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<document id="65E97F57D98FEE523F1D96B8A27A95EB" ID-DOI="10.11646/zootaxa.4789.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3887658" ID-ZooBank="688622C6-CD9A-467C-A657-D68FDF40144E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591774606122" checkinUser="plazi" docAuthor="Derafshan, Hossein Ali, Olmi, Massimo, Moghaddam, Mostafa Ghafouri &amp; Rakhshani, Ehsan" docDate="2020" docId="03B9E5513030AA6CFF07FE84FA43DED5" docLanguage="en" docName="zootaxa.4789.2.3.pdf" docOrigin="Zootaxa 4789 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonatopodinae Kieffer 1906" docType="key" docVersion="3" lastPageNumber="373" masterDocId="FF809D293032AA6EFF90FFC9FFDFDD74" masterDocTitle="Taxonomy of Iranian Gonatopodinae (Hymenoptera, Dryinidae), with description of a new species" masterLastPageNumber="402" masterPageNumber="371" pageNumber="373" updateTime="1700276522446" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="AD1B55D7487C2DC1984D07A7B75BA783" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8F3AD3951BAED92314876D1630917F7C">
<mods:title id="E4FE0AB7826273017363DAD9DD209924">Taxonomy of Iranian Gonatopodinae (Hymenoptera, Dryinidae), with description of a new species</mods:title>
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<mods:name id="D9E92752346B0A0C3E088A50063AA0B5" type="personal">
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<mods:namePart id="7214FBE65947A4D0B650E1D027FFD5F3">Derafshan, Hossein Ali</mods:namePart>
<mods:affiliation id="966FDDACF7CEC712C83575280BC16566">Department of Plant Protection, College of Agriculture, University of Zabol, P. O. Box 98615 - 538, I. R. Iran. &amp; Derafshan 69 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3361 - 9697</mods:affiliation>
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<mods:namePart id="F0CB7B7F568E6CFFC9B6B07B5ECB33CC">Olmi, Massimo</mods:namePart>
<mods:affiliation id="E562F3453A0C5088DC3092A34338910E">Tropical Entomology Research Center, Via De Gasperi 10, I- 01100 Viterbo, Italy</mods:affiliation>
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<mods:namePart id="2E19339C14F48A77B60197E62D865A99">Moghaddam, Mostafa Ghafouri</mods:namePart>
<mods:affiliation id="2486ED52389F26E889295ACA53C81A72">Department of Plant Protection, College of Agriculture, University of Zabol, P. O. Box 98615 - 538, I. R. Iran. &amp; ghafourim @ uoz. ac. ir; msc. ghafouri @ gmail. com; https: // orcid. org / 0000 - 0002 - 1942 - 9689</mods:affiliation>
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<mods:namePart id="73BE824523DC568D3358BC92CEDAEB4A">Rakhshani, Ehsan</mods:namePart>
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<mods:title id="043BC2C10B9EEC69574B8DF22E86CA1F">Zootaxa</mods:title>
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<mods:date id="AFCAC39C96B48CCD36CB5B217A09D798">2020</mods:date>
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<mods:number id="6F9E522D1A2C00957C7B412186DAE9E7">2020-06-09</mods:number>
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<mods:number id="EE0872782E8926CA51189C167B8E9AD2">4789</mods:number>
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<mods:classification id="31313A01A674B639E7B9B7DF713D1ED0">journal article</mods:classification>
<mods:identifier id="4ACBCD3BD6AC7012C1FDEE0D6CDB713A" type="DOI">10.11646/zootaxa.4789.2.3</mods:identifier>
<mods:identifier id="513C903BFF5614E2DC4B5DC52485EBF4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B9E5513030AA6CFF07FE84FA43DED5" LSID="urn:lsid:plazi:treatment:03B9E5513030AA6CFF07FE84FA43DED5" httpUri="http://treatment.plazi.org/id/03B9E5513030AA6CFF07FE84FA43DED5" lastPageNumber="373" pageId="2" pageNumber="373">
<subSubSection id="C30A07CC3030AA6CFF07FE84FF31DCDB" pageId="2" pageNumber="373" type="nomenclature">
<paragraph id="8BAF54473030AA6CFF07FE84FD29DC13" blockId="2.[151,758,333,359]" box="[151,758,333,359]" pageId="2" pageNumber="373">
<heading id="D0E7E32B3030AA6CFF07FE84FD29DC13" bold="true" box="[151,758,333,359]" fontSize="11" level="1" pageId="2" pageNumber="373" reason="1">
<emphasis id="B96488553030AA6CFF07FE84FD29DC13" bold="true" box="[151,758,333,359]" pageId="2" pageNumber="373">
Key to species of Iranian
<taxonomicName id="4C102FC43030AA6CFE44FE84FD29DC13" authority="Kieffer" authorityName="Kieffer" authorityYear="1906" box="[468,758,333,359]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="subFamily" subFamily="Gonatopodinae">Gonatopodinae Kieffer</taxonomicName>
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<paragraph id="8BAF54473030AA6CFF07FE5CFF31DCDB" blockId="2.[151,1436,405,929]" box="[151,238,405,431]" pageId="2" pageNumber="373">
<heading id="D0E7E32B3030AA6CFF07FE5CFF31DCDB" bold="true" box="[151,238,405,431]" fontSize="11" level="1" pageId="2" pageNumber="373" reason="1">
<emphasis id="B96488553030AA6CFF07FE5CFF31DCDB" bold="true" box="[151,238,405,431]" pageId="2" pageNumber="373">Female</emphasis>
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<subSubSection id="C30A07CC3030AA6CFF07FE7FFA43DED5" pageId="2" pageNumber="373" type="key">
<keyStep id="30E44CE23030AA6CFF07FE7FFA43DF71" pageId="2" pageNumber="373">
<paragraph id="8BAF54473030AA6CFF07FE7FFA43DC9D" blockId="2.[151,1436,405,929]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FE7FFA43DC9D" pageId="2" pageNumber="373">
1. Pronotum not crossed by transversal furrow or very slightly impressed, palpal formula 2/1.....................................................................................................
<taxonomicName id="4C102FC43030AA6CFBC8FE1AFA43DC9D" authority="Perkins" authorityName="R. Perkins" authorityYear="1905" box="[1112,1436,467,489]" class="Insecta" family="Dryinidae" genus="Haplogonatopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B96488553030AA6CFBC8FE1AFA91DC9D" bold="true" box="[1112,1358,467,489]" italics="true" pageId="2" pageNumber="373">Haplogonatopus apicalis</emphasis>
Perkins
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</keyLead>
</paragraph>
<paragraph id="8BAF54473030AA6CFF07FE39FA43DF71" blockId="2.[151,1436,405,929]" box="[151,1436,496,518]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FE39FA43DF71" box="[151,1436,496,518]" pageId="2" pageNumber="373">
<emphasis id="B96488553030AA6CFF07FE39FF40DF71" bold="true" box="[151,159,496,517]" pageId="2" pageNumber="373">-</emphasis>
Pronotum crossed by transversal furrow (
<figureCitation id="132B48C23030AA6CFDFEFE39FD61DF72" box="[622,702,496,518]" captionStart="FIGURE 6" captionStartId="9.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@9.[151,1436,181,1872]" captionTargetPageId="9" captionText="FIGURE 6. Gonatopus camelinus Kieffer, 1904, from Iran, Zabol-Nimrooz: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887671" httpUri="https://zenodo.org/record/3887671/files/figure.png" pageId="2" pageNumber="373">Figs 6D</figureCitation>
,
<figureCitation id="132B48C23030AA6CFD58FE39FD3DDF71" box="[712,738,496,517]" captionStart="FIGURE 8" captionStartId="11.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@11.[151,1436,181,1872]" captionTargetPageId="11" captionText="FIGURE 8. Gonatopus clavipes (Thunberg, 1827), female from Iran, Razavi Khorasan-Torghabeh: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887675" httpUri="https://zenodo.org/record/3887675/files/figure.png" pageId="2" pageNumber="373">8F</figureCitation>
,
<figureCitation id="132B48C23030AA6CFD7CFE39FCCEDF71" box="[748,785,496,517]" captionStart="FIGURE 10" captionStartId="14.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@14.[151,1436,181,1872]" captionTargetPageId="14" captionText="FIGURE 10. Gonatopus distinguendus Kieffer, 1905, female from Italy, Abruzzo Chieti-Pretoro: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887679" httpUri="https://zenodo.org/record/3887679/files/figure.png" pageId="2" pageNumber="373">10F</figureCitation>
,
<figureCitation id="132B48C23030AA6CFC8BFE39FC9EDF71" box="[795,833,496,517]" captionStart="FIGURE 13" captionStartId="17.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@17.[151,1436,181,1872]" captionTargetPageId="17" captionText="FIGURE 13. Gonatopus lunatus Klug, 1810, female from Iran, Khash-Darre’gol: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887685" httpUri="https://zenodo.org/record/3887685/files/figure.png" pageId="2" pageNumber="373">13F</figureCitation>
,
<figureCitation id="132B48C23030AA6CFCDBFE39FCA9DF72" box="[843,886,496,518]" captionStart="FIGURE 15" captionStartId="19.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@19.[151,1436,181,1872]" captionTargetPageId="19" captionText="FIGURE 15. Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., female paratype: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887689" httpUri="https://zenodo.org/record/3887689/files/figure.png" pageId="2" pageNumber="373">15D</figureCitation>
,
<figureCitation id="132B48C23030AA6CFC10FE39FC77DF72" box="[896,936,496,518]" captionStart="FIGURE 17" captionStartId="21.[151,250,1898,1923]" captionTargetBox="[151,1435,181,1872]" captionTargetId="figure@21.[151,1435,181,1872]" captionTargetPageId="21" captionText="FIGURE 17. Color variation Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., female paratype: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887693" httpUri="https://zenodo.org/record/3887693/files/figure.png" pageId="2" pageNumber="373">17F</figureCitation>
), palpal formula never 2/1...................
<emphasis id="B96488553030AA6CFA00FE39FA43DF71" bold="true" box="[1424,1436,496,517]" pageId="2" pageNumber="373">2</emphasis>
</keyLead>
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<keyStep id="30E44CE23030AA6CFF07FDC7FA43DF34" pageId="2" pageNumber="373">
<paragraph id="8BAF54473030AA6CFF07FDC7FA43DF50" blockId="2.[151,1436,405,929]" box="[151,1436,525,548]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FDC7FA43DF50" box="[151,1436,525,548]" pageId="2" pageNumber="373">
2. Macropterous form (
<figureCitation id="132B48C23030AA6CFE39FDC7FD9ADF57" box="[425,581,526,548]" captionStart="FIGURE 3" captionStartId="5.[151,250,1895,1920]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure@5.[151,1436,181,1870]" captionTargetPageId="5" captionText="FIGURE 3. Echthrodelphax tauricus Ponomarenko, 1970, female from Iran, Zabol-Nimrooz: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Pronotum, dorsal view; F. Forewing; G. Mesonotum and metapectal-propodeal complex, dorsal view; H. Metasoma, lateral view; I. Chela." figureDoi="http://doi.org/10.5281/zenodo.3887665" httpUri="https://zenodo.org/record/3887665/files/figure.png" pageId="2" pageNumber="373">Figs 3D, 3F, 3G</figureCitation>
,
<figureCitation id="132B48C23030AA6CFDDFFDC4FDB2DF57" box="[591,621,525,547]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1947]" captionTargetBox="[151,1436,181,1897]" captionTargetId="figure@7.[151,1436,181,1897]" captionTargetPageId="7" captionText="FIGURE 5. General habitus of Echthrodelphax tauricus Ponomarenko, 1970, from Iran, Zabol-Nimrooz: A. Female; B. Male." figureDoi="http://doi.org/10.5281/zenodo.3887669" httpUri="https://zenodo.org/record/3887669/files/figure.png" pageId="2" pageNumber="373">5A</figureCitation>
)...................................
<taxonomicName id="4C102FC43030AA6CFB8AFDC4FA43DF50" authority="Ponomarenko" authorityName="Ponomarenko" authorityYear="1970" box="[1050,1436,525,548]" class="Insecta" family="Dryinidae" genus="Echthrodelphax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="tauricus">
<emphasis id="B96488553030AA6CFB8AFDC4FACEDF57" bold="true" box="[1050,1297,525,547]" italics="true" pageId="2" pageNumber="373">Echthrodelphax tauricus</emphasis>
Ponomarenko
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF54473030AA6CFF07FDE2FA43DF34" blockId="2.[151,1436,405,929]" box="[151,1436,554,577]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FDE2FA43DF34" box="[151,1436,554,577]" pageId="2" pageNumber="373">
<emphasis id="B96488553030AA6CFF07FDE2FF40DF34" bold="true" box="[151,159,555,576]" pageId="2" pageNumber="373">-</emphasis>
Apterous form (
<figureCitation id="132B48C23030AA6CFEEBFDE2FE65DF34" box="[379,442,555,577]" captionStart="FIGURE 7" captionStartId="10.[151,250,1757,1782]" captionTargetBox="[151,1436,181,1732]" captionTargetId="figure@10.[151,1436,181,1732]" captionTargetPageId="10" captionText="FIGURE 7. General habitus of Gonatopus camelinus Kieffer, 1904, from Iran, Zabol-Nimrooz: A. Female; B. Color variety." figureDoi="http://doi.org/10.5281/zenodo.3887673" httpUri="https://zenodo.org/record/3887673/files/figure.png" pageId="2" pageNumber="373">Figs 7</figureCitation>
,
<figureCitation id="132B48C23030AA6CFE54FDE2FE0DDF34" box="[452,466,555,576]" captionStart="FIGURE 9" captionStartId="12.[151,250,983,1008]" captionTargetBox="[151,1436,181,958]" captionTargetId="figure@12.[151,1436,181,958]" captionTargetPageId="12" captionText="FIGURE 9. General habitus of Gonatopus clavipes (Thunberg, 1827), female from Iran, Razavi Khorasan-Torghabeh." figureDoi="http://doi.org/10.5281/zenodo.3887677" httpUri="https://zenodo.org/record/3887677/files/figure.png" pageId="2" pageNumber="373">9</figureCitation>
,
<figureCitation id="132B48C23030AA6CFE4CFDE2FE2ADF34" box="[476,501,555,576]" captionStart="FIGURE 11" captionStartId="15.[151,250,947,972]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure@15.[151,1436,181,921]" captionTargetPageId="15" captionText="FIGURE 11. General habitus of Gonatopus distinguendus Kieffer, 1905, from Italy, Abruzzo Chieti-Pretoro." figureDoi="http://doi.org/10.5281/zenodo.3887681" httpUri="https://zenodo.org/record/3887681/files/figure.png" pageId="2" pageNumber="373">11</figureCitation>
,
<figureCitation id="132B48C23030AA6CFE6FFDE2FDC6DF34" box="[511,537,555,576]" captionStart="FIGURE 12" captionStartId="15.[151,250,1798,1823]" captionTargetBox="[159,1430,996,1749]" captionTargetId="figure@15.[151,1436,994,1773]" captionTargetPageId="15" captionText="FIGURE 12. General habitus of Gonatopus iranicus Olmi, 1984, female holotype, from Iran, Zanjan-Ziaran." figureDoi="http://doi.org/10.5281/zenodo.3887683" httpUri="https://zenodo.org/record/3887683/files/figure.png" pageId="2" pageNumber="373">12</figureCitation>
,
<figureCitation id="132B48C23030AA6CFDB3FDE2FDE2DF34" box="[547,573,555,576]" captionStart="FIGURE 14" captionStartId="18.[151,250,1122,1147]" captionTargetBox="[151,1436,181,1096]" captionTargetId="figure@18.[151,1436,181,1096]" captionTargetPageId="18" captionText="FIGURE 14. General habitus of Gonatopus lunatus Klug, 1810, female from Iran, Khash-Darre’gol." figureDoi="http://doi.org/10.5281/zenodo.3887687" httpUri="https://zenodo.org/record/3887687/files/figure.png" pageId="2" pageNumber="373">14</figureCitation>
,
<figureCitation id="132B48C23030AA6CFDD7FDE2FDBFDF35" box="[583,608,555,577]" captionStart="FIGURE 16" captionStartId="20.[151,250,1640,1665]" captionTargetBox="[151,1436,181,1614]" captionTargetId="figure@20.[151,1436,181,1614]" captionTargetPageId="20" captionText="FIGURE 16. General habitus of Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., female holotype: A. Dorsal view; B. Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3887691" httpUri="https://zenodo.org/record/3887691/files/figure.png" pageId="2" pageNumber="373">16</figureCitation>
) [
<taxonomicName id="4C102FC43030AA6CFDE5FDE2FD3EDF34" ID-CoL="8VXXM" authorityName="Ljungh" authorityYear="1810" box="[629,737,555,576]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="genus">
<emphasis id="B96488553030AA6CFDE5FDE2FD3EDF34" bold="true" box="[629,737,555,576]" italics="true" pageId="2" pageNumber="373">Gonatopus</emphasis>
</taxonomicName>
]........................................................
<emphasis id="B96488553030AA6CFA00FDE2FA43DF34" bold="true" box="[1424,1436,555,576]" pageId="2" pageNumber="373">3</emphasis>
</keyLead>
</paragraph>
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<keyStep id="30E44CE23030AA6CFF07FD81FA43DF0E" pageId="2" pageNumber="373">
<paragraph id="8BAF54473030AA6CFF07FD81FA43DF2A" blockId="2.[151,1436,405,929]" box="[151,1436,584,606]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FD81FA43DF2A" box="[151,1436,584,606]" pageId="2" pageNumber="373">
3. Enlarged claw with lamellae, without peg-like setae (
<figureCitation id="132B48C23030AA6CFD70FD81FCF6DF2A" box="[736,809,584,606]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[157,1435,963,1910]" captionTargetId="figure@3.[151,1436,957,1916]" captionTargetPageId="3" captionText="FIGURE 1. Redrawn chelae of Gonatopodinae Kieffer (from Olmi, 1984): A. Echthrodelphax tauricus Ponomarenko, 1970, from Sciolze (Italy); B. Gonatopus camelinus Kieffer, 1904, from S. Cruz de la Palma (Canaries); C. Gonatopus clavipes (Thunberg, 1827), from Kuusamo (Russia); D. Gonatopus distinguendus Kieffer, 1905, from El Pardo (Spain)." figureDoi="http://doi.org/10.5281/zenodo.3887660" httpUri="https://zenodo.org/record/3887660/files/figure.png" pageId="2" pageNumber="373">Fig. 1B</figureCitation>
)...........................
<taxonomicName id="4C102FC43030AA6CFBEAFD81FA43DF2A" authority="Kieffer" authorityName="Kieffer" authorityYear="1904" box="[1146,1436,584,606]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="camelinus">
<emphasis id="B96488553030AA6CFBEAFD81FA8EDF2A" bold="true" box="[1146,1361,584,606]" italics="true" pageId="2" pageNumber="373">Gonatopus camelinus</emphasis>
Kieffer
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<keyLead id="30E1F7723030AA6CFF07FDAFFA43DF0E" box="[151,1436,613,636]" pageId="2" pageNumber="373">
- Enlarged claw without lamellae, with setae or peg-like setae (
<figureCitation id="132B48C23030AA6CFCA0FDAFFCB5DF0F" box="[816,874,614,636]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[157,1435,963,1910]" captionTargetId="figure@3.[151,1436,957,1916]" captionTargetPageId="3" captionText="FIGURE 1. Redrawn chelae of Gonatopodinae Kieffer (from Olmi, 1984): A. Echthrodelphax tauricus Ponomarenko, 1970, from Sciolze (Italy); B. Gonatopus camelinus Kieffer, 1904, from S. Cruz de la Palma (Canaries); C. Gonatopus clavipes (Thunberg, 1827), from Kuusamo (Russia); D. Gonatopus distinguendus Kieffer, 1905, from El Pardo (Spain)." figureDoi="http://doi.org/10.5281/zenodo.3887660" httpUri="https://zenodo.org/record/3887660/files/figure.png" pageId="2" pageNumber="373">Figs 1</figureCitation>
C–1D, 2A–C)..................................
<emphasis id="B96488553030AA6CFA00FDACFA43DF0E" bold="true" box="[1424,1436,613,634]" pageId="2" pageNumber="373">4</emphasis>
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<keyStep id="30E44CE23030AA6CFF07FD4AFA43DFC1" pageId="2" pageNumber="373">
<paragraph id="8BAF54473030AA6CFF07FD4AFA43DFEC" blockId="2.[151,1436,405,929]" box="[151,1436,643,665]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FD4AFA43DFEC" box="[151,1436,643,665]" pageId="2" pageNumber="373">
4. Occipital carina very long (
<figureCitation id="132B48C23030AA6CFE78FD4AFD9CDFED" box="[488,579,643,665]" captionStart="FIGURE 15" captionStartId="19.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@19.[151,1436,181,1872]" captionTargetPageId="19" captionText="FIGURE 15. Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., female paratype: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887689" httpUri="https://zenodo.org/record/3887689/files/figure.png" pageId="2" pageNumber="373">Figs 15C</figureCitation>
,
<figureCitation id="132B48C23030AA6CFDDDFD4AFDA7DFED" box="[589,632,643,665]" captionStart="FIGURE 16" captionStartId="20.[151,250,1640,1665]" captionTargetBox="[151,1436,181,1614]" captionTargetId="figure@20.[151,1436,181,1614]" captionTargetPageId="20" captionText="FIGURE 16. General habitus of Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., female holotype: A. Dorsal view; B. Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3887691" httpUri="https://zenodo.org/record/3887691/files/figure.png" pageId="2" pageNumber="373">16A</figureCitation>
,
<figureCitation id="132B48C23030AA6CFD12FD4AFD74DFED" box="[642,683,643,665]" captionStart="FIGURE 17" captionStartId="21.[151,250,1898,1923]" captionTargetBox="[151,1435,181,1872]" captionTargetId="figure@21.[151,1435,181,1872]" captionTargetPageId="21" captionText="FIGURE 17. Color variation Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., female paratype: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887693" httpUri="https://zenodo.org/record/3887693/files/figure.png" pageId="2" pageNumber="373">17C</figureCitation>
).......................
<taxonomicName id="4C102FC43030AA6CFC54FD4AFA96DFED" authority="Rakhshani &amp; Olmi" authorityName="Rakhshani &amp; Olmi" authorityYear="2020" box="[964,1353,643,665]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="opsiicida" status="sp. nov.">
<emphasis id="B96488553030AA6CFC54FD4AFB51DFED" bold="true" box="[964,1166,643,665]" italics="true" pageId="2" pageNumber="373">Gonatopus opsiicida</emphasis>
Rakhshani &amp; Olmi
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<taxonomicNameLabel id="A257352E3030AA6CFAC0FD4AFA43DFEC" box="[1360,1436,643,664]" pageId="2" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF54473030AA6CFF07FD69FA43DFC1" blockId="2.[151,1436,405,929]" box="[151,1436,672,694]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FD69FA43DFC1" box="[151,1436,672,694]" pageId="2" pageNumber="373">
<emphasis id="B96488553030AA6CFF07FD69FF40DFC1" bold="true" box="[151,159,672,693]" pageId="2" pageNumber="373">-</emphasis>
Occipital carina absent or reduced to short tracks on sides of lateral ocelli (
<figureCitation id="132B48C23030AA6CFC20FD69FC21DFC1" box="[944,1022,672,694]" captionStart="FIGURE 8" captionStartId="11.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@11.[151,1436,181,1872]" captionTargetPageId="11" captionText="FIGURE 8. Gonatopus clavipes (Thunberg, 1827), female from Iran, Razavi Khorasan-Torghabeh: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887675" httpUri="https://zenodo.org/record/3887675/files/figure.png" pageId="2" pageNumber="373">Figs 8C</figureCitation>
,
<figureCitation id="132B48C23030AA6CFB98FD69FBEDDFC1" box="[1032,1074,672,693]" captionStart="FIGURE 10" captionStartId="14.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@14.[151,1436,181,1872]" captionTargetPageId="14" captionText="FIGURE 10. Gonatopus distinguendus Kieffer, 1905, female from Italy, Abruzzo Chieti-Pretoro: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887679" httpUri="https://zenodo.org/record/3887679/files/figure.png" pageId="2" pageNumber="373">10C</figureCitation>
,
<figureCitation id="132B48C23030AA6CFBACFD69FB89DFC1" box="[1084,1110,672,693]" captionStart="FIGURE 12" captionStartId="15.[151,250,1798,1823]" captionTargetBox="[159,1430,996,1749]" captionTargetId="figure@15.[151,1436,994,1773]" captionTargetPageId="15" captionText="FIGURE 12. General habitus of Gonatopus iranicus Olmi, 1984, female holotype, from Iran, Zanjan-Ziaran." figureDoi="http://doi.org/10.5281/zenodo.3887683" httpUri="https://zenodo.org/record/3887683/files/figure.png" pageId="2" pageNumber="373">12</figureCitation>
,
<figureCitation id="132B48C23030AA6CFBF0FD69FB56DFC2" box="[1120,1161,672,694]" captionStart="FIGURE 13" captionStartId="17.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@17.[151,1436,181,1872]" captionTargetPageId="17" captionText="FIGURE 13. Gonatopus lunatus Klug, 1810, female from Iran, Khash-Darre’gol: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887685" httpUri="https://zenodo.org/record/3887685/files/figure.png" pageId="2" pageNumber="373">13C</figureCitation>
).....................
<emphasis id="B96488553030AA6CFA00FD69FA43DFC1" bold="true" box="[1424,1436,672,693]" pageId="2" pageNumber="373">5</emphasis>
</keyLead>
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<paragraph id="8BAF54473030AA6CFF07FD74FA43DFA0" blockId="2.[151,1436,405,929]" box="[151,1436,701,724]" pageId="2" pageNumber="373">
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5. Protarsomere 5 with medial lamellae situated on distinct prominence (
<figureCitation id="132B48C23030AA6CFC13FD77FC11DFA0" box="[899,974,702,724]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[157,1435,963,1910]" captionTargetId="figure@3.[151,1436,957,1916]" captionTargetPageId="3" captionText="FIGURE 1. Redrawn chelae of Gonatopodinae Kieffer (from Olmi, 1984): A. Echthrodelphax tauricus Ponomarenko, 1970, from Sciolze (Italy); B. Gonatopus camelinus Kieffer, 1904, from S. Cruz de la Palma (Canaries); C. Gonatopus clavipes (Thunberg, 1827), from Kuusamo (Russia); D. Gonatopus distinguendus Kieffer, 1905, from El Pardo (Spain)." figureDoi="http://doi.org/10.5281/zenodo.3887660" httpUri="https://zenodo.org/record/3887660/files/figure.png" pageId="2" pageNumber="373">Fig. 1D</figureCitation>
)..........
<taxonomicName id="4C102FC43030AA6CFBC5FD77FA43DFA0" authority="Kieffer" authorityName="Kieffer" authorityYear="1905" box="[1109,1436,701,724]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="distinguendus">
<emphasis id="B96488553030AA6CFBC5FD77FA8DDFA7" bold="true" box="[1109,1362,701,723]" italics="true" pageId="2" pageNumber="373">Gonatopus distinguendus</emphasis>
Kieffer
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<paragraph id="8BAF54473030AA6CFF07FD12FA43DF84" blockId="2.[151,1436,405,929]" box="[151,1436,731,753]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FD12FA43DF84" box="[151,1436,731,753]" pageId="2" pageNumber="373">
- Protarsomere 5 with medial lamellae not situated on prominence (
<figureCitation id="132B48C23030AA6CFCCAFD12FC77DF84" box="[858,936,731,753]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[157,1435,963,1910]" captionTargetId="figure@3.[151,1436,957,1916]" captionTargetPageId="3" captionText="FIGURE 1. Redrawn chelae of Gonatopodinae Kieffer (from Olmi, 1984): A. Echthrodelphax tauricus Ponomarenko, 1970, from Sciolze (Italy); B. Gonatopus camelinus Kieffer, 1904, from S. Cruz de la Palma (Canaries); C. Gonatopus clavipes (Thunberg, 1827), from Kuusamo (Russia); D. Gonatopus distinguendus Kieffer, 1905, from El Pardo (Spain)." figureDoi="http://doi.org/10.5281/zenodo.3887660" httpUri="https://zenodo.org/record/3887660/files/figure.png" pageId="2" pageNumber="373">Figs 1C</figureCitation>
,
<figureCitation id="132B48C23030AA6CFC23FD12FC60DF85" box="[947,959,731,753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1576]" captionTargetBox="[160,1435,190,1524]" captionTargetId="figure@4.[151,1436,181,1526]" captionTargetPageId="4" captionText="FIGURE 2. Redrawn chelae of Gonatopodinae (from Olmi, 1984): A. Gonatopus iranicus Olmi, 1984, holotype; B. Gonatopus lunatus Klug, 1810, from Fontainebleau (France); C. Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., holotype; D. Haplogonatopus apicalis Perkins, 1905, from Kalugomuwa (Sri Lanka)." figureDoi="http://doi.org/10.5281/zenodo.3887662" httpUri="https://zenodo.org/record/3887662/files/figure.png" pageId="2" pageNumber="373">2</figureCitation>
A–2B).................................
<emphasis id="B96488553030AA6CFA00FD12FA43DF84" bold="true" box="[1424,1436,731,752]" pageId="2" pageNumber="373">6</emphasis>
</keyLead>
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<keyStep id="30E44CE23030AA6CFF07FD31FA43DE11" pageId="2" pageNumber="373">
<paragraph id="8BAF54473030AA6CFF07FD31FA43DE58" blockId="2.[151,1436,405,929]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FD31FA43DE58" pageId="2" pageNumber="373">
6. Protarsomere 5 with two rows of numerous lamellae both situated in the proximal and distal half (
<figureCitation id="132B48C23030AA6CFB1EFD31FB05DE7A" box="[1166,1242,760,782]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1576]" captionTargetBox="[160,1435,190,1524]" captionTargetId="figure@4.[151,1436,181,1526]" captionTargetPageId="4" captionText="FIGURE 2. Redrawn chelae of Gonatopodinae (from Olmi, 1984): A. Gonatopus iranicus Olmi, 1984, holotype; B. Gonatopus lunatus Klug, 1810, from Fontainebleau (France); C. Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., holotype; D. Haplogonatopus apicalis Perkins, 1905, from Kalugomuwa (Sri Lanka)." figureDoi="http://doi.org/10.5281/zenodo.3887662" httpUri="https://zenodo.org/record/3887662/files/figure.png" pageId="2" pageNumber="373">Fig. 2A</figureCitation>
)..............................................................................................
<taxonomicName id="4C102FC43030AA6CFB31FCDFFA43DE58" authority="Olmi" authorityName="Olmi" authorityYear="1984" box="[1185,1436,790,812]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="iranicus">
<emphasis id="B96488553030AA6CFB31FCDFFABADE5F" bold="true" box="[1185,1381,790,811]" italics="true" pageId="2" pageNumber="373">Gonatopus iranicus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAF54473030AA6CFF07FCFAFA43DE11" blockId="2.[151,1436,405,929]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FCFAFA43DE11" pageId="2" pageNumber="373">
<emphasis id="B96488553030AA6CFF07FCFAFF40DE3C" bold="true" box="[151,159,819,840]" pageId="2" pageNumber="373">-</emphasis>
Protarsomere 5 with one row of lamellae; occasionally with two rows of lamellae, then one row very short and situated in proximal half (
<figureCitation id="132B48C23030AA6CFEE1FC99FE60DE11" box="[369,447,848,870]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1966]" captionTargetBox="[157,1435,963,1910]" captionTargetId="figure@3.[151,1436,957,1916]" captionTargetPageId="3" captionText="FIGURE 1. Redrawn chelae of Gonatopodinae Kieffer (from Olmi, 1984): A. Echthrodelphax tauricus Ponomarenko, 1970, from Sciolze (Italy); B. Gonatopus camelinus Kieffer, 1904, from S. Cruz de la Palma (Canaries); C. Gonatopus clavipes (Thunberg, 1827), from Kuusamo (Russia); D. Gonatopus distinguendus Kieffer, 1905, from El Pardo (Spain)." figureDoi="http://doi.org/10.5281/zenodo.3887660" httpUri="https://zenodo.org/record/3887660/files/figure.png" pageId="2" pageNumber="373">Figs 1C</figureCitation>
,
<figureCitation id="132B48C23030AA6CFE59FC99FE39DE12" box="[457,486,848,870]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1576]" captionTargetBox="[160,1435,190,1524]" captionTargetId="figure@4.[151,1436,181,1526]" captionTargetPageId="4" captionText="FIGURE 2. Redrawn chelae of Gonatopodinae (from Olmi, 1984): A. Gonatopus iranicus Olmi, 1984, holotype; B. Gonatopus lunatus Klug, 1810, from Fontainebleau (France); C. Gonatopus opsiicida Rakhshani &amp; Olmi sp. nov., holotype; D. Haplogonatopus apicalis Perkins, 1905, from Kalugomuwa (Sri Lanka)." figureDoi="http://doi.org/10.5281/zenodo.3887662" httpUri="https://zenodo.org/record/3887662/files/figure.png" pageId="2" pageNumber="373">2B</figureCitation>
).............................................................................
<emphasis id="B96488553030AA6CFA00FC99FA43DE11" bold="true" box="[1424,1436,848,869]" pageId="2" pageNumber="373">7</emphasis>
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<keyStep id="30E44CE23030AA6CFF07FCA7FA43DED5" pageId="2" pageNumber="373">
<paragraph id="8BAF54473030AA6CFF07FCA7FA43DEF0" blockId="2.[151,1436,405,929]" box="[151,1436,877,900]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FCA7FA43DEF0" box="[151,1436,877,900]" pageId="2" pageNumber="373">
7. Mesoscutum very slender, with two lateral pointed apophyses (
<figureCitation id="132B48C23030AA6CFCAFFCA7FC4ADEF7" box="[831,917,878,900]" captionStart="FIGURE 13" captionStartId="17.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@17.[151,1436,181,1872]" captionTargetPageId="17" captionText="FIGURE 13. Gonatopus lunatus Klug, 1810, female from Iran, Khash-Darre’gol: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887685" httpUri="https://zenodo.org/record/3887685/files/figure.png" pageId="2" pageNumber="373">Figs 13F</figureCitation>
,
<figureCitation id="132B48C23030AA6CFC0FFCA7FC67DEF0" box="[927,952,878,900]" captionStart="FIGURE 14" captionStartId="18.[151,250,1122,1147]" captionTargetBox="[151,1436,181,1096]" captionTargetId="figure@18.[151,1436,181,1096]" captionTargetPageId="18" captionText="FIGURE 14. General habitus of Gonatopus lunatus Klug, 1810, female from Iran, Khash-Darre’gol." figureDoi="http://doi.org/10.5281/zenodo.3887687" httpUri="https://zenodo.org/record/3887687/files/figure.png" pageId="2" pageNumber="373">14</figureCitation>
)...................
<taxonomicName id="4C102FC43030AA6CFB39FCA7FA43DEF0" authority="Klug" authorityName="Klug" box="[1193,1436,877,900]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="lunatus">
<emphasis id="B96488553030AA6CFB39FCA7FAB9DEF7" bold="true" box="[1193,1382,877,899]" italics="true" pageId="2" pageNumber="373">Gonatopus lunatus</emphasis>
Klug
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<paragraph id="8BAF54473030AA6CFF07FC42FA43DED5" blockId="2.[151,1436,405,929]" box="[151,1436,907,929]" pageId="2" pageNumber="373">
<keyLead id="30E1F7723030AA6CFF07FC42FA43DED5" box="[151,1436,907,929]" pageId="2" pageNumber="373">
- Mesoscutum wide and hump, without lateral pointed apophyses (
<figureCitation id="132B48C23030AA6CFCC3FC42FC19DED4" box="[851,966,907,929]" captionStart="FIGURE 8" captionStartId="11.[151,250,1898,1923]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure@11.[151,1436,181,1872]" captionTargetPageId="11" captionText="FIGURE 8. Gonatopus clavipes (Thunberg, 1827), female from Iran, Razavi Khorasan-Torghabeh: A. Head, lateral view; B. Head, frontal view; C. Head, dorsal view; D. Mesosoma, lateral view; E. Chela, F. Mesosoma, dorsal view; G. Metasoma, lateral view; H. Metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3887675" httpUri="https://zenodo.org/record/3887675/files/figure.png" pageId="2" pageNumber="373">Figs 8D, 8F</figureCitation>
,
<figureCitation id="132B48C23030AA6CFC40FC42FC02DED5" box="[976,989,907,929]" captionStart="FIGURE 9" captionStartId="12.[151,250,983,1008]" captionTargetBox="[151,1436,181,958]" captionTargetId="figure@12.[151,1436,181,958]" captionTargetPageId="12" captionText="FIGURE 9. General habitus of Gonatopus clavipes (Thunberg, 1827), female from Iran, Razavi Khorasan-Torghabeh." figureDoi="http://doi.org/10.5281/zenodo.3887677" httpUri="https://zenodo.org/record/3887677/files/figure.png" pageId="2" pageNumber="373">9</figureCitation>
)..........
<taxonomicName id="4C102FC43030AA6CFBF9FC42FA43DED5" authority="(Thunberg)" baseAuthorityName="Thunberg" baseAuthorityYear="1827" box="[1129,1436,907,929]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="373" phylum="Tracheophyta" rank="species" species="clavipes">
<emphasis id="B96488553030AA6CFBF9FC42FAF5DED5" bold="true" box="[1129,1322,907,929]" italics="true" pageId="2" pageNumber="373">Gonatopus clavipes</emphasis>
(Thunberg)
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</keyStep>
</subSubSection>
</treatment>
</document>
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