520]" captionTargetId-6="figure@13.[168,1453,323,1581]" captionTargetId-7="figure@14.[414,1185,485,1528]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="10" captionTargetPageId-4="12" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="14" captionText-0="FIGURE 3 A. Patagolycus melastomus gen. et sp. nov., UAB. ZM 2 (holotype), 437 mm TL male, from SW Atlantic Ocean. Left lateral view showing general shape and body colour." captionText-1="FIGURE 4. Neurocranium of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; EXO, exoccipital; FR, frontal; INT, intercalar; LAT ETM, lateral ethmoid; MES, mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V, vomer." captionText-2="FIGURE 5. Left lateral view of the lower jaw of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male) with submental crest. ANG, angular; DENT, dentary; RET, retroarticular; SC, submental crest." captionText-3="FIGURE 6. First right gill arch of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male) with scalloped gill rakers." captionText-4="FIGURE 7. Left suspensorium and preopercle (PO) of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). EC, ectopterygoid; HY, hyomandibula; MS, mesopterygoid; MT, metapterygoid; PAL, palatine; QD, quadrate; SY, symplectic; 6, 7 and 8, preopercular pores." captionText-5="FIGURE 8. Left hyoid bar of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). BR, branchiostegal rays; CH, ceratohyal; EH, epihyal; HHD, dorsal hypohyal; HHV, ventral hypohyal; IH, interhyal." captionText-6="FIGURE 9. Left lateral view of the pectoral girdle of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). CL, cleithrum; CO, coracoid; PC, postcleithrum; PT, posttemporal; AC, actinosts (= radials); SCA, scapula; SCL, supracleithrum." captionText-7="FIGURE 10. Interrelationships among the Magellanic endemic lycodine genera. Consensus tree resulting from the parsimony analysis of 78 morphological characters (Length = 105; CI = 0.53; RI = 0.50). Bootstrap support values (rep = 1000) are indicated on the nodes. White boxes on branches: homoplastic apomorphies; black boxes: synapomorphies." httpUri-0="https://zenodo.org/record/280873/files/figure.png" httpUri-1="https://zenodo.org/record/280875/files/figure.png" httpUri-2="https://zenodo.org/record/280876/files/figure.png" httpUri-3="https://zenodo.org/record/280877/files/figure.png" httpUri-4="https://zenodo.org/record/280878/files/figure.png" httpUri-5="https://zenodo.org/record/280879/files/figure.png" httpUri-6="https://zenodo.org/record/280880/files/figure.png" httpUri-7="https://zenodo.org/record/280881/files/figure.png" pageId="7" pageNumber="8">Figs. 310</figureCitation>
; Tables 1, 2 &amp; 4)
</paragraph>
</subSubSection>
<subSubSection id="C36C654B8D1ED646FF48E2ADFE4CF8B7" box="[151,472,1790,1816]" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BC936C08D1ED646FF48E2ADFE4CF8B7" blockId="7.[151,1437,1790,2031]" box="[151,472,1790,1816]" pageId="7" pageNumber="8">
<emphasis id="B902EAD28D1ED646FF48E2ADFE85F8B8" bold="true" box="[151,273,1790,1815]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<collectingCountry id="F36176508D1ED646FEC7E353FEA3F8B7" box="[280,311,1792,1816]" name="American Samoa" pageId="7" pageNumber="8">As</collectingCountry>
for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C36C654B8D1ED64FFF18E371FD9FFE90" lastPageId="14" lastPageNumber="15" pageId="7" pageNumber="8" type="description">
<paragraph id="8BC936C08D1ED646FF18E371FBEFF840" blockId="7.[151,1437,1790,2031]" pageId="7" pageNumber="8">
<emphasis id="B902EAD28D1ED646FF18E371FECDF894" bold="true" box="[199,345,1826,1851]" pageId="7" pageNumber="8">Description.</emphasis>
Body robust, short, ovoid in cross section. Tail laterally compressed especially posteriorly. Head robust, not depressed, as wide as high in both males and females; snout gently sloping. Mouth inferior; lips without lateral lobes. Snout blunt, upper jaw slightly protruding, end of maxilla extending to posterior part of pupil in both males and females. Eyes slightly ellipsoid entering dorsal profile of head. Nasal tube pigmented at base, not reaching upper lip when depressed forward. Gill slit well developed, extending ventrally to below ventral edge of pectoral-fin base, but above pelvic fin insertion. Opercular lobe short, triangular (
<figureCitation id="134D2A458D1ED646FC35E384FBACF840" box="[1002,1080,2007,2031]" captionStart="FIGURE 3 A" captionStartId="7.[151,250,964,987]" captionTargetBox="[362,1226,704,935]" captionTargetId="figure@7.[354,1233,697,943]" captionTargetPageId="7" captionText="FIGURE 3 A. Patagolycus melastomus gen. et sp. nov., UAB. ZM 2 (holotype), 437 mm TL male, from SW Atlantic Ocean. Left lateral view showing general shape and body colour." httpUri="https://zenodo.org/record/280873/files/figure.png" pageId="7" pageNumber="8">Fig. 3a</figureCitation>
&amp;
<figureCitation id="134D2A458D1ED646FB83E384FBF9F840" box="[1116,1133,2007,2031]" captionStart="FIGURE 3 B" captionStartId="7.[151,250,1554,1577]" captionTargetBox="[369,1211,1069,1533]" captionTargetId="figure@7.[356,1233,1067,1535]" captionTargetPageId="7" captionText="FIGURE 3 B. Left lateral view of head and trunk showing cephalic pore pattern, dark band between the anterior edge of the eye and the nostril tube, lateral line, and pectoral-fin shape." httpUri="https://zenodo.org/record/280874/files/figure.png" pageId="7" pageNumber="8">b</figureCitation>
).
</paragraph>
<caption id="DF0966488D10D648FF48E197FCA9F999" httpUri="https://zenodo.org/record/280875/files/figure.png" pageId="9" pageNumber="10" targetBox="[162,1421,211,1450]" targetPageId="9">
<paragraph id="8BC936C08D10D648FF48E197FCA9F999" blockId="9.[151,1436,1476,1590]" pageId="9" pageNumber="10">
<emphasis id="B902EAD28D10D648FF48E197FE9AFA75" bold="true" box="[151,270,1476,1499]" pageId="9" pageNumber="10">FIGURE 4</emphasis>
. Neurocranium of
<taxonomicName id="4C764D438D10D648FE0AE196FD46FA74" box="[469,722,1477,1499]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D10D648FE0AE196FD46FA74" box="[469,722,1477,1499]" italics="true" pageId="9" pageNumber="10">Patagolycus melastomus</emphasis>
</taxonomicName>
gen. et sp. nov. (UAB.P24, 395 mm TL male). Lateral view (
<emphasis id="B902EAD28D10D648FA85E196FAF2FA75" bold="true" box="[1370,1382,1477,1498]" pageId="9" pageNumber="10">a</emphasis>
) and dorsal view (
<emphasis id="B902EAD28D10D648FEC2E1B7FEBFFA56" bold="true" box="[285,299,1508,1529]" pageId="9" pageNumber="10">b</emphasis>
). BAS, basioccipital; EPO, epioccipital; EXO, exoccipital; FR, frontal; INT, intercalar; LAT ETM, lateral ethmoid; MES, mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V, vomer.
</paragraph>
</caption>
<paragraph id="8BC936C08D10D648FF16E230FE1CF8A4" blockId="9.[151,1436,1635,2019]" pageId="9" pageNumber="10">
Cephalic lateralis pore system with small pores. Nasal pores 3, one located anteromesial to nasal tube, two other posterodorsal to the nasal tube and the posterior one very small. Postorbital pores 1 and 4 present in all. Suborbital pores 6+1, in a reversed L-shaped pattern. Preoperculomandibular pores 8 (
<figureCitation id="134D2A458D10D648FBB1E2F8FB52F96C" box="[1134,1222,1707,1731]" captionStart-0="FIGURE 3 A" captionStart-1="FIGURE 3 B" captionStartId-0="7.[151,250,964,987]" captionStartId-1="7.[151,250,1554,1577]" captionTargetBox-0="[362,1226,704,935]" captionTargetBox-1="[369,1211,1069,1533]" captionTargetId-0="figure@7.[354,1233,697,943]" captionTargetId-1="figure@7.[356,1233,1067,1535]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURE 3 A. Patagolycus melastomus gen. et sp. nov., UAB. ZM 2 (holotype), 437 mm TL male, from SW Atlantic Ocean. Left lateral view showing general shape and body colour." captionText-1="FIGURE 3 B. Left lateral view of head and trunk showing cephalic pore pattern, dark band between the anterior edge of the eye and the nostril tube, lateral line, and pectoral-fin shape." httpUri-0="https://zenodo.org/record/280873/files/figure.png" httpUri-1="https://zenodo.org/record/280874/files/figure.png" pageId="9" pageNumber="10">Fig. 3b</figureCitation>
). Interorbital pore absent. Supratemporal commissure and occipital pores absent. Lateral line mediolateral, extending from postorbital pore 4 to near tail tip.
</paragraph>
<paragraph id="8BC936C08D10D64BFF16E344FEA8FF7B" blockId="9.[151,1436,1635,2019]" lastBlockId="10.[151,1435,151,212]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Pectoral-fin origin at body midline; pectoral-fin base extending ventrally to abdomen; posterior margin of pectoral-fin ovoid; middle rays longest; 68 ventralmost rays thickened and exerted at tips (
<figureCitation id="134D2A458D10D648FB4AE368FB73F8FC" box="[1173,1255,1851,1875]" captionStart-0="FIGURE 3 A" captionStart-1="FIGURE 3 B" captionStartId-0="7.[151,250,964,987]" captionStartId-1="7.[151,250,1554,1577]" captionTargetBox-0="[362,1226,704,935]" captionTargetBox-1="[369,1211,1069,1533]" captionTargetId-0="figure@7.[354,1233,697,943]" captionTargetId-1="figure@7.[356,1233,1067,1535]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURE 3 A. Patagolycus melastomus gen. et sp. nov., UAB. ZM 2 (holotype), 437 mm TL male, from SW Atlantic Ocean. Left lateral view showing general shape and body colour." captionText-1="FIGURE 3 B. Left lateral view of head and trunk showing cephalic pore pattern, dark band between the anterior edge of the eye and the nostril tube, lateral line, and pectoral-fin shape." httpUri-0="https://zenodo.org/record/280873/files/figure.png" httpUri-1="https://zenodo.org/record/280874/files/figure.png" pageId="9" pageNumber="10">Fig. 3b</figureCitation>
). Flesh and skin firm; skin covering vertical fins. Scales relatively large, circular, non-overlapping, covering entire body, proximal two thirds of pectoral-fin base and axil, abdomen, tail, nape, posterior part of the interorbital space, cheeks, opercle, and vertical fins to nearly it margin. Two juvenile, BMNH 1936.8.26.984987:(984.2),
<quantity id="4C8E9B258D10D648FB5CE3F4FB76F810" box="[1155,1250,1959,1983]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" pageId="9" pageNumber="10" unit="mm" value="107.0">107 mm</quantity>
TL and BMNH 1936.8.26.984987: (984.3),
<quantity id="4C8E9B258D10D648FE34E398FDD9F84C" box="[491,589,1995,2019]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="9" pageNumber="10" unit="mm" value="103.0">103 mm</quantity>
TL, also have extensive squamation, including nape and upper part of opercle. The
<quantity id="4C8E9B258D13D64BFEEBE4C4FE1CFF00" box="[308,392,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="10" pageNumber="11" unit="mm" value="94.0">94 mm</quantity>
TL specimen, BMNH 1936.8.26.984987:(984.4), has scales throughout its body, but a scaleless head.
</paragraph>
<caption id="DF0966488D13D64BFF48E704FBB4FC23" httpUri="https://zenodo.org/record/280876/files/figure.png" pageId="10" pageNumber="11" targetBox="[165,1416,262,819]" targetPageId="10">
<paragraph id="8BC936C08D13D64BFF48E704FBB4FC23" blockId="10.[151,1436,855,908]" pageId="10" pageNumber="11">
<emphasis id="B902EAD28D13D64BFF48E704FE87FCC2" bold="true" box="[151,275,855,878]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
Left lateral view of the lower jaw of
<taxonomicName id="4C764D438D13D64BFD4CE70BFC1AFCC1" box="[659,910,856,878]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D13D64BFD4CE70BFC1AFCC1" box="[659,910,856,878]" italics="true" pageId="10" pageNumber="11">Patagolycus melastomus</emphasis>
</taxonomicName>
gen. et sp. nov. (UAB.P24, 395 mm TL male) with submental crest. ANG, angular; DENT, dentary; RET, retroarticular; SC, submental crest.
</paragraph>
</caption>
<caption id="DF0966488D13D64BFF48E2E4FF48F943" httpUri="https://zenodo.org/record/280877/files/figure.png" pageId="10" pageNumber="11" targetBox="[153,1431,965,1689]" targetPageId="10">
<paragraph id="8BC936C08D13D64BFF48E2E4FF48F943" blockId="10.[151,1436,1719,1772]" pageId="10" pageNumber="11">
<emphasis id="B902EAD28D13D64BFF48E2E4FE99F963" bold="true" box="[151,269,1719,1742]" pageId="10" pageNumber="11">FIGURE 6</emphasis>
. First right gill arch of
<taxonomicName id="4C764D438D13D64BFE20E2E4FD6FF962" box="[511,763,1719,1741]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D13D64BFE20E2E4FD6FF962" box="[511,763,1719,1741]" italics="true" pageId="10" pageNumber="11">Patagolycus melastomus</emphasis>
</taxonomicName>
gen. et sp. nov. (UAB.P24, 395 mm TL male) with scalloped gill rakers.
</paragraph>
</caption>
<paragraph id="8BC936C08D13D64AFF16E340FF68FF58" blockId="10.[151,1437,1811,2015]" lastBlockId="11.[151,1436,151,247]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Neurocranium well ossified, narrowed (
<figureCitation id="134D2A458D13D64BFD51E340FD46F884" box="[654,722,1811,1835]" captionStart="FIGURE 4" captionStartId="9.[151,250,1476,1499]" captionTargetBox="[162,1421,211,1450]" captionTargetId="figure@9.[151,1436,201,1463]" captionTargetPageId="9" captionText="FIGURE 4. Neurocranium of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; EXO, exoccipital; FR, frontal; INT, intercalar; LAT ETM, lateral ethmoid; MES, mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V, vomer." httpUri="https://zenodo.org/record/280875/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
a &amp; b). Frontal and parasphenoid well separated by pterosphenoid (
<figureCitation id="134D2A458D13D64BFF06E364FE8AF8E0" box="[217,286,1847,1871]" captionStart="FIGURE 4" captionStartId="9.[151,250,1476,1499]" captionTargetBox="[162,1421,211,1450]" captionTargetId="figure@9.[151,1436,201,1463]" captionTargetPageId="9" captionText="FIGURE 4. Neurocranium of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; EXO, exoccipital; FR, frontal; INT, intercalar; LAT ETM, lateral ethmoid; MES, mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V, vomer." httpUri="https://zenodo.org/record/280875/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
a); sphenotic broadly articulating with parietal (
<figureCitation id="134D2A458D13D64BFCE9E364FCE8F8E0" box="[822,892,1847,1871]" captionStart="FIGURE 4" captionStartId="9.[151,250,1476,1499]" captionTargetBox="[162,1421,211,1450]" captionTargetId="figure@9.[151,1436,201,1463]" captionTargetPageId="9" captionText="FIGURE 4. Neurocranium of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; EXO, exoccipital; FR, frontal; INT, intercalar; LAT ETM, lateral ethmoid; MES, mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V, vomer." httpUri="https://zenodo.org/record/280875/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
b). Ascending rami of parasphenoid wing reaching mid-height of the trigeminofacialis foramen. Parasphenoid and prootic juncture, as well as prootic and pterotic juncture strongly interdigitating. Intercalar large, posteriorly set. Frontal ramus long, convex, with an anterior foramen in the interorbital space; frontal corner squared. Anterior portion of frontals fused with no trace of a suture, posterior portion showing a superficial suture; posterolateral edge of the frontals retreat. Supraoccipital wide, with a well developed median crest posteriorly; supraoccipital excluded from exoccipital by epioccipital; no supratemporal commissure across parietals. Ethmoid cartilage protruding well into orbital fenestra, with an anterior foramen.
</paragraph>
<paragraph id="8BC936C08D12D64AFF16E116FDDEF9D2" blockId="11.[151,1437,1349,2022]" pageId="11" pageNumber="12">
Teeth in jaws, vomer and palate conical. Upper jaw with 23 (in males), to 34 (in females) rows near symphysis merging into single posterior row; first premaxillary tooth is canine-like in adult males. Lower jaw with 34 (in males), to 45 (in females) irregular rows in the anterior part and single row in the posterior part; teeth of outermost rows larger than the inner rows ones. In the middle of the lower jaw, a tooth is distinctly enlarged in males (
<figureCitation id="134D2A458D12D64AFF40E186FF71FA42" box="[159,229,1493,1517]" captionStart="FIGURE 5" captionStartId="10.[151,250,855,878]" captionTargetBox="[165,1416,262,819]" captionTargetId="figure@10.[151,1436,247,835]" captionTargetPageId="10" captionText="FIGURE 5. Left lateral view of the lower jaw of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male) with submental crest. ANG, angular; DENT, dentary; RET, retroarticular; SC, submental crest." httpUri="https://zenodo.org/record/280876/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
). A patch of 47 vomerine teeth. Palatine teeth in a single row of 37 teeth. Lower jaw with reduced submental cartilaginous crests, not fused anteriorly (
<figureCitation id="134D2A458D12D64AFD16E1AAFC9AF9BE" box="[713,782,1529,1553]" captionStart="FIGURE 5" captionStartId="10.[151,250,855,878]" captionTargetBox="[165,1416,262,819]" captionTargetId="figure@10.[151,1436,247,835]" captionTargetPageId="10" captionText="FIGURE 5. Left lateral view of the lower jaw of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male) with submental crest. ANG, angular; DENT, dentary; RET, retroarticular; SC, submental crest." httpUri="https://zenodo.org/record/280876/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
). Oral valve nearly reaching anterior edge of vomer and well separated from the palate laterally. Pyloric caeca two small nubs. Gill rakers 23+1113=1315, stout and scalloped (
<figureCitation id="134D2A458D12D64AFECDE212FEC3F9F6" box="[274,343,1601,1625]" captionStart="FIGURE 6" captionStartId="10.[151,250,1719,1742]" captionTargetBox="[153,1431,965,1689]" captionTargetId="figure@10.[151,1436,961,1698]" captionTargetPageId="10" captionText="FIGURE 6. First right gill arch of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male) with scalloped gill rakers." httpUri="https://zenodo.org/record/280877/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
); in specimens preserved for a long periode of time, the indentations are likely to get damaged. Pseudobranch filaments 67, elongate.
</paragraph>
<paragraph id="8BC936C08D12D64AFF16E2DAFC88F89E" blockId="11.[151,1437,1349,2022]" pageId="11" pageNumber="12">
Palatopterygoid series well developed (
<figureCitation id="134D2A458D12D64AFD5BE2DAFD5DF90E" box="[644,713,1673,1697]" captionStart="FIGURE 7" captionStartId="12.[151,250,922,945]" captionTargetBox="[151,1419,193,894]" captionTargetId="figure@12.[151,1436,193,901]" captionTargetPageId="12" captionText="FIGURE 7. Left suspensorium and preopercle (PO) of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). EC, ectopterygoid; HY, hyomandibula; MS, mesopterygoid; MT, metapterygoid; PAL, palatine; QD, quadrate; SY, symplectic; 6, 7 and 8, preopercular pores." httpUri="https://zenodo.org/record/280878/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
), with mesopterygoid overlapping more than half dorsal surface of quadrate and ectopterygoid overlapping half anterior surface of quadrate. Metapterygoid large. Posterior ramus of hyomandibula elongate. Hyoid bar with ceratohyal epihyal joint with bone interdigitating dorsally (
<figureCitation id="134D2A458D12D64AFACEE282FAC1F946" box="[1297,1365,1745,1769]" captionStart="FIGURE 8" captionStartId="12.[151,250,1541,1564]" captionTargetBox="[164,1414,1070,1518]" captionTargetId="figure@12.[151,1436,1056,1520]" captionTargetPageId="12" captionText="FIGURE 8. Left hyoid bar of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). BR, branchiostegal rays; CH, ceratohyal; EH, epihyal; HHD, dorsal hypohyal; HHV, ventral hypohyal; IH, interhyal." httpUri="https://zenodo.org/record/280879/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
). Five branchiostegal rays: first, slender, attached on medial side of ceratohyal; remainder four thickened, 2 articulating on outer side of ceratohyal and 2 on outer side of epihyal.
</paragraph>
<paragraph id="8BC936C08D12D64AFF16E36EFC2FF84A" blockId="11.[151,1437,1349,2022]" pageId="11" pageNumber="12">
Pectoral girdle (
<figureCitation id="134D2A458D12D64AFE5DE36EFE5CF8FA" box="[386,456,1853,1877]" captionStart="FIGURE 9" captionStartId="13.[151,250,1595,1618]" captionTargetBox="[168,1429,324,1565]" captionTargetId="figure@13.[168,1453,323,1581]" captionTargetPageId="13" captionText="FIGURE 9. Left lateral view of the pectoral girdle of Patagolycus melastomus gen. et sp. nov. (UAB. P 24, 395 mm TL male). CL, cleithrum; CO, coracoid; PC, postcleithrum; PT, posttemporal; AC, actinosts (= radials); SCA, scapula; SCL, supracleithrum." httpUri="https://zenodo.org/record/280880/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
) with a strong posttemporal bearing a well-developed ventral ramus. Supracleithrum with a posteriorly-directed prong. Scapular foramen enclosed by bone; prominent postero-dorsal scapular strut. Coracoid with a well-developed posterior strut and a foramen. Radials (=actinosts) 4, the uppermost smaller. Four foramina in the cartilaginous basal plate: one between each two radials, and another between radial 1 and scapula. Postcleithrum present. Pelvic-fin rays joined, ensheathed by the dermis.
</paragraph>
<caption id="DF0966488D15D64DFF48E7C9FE58FC41" httpUri="https://zenodo.org/record/280878/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1419,193,894]" targetPageId="12">
<paragraph id="8BC936C08D15D64DFF48E7C9FE58FC41" blockId="12.[151,1437,922,1006]" pageId="12" pageNumber="13">
<emphasis id="B902EAD28D15D64DFF48E7C9FE86FC1F" bold="true" box="[151,274,922,945]" pageId="12" pageNumber="13">FIGURE 7.</emphasis>
Left suspensorium and preopercle (PO) of
<taxonomicName id="4C764D438D15D64DFD17E7C8FC57FC1E" box="[712,963,923,945]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D15D64DFD17E7C8FC57FC1E" box="[712,963,923,945]" italics="true" pageId="12" pageNumber="13">Patagolycus melastomus</emphasis>
</taxonomicName>
gen. et sp. nov. (UAB.P24, 395 mm TL male). EC, ectopterygoid; HY, hyomandibula; MS, mesopterygoid; MT, metapterygoid; PAL, palatine; QD, quadrate; SY, symplectic; 6, 7 and 8, preopercular pores.
</paragraph>
</caption>
<caption id="DF0966488D15D64DFF48E256FBACF995" httpUri="https://zenodo.org/record/280879/files/figure.png" pageId="12" pageNumber="13" targetBox="[164,1414,1070,1518]" targetPageId="12">
<paragraph id="8BC936C08D15D64DFF48E256FBACF995" blockId="12.[151,1436,1541,1594]" pageId="12" pageNumber="13">
<emphasis id="B902EAD28D15D64DFF48E256FE86F9B4" bold="true" box="[151,274,1541,1564]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
Left hyoid bar of
<taxonomicName id="4C764D438D15D64DFE16E255FD50F9B3" box="[457,708,1542,1564]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D15D64DFE16E255FD50F9B3" box="[457,708,1542,1564]" italics="true" pageId="12" pageNumber="13">Patagolycus melastomus</emphasis>
</taxonomicName>
gen. et sp. nov. (UAB.P24, 395 mm TL male). BR, branchiostegal rays; CH, ceratohyal; EH, epihyal; HHD, dorsal hypohyal; HHV, ventral hypohyal; IH, interhyal.
</paragraph>
</caption>
<paragraph id="8BC936C08D15D64DFF16E234FEBDF8BF" blockId="12.[151,1437,1639,2023]" pageId="12" pageNumber="13">Vertebrae asymmetrical, 2021+6572=8693. Last precaudal vertebra associated with dorsal-fin rays 1820. Dorsal-fin origin associated with vertebra 4 with no free pterygiophores. Dorsal-fin rays 8389. Anal-fin rays 67 73. Terminal dorsal-fin ray associated with second preural vertebra. Terminal anal-fin ray associated with second preural vertebra. One epural. Caudal-fin rays 911, with 12 epural, four upper hypural and four or five lower hypural rays.</paragraph>
<paragraph id="8BC936C08D15D64CFF16E348FE39FEB3" blockId="12.[151,1437,1639,2023]" lastBlockId="13.[151,1436,151,284]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Fresh colouration. Medium to dark brown ground colour with 45 wide and darker vertical cross-bars from dorsal profile to well below body mid-line. Numerous circular and subcircular white spots on head, body, tail and pectoral fins; suborbital area without spots. Dorsal-fin with some slightly elongated white blotches. Edge of vertical fins black; anal-fin edge white in the male
<typeStatus id="54CD88628D15D64DFD41E3D4FC90F830" box="[670,772,1927,1951]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
. Edge of pectoral fins, ventral part of head, opercular edge, ventral fins and abdomen, white or dull white. A darker band across the snout between the anterior edge of the eye and the nasal tube (
<figureCitation id="134D2A458D15D64DFEA5E39CFE42F848" box="[378,470,1999,2023]" captionStart="FIGURE 3 A" captionStartId="7.[151,250,964,987]" captionTargetBox="[362,1226,704,935]" captionTargetId="figure@7.[354,1233,697,943]" captionTargetPageId="7" captionText="FIGURE 3 A. Patagolycus melastomus gen. et sp. nov., UAB. ZM 2 (holotype), 437 mm TL male, from SW Atlantic Ocean. Left lateral view showing general shape and body colour." httpUri="https://zenodo.org/record/280873/files/figure.png" pageId="12" pageNumber="13">Figs. 3a</figureCitation>
&amp;
<figureCitation id="134D2A458D15D64DFE21E39CFD9BF848" box="[510,527,1999,2023]" captionStart="FIGURE 3 B" captionStartId="7.[151,250,1554,1577]" captionTargetBox="[369,1211,1069,1533]" captionTargetId="figure@7.[356,1233,1067,1535]" captionTargetPageId="7" captionText="FIGURE 3 B. Left lateral view of head and trunk showing cephalic pore pattern, dark band between the anterior edge of the eye and the nostril tube, lateral line, and pectoral-fin shape." httpUri="https://zenodo.org/record/280874/files/figure.png" pageId="12" pageNumber="13">b</figureCitation>
). Lips pale; orobranchial cavity and peritoneum black. Coloration in preserved juvenile specimens of
<taxonomicName id="4C764D438D14D64CFE41E4CBFD23FF1F" box="[414,695,152,176]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D14D64CFE41E4CBFD23FF1F" box="[414,695,152,176]" italics="true" pageId="13" pageNumber="14">Patagolycus melastomus</emphasis>
</taxonomicName>
. Dorsolateral part of body mid-brown with dull white rounded spots; ventrolateral part lighter; dorsolateral part of head uniform mid-brown except a darker band on snout between the anterior edge of the eye and the nasal tube; ventrolateral part light. Edge of dorsal fin with 34 darker spots on its anterior half.
</paragraph>
<caption id="DF0966488D14D64CFF48E268FF6BF920" httpUri="https://zenodo.org/record/280880/files/figure.png" pageId="13" pageNumber="14" targetBox="[168,1429,324,1565]" targetPageId="13">
<paragraph id="8BC936C08D14D64CFF48E268FF6BF920" blockId="13.[151,1436,1595,1679]" pageId="13" pageNumber="14">
<emphasis id="B902EAD28D14D64CFF48E268FE87F9FE" bold="true" box="[151,275,1595,1618]" pageId="13" pageNumber="14">FIGURE 9.</emphasis>
Left lateral view of the pectoral girdle of
<taxonomicName id="4C764D438D14D64CFD1FE26FFC2FF9FD" box="[704,955,1596,1618]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D14D64CFD1FE26FFC2FF9FD" box="[704,955,1596,1618]" italics="true" pageId="13" pageNumber="14">Patagolycus melastomus</emphasis>
</taxonomicName>
gen. et sp. nov. (UAB.P24, 395 mm TL male). CL, cleithrum; CO, coracoid; PC, postcleithrum; PT, posttemporal; AC, actinosts (=radials); SCA, scapula; SCL, supracleithrum.
</paragraph>
</caption>
<paragraph id="8BC936C08D14D64CFF16E2EFFDDAF97B" blockId="13.[151,1436,1724,2037]" box="[201,590,1724,1749]" pageId="13" pageNumber="14">
<emphasis id="B902EAD28D14D64CFF16E2EFFE7AF97A" bold="true" box="[201,494,1724,1749]" pageId="13" pageNumber="14">Counts and proportions.</emphasis>
Table 1.
</paragraph>
<paragraph id="8BC936C08D14D64FFF16E28CFD9FFE90" blockId="13.[151,1436,1724,2037]" lastBlockId="14.[151,1436,151,463]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B902EAD28D14D64CFF16E28CFEF9F957" bold="true" box="[201,365,1759,1784]" pageId="13" pageNumber="14">Reproduction</emphasis>
. Despite the reduced sample used, head length, head width, snout length as well as dorsal-fin height, are proportionally higher in the male
<typeStatus id="54CD88628D14D64CFD45E357FD68F8B3" box="[666,764,1796,1820]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(the larger specimen) than in both male and females
<typeStatus id="54CD88628D14D64CFABDE356FF46F8EE" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
indicating both sexual dimorphism and ontogenic change (Table 1). In the single ovary of three female
<typeStatus id="54CD88628D14D64CFABDE37AFF46F8CA" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
two distinct clutches of oocytes can be distinguished: a large clutch of oocytes with a diameter between
<quantity id="4C8E9B258D14D64CFA81E31FFF6BF827" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.42" metricValueMin="1.33" pageId="13" pageNumber="14" unit="mm" value="1.375" valueMax="1.42" valueMin="1.33">1.33 1.42 mm</quantity>
, and another with a diameter of about
<quantity id="4C8E9B258D14D64CFD1EE323FCBEF827" box="[705,810,1904,1928]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="13" pageNumber="14" unit="mm" value="0.72">0.72 mm</quantity>
. The single ovary of
<taxonomicName id="4C764D438D14D64CFBFDE323FAADF827" box="[1058,1337,1904,1928]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="melastomus" status="sp. nov.">
<emphasis id="B902EAD28D14D64CFBFDE323FAADF827" box="[1058,1337,1904,1928]" italics="true" pageId="13" pageNumber="14">Patagolycus melastomus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157A98D14D64CFA9BE322FA08F827" box="[1348,1436,1905,1928]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
belong to the cystovarian
<typeStatus id="54CD88628D14D64CFE62E3C6FE64F802" box="[445,496,1941,1965]" pageId="13" pageNumber="14">type</typeStatus>
, the most common
<typeStatus id="54CD88628D14D64CFD10E3C6FD6BF802" box="[719,767,1941,1965]" pageId="13" pageNumber="14">type</typeStatus>
in Teleosts, according to the classification of
<bibRefCitation id="EFE74B318D14D64CFAD5E3C6FA0CF803" author="Hoar" box="[1290,1432,1940,1964]" pageId="13" pageNumber="14" refString="Hoar, W. S. (1969) Reproduction. In Hoar, W. S. &amp; Randall, D. L. (Eds.), Fish physiology. Academic Press, London, v. 3, pp. 1 - 72." type="journal article" year="1969">Hoar (1969)</bibRefCitation>
. Ovaries in the female specimens of the new species are unripe, they are in a maturing stage, with oocytes visible to the naked eye. It is known that, in general, mature zoarcid eggs are large, about
<quantity id="4C8E9B258D14D64CFBF2E38FFB1AF85B" box="[1069,1166,2012,2036]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="13" pageNumber="14" unit="mm" value="6.5" valueMax="9.0" valueMin="4.0">49 mm</quantity>
in diameter (
<bibRefCitation id="EFE74B318D14D64FFAF9E38FFF44FF00" author="Anderson" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" refString="Anderson, M. E. (1984) Zoarcidae: Development and relationships. In Moser, H. G. (Ed.), Ontogeny and systematics of fishes. American Society of Ichthyologists and Herpetologists, Special Publication, 1, 578 - 582." type="journal article" year="1984">Anderson, 1984</bibRefCitation>
) and those of the Patagonian species confirm this rule:
<bibRefCitation id="EFE74B318D17D64FFC9CE4C4FB9DFF00" author="Gosztonyi" box="[835,1033,151,175]" pageId="14" pageNumber="15" refString="Gosztonyi, A. E. (1977) Results of the research cruises of FRV &quot; Walther Herwig &quot; to South America. XLVIII. Revision of the South American Zoarcidae (Osteichthyes, Blennioidei) with the description of three new genera and five new species. Archiv fur Fischereiwissenschaft, 27, 191 - 249." type="journal article" year="1977">Gosztonyi (1977)</bibRefCitation>
reports 5.0
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in diameter for
<taxonomicName id="4C764D438D17D64FFF48E4EFFE20FF7B" box="[151,436,188,212]" class="Actinopterygii" family="Zoarcidae" genus="Argentinolycus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B902EAD28D17D64FFF48E4EFFE20FF7B" box="[151,436,188,212]" italics="true" pageId="14" pageNumber="15">Argentinolycus elongatus</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C764D438D17D64FFE3CE4EFFD5FFF7B" box="[483,715,188,212]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B902EAD28D17D64FFE3CE4EFFD5FFF7B" box="[483,715,188,212]" italics="true" pageId="14" pageNumber="15">Iluocoetes elongatus</emphasis>
</taxonomicName>
),
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for
<taxonomicName id="4C764D438D17D64FFC48E4EFFB03FF7B" box="[919,1175,188,212]" class="Actinopterygii" family="Zoarcidae" genus="Austrolycus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="laticinctus">
<emphasis id="B902EAD28D17D64FFC48E4EFFB03FF7B" box="[919,1175,188,212]" italics="true" pageId="14" pageNumber="15">Austrolycus laticinctus</emphasis>
</taxonomicName>
, 5.0 mm for
<taxonomicName id="4C764D438D17D64FFAF5E4EFFF7AFF57" class="Actinopterygii" family="Zoarcidae" genus="Dadyanos" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="insignis">
<emphasis id="B902EAD28D17D64FFAF5E4EFFF7AFF57" italics="true" pageId="14" pageNumber="15">Dadyanos insignis</emphasis>
</taxonomicName>
and
<quantity id="4C8E9B258D17D64FFEFAE48CFEEFFF58" box="[293,379,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="15" unit="mm" value="4.5">4.5 mm</quantity>
for
<taxonomicName id="4C764D438D17D64FFE75E4B3FD07FF57" box="[426,659,224,248]" class="Actinopterygii" family="Zoarcidae" genus="Phucocoetes" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="latitans">
<emphasis id="B902EAD28D17D64FFE75E4B3FD07FF57" box="[426,659,224,248]" italics="true" pageId="14" pageNumber="15">Phucocoetes latitans</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFE74B318D17D64FFD7DE48CFC3AFF58" box="[674,942,223,248]" pageId="14" pageNumber="15" refString="Matallanas, J., Rucabado, J., Lloris, D. &amp; Olivar, P. (1990) Early stages of development and reproductive biology of the South- American eelpout Austrolycus depressiceps Regan, 1913 (Teleostei: Zoarcidae). Scientia Marina, 54, 257 - 261." type="journal article">
Matallanas
<emphasis id="B902EAD28D17D64FFCFBE4B2FCC9FF57" box="[804,861,224,248]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
(1990)
</bibRefCitation>
described a demersal egg cluster of
<taxonomicName id="4C764D438D17D64FFA96E4B3FEF1FEB3" class="Actinopterygii" family="Zoarcidae" genus="Austrolycus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="depressiceps">
<emphasis id="B902EAD28D17D64FFA96E4B3FEF1FEB3" italics="true" pageId="14" pageNumber="15">Austrolycus depressiceps</emphasis>
</taxonomicName>
, obtained in the Beagle Channel, formed by 465 spherical eggs with
<quantity id="4C8E9B258D17D64FFBBBE557FB7DFEB3" box="[1124,1257,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="9.8" metricValueMin="9.2" pageId="14" pageNumber="15" unit="mm" value="9.5" valueMax="9.8" valueMin="9.2">9.29.8 mm</quantity>
in diameter, the largest reported size for zoarcids.
</paragraph>
</subSubSection>
<subSubSection id="C36C654B8D17D64FFF16E518FF71FE28" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BC936C08D17D64FFF16E518FF71FE28" blockId="14.[151,1436,151,463]" pageId="14" pageNumber="15">
<emphasis id="B902EAD28D17D64FFF16E518FECFFECB" bold="true" box="[201,347,331,356]" pageId="14" pageNumber="15">Distribution</emphasis>
. Southwest Atlantic Ocean between 45º28
<geoCoordinate id="EE4250078D17D64FFCBBE51FFC29FECB" box="[868,957,332,356]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-51.45">51º27S</geoCoordinate>
and 60º21
<geoCoordinate id="EE4250078D17D64FFBBDE51FFB53FECB" box="[1122,1223,332,356]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-68.833336">68º50W</geoCoordinate>
, at depths of
<quantity id="4C8E9B258D17D64FFABCE51FFF75FE28" metricMagnitude="2" metricUnit="m" metricValue="3.2649999999999997" metricValueMax="4.89" metricValueMin="1.64" pageId="14" pageNumber="15" unit="m" value="326.5" valueMax="489.0" valueMin="164.0">
164
<specimenCount id="9D70FD498D17D64FFF48E53CFF75FE28" box="[151,225,367,391]" pageId="14" pageNumber="15" type="generic">489 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36C654B8D17D64FFF0FE5C7FD90FE60" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BC936C08D17D64FFF0FE5C7FD90FE60" blockId="14.[151,1436,151,463]" pageId="14" pageNumber="15">
<emphasis id="B902EAD28D17D64FFF0FE5C7FEC4FE03" bold="true" box="[208,336,404,428]" pageId="14" pageNumber="15">Etymology</emphasis>
. The specific name,
<taxonomicName id="4C764D438D17D64FFDE1E5C7FD56FE03" box="[574,706,404,428]" class="Actinopterygii" family="Zoarcidae" genus="Patagolycus" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="melastomus">
<emphasis id="B902EAD28D17D64FFDE1E5C7FD56FE03" box="[574,706,404,428]" italics="true" pageId="14" pageNumber="15">melastomus</emphasis>
</taxonomicName>
, from the Greek
<emphasis id="B902EAD28D17D64FFC57E5C7FC5DFE03" box="[904,969,404,428]" italics="true" pageId="14" pageNumber="15">melas</emphasis>
(black, dark) and
<emphasis id="B902EAD28D17D64FFB43E5C6FB4BFE03" box="[1180,1247,405,428]" italics="true" pageId="14" pageNumber="15">stoma</emphasis>
(mouth), for the colour of its orobranchial cavity.
</paragraph>
</subSubSection>
</treatment>
</document>