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<document id="8D6A82C56F05AF616541A9F87D1BC29E" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56790C36517780A3FCC14DE8" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola pontica" docType="treatment" docVersion="14" lastPageNumber="18" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="15" updateTime="1698518020641" updateUser="ExternalLinkService">
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<mods:title id="7B324B08DEE25C6EFE3E1336DA63A048">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="A52F1BFAC3E4AC1F1971BEEB8A3A790D">Kunt, Kadir B.</mods:namePart>
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<paragraph id="8BAF055A56790C29517780A3FC324DCE" blockId="14.[310,1277,1456,1520]" box="[587,1000,1456,1483]" pageId="14" pageNumber="15">
<heading id="D0E7B23656790C29517780A3FC324DCE" box="[587,1000,1456,1483]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C107ED956790C29517780A3FC324DCE" ID-CoL="6X4WX" authority="(Spassky, 1937)" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[587,1000,1456,1483]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856790C29517780A3FCFC4DCE" bold="true" box="[587,806,1456,1483]" italics="true" pageId="14" pageNumber="15">Raveniola pontica</emphasis>
(
<bibRefCitation id="EF8178AB56790C29500980A2FC3A4DCE" author="Spassky S. A." box="[821,992,1457,1483]" journalOrPublisher="Festschrift Embrik Strand" pageId="14" pageNumber="15" pagination="361 - 368" part="3" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." title="Araneae palaearcticae novae. Mygalomorphae. I" type="journal article" year="1937">Spassky, 1937</bibRefCitation>
)
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<subSubSection id="C30A56D156790C29520A80C6FB274DF5" box="[310,1277,1493,1520]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BAF055A56790C29520A80C6FB274DF5" blockId="14.[310,1277,1456,1520]" box="[310,1277,1493,1520]" pageId="14" pageNumber="15">
<figureCitation id="132B19DF56790C29520A80C6FE444DF5" box="[310,414,1493,1520]" captionStart="Figs 1–6" captionStartId="50.[189,241,1734,1760]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@50.[189,1399,265,1680]" captionText="Figs 1–6.Raveniola spp., ♀♀ (1–3, 5–6) and ♂ (4). Chelicerae (1), apical edge of maxilla (2–3), palpal organ (4) and tarsus I (5–6); ventroapical (1), ventral (2–4, 6) and dorsal (5) views. — 1–2. R. micropa (Ausserer, 1871). 3–5. R. pontica (Spassky, 1937). 6. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163424/files/figure.png" pageId="14" pageNumber="15">Figs 3–5</figureCitation>
, 
<figureCitation id="132B19DF56790C29529780C5FE114DF5" box="[427,459,1494,1520]" captionStart="Figs 7–12" captionStartId="51.[189,241,1719,1745]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@51.[189,1399,265,1680]" captionText="Figs 7–12. Raveniola spp., ♀♀. Tarsal organ, legs I (7–8, 10, 12) and IV (9, 11), dorsal views. — 7–9. R. hyrcanica Dunin, 1988. 10. R. pontica (Spassky, 1937). 11. R. micropa (Ausserer, 1871). 12. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163426/files/figure.png" pageId="14" pageNumber="15">10</figureCitation>
, 
<figureCitation id="132B19DF56790C2952EB80C6FE2C4DF5" box="[471,502,1493,1520]" captionStart="Figs 13–18" captionStartId="52.[189,241,1725,1751]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@52.[189,1399,265,1680]" captionText="Figs 13–18. Raveniola spp., ♀♀. Bases of trichobothria, legs I (13, 15–16, 18) and IV (14, 17), dorsal views.—13–14.R.hyrcanicaDunin,1988.15.R.pontica(Spassky,1937).16–17.R.micropa(Ausserer, 1871). 18. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163428/files/figure.png" pageId="14" pageNumber="15">15</figureCitation>
, 
<figureCitation id="132B19DF56790C29513F80C6FDF94DEA" box="[515,547,1493,1519]" captionStart="Figs 25–30" captionStartId="54.[189,241,1739,1765]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@54.[189,1399,265,1680]" captionText="Figs 25–30. Raveniolaspp.,♀♀. Posteriormedian spinnerets (25–27),detailsof spigots (28–30);apical (25–26,28–30)andventral(27)views.—25,29–30.R.hyrcanicaDunin,1988.26.R.pontica(Spassky, 1937). 27–28. R. zaitzevi (Charitonov, 1948)." httpUri="https://zenodo.org/record/1163432/files/figure.png" pageId="14" pageNumber="15">26</figureCitation>
, 
<figureCitation id="132B19DF56790C29511380C5FDA14DF5" box="[559,635,1494,1520]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="14" pageNumber="15">48–49</figureCitation>
, 
<figureCitation id="132B19DF56790C2951BB80C6FD094DF5" box="[647,723,1493,1520]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="14" pageNumber="15">59–60</figureCitation>
, 
<figureCitation id="132B19DF56790C2951E380C5FD254DF5" box="[735,767,1494,1520]" captionStart="Figs 68–73" captionStartId="60.[189,241,1882,1908]" captionTargetBox="[190,1398,265,1816]" captionTargetId="figure@60.[189,1398,265,1816]" captionText="Figs 68–73. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 68. R. adjarica sp. nov. (holotype). 69. R. dunini sp. nov. (holotype). 70. R. hyrcanica Dunin, 1988 (holotype). 71. R. pontica (Spassky, 1937) (Lazarevskoe). 72. R. birecikensis sp. nov. (holotype). 73. Same species as preceding (paratype, Birecik). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163444/files/figure.png" pageId="14" pageNumber="15">71</figureCitation>
, 
<figureCitation id="132B19DF56790C29503780C5FCF14DF5" box="[779,811,1494,1520]" captionStart="Figs 84–89" captionStartId="63.[189,241,1892,1918]" captionTargetBox="[189,1398,266,1873]" captionTargetId="figure@63.[189,1398,265,1878]" captionText="Figs 84–89. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163450/files/figure.png" pageId="14" pageNumber="15">87</figureCitation>
, 
<figureCitation id="132B19DF56790C29500B80C6FCBF4DF5" box="[823,869,1493,1520]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="14" pageNumber="15">105</figureCitation>
, 
<figureCitation id="132B19DF56790C29504D80C5FC7A4DF5" box="[881,928,1494,1520]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="14" pageNumber="15">120</figureCitation>
, 
<figureCitation id="132B19DF56790C29509080C5FC014DF5" box="[940,987,1494,1520]" captionStart="Figs129–132" captionStartId="68.[189,241,1870,1896]" captionTargetBox="[220,1291,268,1789]" captionTargetId="figure@68.[189,1323,265,1791]" captionText="Figs129–132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." httpUri="https://zenodo.org/record/1163460/files/figure.png" pageId="14" pageNumber="15">132</figureCitation>
, 
<figureCitation id="132B19DF56790C2950DB80C6FBCC4DF5" box="[999,1046,1493,1520]" captionStart="Figs 149–152" captionStartId="73.[189,241,1930,1956]" captionTargetBox="[194,1398,266,1866]" captionTargetId="figure@73.[189,1398,265,1875]" captionText="Figs 149–152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163470/files/figure.png" pageId="14" pageNumber="15">152</figureCitation>
, 
<figureCitation id="132B19DF56790C29571D80C5FB514DF5" box="[1057,1163,1494,1520]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="14" pageNumber="15">178–179</figureCitation>
, 
<figureCitation id="132B19DF56790C2957AB80C5FB1C4DF5" box="[1175,1222,1494,1520]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="14" pageNumber="15">202</figureCitation>
, 
<figureCitation id="132B19DF56790C2957ED80C5FB274DF5" box="[1233,1277,1494,1520]" captionStart="Figure 219" captionStartId="85.[189,272,1534,1560]" captionTargetBox="[189,1398,265,1471]" captionTargetId="figure@85.[189,1398,265,1471]" captionText="Figure 219. Localities of Raveniola spp. in the western part of the Caucasus: R. adjarica sp. nov. () and R. pontica (Spassky, 1937) ()." httpUri="https://zenodo.org/record/1163494/files/figure.png" pageId="14" pageNumber="15">219</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156790C295381830BFAAC4F0C" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BAF055A56790C295381830BFD164E36" blockId="14.[189,716,1560,1587]" box="[189,716,1560,1587]" pageId="14" pageNumber="15">
<taxonomicName id="4C107ED956790C295381830BFD4E4E37" authority="Spassky, 1937: 368" authorityName="Spassky" authorityPageNumber="368" authorityYear="1937" box="[189,660,1560,1587]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856790C295381830BFE704E37" box="[189,426,1560,1586]" italics="true" pageId="14" pageNumber="15">Brachythele pontica</emphasis>
<bibRefCitation id="EF8178AB56790C29528D830BFD874E36" author="Spassky S. A." box="[433,605,1560,1587]" journalOrPublisher="Festschrift Embrik Strand" pageId="14" pageNumber="15" pagination="361 - 368" part="3" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." title="Araneae palaearcticae novae. Mygalomorphae. I" type="journal article" year="1937">Spassky, 1937</bibRefCitation>
: 368
</taxonomicName>
(♀).
</paragraph>
<paragraph id="8BAF055A56790C295381834FFE0C4E9E" blockId="14.[189,1399,1627,1801]" pageId="14" pageNumber="15">
<taxonomicName id="4C107ED956790C295381834FFE734E70" authorityName="Spassky" authorityYear="1937" box="[189,425,1628,1654]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856790C295381834FFE734E70" box="[189,425,1628,1654]" italics="true" pageId="14" pageNumber="15">Brachythele pontica</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EF8178AB56790C2952FF8348FCD64E73" author="Spassky S. A. &amp; Minenkova K." box="[451,780,1627,1654]" journalOrPublisher="Trudy Rostovskogo biologicheskogo Obshchestva" pageId="14" pageNumber="15" pagination="138 - 141" part="4" refString="Spassky S. A. &amp; Minenkova K. 1940. New data on spiders of the group Mygalomorphae in the USSR and some conclusions from their geographical distribution. Trudy Rostovskogo biologicheskogo Obshchestva 4: 138 - 141. [In Russian]" title="New data on spiders of the group Mygalomorphae in the USSR and some conclusions from their geographical distribution" type="journal article" year="1940">Spassky &amp; Minenkova 1940</bibRefCitation>
: 140. — 
<bibRefCitation id="EF8178AB56790C29504A834FFBC84E73" author="Roewer C. F." box="[886,1042,1628,1654]" journalOrPublisher="Natura, Bremen" pageId="14" pageNumber="15" part="1" refString="Roewer C. F. 1942. Katalog der Araneae von 1758 bis 1940. Volume 1. Natura, Bremen." title="Katalog der Araneae von 1758 bis 1940" type="journal article" year="1942">Roewer 1942</bibRefCitation>
: 196. — 
<bibRefCitation id="EF8178AB56790C295741834FFAC84E73" author="Bonnet P." box="[1149,1298,1627,1654]" journalOrPublisher="Douladoure, Toulouse" pageId="14" pageNumber="15" pagination="1" part="2" refString="Bonnet P. 1955. Bibliographia araneorum Vol. 2 (1). Douladoure, Toulouse." title="Bibliographia araneorum" type="journal article" year="1955">Bonnet 1955</bibRefCitation>
: 912. — 
<bibRefCitation id="EF8178AB56790C2953D68393FE494E9E" author="Zonstein S. L." box="[234,403,1664,1691]" journalOrPublisher="Trudy zoologicheskogo Instituta AN SSSR" pageId="14" pageNumber="15" pagination="156 - 161" part="139" refString="Zonstein S. L. 1985. Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR. Trudy zoologicheskogo Instituta AN SSSR 139: 156 - 161. [In Russian with English summary]" title="Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR" type="journal article" year="1985">Zonstein 1985</bibRefCitation>
: 159.
</paragraph>
<paragraph id="8BAF055A56790C29538183B6FAAC4F0C" blockId="14.[189,1399,1627,1801]" pageId="14" pageNumber="15">
<taxonomicName id="4C107ED956790C29538183B6FE504EBA" ID-CoL="6X4WX" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[189,394,1701,1727]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856790C29538183B6FE504EBA" box="[189,394,1701,1727]" italics="true" pageId="14" pageNumber="15">Raveniola pontica</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EF8178AB56790C29529683B6FD964EBA" author="Zonstein S. L." box="[426,588,1701,1727]" journalOrPublisher="Zoologicheskii Zhurnal" pageId="14" pageNumber="15" pagination="1013 - 1019" part="66" refString="Zonstein S. L. 1987. A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna. Zoologicheskii Zhurnal 66 (7): 1013 - 1019. [In Russian with English summary]" title="A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna" type="journal article" year="1987">Zonstein 1987</bibRefCitation>
: 1015 — 
<bibRefCitation id="EF8178AB56790C2951FF83B6FC854EBA" author="Platnick N. I." box="[707,863,1701,1727]" journalOrPublisher="A Supplement to Brignoli's A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester and New York" pageId="14" pageNumber="15" pagination="1981 - 1987" refString="Platnick N. I. 1989. Advances in Spider Taxonomy 1981 - 1987: A Supplement to Brignoli's A Catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester and New York." title="Advances in Spider Taxonomy" type="book" year="1989">Platnick 1989</bibRefCitation>
: 90. — 
<bibRefCitation id="EF8178AB56790C2950FD83B6FBAF4EBA" author="Mikhailov K. G." box="[961,1141,1701,1727]" journalOrPublisher="Zoological Museum of the Moscow State University, Moscow" pageId="14" pageNumber="15" refString="Mikhailov K. G. 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum of the Moscow State University, Moscow." title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)" type="book" year="1997">Mikhailov 1997</bibRefCitation>
: 20. — 
<bibRefCitation id="EF8178AB56790C2957EB83B5FE104EE1" author="Ponomarev A. &amp; Chumachenko Y. A." editor="Lebedeva N. V." journalOrPublisher="SSC RAS Publishing, Rostov-on-Don. [In Russian]" pageId="14" pageNumber="15" pagination="151 - 163" part="III" refString="Ponomarev A. V. &amp; Chumachenko Y. A. 2007. Arachnida in ground mesofauna of yew-box grove of the Caucasian biospheric reserve. In: Lebedeva N. V. (ed.) Studies of the Southern Scientific Center of the Russian Academy of Sciences. Issue III: Biodiversity and Transformation of Mountain Ecosystems of Caucasus: 151 - 163. SSC RAS Publishing, Rostov-on-Don. [In Russian]" title="Arachnida in ground mesofauna of yew-box grove of the Caucasian biospheric reserve" type="journal article" volumeTitle="Studies of the Southern Scientific Center of the Russian Academy of Sciences" year="2007">Ponomarev &amp; Chumachenko 2007</bibRefCitation>
: 157. — 
<bibRefCitation id="EF8178AB56790C29510C83D8FC594EE1" author="Ponomarev A. &amp; Mikhailov K. G." box="[560,899,1738,1765]" editor="Lebedeva N. V." journalOrPublisher="SSC RAS Publishing, Rostov-on-Don. [In Russian]" pageId="14" pageNumber="15" pagination="130 - 150" part="III" refString="Ponomarev A. V. &amp; Mikhailov K. G. 2007. Addition to the fauna of spiders (Aranei) of the Russian Caucasus. In: Lebedeva N. V. (ed.) Studies of the Southern Scientific Center of the Russian Academy of Sciences. Issue III: Biodiversity and Transformation of Mountain Ecosystems of Caucasus: 130 - 150. SSC RAS Publishing, Rostov-on-Don. [In Russian]" title="Addition to the fauna of spiders (Aranei) of the Russian Caucasus" type="journal article" volumeTitle="Studies of the Southern Scientific Center of the Russian Academy of Sciences" year="2007">Ponomarev &amp; Mikhailov 2007</bibRefCitation>
: 130. — 
<bibRefCitation id="EF8178AB56790C2950D583D9FAE24EE1" author="Kovblyuk M. M. &amp; Ponomarev A. V." box="[1001,1336,1738,1765]" journalOrPublisher="Caucasian Entomological Bulletin" pageId="14" pageNumber="15" pagination="143 - 154" part="2" refString="Kovblyuk M. M. &amp; Ponomarev A. V. 2008. New and interesting spiders (Aranei: Agelenidae, Corinnidae, Gnaphosidae, Nemesiidae, Thomisidae) from the West Caucasus. Caucasian Entomological Bulletin 4 (2): 143 - 154. [In Russian with English summary]" title="New and interesting spiders (Aranei: Agelenidae, Corinnidae, Gnaphosidae, Nemesiidae, Thomisidae) from the West Caucasus" type="journal article" year="2008">Kovblyuk &amp; Ponomarev 2008</bibRefCitation>
: 151, figs 36–40 (♂). — 
<bibRefCitation id="EF8178AB56790C2952FE83FCFD5B4F0C" author="Ponomarev A. V." box="[450,641,1775,1801]" journalOrPublisher="Caucasian Entomological Bulletin" pageId="14" pageNumber="15" pagination="143 - 146" part="5" refString="Ponomarev A. V. 2009. New species and finds of spiders (Aranei) from the south of Russia and Western Kazakhstan. Caucasian Entomological Bulletin 5 (2): 143 - 146. [In Russian]" title="New species and finds of spiders (Aranei) from the south of Russia and Western Kazakhstan" type="journal article" year="2009">Ponomarev 2009</bibRefCitation>
: 144. — 
<bibRefCitation id="EF8178AB56790C2951D183FCFC004F0C" author="Kovblyuk M. M. &amp; Marusik Y. M. &amp; Ponomarev A. &amp; Gnelitsa V. A. &amp; Nadolny A. A." box="[749,986,1775,1801]" journalOrPublisher="Arthropoda Selecta" pageId="14" pageNumber="15" pagination="21 - 56" part="20" refString="Kovblyuk M. M., Marusik Y. M., Ponomarev A. V., Gnelitsa V. A. &amp; Nadolny A. A. 2011. Spiders (Arachnida: Aranei) of Abkhazia. Arthropoda Selecta 20 (1): 21 - 56. [In Russian with English summary]" title="Spiders (Arachnida: Aranei) of Abkhazia" type="journal article" year="2011">
Kovblyuk 
<emphasis id="B964D94856790C29505883E3FC464F0C" box="[868,924,1775,1801]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2011
</bibRefCitation>
: 37. — 
<bibRefCitation id="EF8178AB56790C29570483FCFAEC4F0C" author="Ponomarev A. &amp; Kovblyuk N. M. &amp; Chumachenko Y. A. &amp; Volkova D. D." box="[1080,1334,1775,1801]" editor="Matishov G. G." journalOrPublisher="SSC RAS Publishing, Rostov-on-Don. [In Russian]" pageId="14" pageNumber="15" pagination="447 - 481" refString="Ponomarev A. V., Kovblyuk N. M., Chumachenko Y. A. &amp; Volkova D. D. 2012. Preliminary data on the fauna of spiders of the Republic of Adygea. In: Matishov G. G. &amp; Khunagov P. O. (eds) Social-Humane and Ecological Problems of Development of Contemporary Adygea: 447 - 481. SSC RAS Publishing, Rostov-on-Don. [In Russian]" title="Preliminary data on the fauna of spiders of the Republic of Adygea" type="book chapter" volumeTitle="Social-Humane and Ecological Problems of Development of Contemporary Adygea" year="2012">
Ponomarev 
<emphasis id="B964D94856790C29578383E3FB2D4F0C" box="[1215,1271,1775,1801]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2012
</bibRefCitation>
: 470.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156790C2953818221FEEF4FEE" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BAF055A56790C2953818221FEEC4F49" blockId="14.[189,310,1842,1868]" box="[189,310,1842,1868]" pageId="14" pageNumber="15">
<heading id="D0E7B23656790C2953818221FEEC4F49" bold="true" box="[189,310,1842,1868]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B964D94856790C2953818221FEEC4F49" bold="true" box="[189,310,1842,1868]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56790C2953818271FEEF4FEE" blockId="14.[189,1398,1890,2027]" pageId="14" pageNumber="15">
This species differs from other members of the same group by having a longer embolus with a rounded subapical keel (
<figureCitation id="132B19DF56790C2952E08294FD584FA4" box="[476,642,1927,1953]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="14" pageNumber="15">Figs 178–179</figureCitation>
; cf. 
<figureCitation id="132B19DF56790C2951898294FC8D4FA4" box="[693,855,1927,1953]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="14" pageNumber="15">Figs 171–177</figureCitation>
) and by possessing long and moderately wide spermathecae, with median and lateral receptacles located close to each other (
<figureCitation id="132B19DF56790C29575982BFFB174FC3" box="[1125,1229,1964,1990]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="14" pageNumber="15">Fig. 202</figureCitation>
; cf. 
<figureCitation id="132B19DF56790C29563D82BFFEF24FEE" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="14" pageNumber="15">Figs 198– 201, 203</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156780C2853818418FCF64C13" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BAF055A56780C2853818418FE704920" blockId="15.[189,426,267,293]" box="[189,426,267,293]" pageId="15" pageNumber="16">
<heading id="D0E7B23656780C2853818418FE704920" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B964D94856780C2853818418FE704920" bold="true" box="[189,426,267,293]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B780F0756780C2853D68429FE134999" ID-GBIF-Occurrence="1806244365" collectingDate="1928-08" collectionCode="ZISP" collectorName="S. A. Spassky &amp; N. Spasskaya" country="Russia" latitude="43.516666" location="Khosta (10 km SSE of Sochi)" longLatPrecision="1143" longitude="39.883335" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<paragraph id="8BAF055A56780C2853D68429FCD54950" blockId="15.[189,1398,314,412]" box="[234,783,314,341]" pageId="15" pageNumber="16">
<typeStatus id="54ABBBF856780C2853D68429FEBC4951" box="[234,358,314,340]" pageId="15" pageNumber="16" type="lectotype">
<emphasis id="B964D94856780C2853D68429FEBC4951" bold="true" box="[234,358,314,340]" pageId="15" pageNumber="16">Lectotype</emphasis>
</typeStatus>
(designated by 
<bibRefCitation id="EF8178AB56780C28511D8429FD104950" author="Zonstein S. L." box="[545,714,314,341]" journalOrPublisher="Trudy zoologicheskogo Instituta AN SSSR" pageId="15" pageNumber="16" pagination="156 - 161" part="139" refString="Zonstein S. L. 1985. Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR. Trudy zoologicheskogo Instituta AN SSSR 139: 156 - 161. [In Russian with English summary]" title="Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR" type="journal article" year="1985">Zonstein 1985</bibRefCitation>
: 159)
</paragraph>
<paragraph id="8BAF055A56780C285381844DFE134999" blockId="15.[189,1398,314,412]" pageId="15" pageNumber="16">
<collectingCountry id="F30745CA56780C285381844DFEF0497D" box="[189,298,350,376]" name="Russia" pageId="15" pageNumber="16">RUSSIA</collectingCountry>
: 
<specimenCount id="9D16CED356780C285205844DFE95497D" box="[313,335,350,376]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
, 
<location id="8ECF538156780C28525E844DFD0D497C" LSID="urn:lsid:plazi:treatment:03B9B44C56790C36517780A3FCC14DE8:8ECF538156780C28525E844DFD0D497C" box="[354,727,350,377]" country="Russia" latitude="43.516666" longLatPrecision="1143" longitude="39.883335" name="Khosta (10 km SSE of Sochi)" pageId="15" pageNumber="16">
Khosta (
<quantity id="4CE8A8BF56780C2852F6844DFDC3497D" box="[458,537,350,376]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="km" value="10.0">10 km</quantity>
SSE of Sochi)
</location>
, 
<geoCoordinate id="EE24639D56780C2851D4844FFC82497D" box="[744,856,348,376]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="43.516666">43°31′ N</geoCoordinate>
, 
<geoCoordinate id="EE24639D56780C285054844FFC0F497D" box="[872,981,348,376]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="39.883335">39°53′ E</geoCoordinate>
, 
<collectingDate id="EFEADA7256780C2850D8844DFBBD497D" box="[996,1127,350,376]" pageId="15" pageNumber="16" value="1928-08">
<date id="FFAE239A56780C2850D8844DFBBD497D" box="[996,1127,350,376]" pageId="15" pageNumber="16" value="1928-08">Aug. 1928</date>
</collectingDate>
, 
<collectorName id="26E5608C56780C285744844DFAC2497D" box="[1144,1304,350,376]" pageId="15" pageNumber="16">S.A. Spassky</collectorName>
and 
<collectorName id="26E5608C56780C285666844DFEED4999" pageId="15" pageNumber="16">N. Spasskaya</collectorName>
leg. (
<collectionCode id="ED019D9F56780C2852468491FE664999" box="[378,444,386,412]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="15" pageNumber="16">ZISP</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BAF055A56780C2853D684D7FCCA49DA" blockId="15.[189,1399,452,657]" box="[234,784,452,479]" pageId="15" pageNumber="16">
<emphasis id="B964D94856780C2853D684D7FE3B49DB" bold="true" box="[234,481,452,478]" pageId="15" pageNumber="16">Additional material</emphasis>
(130 ♂♂, 14 ♀♀, 2 juvs)
</paragraph>
<paragraph id="8BAF055A56780C28538184FBFE0E4A94" blockId="15.[189,1399,452,657]" pageId="15" pageNumber="16">
<materialsCitation id="3B780F0756780C28538184FBFB2F4A07" ID-GBIF-Occurrence="1806244423" box="[189,1269,487,514]" pageId="15" pageNumber="16">GEORGIA/ABKHAZIA: 1 ♀, 2 juvs, Gagry, 1–16 June 1916, D.M. Fedotov leg. (ZMPU);</materialsCitation>
<materialsCitation id="3B780F0756780C2857C284FBFD5E4A23" ID-GBIF-Occurrence="1806244413" pageId="15" pageNumber="16">1 ♂, same locality, no other data (MNHN 17887);</materialsCitation>
<materialsCitation id="3B780F0756780C2851B6871FFE3C4A4C" ID-GBIF-Occurrence="1806244412" pageId="15" pageNumber="16">3 ♂♂, Myussera, 43°09′ N, 40°28′ E, 50–100 m, 5–16 Jul. 1985, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C2852C8873CFE014A68" ID-GBIF-Occurrence="1806244424" pageId="15" pageNumber="16">3 ♂♂, Odishi, 43°05′ N, 41°05′ E, 500–900 m, 23 Aug.–17 Sep. 1988, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C2852DA8740FE0E4A94" ID-GBIF-Occurrence="1806244391" pageId="15" pageNumber="16">9 ♂♂, 1 ♀, Tsebelda, 43°01′ N, 41°17′ E, 450–500 m, 20 Jun.–1 Jul. 1990, A.G. Koval leg. (TAU).</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56780C28538187AAFCF64C13" blockId="15.[189,1399,695,1046]" pageId="15" pageNumber="16">
<materialsCitation id="3B780F0756780C28538187AAFE0C4AF2" ID-GBIF-Occurrence="1806244398" pageId="15" pageNumber="16">RUSSIA: 1 ♂, 1 ♀, Lazarevskoe, 65 km NE of Sochi, 43°55′ N, 39°21′ E, 20 May–23 Jun. 1984, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C2852E387CEFAFE4AF2" ID-GBIF-Occurrence="1806244418" box="[479,1316,733,760]" pageId="15" pageNumber="16">5 ♂♂, same collection data as preceding, 10 Aug.–6 Sep. 1984 (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C28561087CEFB2E4B1E" ID-GBIF-Occurrence="1806244420" pageId="15" pageNumber="16">6 ♂♂, 4 ♀♀, same collection data as preceding, 20 May–5 Jun. 1990, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C28563C8612FB654B3A" ID-GBIF-Occurrence="1806244421" pageId="15" pageNumber="16">26 ♂♂, 3 ♀♀, Kalezh, 44°00′ N, 39°20′ E, 150–400 m, Jun.–Aug. 1984, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C2857F48636FB804B67" ID-GBIF-Occurrence="1806244415" pageId="15" pageNumber="16">2 ♂♂, Solokh- Aul, 43°47′ N, 39°40′ E, 300–400 m, 6–8 Aug. 1984, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C28575F865BFD864B83" ID-GBIF-Occurrence="1806244410" pageId="15" pageNumber="16">6 ♂♂, Sochi, 25 Jul.–5 Sep. 1986, Y.V. Zaets leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C285154867FFE404BAF" ID-GBIF-Occurrence="1806244411" pageId="15" pageNumber="16">1 ♂, Mt Fisht, 43°58′ N, 39°55′ E, 1800 m, 7 Aug. 1988, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C2852998683FD424BCB" ID-GBIF-Occurrence="1806244414" pageId="15" pageNumber="16">31 ♂♂, 4 ♀♀, Mt Pseashkho, Pslukh Gorge, 43°43′ N, 40°25′ E, 1200–1700 m, Jun.–Jul. 1988, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C28519E86A7FD664BF7" ID-GBIF-Occurrence="1806244422" pageId="15" pageNumber="16">8 ♂♂, Mamedia Gorge, 43°50′ N, 39°20′ E, 200–300 m, 23 Aug.–30 Sep. 1988, A.G. Koval leg. (TAU);</materialsCitation>
<materialsCitation id="3B780F0756780C2851FE86CBFCF64C13" ID-GBIF-Occurrence="1806244408" pageId="15" pageNumber="16">28 ♂♂, Psezuapse River valley, 5–15 km E of Lazarevskoe, 100–500 m, 1–25 Sep. 1988, A.G. Koval leg. (TAU).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156780C365381812EFE3A49A4" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="8BAF055A56780C285381812EFE944C52" blockId="15.[189,334,1085,1111]" box="[189,334,1085,1111]" pageId="15" pageNumber="16">
<heading id="D0E7B23656780C285381812EFE944C52" bold="true" box="[189,334,1085,1111]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B964D94856780C285381812EFE944C52" bold="true" box="[189,334,1085,1111]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56780C2853D6817EFDC04C8D" blockId="15.[189,538,1133,1195]" box="[234,538,1133,1160]" pageId="15" pageNumber="16">
<emphasis id="B964D94856780C2853D6817EFEF04C82" bold="true" box="[234,298,1133,1159]" pageId="15" pageNumber="16">Male</emphasis>
(from Lazarevskoe)
</paragraph>
<paragraph id="8BAF055A56780C2853818182FE6D4CAE" blockId="15.[189,538,1133,1195]" box="[189,439,1169,1195]" pageId="15" pageNumber="16">
HABITUS. See 
<figureCitation id="132B19DF56780C2852618182FE6D4CAE" box="[349,439,1169,1195]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="15" pageNumber="16">Fig. 48.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56780C28538181C0FB7D4CE8" blockId="15.[189,1191,1234,1261]" box="[189,1191,1234,1261]" pageId="15" pageNumber="16">MEASUREMENTS. TBL 13.10, CL 5.51, CW 4.78, LL 0.40, LW 0.85, SL 2.48, SW 2.37.</paragraph>
<paragraph id="8BAF055A56780C2853818006FC7F4DE7" blockId="15.[189,1399,1301,1506]" pageId="15" pageNumber="16">COLOUR. Carapace brownish rufous; eye tubercle with dark brown central and two symmetric lateral spots surrounding AMEs and lateral eyes, respectively; chelicerae and whole leg I intensive reddish brown; palps and legs II–IV brownish rufous; sternum, labium and maxillae light yellowish rufous; abdomen dorsally light greyish brown, with brown pattern represented by median lanceolate spot fused and crossed with few transverse fasciae, and with numerous small lighter spots on darker background; spinnerets and ventral surface of abdomen light yellowish grey.</paragraph>
<paragraph id="8BAF055A56780C2853818318FB444E8A" blockId="15.[189,1399,1546,1680]" pageId="15" pageNumber="16">
PROSOMA. Carapace and chelicerae as shown in 
<figureCitation id="132B19DF56780C28503E8319FC864E21" box="[770,860,1546,1572]" captionStart="Figs 68–73" captionStartId="60.[189,241,1882,1908]" captionTargetBox="[190,1398,265,1816]" captionTargetId="figure@60.[189,1398,265,1816]" captionText="Figs 68–73. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 68. R. adjarica sp. nov. (holotype). 69. R. dunini sp. nov. (holotype). 70. R. hyrcanica Dunin, 1988 (holotype). 71. R. pontica (Spassky, 1937) (Lazarevskoe). 72. R. birecikensis sp. nov. (holotype). 73. Same species as preceding (paratype, Birecik). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163444/files/figure.png" pageId="15" pageNumber="16">Fig. 71</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="132B19DF56780C2857EE8319FAE04E20" box="[1234,1338,1546,1573]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="15" pageNumber="16">Fig. 105</figureCitation>
. Eye diameters and interdistances: AME 0.15(0.19), ALE 0.23, PLE 0.15, PME 0.15, AME–AME 0.10(0.06), ALE–AME 0.08(0.06), ALE–PLE 0.05, PLE–PME 0.03, PME–PME 0.35. Each cheliceral furrow with 10 promarginal teeth and 7–8 mesobasal denticles. Maxillae with 7–8 cuspules each.
</paragraph>
<paragraph id="8BAF055A56780C28538183ABFD8D4F64" blockId="15.[189,1399,1720,1889]" pageId="15" pageNumber="16">
LEGS. Tibia and metatarsus I as shown in 
<figureCitation id="132B19DF56780C28518083ABFCFE4ED7" box="[700,804,1720,1746]" captionStart="Figs129–132" captionStartId="68.[189,241,1870,1896]" captionTargetBox="[220,1291,268,1789]" captionTargetId="figure@68.[189,1323,265,1791]" captionText="Figs129–132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." httpUri="https://zenodo.org/record/1163460/files/figure.png" pageId="15" pageNumber="16">Fig. 132</figureCitation>
. Scopula (thinner than in other congeners except for 
<taxonomicName id="4C107ED956780C2853D683CFFEAA4EF0" authorityName="in Uludag Mts" baseAuthorityName="Ausserer" baseAuthorityYear="1871" box="[234,368,1756,1781]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="micropa">
<emphasis id="B964D94856780C2853D683CFFEAA4EF0" box="[234,368,1756,1781]" italics="true" pageId="15" pageNumber="16">R. micropa</emphasis>
</taxonomicName>
): distal on metatarsi I–II, entire on tarsus I; divided by setae on tarsus II; only few scopuliform hairs on tarsi III–IV. Trichobothria: 2 rows of 10–11 each on tibiae, 11–13 on metatarsi, 11–14 on tarsi, 8–9 on cymbium. Paired claws: inner and outer margins with 7–8 teeth each on tarsi I–II, with 6–7 teeth each on tarsi III–IV.
</paragraph>
<paragraph id="8BAF055A56780C375381829AFBF64976" blockId="15.[189,1399,1929,2028]" lastBlockId="16.[189,1399,267,372]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">SPINATION. Palp: femur d1–1–1, pd1; tibia d1–1, p1–1–1, r1, pv1–1–1, r1, v1–1; cymbium d6(7). Leg I: femur d1–1–1–1, pd1; tibia p1–1, pv1(0), rv1(0)–1-m–m. Leg II: femur d1–1–1–1, pd1–1–1; patella p1; tibia p1–1–1, v2–2–3; metatarsus p1, v2–2–2(3). Leg III: femur d1–1–1–1, pd1–1–1, rd1(0)–1(0)–1; patella p1–1, r1; tibia d1–1, p1–1, r1–1, v2–2–2(3); metatarsus d1–1–1, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur d1–1–1–1, pd1–0–1, rd1–1; patella p1, r1; tibia d1–1, p1–1–1, r1–1–1, v2–2–3; metatarsus d1–1–1–1, p1–1–1, r1–1–1, v2–1–2–3. Metatarsus I and patella I aspinose.</paragraph>
<paragraph id="8BAF055A56670C375381848CFC354A03" blockId="16.[189,1398,414,519]" pageId="16" pageNumber="17">
PALP. Tibia, cymbium and palpal organ as shown in 
<figureCitation id="132B19DF56670C375012848DFC4E49BC" box="[814,916,414,441]" captionStart="Figs 149–152" captionStartId="73.[189,241,1930,1956]" captionTargetBox="[194,1398,266,1866]" captionTargetId="figure@73.[189,1398,265,1875]" captionText="Figs 149–152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163470/files/figure.png" pageId="16" pageNumber="17">Fig. 152</figureCitation>
. Surface of bulb with numerous shallow wrinkles (
<figureCitation id="132B19DF56670C37520E84D6FEA149E5" box="[306,379,453,480]" captionStart="Figs 1–6" captionStartId="50.[189,241,1734,1760]" captionTargetBox="[189,1399,265,1680]" captionTargetId="figure@50.[189,1399,265,1680]" captionText="Figs 1–6.Raveniola spp., ♀♀ (1–3, 5–6) and ♂ (4). Chelicerae (1), apical edge of maxilla (2–3), palpal organ (4) and tarsus I (5–6); ventroapical (1), ventral (2–4, 6) and dorsal (5) views. — 1–2. R. micropa (Ausserer, 1871). 3–5. R. pontica (Spassky, 1937). 6. R. hyrcanica Dunin, 1988." httpUri="https://zenodo.org/record/1163424/files/figure.png" pageId="16" pageNumber="17">Fig. 4</figureCitation>
). Unlike in other members of same group, entire embolus relatively long, but its apical part between keel and tip apex very short and narrow (
<figureCitation id="132B19DF56670C37500084FFFC384A03" box="[828,994,492,518]" captionStart="Figs 171–179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171–179. Raveniola hyrcanica species group, holotypic (171–177) and conspecific (178–179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171–173. R. adjarica sp. nov. 174–175. R. dunini sp. nov. 176– 177. R. hyrcanica Dunin, 1988. 178–179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="16" pageNumber="17">Figs 178–179</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56670C3753818721FB794A77" blockId="16.[189,1398,561,627]" pageId="16" pageNumber="17">SPINNERETS. PMS: length 0.45; diameter 0.27. PLS: maximal diameter 0.40; length of basal, medial and apical segments 0.63, 0.36, 0.48; total length 1.47; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A56670C375381878DFE354ABD" blockId="16.[189,495,670,696]" box="[189,495,670,696]" pageId="16" pageNumber="17">LEG MEASUREMENTS. ♂(♀).</paragraph>
<paragraph id="8BAF055A56670C37525487CFFAB34BC3" pageId="16" pageNumber="17">
<table id="F910F7FA5667F3D853F487C8FAB34BC3" box="[200,1385,731,966]" colsContinueFrom="11.[200,1386,438,673]" colsContinueIn="25.[200,1335,1781,2016]" gridcols="7" gridrows="6" pageId="16" pageNumber="17" rowsContinueIn="18.[200,1335,1180,1415]">
<tr id="352007185667F3D853F487C8FAB34AF3" box="[200,1385,731,758]" gridrow="0" pageId="16" pageNumber="17" rowspan-0="1">
<th id="76F16E645667F3D8527287C8FE154AF3" box="[334,463,731,758]" gridcol="1" gridrow="0" pageId="16" pageNumber="17">Femur</th>
<th id="76F16E645667F3D8513F87C8FD5F4AF3" box="[515,645,731,758]" gridcol="2" gridrow="0" pageId="16" pageNumber="17">Patella</th>
<th id="76F16E645667F3D8518587C8FCE14AF3" box="[697,827,731,758]" gridcol="3" gridrow="0" pageId="16" pageNumber="17">Tibia</th>
<th id="76F16E645667F3D8505287C8FC2A4AF3" box="[878,1008,731,758]" gridcol="4" gridrow="0" pageId="16" pageNumber="17">Metatarsus</th>
<th id="76F16E645667F3D8571887C8FB7C4AF3" box="[1060,1190,731,758]" gridcol="5" gridrow="0" pageId="16" pageNumber="17">Tarsus</th>
<th id="76F16E645667F3D857F687C8FAB34AF3" box="[1226,1385,731,758]" gridcol="6" gridrow="0" pageId="16" pageNumber="17">Total</th>
</tr>
<tr id="352007185667F3D853F48616FAB34B25" box="[200,1385,773,800]" gridrow="1" pageId="16" pageNumber="17">
<th id="76F16E645667F3D853F48616FEC64B25" box="[200,284,773,800]" gridcol="0" gridrow="1" pageId="16" pageNumber="17">Palp</th>
<td id="76F16E645667F3D852728616FE154B25" box="[334,463,773,800]" gridcol="1" gridrow="1" pageId="16" pageNumber="17">2.85 (2.65)</td>
<td id="76F16E645667F3D8513F8616FD5F4B25" box="[515,645,773,800]" gridcol="2" gridrow="1" pageId="16" pageNumber="17">1.70 (1.60)</td>
<td id="76F16E645667F3D851858616FCE14B25" box="[697,827,773,800]" gridcol="3" gridrow="1" pageId="16" pageNumber="17">2.28 (1.87)</td>
<td id="76F16E645667F3D850528616FC2A4B25" box="[878,1008,773,800]" gridcol="4" gridrow="1" pageId="16" pageNumber="17">–</td>
<td id="76F16E645667F3D857188616FB7C4B25" box="[1060,1190,773,800]" gridcol="5" gridrow="1" pageId="16" pageNumber="17">1.12 (1.72)</td>
<td id="76F16E645667F3D857F68616FAB34B25" box="[1226,1385,773,800]" gridcol="6" gridrow="1" pageId="16" pageNumber="17">7.95 (7.84)</td>
</tr>
<tr id="352007185667F3D853F4863DFAB34B4C" box="[200,1385,814,841]" gridrow="2" pageId="16" pageNumber="17">
<th id="76F16E645667F3D853F4863DFEC64B4C" box="[200,284,814,841]" gridcol="0" gridrow="2" pageId="16" pageNumber="17">Leg I</th>
<td id="76F16E645667F3D85272863DFE154B4C" box="[334,463,814,841]" gridcol="1" gridrow="2" pageId="16" pageNumber="17">4.57 (3.57)</td>
<td id="76F16E645667F3D8513F863DFD5F4B4C" box="[515,645,814,841]" gridcol="2" gridrow="2" pageId="16" pageNumber="17">2.78 (2.28)</td>
<td id="76F16E645667F3D85185863DFCE14B4C" box="[697,827,814,841]" gridcol="3" gridrow="2" pageId="16" pageNumber="17">3.65 (2.77)</td>
<td id="76F16E645667F3D85052863DFC2A4B4C" box="[878,1008,814,841]" gridcol="4" gridrow="2" pageId="16" pageNumber="17">3.73 (2.13)</td>
<td id="76F16E645667F3D85718863DFB7C4B4C" box="[1060,1190,814,841]" gridcol="5" gridrow="2" pageId="16" pageNumber="17">2.35 (1.65)</td>
<td id="76F16E645667F3D857F6863DFAB34B4C" box="[1226,1385,814,841]" gridcol="6" gridrow="2" pageId="16" pageNumber="17">17.08 (12.40)</td>
</tr>
<tr id="352007185667F3D853F4864BFAB34B76" box="[200,1385,856,883]" gridrow="3" pageId="16" pageNumber="17">
<th id="76F16E645667F3D853F4864BFEC64B76" box="[200,284,856,883]" gridcol="0" gridrow="3" pageId="16" pageNumber="17">Leg II</th>
<td id="76F16E645667F3D85272864BFE154B76" box="[334,463,856,883]" gridcol="1" gridrow="3" pageId="16" pageNumber="17">3.98 (3.23)</td>
<td id="76F16E645667F3D8513F864BFD5F4B76" box="[515,645,856,883]" gridcol="2" gridrow="3" pageId="16" pageNumber="17">2.37 (2.05)</td>
<td id="76F16E645667F3D85185864BFCE14B76" box="[697,827,856,883]" gridcol="3" gridrow="3" pageId="16" pageNumber="17">3.18 (2.37)</td>
<td id="76F16E645667F3D85052864BFC2A4B76" box="[878,1008,856,883]" gridcol="4" gridrow="3" pageId="16" pageNumber="17">3.23 (2.15)</td>
<td id="76F16E645667F3D85718864BFB7C4B76" box="[1060,1190,856,883]" gridcol="5" gridrow="3" pageId="16" pageNumber="17">2.07 (1.70)</td>
<td id="76F16E645667F3D857F6864BFAB34B76" box="[1226,1385,856,883]" gridcol="6" gridrow="3" pageId="16" pageNumber="17">14.83 (11.50)</td>
</tr>
<tr id="352007185667F3D853F48691FAB34B98" box="[200,1385,898,925]" gridrow="4" pageId="16" pageNumber="17">
<th id="76F16E645667F3D853F48691FEC64B98" box="[200,284,898,925]" gridcol="0" gridrow="4" pageId="16" pageNumber="17">Leg III</th>
<td id="76F16E645667F3D852728691FE154B98" box="[334,463,898,925]" gridcol="1" gridrow="4" pageId="16" pageNumber="17">3.47 (3.10)</td>
<td id="76F16E645667F3D8513F8691FD5F4B98" box="[515,645,898,925]" gridcol="2" gridrow="4" pageId="16" pageNumber="17">2.05 (1.88)</td>
<td id="76F16E645667F3D851858691FCE14B98" box="[697,827,898,925]" gridcol="3" gridrow="4" pageId="16" pageNumber="17">2.77 (2.15)</td>
<td id="76F16E645667F3D850528691FC2A4B98" box="[878,1008,898,925]" gridcol="4" gridrow="4" pageId="16" pageNumber="17">3.55 (2.67)</td>
<td id="76F16E645667F3D857188691FB7C4B98" box="[1060,1190,898,925]" gridcol="5" gridrow="4" pageId="16" pageNumber="17">2.10 (1.75)</td>
<td id="76F16E645667F3D857F68691FAB34B98" box="[1226,1385,898,925]" gridcol="6" gridrow="4" pageId="16" pageNumber="17">13.94 (11.55)</td>
</tr>
<tr id="352007185667F3D853F486B8FAB34BC3" box="[200,1385,939,966]" gridrow="5" pageId="16" pageNumber="17">
<th id="76F16E645667F3D853F486B8FEC64BC3" box="[200,284,939,966]" gridcol="0" gridrow="5" pageId="16" pageNumber="17">Leg IV</th>
<td id="76F16E645667F3D8527286B8FE154BC3" box="[334,463,939,966]" gridcol="1" gridrow="5" pageId="16" pageNumber="17">4.48 (3.80)</td>
<td id="76F16E645667F3D8513F86B8FD5F4BC3" box="[515,645,939,966]" gridcol="2" gridrow="5" pageId="16" pageNumber="17">2.30 (2.12)</td>
<td id="76F16E645667F3D8518586B8FCE14BC3" box="[697,827,939,966]" gridcol="3" gridrow="5" pageId="16" pageNumber="17">3.60 (3.27)</td>
<td id="76F16E645667F3D8505286B8FC2A4BC3" box="[878,1008,939,966]" gridcol="4" gridrow="5" pageId="16" pageNumber="17">5.12 (4.17)</td>
<td id="76F16E645667F3D8571886B8FB7C4BC3" box="[1060,1190,939,966]" gridcol="5" gridrow="5" pageId="16" pageNumber="17">2.37 (2.07)</td>
<td id="76F16E645667F3D857F686B8FAB34BC3" box="[1226,1385,939,966]" gridcol="6" gridrow="5" pageId="16" pageNumber="17">17.87 (15.43)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A56670C3753D686E1FDEF4C08" blockId="16.[189,565,1010,1075]" box="[234,565,1010,1037]" pageId="16" pageNumber="17">
<emphasis id="B964D94856670C3753D686E1FE9F4C09" bold="true" box="[234,325,1010,1036]" pageId="16" pageNumber="17">Female</emphasis>
(from Lazarevskoe)
</paragraph>
<paragraph id="8BAF055A56670C375381810AFE6D4C36" blockId="16.[189,565,1010,1075]" box="[189,439,1049,1075]" pageId="16" pageNumber="17">
HABITUS. See 
<figureCitation id="132B19DF56670C375261810AFE6D4C36" box="[349,439,1049,1075]" captionStart="Figs43–51" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs43–51. Raveniolaspp.,habitusof ♂♂(43,45–46,48,50–51)and ♀♀(44,47,49),dorsalviews.— 43–44. R. adjarica sp. nov. 45. R. dunini sp. nov. 46–47. R. hyrcanica Dunin, 1988. 48–49. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="16" pageNumber="17">Fig. 49.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56670C375381814CFB7D4C7C" blockId="16.[189,1191,1118,1145]" box="[189,1191,1118,1145]" pageId="16" pageNumber="17">MEASUREMENTS. TBL 16.15, CL 5.05, CW 4.35, LL 0.44, LW 1.05, SL 2.33, SW 2.35.</paragraph>
<paragraph id="8BAF055A56670C37538181B0FE714CB8" blockId="16.[189,427,1187,1214]" box="[189,427,1187,1214]" pageId="16" pageNumber="17">COLOUR. As in male.</paragraph>
<paragraph id="8BAF055A56670C37538181FAFB554D72" blockId="16.[189,1399,1256,1400]" pageId="16" pageNumber="17">
PROSOMA. Carapace and chelicerae as shown in 
<figureCitation id="132B19DF56670C37503E81FBFC864D07" box="[770,860,1256,1282]" captionStart="Figs 84–89" captionStartId="63.[189,241,1892,1918]" captionTargetBox="[189,1398,266,1873]" captionTargetId="figure@63.[189,1398,265,1878]" captionText="Figs 84–89. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163450/files/figure.png" pageId="16" pageNumber="17">Fig. 87</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="132B19DF56670C3757EE81FBFAE04D07" box="[1234,1338,1256,1282]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="16" pageNumber="17">Fig. 120</figureCitation>
. Eye diameters and interdistances: AME 0.14(0.17), ALE 0.25, PLE 0.13, PME 0.13, AME–AME 0.10(0.07), ALE–AME 0.07(0.06), ALE–PLE 0.06, PLE–PME 0.03, PME–PME 0.38. Each cheliceral furrow with 9 promarginal teeth and 5–6 mesobasal denticles. Maxillae with 7–8 cuspules each.
</paragraph>
<paragraph id="8BAF055A56670C37538180B0FE814E34" blockId="16.[189,1399,1442,1585]" pageId="16" pageNumber="17">LEGS. Scopula: distal on metatarsi I–II, divided on tarsus I and palpal tarsus, widely divided on tarsus II, elsewhere absent. Trichobothria: 2 rows of 8–9 each on tibiae, 10–14 on metatarsi, 11–12 on tarsi, 9 on palpal tarsus. Paired claws: outer and inner margins with 5–7 teeth each. Palpal claw with 4 teeth on inner margin.</paragraph>
<paragraph id="8BAF055A56670C375381834EFD804F3C" blockId="16.[189,1399,1628,1849]" pageId="16" pageNumber="17">SPINATION. All femora with one basodorsal spine and 2–3 dorsal bristles; palpal patella and patellae I–II aspinose. Palp: femur d1, pd1; tibia p1–1, v2–1–3; tarsus d5(6). Leg I: femur d1, pd1; tibia p1–1, v2–2–2; metatarsus v2–2–2. Leg II: femur d1, pd1; tibia p1–1, v2–2–2; metatarsus p1, v2–2–2. Leg III: femur d1, pd1, rd1–1; patella p1, r1; tibia d1, p1–1, r1–1, v2–2–3; metatarsus d1–1, p1–1–1, r1–1– 1–1, v2–2–3. Leg IV: femur d1, rd1; patella p1, r1; tibia d1, p1–1, r1–1, v2–2–2(3); metatarsus d1–1, p1–1–1–1, r1–1–1–1–1, v2–2–2–3.</paragraph>
<paragraph id="8BAF055A56670C3753818276FADB4F7A" blockId="16.[189,1281,1892,1919]" box="[189,1281,1892,1919]" pageId="16" pageNumber="17">
SPERMATHECAE. Moderately long and wide, with receptacles close set to each other (
<figureCitation id="132B19DF56670C3757B18277FB2F4F7A" box="[1165,1269,1892,1919]" captionStart="Figs 198–203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198–203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="16" pageNumber="17">Fig. 202</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56670C37538182B9FB794FEE" blockId="16.[189,1398,1962,2027]" pageId="16" pageNumber="17">SPINNERETS. PMS: length 0.44; diameter 0.27. PLS: maximal diameter 0.48; length of basal, medial and apical segments 0.85, 0.35, 0.38; total length 1.58; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A56660C3653818418FEE94920" blockId="17.[189,307,267,293]" box="[189,307,267,293]" pageId="17" pageNumber="18">
<heading id="D0E7B23656660C3653818418FEE94920" bold="true" box="[189,307,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B964D94856660C3653818418FEE94920" bold="true" box="[189,307,267,293]" pageId="17" pageNumber="18">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56660C365381842FFE3A49A4" blockId="17.[189,1398,315,417]" pageId="17" pageNumber="18">Carapace length in males varies from 4.40 to 5.75, in females from 4.35 to 5.93 mm. In some specimens the distal abdominal pattern is almost indiscernible. The copulative organs of both sexes do not show any significant variation.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156660C36538184D8FE1B4AD4" pageId="17" pageNumber="18" type="biology_ecology">
<paragraph id="8BAF055A56660C36538184D8FEFA49E0" blockId="17.[189,288,459,485]" box="[189,288,459,485]" pageId="17" pageNumber="18">
<heading id="D0E7B23656660C36538184D8FEFA49E0" bold="true" box="[189,288,459,485]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B964D94856660C36538184D8FEFA49E0" bold="true" box="[189,288,459,485]" pageId="17" pageNumber="18">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56660C36538184E8FE1B4AD4" blockId="17.[189,1399,507,721]" pageId="17" pageNumber="18">
Judging from sample data and collecting localities, 
<taxonomicName id="4C107ED956660C36501E84EFFC414A10" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[802,923,508,533]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856660C36501E84EFFC414A10" box="[802,923,508,533]" italics="true" pageId="17" pageNumber="18">R. pontica</emphasis>
</taxonomicName>
is a very common spider species which occurs in different types of subcoastal and montane broad-leaved forests (see 
<figureCitation id="132B19DF56660C3657518732FACC4A3E" box="[1133,1302,544,571]" captionStart="Figs 223–229" captionStartId="88.[189,241,1652,1678]" captionTargetBox="[189,1399,306,1600]" captionTargetId="figure@88.[189,1399,265,1600]" captionText="Figs 223–229. Habitats of Raveniola spp. — 223–224. Azerbaijan, southern macroslope of Caucasus Major,forested midlands inBelokany area, betweenLagodekhi,Georgia [thetypelocality of R. zaitzevi (Charitonov, 1948)] and Zakataly, Azerbaijan. 225–227. Abkhazia, canyon of the Bzyb River, forested biotopes inhabited by R.pontica (Spassky, 1937). 228–229.Azerbaijan, foothillsof TalyshMts,Hyrcan Reserve, humid localities typical for R. hyrcanica Dunin, 1988. Images courtesy of Natasha Snegovaya (223–224, 228–229) and Mykola Kovblyuk (225–227)." httpUri="https://zenodo.org/record/1163502/files/figure.png" pageId="17" pageNumber="18">Figs 225–227</figureCitation>
). At the upper limit of their distribution (1800 m), the spiders have been found inhabiting subalpine meadows above the mid-mountain forest zone. The relatively rarely sampled females are believed to hide in cavities under stones, logs, trunks of fallen trees, etc., whereas the vagrant males have been collected mostly in pitfall traps.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156660C36538187E9FC2A4C22" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BAF055A56660C36538187E9FE8C4B11" blockId="17.[189,342,762,788]" box="[189,342,762,788]" pageId="17" pageNumber="18">
<heading id="D0E7B23656660C36538187E9FE8C4B11" bold="true" box="[189,342,762,788]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B964D94856660C36538187E9FE8C4B11" bold="true" box="[189,342,762,788]" pageId="17" pageNumber="18">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56660C3653818638FC2A4C22" blockId="17.[189,1399,811,1063]" pageId="17" pageNumber="18">
The currently known area inhabited by this species is shown in 
<figureCitation id="132B19DF56660C3650A58638FC274B40" box="[921,1021,811,837]" captionStart="Figure 219" captionStartId="85.[189,272,1534,1560]" captionTargetBox="[189,1398,265,1471]" captionTargetId="figure@85.[189,1398,265,1471]" captionText="Figure 219. Localities of Raveniola spp. in the western part of the Caucasus: R. adjarica sp. nov. () and R. pontica (Spassky, 1937) ()." httpUri="https://zenodo.org/record/1163494/files/figure.png" pageId="17" pageNumber="18">Fig. 219</figureCitation>
. It includes Abkhazia and the far south of Russia, i.e., the Black Sea coast and the adjacent NW-Caucasus area from Bolshoy Utrish Cape in the north-west (44°45′ N, 37°24′ E) to the surroundings of Sokhumi (Sukhum) (43°00′ N, 41°00′ E) in the south and Maykop (43°31′ N, 39°52′ E) in the east (according to 
<bibRefCitation id="EF8178AB56660C3650E6868FFAB54BB3" author="Ponomarev A. &amp; Chumachenko Y. A." box="[986,1391,923,950]" editor="Lebedeva N. V." journalOrPublisher="SSC RAS Publishing, Rostov-on-Don. [In Russian]" pageId="17" pageNumber="18" pagination="151 - 163" part="III" refString="Ponomarev A. V. &amp; Chumachenko Y. A. 2007. Arachnida in ground mesofauna of yew-box grove of the Caucasian biospheric reserve. In: Lebedeva N. V. (ed.) Studies of the Southern Scientific Center of the Russian Academy of Sciences. Issue III: Biodiversity and Transformation of Mountain Ecosystems of Caucasus: 151 - 163. SSC RAS Publishing, Rostov-on-Don. [In Russian]" title="Arachnida in ground mesofauna of yew-box grove of the Caucasian biospheric reserve" type="journal article" volumeTitle="Studies of the Southern Scientific Center of the Russian Academy of Sciences" year="2007">Ponomarev &amp; Chumachenko 2007</bibRefCitation>
; 
<bibRefCitation id="EF8178AB56660C36538186D1FDFA4BDE" author="Ponomarev A. &amp; Mikhailov K. G." box="[189,544,961,988]" editor="Lebedeva N. V." journalOrPublisher="SSC RAS Publishing, Rostov-on-Don. [In Russian]" pageId="17" pageNumber="18" pagination="130 - 150" part="III" refString="Ponomarev A. V. &amp; Mikhailov K. G. 2007. Addition to the fauna of spiders (Aranei) of the Russian Caucasus. In: Lebedeva N. V. (ed.) Studies of the Southern Scientific Center of the Russian Academy of Sciences. Issue III: Biodiversity and Transformation of Mountain Ecosystems of Caucasus: 130 - 150. SSC RAS Publishing, Rostov-on-Don. [In Russian]" title="Addition to the fauna of spiders (Aranei) of the Russian Caucasus" type="journal article" volumeTitle="Studies of the Southern Scientific Center of the Russian Academy of Sciences" year="2007">Ponomarev &amp; Mikhailov 2007</bibRefCitation>
; 
<bibRefCitation id="EF8178AB56660C36511286D2FC574BDE" author="Kovblyuk M. M. &amp; Ponomarev A. V." box="[558,909,961,988]" journalOrPublisher="Caucasian Entomological Bulletin" pageId="17" pageNumber="18" pagination="143 - 154" part="2" refString="Kovblyuk M. M. &amp; Ponomarev A. V. 2008. New and interesting spiders (Aranei: Agelenidae, Corinnidae, Gnaphosidae, Nemesiidae, Thomisidae) from the West Caucasus. Caucasian Entomological Bulletin 4 (2): 143 - 154. [In Russian with English summary]" title="New and interesting spiders (Aranei: Agelenidae, Corinnidae, Gnaphosidae, Nemesiidae, Thomisidae) from the West Caucasus" type="journal article" year="2008">Kovblyuk &amp; Ponomarev 2008</bibRefCitation>
and the aforementioned label data). When describing this species, 
<bibRefCitation id="EF8178AB56660C3652E286F5FD424C04" author="Spassky S. A." box="[478,664,998,1025]" journalOrPublisher="Festschrift Embrik Strand" pageId="17" pageNumber="18" pagination="361 - 368" part="3" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." title="Araneae palaearcticae novae. Mygalomorphae. I" type="journal article" year="1937">Spassky (1937)</bibRefCitation>
also mentioned a female that had been collected at Batumi. However, this record in fact refers to 
<taxonomicName id="4C107ED956660C365148811EFD2D4C23" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[628,759,1036,1062]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="adjarica" status="sp. nov.">
<emphasis id="B964D94856660C365148811EFD2D4C23" box="[628,759,1036,1062]" italics="true" pageId="17" pageNumber="18">R. adjarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356660C3651C3811EFC804C22" box="[767,858,1037,1063]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
(see below).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156660C3653818143FCC14DE8" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BAF055A56660C3653818143FEF54C6F" blockId="17.[189,303,1104,1130]" box="[189,303,1104,1130]" pageId="17" pageNumber="18">
<heading id="D0E7B23656660C3653818143FEF54C6F" bold="true" box="[189,303,1104,1130]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B964D94856660C3653818143FEF54C6F" bold="true" box="[189,303,1104,1130]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56660C3653818193FCC14DE8" blockId="17.[189,1399,1152,1517]" pageId="17" pageNumber="18">
Although males of 
<taxonomicName id="4C107ED956660C3652A28193FDAE4C9F" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[414,628,1152,1178]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856660C3652A28193FDAE4C9F" box="[414,628,1152,1178]" italics="true" pageId="17" pageNumber="18">Raveniola pontica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C107ED956660C3651918192FCEB4C9F" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[685,817,1152,1178]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="adjarica" status="sp. nov.">
<emphasis id="B964D94856660C3651918192FCEB4C9F" box="[685,817,1152,1178]" italics="true" pageId="17" pageNumber="18">R. adjarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356660C3650048192FC494C9E" box="[824,915,1153,1179]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
are well distinguishable from each other, the conspecific females could not be identified as belonging to either of these species without dissecting and examining the vulvae. This may explain why Spassky assigned both females at his disposal to the same species. It is much more difficult to understand, however, why both spiders collected from quite different localities were placed in the same vial. In addition, both females are of approximately equal size! As the material had been desiccated at least once and then macerated, the identification of these specimens has become problematic. Some time was devoted to understanding which of them was collected at Khosta and which at Batumi. Since the Khosta specimen was mentioned first and its characters were found to correspond better to the original description, it was designated as the lectotype (
<bibRefCitation id="EF8178AB56660C3653FB80C1FCD54DE8" author="Zonstein S. L." box="[199,783,1490,1517]" journalOrPublisher="Trudy zoologicheskogo Instituta AN SSSR" pageId="17" pageNumber="18" pagination="156 - 161" part="139" refString="Zonstein S. L. 1985. Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR. Trudy zoologicheskogo Instituta AN SSSR 139: 156 - 161. [In Russian with English summary]" title="Preliminary data on the fauna of the spider suborder Mygalomorphae from the USSR" type="journal article" year="1985">Zonstein 1985; erroneously called the “holotype”</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>