278 lines
37 KiB
XML
278 lines
37 KiB
XML
<document id="EC5D2BB0EE031789CDE5A8CD75DC314C" ID-CLB-Dataset="46751" ID-DOI="10.11646/zootaxa.3753.2.8" ID-GBIF-Dataset="6e36efa8-ad13-4a14-964e-28908950dad1" ID-ISSN="1175-5326" ID-Zenodo-Dep="227808" ID-ZooBank="3161924A-23A6-4FB7-83FD-8614D4C36D85" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460364315575" checkinUser="plazi" docAuthor="Laššová, Kristína, Čiampor, Fedor & Čiamporová-Zaťovičová, Zuzana" docDate="2014" docId="03B9B015FFCCFFF0CEF077926B22AEAE" docLanguage="en" docName="zt03753p195.pdf" docOrigin="Zootaxa 3753 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexanchorus angeli Laššová, Čiampor & Čiamporová-Zaťovičová, 2014, n. sp." docType="treatment" docVersion="8" lastPageNumber="191" masterDocId="FF80C86DFFCDFFF4CE67743D6A0AAE56" masterDocTitle="Two new Larainae species from Guayana region, Venezuela (Coleoptera: Elmidae)" masterLastPageNumber="195" masterPageNumber="187" pageNumber="188" updateTime="1698295006129" updateUser="ExternalLinkService">
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<mods:mods id="519B9B33D2C8378D1D63ECA36FE40FC4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B135F335C5AB3D62C69BCFA7342FAF9C">
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<mods:title id="ABD88C8ECB03473ED502D4D4338E62EC">Two new Larainae species from Guayana region, Venezuela (Coleoptera: Elmidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B2C01D885050B11B4D1D1BA2E13D8AA6" type="personal">
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<mods:namePart id="18B56E86904262F2C754069740381DF1">Laššová, Kristína</mods:namePart>
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<mods:namePart id="0191EC55B4628F877D5F46A355F6BED9">Čiampor, Fedor</mods:namePart>
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</mods:name>
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<mods:name id="16D38C05C3820C55AAEE4EF9D8207124" type="personal">
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<mods:roleTerm id="D17514ECE768B2930B2971CC828D5D93">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EDB2AA6D5B5145913A7ADB39B377D4A4">Čiamporová-Zaťovičová, Zuzana</mods:namePart>
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<mods:title id="E4F49FD9A41A48F3D02B70A44C86BECE">Zootaxa</mods:title>
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<mods:part id="F365103370C96E669DC3176EA22E0F41">
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<mods:date id="145B4203F5DECA874381E33A841C3C13">2014</mods:date>
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<mods:number id="BADF6C4B7A8DAA3D40E37E6ECC5222D1">3753</mods:number>
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<treatment id="03B9B015FFCCFFF0CEF077926B22AEAE" ID-DOI="http://doi.org/10.5281/zenodo.6126867" ID-GBIF-Taxon="119410391" ID-Zenodo-Dep="6126867" LSID="urn:lsid:plazi:treatment:03B9B015FFCCFFF0CEF077926B22AEAE" httpUri="http://treatment.plazi.org/id/03B9B015FFCCFFF0CEF077926B22AEAE" lastPageId="4" lastPageNumber="191" pageId="1" pageNumber="188">
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<subSubSection id="C30A5288FFCCFFF5CEF077926B0CADBA" pageId="1" pageNumber="188" type="nomenclature">
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<paragraph id="8BAF0103FFCCFFF5CEF077926BC2AD9E" blockId="1.[151,456,943,1005]" box="[151,456,943,968]" pageId="1" pageNumber="188">
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<emphasis id="B964DD11FFCCFFF5CEF077926BC2AD9E" bold="true" box="[151,456,943,968]" pageId="1" pageNumber="188">
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<taxonomicName id="4C107A80FFCCFFF5CEF077926B89AD9E" ID-CoL="3LGJG" box="[151,387,943,968]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
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<emphasis id="B964DD11FFCCFFF5CEF077926B89AD9E" bold="true" box="[151,387,943,968]" italics="true" pageId="1" pageNumber="188">Hexanchorus angeli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A257606AFFCCFFF5CFEE77926BC2AD9E" box="[393,456,943,968]" pageId="1" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAF0103FFCCFFF5CEF077E96B0CADBA" blockId="1.[151,456,943,1005]" box="[151,262,980,1005]" pageId="1" pageNumber="188">
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<figureCitation id="132B1D86FFCCFFF5CEF077E96B0CADBA" box="[151,262,980,1005]" captionStart-0="FIGURES 1, 2" captionStart-1="FIGURE 3" captionStart-2="FIGURES 4 – 12" captionStart-3="FIGURES 13 – 16" captionStartId-0="2.[151,264,1409,1432]" captionStartId-1="3.[151,250,1411,1433]" captionStartId-2="4.[151,264,1723,1746]" captionStartId-3="5.[151,264,781,804]" captionTargetBox-0="[183,1407,556,1377]" captionTargetBox-1="[152,1436,181,1382]" captionTargetBox-2="[159,1433,313,1693]" captionTargetBox-3="[151,1436,193,759]" captionTargetId-0="figure@2.[174,1413,553,1388]" captionTargetId-1="figure@3.[151,1436,181,1389]" captionTargetId-2="figure@4.[151,1437,303,1702]" captionTargetId-3="figure@5.[151,1436,193,760]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 1, 2. Habitus of Hexanchorus angeli n. sp.: 1) holotype, male, 2) paratype female." captionText-1="FIGURE 3. Hexanchorus angeli n. sp. male and female ventral view." captionText-2="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." captionText-3="FIGURES 13 – 16. Type localities: 13, 14) Río Churún, type locality of Hexanchorus angeli n. sp.; 15, 16) small stream near Cerro Autana, type locality of Hypsilara autanai n. sp." httpUri-0="https://zenodo.org/record/227809/files/figure.png" httpUri-1="https://zenodo.org/record/227810/files/figure.png" httpUri-2="https://zenodo.org/record/227811/files/figure.png" httpUri-3="https://zenodo.org/record/227812/files/figure.png" pageId="1" pageNumber="188">Figs 1–14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A5288FFCCFFF5CEF0702168C4AA93" pageId="1" pageNumber="188" type="materials_examined">
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<paragraph id="8BAF0103FFCCFFF5CEF0702169CEAA0E" blockId="1.[151,1436,1052,2013]" pageId="1" pageNumber="188">
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<emphasis id="B964DD11FFCCFFF5CEF070216B39AA63" bold="true" box="[151,307,1052,1077]" pageId="1" pageNumber="188">
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<typeStatus id="54ABBFA1FFCCFFF5CEF070216AD8AA63" box="[151,210,1052,1077]" pageId="1" pageNumber="188">Type</typeStatus>
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locality
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</emphasis>
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: environment of the Río Churun, large stream ca.
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wide and mostly about
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deep with orange-brown coloured humic water (specimens were collected at light).
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</paragraph>
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<paragraph id="8BAF0103FFCCFFF5CEA0705968C4AA93" blockId="1.[151,1436,1052,2013]" pageId="1" pageNumber="188">
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<emphasis id="B964DD11FFCCFFF5CEA070596B67AA2B" bold="true" box="[199,365,1124,1149]" pageId="1" pageNumber="188">
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<typeStatus id="54ABBFA1FFCCFFF5CEA070596B08AA2B" box="[199,258,1124,1149]" pageId="1" pageNumber="188">Type</typeStatus>
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material
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</emphasis>
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:
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<emphasis id="B964DD11FFCCFFF5CF1C70596BE8AA2B" bold="true" box="[379,482,1124,1149]" pageId="1" pageNumber="188">
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<typeStatus id="54ABBFA1FFCCFFF5CF1C70596BE8AA2B" box="[379,482,1124,1149]" pageId="1" pageNumber="188" type="holotype">holotype</typeStatus>
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</emphasis>
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male (
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<collectionCode id="ED0199C6FFCCFFF5CC4970596877AA2A" box="[558,637,1124,1148]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="188">NMW</collectionCode>
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): ”
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<collectingCountry id="F3074193FFCCFFF5CCFF7059691AAA2B" box="[664,784,1124,1149]" name="Venezuela" pageId="1" pageNumber="188">Venezuela</collectingCountry>
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, Auyán Tepui env., Salto Angel camp, Río Churun env., at light,
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<quantity id="4CE8ACE6FFCCFFF5CEB870BA6B26AAC9" box="[223,300,1159,1184]" metricMagnitude="2" metricUnit="m" metricValue="5.06" pageId="1" pageNumber="188" unit="m" value="506.0">506 m</quantity>
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||
a.s.l.,
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<geoCoordinate id="EE2467C4FFCCFFF5CFE470BA6821AAC9" box="[387,555,1159,1184]" direction="north" orientation="latitude" pageId="1" pageNumber="188" precision="1" value="5.9765277">05°58'35.5'' N</geoCoordinate>
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,
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<geoCoordinate id="EE2467C4FFCCFFF5CC5A70BA68FDAAC9" box="[573,759,1159,1184]" direction="west" orientation="longitude" pageId="1" pageNumber="188" precision="1" value="-62.51614">062°30'58.1'' W</geoCoordinate>
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,
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<date id="FFAE27C3FFCCFFF5CD6F70BA6977AAF6" box="[776,893,1159,1184]" pageId="1" pageNumber="188" value="2012-05-12">5.12.2012</date>
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”,
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<emphasis id="B964DD11FFCCFFF5CDFE70BA6E05AAF6" bold="true" box="[921,1039,1159,1184]" pageId="1" pageNumber="188">
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<typeStatus id="54ABBFA1FFCCFFF5CDFE70BA6E05AAF6" box="[921,1039,1159,1184]" pageId="1" pageNumber="188" type="paratype">paratypes</typeStatus>
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||
</emphasis>
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||
(
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<collectionCode id="ED0199C6FFCCFFF5CA4270B56E6BAAF6" box="[1061,1121,1160,1184]" httpUri="http://grbio.org/cool/e3tf-9780" name="Colecao de Culturas de Basidiomicetos" pageId="1" pageNumber="188">CCB</collectionCode>
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,
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||
<collectionCode id="ED0199C6FFCCFFF5CA1470B56EB7AAF6" LSID="urn:lsid:biocol.org:col:33834" box="[1139,1213,1160,1184]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="188">MIZA</collectionCode>
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,
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<collectionCode id="ED0199C6FFCCFFF5CAA870B56F1DAAC9" box="[1231,1303,1160,1183]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="188">NMW</collectionCode>
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,
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<collectionCode id="ED0199C6FFCCFFF5CB4E70B56F62AAF6" box="[1321,1384,1160,1184]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="188">ZSM</collectionCode>
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):
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<specimenCount id="9D16CA8AFFCCFFF5CBE670B56B06AA93" pageId="1" pageNumber="188" type="generic">11 specimens</specimenCount>
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with the same locality data as
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<typeStatus id="54ABBFA1FFCCFFF5CC03709168C0AA93" box="[612,714,1196,1221]" pageId="1" pageNumber="188" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A5288FFCCFFF5CEA070F26EB7AB2E" pageId="1" pageNumber="188" type="diagnosis">
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<paragraph id="8BAF0103FFCCFFF5CEA070F26EB7AB2E" blockId="1.[151,1436,1052,2013]" pageId="1" pageNumber="188">
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<emphasis id="B964DD11FFCCFFF5CEA070F26B4BAAB1" bold="true" box="[199,321,1231,1256]" pageId="1" pageNumber="188">Diagnosis.</emphasis>
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<taxonomicName id="4C107A80FFCCFFF5CF2E70ED6826AABE" box="[329,556,1232,1256]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
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<emphasis id="B964DD11FFCCFFF5CF2E70ED6826AABE" box="[329,556,1232,1256]" italics="true" pageId="1" pageNumber="188">Hexanchorus angeli</emphasis>
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</taxonomicName>
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<emphasis id="B964DD11FFCCFFF5CC5370F2687EAABE" bold="true" box="[564,628,1231,1256]" pageId="1" pageNumber="188">
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||
<taxonomicNameLabel id="A257606AFFCCFFF5CC5370F2687EAABE" box="[564,628,1231,1256]" pageId="1" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
is externally similar to all other known
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||
<taxonomicName id="4C107A80FFCCFFF5CA5A70ED6ED8AABE" box="[1085,1234,1232,1256]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B964DD11FFCCFFF5CA5A70ED6ED8AABE" box="[1085,1234,1232,1256]" italics="true" pageId="1" pageNumber="188">Hexanchorus</emphasis>
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||
</taxonomicName>
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species. It differs in combination of the following characters: 1) presence of a greenish iridescence on dorsal surface, 2) without median longitudinal sulcus at base of pronotum, 3) elytral apices feebly (males) or distinctly (females) upturned, 4) third ventrite of females with distinct median process, 5) ventrite 5 slightly emarginated apically in males, 6) aedeagus with median lobe short, narrow apically, parameres long abruptly constricted basally.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30A5288FFCCFFF7CEA071B96B1CA83F" lastPageId="3" lastPageNumber="190" pageId="1" pageNumber="188" type="description">
|
||
<paragraph id="8BAF0103FFCCFFF5CEA071B96EBEA85E" blockId="1.[151,1436,1052,2013]" pageId="1" pageNumber="188">
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||
<emphasis id="B964DD11FFCCFFF5CEA071B96B53ABCB" bold="true" box="[199,345,1412,1437]" pageId="1" pageNumber="188">Description.</emphasis>
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Body elongate, subparallel (
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||
<figureCitation id="132B1D86FFCCFFF5CCCE71B9691FABCA" box="[681,789,1412,1437]" captionStart="FIGURES 1, 2" captionStartId="2.[151,264,1409,1432]" captionTargetBox="[183,1407,556,1377]" captionTargetId="figure@2.[174,1413,553,1388]" captionTargetPageId="2" captionText="FIGURES 1, 2. Habitus of Hexanchorus angeli n. sp.: 1) holotype, male, 2) paratype female." httpUri="https://zenodo.org/record/227809/files/figure.png" pageId="1" pageNumber="188">Figs 1, 2</figureCitation>
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); length
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||
<quantity id="4CE8ACE6FFCCFFF5CDE771B969E1ABCA" box="[896,1003,1412,1437]" metricMagnitude="-3" metricUnit="m" metricValue="3.16" pageId="1" pageNumber="188" unit="mm" value="3.16">3.16 mm</quantity>
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; width
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<quantity id="4CE8ACE6FFCCFFF5CA2271B96EBAABCA" box="[1093,1200,1412,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="188" unit="mm" value="1.15">1.15 mm</quantity>
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; dorsum moderately convex, dorsal surface with very dense short recumbent setae with greenish iridescence and more sparse, longer, dark, semierected setae. Body black, scape and pedicel dark brown, maxillae, trochanters and basal portion of femora pale brown, tarsal claws reddish-brown. Venter dark with light yellowish-brown setae.
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</paragraph>
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<paragraph id="8BAF0103FFCCFFF5CEA072296B24A97E" blockId="1.[151,1436,1052,2013]" pageId="1" pageNumber="188">
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||
<emphasis id="B964DD11FFCCFFF5CEA072296B04A87B" bold="true" box="[199,270,1556,1581]" pageId="1" pageNumber="188">Head.</emphasis>
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Head partly retractable into prothorax. Labrum densely setose, with largely rounded anterior angles, feebly emarginate anteromedially; clypeus densely setose, shorter and narrower than labrum, about 3.0 times as wide as long, frontoclypeal suture visible, almost straight. Eyes well developed, suboval in lateral view, protruding from head outline, bordered with long black curved setae arising near dorsal and ventral side and extend toward middle of eye, cranial surface not raised around eyes. Frons between eyes flat. Antennae eleven segmented, densely setose, scape curved, about 2.0 times as long as pedicel, remaining segments about three times longer than scape and pedicel combined, segments 3–10 short, subtriangular, terminal segment subglobular with slightly pointed apex.
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</paragraph>
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<paragraph id="8BAF0103FFCCFFF6CEA073096E37AFA3" blockId="1.[151,1436,1052,2013]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="189" pageId="1" pageNumber="188">
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<emphasis id="B964DD11FFCCFFF5CEA073096B2CA91B" bold="true" box="[199,294,1844,1869]" pageId="1" pageNumber="188">Thorax.</emphasis>
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Pronotum widest behind middle, PW:
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, PL:
|
||
<quantity id="4CE8ACE6FFCCFFF5CDEA730969FDA91A" box="[909,1015,1844,1868]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="188" unit="mm" value="0.78">0.78 mm</quantity>
|
||
; partly shiny with dense smaller and larger setigerous punctures; disc convex, divided by broad transversal U-shaped depression in the first third; triangular median depression at base shallow, reaching basal third of pronotum. Sides of pronotum convex in posterior 2/3, slightly constricted before base, feebly convex in anterior third before transverse constriction; lateral margins narrowly rimmed, posterior angles orthogonal, anterior angles not produced. Lateral sides decliving to the ventral side. Hypomeron narrow, sinuate. Prosternal process narrow, triangular, with apex rounded; prosternum in front of coxae extremely narrow, reduced. Mesoventrite short and wide, disc narrowed posteriad, posteromedially with deep triangular impression for reception of prosternal process. Metaventrite wide, metaventral discrimen thin, present along basal 2/3; disc raised on sides, surface reticulate and densely setose (
|
||
<figureCitation id="132B1D86FFCFFFF6CA2D75396E9AAF4A" box="[1098,1168,260,285]" captionStart="FIGURE 3" captionStartId="3.[151,250,1411,1433]" captionTargetBox="[152,1436,181,1382]" captionTargetId="figure@3.[151,1436,181,1389]" captionTargetPageId="3" captionText="FIGURE 3. Hexanchorus angeli n. sp. male and female ventral view." httpUri="https://zenodo.org/record/227810/files/figure.png" pageId="2" pageNumber="189">Fig. 3</figureCitation>
|
||
). Elytra (EL:
|
||
<quantity id="4CE8ACE6FFCFFFF6CB5775396F9DAF4A" box="[1328,1431,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.42" pageId="2" pageNumber="189" unit="mm" value="2.42">2.42 mm</quantity>
|
||
, EW:
|
||
<quantity id="4CE8ACE6FFCFFFF6CEB1751A6B49AF16" box="[214,323,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="189" unit="mm" value="1.15">1.15 mm</quantity>
|
||
), in anterior 2/3 parallel, then tapering toward apices, disc moderately convex, sides strongly declivous; surface glabrous with very dense short setae with greenish iridescence and sparser longer semierect hairs; elytral margin only very narrowly rimmed; humeri rounded, slightly produced; epipleuron widest in anterior third, then reduced, very thin. Each elytron with ten rows of small punctures, striae impressed on disc, impression diminishes toward lateral margin and elytral apex. Scutellum wide, subtriangular, anterior portion flat, apical triangular part raised. Legs moderately long, mesotibia markedly flattened, longest.
|
||
</paragraph>
|
||
<caption id="DF6F518BFFCFFFF6CEF071BC6E47ABCE" box="[151,1101,1409,1432]" httpUri="https://zenodo.org/record/227809/files/figure.png" pageId="2" pageNumber="189" targetBox="[183,1407,556,1377]" targetPageId="2">
|
||
<paragraph id="8BAF0103FFCFFFF6CEF071BC6E47ABCE" blockId="2.[151,1101,1409,1432]" box="[151,1101,1409,1432]" pageId="2" pageNumber="189">
|
||
<emphasis id="B964DD11FFCFFFF6CEF071BC6B30ABC1" bold="true" box="[151,314,1409,1432]" pageId="2" pageNumber="189">FIGURES 1, 2.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C107A80FFCFFFF6CFD671BF6877ABC1" box="[433,637,1410,1431]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
|
||
<emphasis id="B964DD11FFCFFFF6CFD671BF6877ABC1" box="[433,637,1410,1431]" italics="true" pageId="2" pageNumber="189">Hexanchorus angeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B964DD11FFCFFFF6CCE371BF68B7ABC1" bold="true" box="[644,701,1410,1431]" pageId="2" pageNumber="189">
|
||
<taxonomicNameLabel id="A257606AFFCFFFF6CCE371BF68B7ABC1" box="[644,701,1410,1431]" pageId="2" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 1) holotype, male, 2) paratype female.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF0103FFCFFFF6CEA071826B12A986" blockId="2.[151,1436,1471,2000]" pageId="2" pageNumber="189">
|
||
<emphasis id="B964DD11FFCFFFF6CEA071826B49AB8E" bold="true" box="[199,323,1471,1496]" pageId="2" pageNumber="189">Abdomen.</emphasis>
|
||
Ventrites densely setose (
|
||
<figureCitation id="132B1D86FFCFFFF6CC05718268ACAB81" box="[610,678,1471,1496]" captionStart="FIGURE 3" captionStartId="3.[151,250,1411,1433]" captionTargetBox="[152,1436,181,1382]" captionTargetId="figure@3.[151,1436,181,1389]" captionTargetPageId="3" captionText="FIGURE 3. Hexanchorus angeli n. sp. male and female ventral view." httpUri="https://zenodo.org/record/227810/files/figure.png" pageId="2" pageNumber="189">Fig. 3</figureCitation>
|
||
). Intercoxal process of first ventrite with apex rounded, lateral sides not raised, sublateral carinae absent; distal margins of first and second ventrite feebly sinuate; third ventrite simple in males, in females with distinct median process (
|
||
<figureCitation id="132B1D86FFCFFFF6CCAF723A6911A849" box="[712,795,1543,1568]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 12</figureCitation>
|
||
). Ventrite 5 with distal margin emarginated in males (
|
||
<figureCitation id="132B1D86FFCFFFF6CB09723A6AADA815" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 7</figureCitation>
|
||
), simple in females (
|
||
<figureCitation id="132B1D86FFCFFFF6CFF372166BEDA815" box="[404,487,1579,1604]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 10</figureCitation>
|
||
). Sternite 8 weakly sclerotized and reduced as two darkened spots in males (
|
||
<figureCitation id="132B1D86FFCFFFF6CB2E72166F86A815" box="[1353,1420,1579,1604]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 8</figureCitation>
|
||
), deeply emarginate in females (
|
||
<figureCitation id="132B1D86FFCFFFF6CF987272685EA831" box="[511,596,1615,1640]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 11</figureCitation>
|
||
). The ninth segment and spiculum gastrale extremely elongate (
|
||
<figureCitation id="132B1D86FFCFFFF6CB2272726F87A83E" box="[1349,1421,1615,1640]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 6</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="132B1D86FFCFFFF6CF7D724E6B80A8DA" box="[282,394,1651,1676]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Figs 4, 5</figureCitation>
|
||
) elongate, penis with fibula feebly sclerotized, better visible from lateral view, corona membranous; parameres slightly shorter than penis, in lateral view widest in middle, moderately narrowed in basal half, distal half moderately narrowed toward rounded apex, in ventral view moderately narrowed in apical ca. 0.4, in basal 0.6 parallel-sided, base constricted; penis in ventral view subparallel in basal 0.6 with distinct basal apophyses, apical 0.4 narrowed toward thin rounded apex, in lateral view slender, sinuate, with widened basal 2/3. Phallobase parallelsided, curved in lateral view. Penis and parameres with sparse fine spines. Ovipositor (
|
||
<figureCitation id="132B1D86FFCFFFF6CB29731A6F99A969" box="[1358,1427,1831,1856]" captionStart="FIGURES 4 – 12" captionStartId="4.[151,264,1723,1746]" captionTargetBox="[159,1433,313,1693]" captionTargetId="figure@4.[151,1437,303,1702]" captionTargetPageId="4" captionText="FIGURES 4 – 12. Hexanchorus angeli n. sp.: 4) aedeagus lateral view; 5) aedeagus ventral view; 6) male ninth segment and spiculum gastrale; 7) ventrite 5 male; 8) sternite 8 male; 9) ovipositor; 10) ventrite 5 female; 11) sternite 8 female; 12) ventrite 3 female. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/227811/files/figure.png" pageId="2" pageNumber="189">Fig. 9</figureCitation>
|
||
) with stylus (terminal segment) short, slightly widened basally; preterminal segment short and robust, about 2.7 times as long as stylus, apically with numerous curved blunt and acuminate spines or sensilla; basal segment with baculus membranous, ca. 0.8 times as long as preterminal and distal segments combined, baculus straight, well sclerotized.
|
||
</paragraph>
|
||
<caption id="DF6F518BFFCEFFF7CEF071BE6956ABCF" box="[151,860,1411,1433]" httpUri="https://zenodo.org/record/227810/files/figure.png" pageId="3" pageNumber="190" targetBox="[152,1436,181,1382]" targetPageId="3">
|
||
<paragraph id="8BAF0103FFCEFFF7CEF071BE6956ABCF" blockId="3.[151,860,1411,1433]" box="[151,860,1411,1433]" pageId="3" pageNumber="190">
|
||
<emphasis id="B964DD11FFCEFFF7CEF071BE6B19ABCE" bold="true" box="[151,275,1411,1433]" pageId="3" pageNumber="190">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C107A80FFCEFFF7CF7E71BE6BEFABCE" box="[281,485,1411,1432]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
|
||
<emphasis id="B964DD11FFCEFFF7CF7E71BE6BEFABCE" box="[281,485,1411,1432]" italics="true" pageId="3" pageNumber="190">Hexanchorus angeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B964DD11FFCEFFF7CF8B71BE682FABCE" bold="true" box="[492,549,1411,1432]" pageId="3" pageNumber="190">
|
||
<taxonomicNameLabel id="A257606AFFCEFFF7CF8B71BE682FABCE" box="[492,549,1411,1432]" pageId="3" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male and female ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAF0103FFCEFFF7CEA071FD6B1CA83F" blockId="3.[151,1436,1472,2037]" pageId="3" pageNumber="190">
|
||
<emphasis id="B964DD11FFCEFFF7CEA071FD6BBBAB8F" bold="true" box="[199,433,1472,1497]" pageId="3" pageNumber="190">Sexual dimorphism</emphasis>
|
||
. Both sexes are generally similar in shape and size, however males are usually slightly smaller. From females, males can be easily distinguished by elytral apices not produced, by ventrite 3 without distal process, by shape of distal margin of ventrite 5, by ventrites 1–2 concave, not convex. Pronotum of males with feeble median triangular depression in basal third, medio basal portion of female pronotum without visible depression.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A5288FFCEFFF7CEA072496E3EA8DB" box="[199,1076,1652,1677]" pageId="3" pageNumber="190" type="distribution">
|
||
<paragraph id="8BAF0103FFCEFFF7CEA072496E3EA8DB" blockId="3.[151,1436,1472,2037]" box="[199,1076,1652,1677]" pageId="3" pageNumber="190">
|
||
<emphasis id="B964DD11FFCEFFF7CEA072496B53A8DB" bold="true" box="[199,345,1652,1677]" pageId="3" pageNumber="190">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="4C107A80FFCEFFF7CF0072496843A8DA" box="[359,585,1652,1676]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
|
||
<emphasis id="B964DD11FFCEFFF7CF0072496843A8DA" box="[359,585,1652,1676]" italics="true" pageId="3" pageNumber="190">Hexanchorus angeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B964DD11FFCEFFF7CC3772496885A8DB" bold="true" box="[592,655,1652,1677]" pageId="3" pageNumber="190">
|
||
<taxonomicNameLabel id="A257606AFFCEFFF7CC3772496885A8DB" box="[592,655,1652,1677]" pageId="3" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="54ABBFA1FFCEFFF7CDC4724869D9A8DB" box="[931,979,1653,1677]" pageId="3" pageNumber="190">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A5288FFCEFFF7CEA072A56BA9A9A3" pageId="3" pageNumber="190" type="discussion">
|
||
<paragraph id="8BAF0103FFCEFFF7CEA072A56903A917" blockId="3.[151,1436,1472,2037]" pageId="3" pageNumber="190">
|
||
<emphasis id="B964DD11FFCEFFF7CEA072A56B3EA8E7" bold="true" box="[199,308,1688,1713]" pageId="3" pageNumber="190">Remarks</emphasis>
|
||
. One female of
|
||
<taxonomicName id="4C107A80FFCEFFF7CF8F72A56877A8E6" box="[488,637,1688,1712]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B964DD11FFCEFFF7CF8F72A56877A8E6" box="[488,637,1688,1712]" italics="true" pageId="3" pageNumber="190">Hexanchorus</emphasis>
|
||
</taxonomicName>
|
||
was found also in Río Tarota (Gran Sabana), more than
|
||
<quantity id="4CE8ACE6FFCEFFF7CA9272A56F40A8E7" box="[1269,1354,1688,1713]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="190" unit="km" value="120.0">120 km</quantity>
|
||
distant from the
|
||
<typeStatus id="54ABBFA1FFCEFFF7CE9972806B24A883" box="[254,302,1725,1749]" pageId="3" pageNumber="190">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C107A80FFCEFFF7CFD772806813A882" box="[432,537,1724,1748]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
|
||
<emphasis id="B964DD11FFCEFFF7CFD772806813A882" box="[432,537,1724,1748]" italics="true" pageId="3" pageNumber="190">H. angeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B964DD11FFCEFFF7CC477281686AA883" bold="true" box="[544,608,1724,1749]" pageId="3" pageNumber="190">
|
||
<taxonomicNameLabel id="A257606AFFCEFFF7CC477281686AA883" box="[544,608,1724,1749]" pageId="3" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
This female is morphologically very similar to the females of the species described herein. However, observed 4 % molecular distance, measured as uncorrected p-distance on 816bp long fragment of the mitochondrial gene for cytochrome oxidase c subunit I (tree not shown), suggest that the female from the Gran Sabana should belong to another species.
|
||
</paragraph>
|
||
<paragraph id="8BAF0103FFCEFFF7CEA073716BA9A9A3" blockId="3.[151,1436,1472,2037]" pageId="3" pageNumber="190">
|
||
<emphasis id="B964DD11FFCEFFF7CEA073716B29A933" bold="true" box="[199,291,1868,1893]" pageId="3" pageNumber="190">Habitat</emphasis>
|
||
. The specimens were caught on light, so we can only suggest that
|
||
<taxonomicName id="4C107A80FFCEFFF7CA7A73706E8DA932" box="[1053,1159,1868,1892]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="angeli" status="sp. nov.">
|
||
<emphasis id="B964DD11FFCEFFF7CA7A73706E8DA932" box="[1053,1159,1868,1892]" italics="true" pageId="3" pageNumber="190">H. angeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B964DD11FFCEFFF7CAE873716EDAA933" bold="true" box="[1167,1232,1868,1893]" pageId="3" pageNumber="190">
|
||
<taxonomicNameLabel id="A257606AFFCEFFF7CAE873716EDAA933" box="[1167,1232,1868,1893]" pageId="3" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
adults live in and around rapids of the Río Churún, as do most of other related
|
||
<taxonomicName id="4C107A80FFCEFFF7CD27734D69A8A9DF" box="[832,930,1904,1929]" class="Insecta" family="Elmidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="subFamily" subFamily="Larainae">Larainae</taxonomicName>
|
||
species, and their larvae develop in the same river or their tributaries. At the place of collecting, the river was about
|
||
<quantity id="4CE8ACE6FFCEFFF7CDCC73A969EEA9FA" box="[939,996,1940,1965]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="190" unit="m" value="15.0">15 m</quantity>
|
||
wide, relatively shallow. Substrate was formed by gravel with large stones and boulders weekly covered by periphyton (
|
||
<figureCitation id="132B1D86FFCEFFF7CA7873856EA2A986" box="[1055,1192,1976,2001]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,781,804]" captionTargetBox="[151,1436,193,759]" captionTargetId="figure@5.[151,1436,193,760]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Type localities: 13, 14) Río Churún, type locality of Hexanchorus angeli n. sp.; 15, 16) small stream near Cerro Autana, type locality of Hypsilara autanai n. sp." httpUri="https://zenodo.org/record/227812/files/figure.png" pageId="3" pageNumber="190">Figs. 13, 14</figureCitation>
|
||
). Water was naturally humic, brown coloured.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30A5288FFC9FFF0CEA074AA6B22AEAE" pageId="4" pageNumber="191" type="etymology">
|
||
<paragraph id="8BAF0103FFC9FFF0CEA074AA6B22AEAE" blockId="4.[151,1436,151,248]" pageId="4" pageNumber="191">
|
||
<emphasis id="B964DD11FFC9FFF0CEA074AA6B4CAEE6" bold="true" box="[199,326,151,176]" pageId="4" pageNumber="191">Etymology</emphasis>
|
||
. The species is named after Jimmie Angel, an American aviator who was the first man flying over the Angel falls. These waterfalls are the highest in the World, falling from the slopes of the Auyán tepui close to the
|
||
<typeStatus id="54ABBFA1FFC9FFF0CEF074DD6ACDAEAE" box="[151,199,224,248]" pageId="4" pageNumber="191">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |