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<document id="770C672C9B5ABD4AA2B99CCDDFF8282D" ID-DOI="10.5281/zenodo.13154915" ID-ISSN="0068-547X" ID-Zenodo-Dep="13154915" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722537880974" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Angel, Kathleen W." docDate="2015" docId="03B03365F917615BFFACFBC5B7E9F96F" docLanguage="en" docName="ProcCalifAcadSci.62.14.381-396.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (14)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Scaphinotus (Pseudonomaretus) relictus" docType="treatment" docVersion="2" lastPageNumber="393" masterDocId="FF894B1DF91B6157FFD7FFF5B600FFD2" masterDocTitle="A Taxonomic Review of the Northwestern Nearctic Subgenus Pseudonomaretus Roeschke 1907, including Description of the New Species Scaphinotus hoodooensis (Coleoptera: Carabidae: Cychrini) from the Bitterroot Mountains of Montana" masterLastPageNumber="396" masterPageNumber="381" pageNumber="393" updateTime="1722543448646" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5F1F897D1433B2548BF87E7234B65BB8">A Taxonomic Review of the Northwestern Nearctic Subgenus Pseudonomaretus Roeschke 1907, including Description of the New Species Scaphinotus hoodooensis (Coleoptera: Carabidae: Cychrini) from the Bitterroot Mountains of Montana</mods:title>
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<mods:namePart id="8B9D3F380CE86EEA81A76E162E573B4B">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="7C198AF3F96389A472CE8C6DA0193006">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A. Email: dkavanaugh @ calacademy. org.</mods:affiliation>
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<mods:namePart id="B03BEE4C62380100E82B1D4239A1A6A7">Angel, Kathleen W.</mods:namePart>
<mods:affiliation id="783AC542EEDC89592939E3D7A5EC8F11">Department of Entomology, University of Wisconsin, Madison, WI 53706</mods:affiliation>
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<mods:title id="F1AB81603C9F9E1E36CBA694AF869647">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="B13D6AD5ACD9EA38F79B57B4A1738F59">2015</mods:date>
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<mods:number id="6593AE81AE243B36D8FA0A970D21FE1F">2015-08-14</mods:number>
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<mods:number id="989A36CDAD9E79EAF1648A7CE47B8BD6">62</mods:number>
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<mods:number id="5ECCFADB4CA82B7E3AF0355940E4F397">14</mods:number>
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<mods:classification id="0EC9D3C1E0B32C0581E1056B51751812">journal article</mods:classification>
<mods:identifier id="8359F5A99F3FE3548F25124FA4BD2670" type="DOI">10.5281/zenodo.13154915</mods:identifier>
<mods:identifier id="BF3FCDA49EDAA898AF092AA2508BA26F" type="ISSN">0068-547X</mods:identifier>
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<treatment id="03B03365F917615BFFACFBC5B7E9F96F" LSID="urn:lsid:plazi:treatment:03B03365F917615BFFACFBC5B7E9F96F" httpUri="http://treatment.plazi.org/id/03B03365F917615BFFACFBC5B7E9F96F" lastPageNumber="393" pageId="12" pageNumber="393">
<subSubSection id="C303D1F8F917615BFFACFBC5B50EFB9B" box="[123,782,1072,1097]" pageId="12" pageNumber="393" type="nomenclature">
<paragraph id="8BA68273F917615BFFACFBC5B50EFB9B" blockId="12.[123,782,1072,1127]" box="[123,782,1072,1097]" pageId="12" pageNumber="393">
<heading id="D0EE351FF917615BFFACFBC5B50EFB9B" box="[123,782,1072,1097]" fontSize="10" level="2" pageId="12" pageNumber="393" reason="4">
<taxonomicName id="4C19F9F0F917615BFFACFBC5B50EFB9B" ID-CoL="6Y5HK" authority="(G. H. Horn) 1881" baseAuthorityName="G. H. Horn" baseAuthorityYear="1881" box="[123,782,1072,1097]" class="Insecta" family="Carabidae" genus="Scaphinotus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="relictus" subGenus="Pseudonomaretus">
<emphasis id="B96D5E61F917615BFFACFBC5B50EFB9B" bold="true" box="[123,782,1072,1097]" pageId="12" pageNumber="393">
<emphasis id="B96D5E61F917615BFFACFBC5B703FB9A" bold="true" box="[123,259,1072,1096]" italics="true" pageId="12" pageNumber="393">Scaphinotus</emphasis>
(
<emphasis id="B96D5E61F917615BFEC4FBC5B7D8FB9A" bold="true" box="[275,472,1072,1096]" italics="true" pageId="12" pageNumber="393">Pseudonomaretus</emphasis>
)
<emphasis id="B96D5E61F917615BFE3EFBC5B43AFB9A" bold="true" box="[489,570,1072,1096]" italics="true" pageId="12" pageNumber="393">relictus</emphasis>
(G. H. Horn) 1881
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C303D1F8F917615BFFACFBA4B7E9F96F" pageId="12" pageNumber="393" type="description">
<paragraph id="8BA68273F917615BFFACFBA4B743FBB4" blockId="12.[123,782,1072,1127]" box="[123,323,1105,1127]" pageId="12" pageNumber="393">
<figureCitation id="13229EF6F917615BFFACFBA4B6E1FBB5" box="[123,225,1105,1127]" captionStart="FIGURE" captionStartId="6.[155,167,1287,1306]" captionText="FIGURE 2. Digital photographs of labrum of Scaphinotus (Pseudonomaretus) species, dorsal aspect; a, S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana); b, S. merkelii (G. Horn) (9 km SE of Low- ell, Idaho County, Idaho); c, S. regularis (LeConte) (Hayden Creek, Kootenai County, Washington); d, S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho); e, S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 0.5 mm." pageId="12" pageNumber="393">Figures 2d</figureCitation>
,
<figureCitation id="13229EF6F917615BFF39FBA4B706FBB5" box="[238,262,1105,1127]" captionStart="FIGURE" captionStartId="7.[155,167,1508,1527]" captionTargetBox="[120,1176,247,1459]" captionTargetId="figure-25@7.[120,1176,216,1493]" captionTargetPageId="7" captionText="FIGURE 3. Digital photographs of right antenna, antennomeres 1 to 5, of Scaphinotus (Pseudonomaretus) species, dorsal aspect, with antennomeres identified by number; a, S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana); b, S. merkelii (G. Horn) (St. Joe National Forest, Idaho); c, S. regularis (LeConte) (Hayden Creek, Kootenai County, Washington); d, S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho); e, S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13154919" httpUri="https://zenodo.org/record/13154919/files/figure.png" pageId="12" pageNumber="393">3d</figureCitation>
,
<figureCitation id="13229EF6F917615BFEC5FBA4B71EFBB5" box="[274,286,1105,1127]" captionStart="FIGURES" captionStartId="9.[155,167,1578,1597]" captionTargetBox="[208,1099,216,1560]" captionTargetId="figure-25@9.[208,1097,216,1560]" captionTargetPageId="9" captionText="FIGURES 4–7. Digital photographs of male left protarsus of Scaphinotus (Pseudonomaretus) species, with tarsomeres identified by number; a, dorsal aspect; b, ventral aspect; as = pad of adhesive setae. Fig. 4. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 5. S. merkelii (G. Horn) (9 km SE of Lowell, Idaho County, Idaho). Fig. 6. S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho). Fig. 7. S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13154923" httpUri="https://zenodo.org/record/13154923/files/figure.png" pageId="12" pageNumber="393">6</figureCitation>
,
<figureCitation id="13229EF6F917615BFEFCFBA4B743FBB4" box="[299,323,1105,1126]" captionStart="FIGURES" captionStartId="11.[155,167,1575,1594]" captionText="FIGURES 8–12. Digital photographs of median lobe of male aedeagus of Scaphinotus (Pseudonomaretus) species; a, dor- sal aspect; b, left lateral aspect; is = internal sac. Fig. 8. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 9. S. merkelii (G. Horn) (Moscow Mountain, Latah County, Idaho). Fig. 10. S. mannii Wick- ham (Wawawai, Whitman County, Washington). Fig. 11. S. regularis (LeConte) (Moscow Mountain, Latah County, Idaho). Fig. 12. S. relictus (G. Horn) (Mount Spokane, Spokane County, Washington). Scale lines = 1.0 mm." pageId="12" pageNumber="393">12</figureCitation>
</paragraph>
<paragraph id="8BA68273F917615BFF7CFB76B4CFF98E" blockId="12.[123,1174,1154,1726]" pageId="12" pageNumber="393">
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<emphasis id="B96D5E61F917615BFF7CFB76B742FB4B" bold="true" box="[171,322,1155,1179]" pageId="12" pageNumber="393">RECOGNITION</emphasis>
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.— Adults of this species can be distinguished from those of other species of
<taxonomicName id="4C19F9F0F917615BFFACFB56B739FB69" authorityName="Roeschke" authorityYear="1907" box="[123,313,1187,1211]" pageId="12" pageNumber="394" rank="subGenus" subGenus="Pseudonomaretus">
<emphasis id="B96D5E61F917615BFFACFB56B739FB69" box="[123,313,1187,1211]" italics="true" pageId="12" pageNumber="393">Pseudonomaretus</emphasis>
</taxonomicName>
by the following combination of character states: SBL = 13.0 mm or more (males 13.0–16.
<quantity id="4CE12F96F917615BFEF1FB36B763FB08" box="[294,355,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="393" unit="mm" value="6.0">6 mm</quantity>
, females 14.7–20.
<quantity id="4CE12F96F917615BFDFEFB36B466FB08" box="[553,614,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="393" unit="mm" value="3.0">3 mm</quantity>
); elytra moderately shiny, microsculpture shallowly impressed, without metallic reflection; head with a distinct transverse impression at level of posterior margin of eyes clearly delineating frontal and occipital regions; labrum (
<figureCitation id="13229EF6F917615BFC6AFAF6B20AFAC9" box="[957,1034,1283,1307]" captionStart="FIGURE" captionStartId="6.[155,167,1287,1306]" captionText="FIGURE 2. Digital photographs of labrum of Scaphinotus (Pseudonomaretus) species, dorsal aspect; a, S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana); b, S. merkelii (G. Horn) (9 km SE of Low- ell, Idaho County, Idaho); c, S. regularis (LeConte) (Hayden Creek, Kootenai County, Washington); d, S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho); e, S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 0.5 mm." pageId="12" pageNumber="393">Fig. 2d</figureCitation>
) with lateral lobes moderate in length and width; antennomere 4 (
<figureCitation id="13229EF6F917615BFD62FAD6B501FAE9" box="[693,769,1315,1339]" captionStart="FIGURE" captionStartId="7.[155,167,1508,1527]" captionTargetBox="[120,1176,247,1459]" captionTargetId="figure-25@7.[120,1176,216,1493]" captionTargetPageId="7" captionText="FIGURE 3. Digital photographs of right antenna, antennomeres 1 to 5, of Scaphinotus (Pseudonomaretus) species, dorsal aspect, with antennomeres identified by number; a, S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana); b, S. merkelii (G. Horn) (St. Joe National Forest, Idaho); c, S. regularis (LeConte) (Hayden Creek, Kootenai County, Washington); d, S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho); e, S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13154919" httpUri="https://zenodo.org/record/13154919/files/figure.png" pageId="12" pageNumber="393">Fig. 3d</figureCitation>
) not pubescent, with only a very few scattered setae in addition to apical whorl of setae; pronotum with basolateral pair of setiferous pores present; male protarsomere 1 (
<figureCitation id="13229EF6F917615BFDD9FA91B45BFAAE" box="[526,603,1380,1404]" captionStart="FIGURES" captionStartId="9.[155,167,1578,1597]" captionTargetBox="[208,1099,216,1560]" captionTargetId="figure-25@9.[208,1097,216,1560]" captionTargetPageId="9" captionText="FIGURES 4–7. Digital photographs of male left protarsus of Scaphinotus (Pseudonomaretus) species, with tarsomeres identified by number; a, dorsal aspect; b, ventral aspect; as = pad of adhesive setae. Fig. 4. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 5. S. merkelii (G. Horn) (9 km SE of Lowell, Idaho County, Idaho). Fig. 6. S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho). Fig. 7. S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13154923" httpUri="https://zenodo.org/record/13154923/files/figure.png" pageId="12" pageNumber="393">Fig. 6b</figureCitation>
) with pad of adhesive setae ventrally over virtually entire ventral surface, protarsomere 4 without a pad; elytra with 13–16 apparent striae, with at least lateral striae disrupted by punctures in and between striae, elytral intervals 4 and 8 each with two to six discal setiferous pores; median lobe of male aedeagus (
<figureCitation id="13229EF6F917615BFCDFFA31B554FA0E" box="[776,852,1476,1500]" captionStart="FIGURES" captionStartId="11.[155,167,1575,1594]" captionText="FIGURES 8–12. Digital photographs of median lobe of male aedeagus of Scaphinotus (Pseudonomaretus) species; a, dor- sal aspect; b, left lateral aspect; is = internal sac. Fig. 8. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 9. S. merkelii (G. Horn) (Moscow Mountain, Latah County, Idaho). Fig. 10. S. mannii Wick- ham (Wawawai, Whitman County, Washington). Fig. 11. S. regularis (LeConte) (Moscow Mountain, Latah County, Idaho). Fig. 12. S. relictus (G. Horn) (Mount Spokane, Spokane County, Washington). Scale lines = 1.0 mm." pageId="12" pageNumber="393">Fig. 12</figureCitation>
) moderate in thickness and of nearly equal thickness throughout, ventral margin arcuate in basal one-fourth, straight in middle half, and slightly deflected ventrally in apical one-fourth in lateral aspect (
<figureCitation id="13229EF6F917615BFC70F9F1B201F9CE" box="[935,1025,1540,1565]" captionStart="FIGURES" captionStartId="11.[155,167,1575,1594]" captionText="FIGURES 8–12. Digital photographs of median lobe of male aedeagus of Scaphinotus (Pseudonomaretus) species; a, dor- sal aspect; b, left lateral aspect; is = internal sac. Fig. 8. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 9. S. merkelii (G. Horn) (Moscow Mountain, Latah County, Idaho). Fig. 10. S. mannii Wick- ham (Wawawai, Whitman County, Washington). Fig. 11. S. regularis (LeConte) (Moscow Mountain, Latah County, Idaho). Fig. 12. S. relictus (G. Horn) (Mount Spokane, Spokane County, Washington). Scale lines = 1.0 mm." pageId="12" pageNumber="393">Fig. 12b</figureCitation>
), widest near basal one-fourth of shaft length, straight, gradually tapered to a flattened point apically in dorsal aspect (
<figureCitation id="13229EF6F917615BFF19F9B0B726F98E" box="[206,294,1605,1629]" captionStart="FIGURES" captionStartId="11.[155,167,1575,1594]" captionText="FIGURES 8–12. Digital photographs of median lobe of male aedeagus of Scaphinotus (Pseudonomaretus) species; a, dor- sal aspect; b, left lateral aspect; is = internal sac. Fig. 8. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 9. S. merkelii (G. Horn) (Moscow Mountain, Latah County, Idaho). Fig. 10. S. mannii Wick- ham (Wawawai, Whitman County, Washington). Fig. 11. S. regularis (LeConte) (Moscow Mountain, Latah County, Idaho). Fig. 12. S. relictus (G. Horn) (Mount Spokane, Spokane County, Washington). Scale lines = 1.0 mm." pageId="12" pageNumber="393">Fig. 12a</figureCitation>
), armature of internal sac as in
<figureCitation id="13229EF6F917615BFDABF9B0B4CFF98E" box="[636,719,1605,1629]" captionStart="FIGURES" captionStartId="11.[155,167,1575,1594]" captionText="FIGURES 8–12. Digital photographs of median lobe of male aedeagus of Scaphinotus (Pseudonomaretus) species; a, dor- sal aspect; b, left lateral aspect; is = internal sac. Fig. 8. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 9. S. merkelii (G. Horn) (Moscow Mountain, Latah County, Idaho). Fig. 10. S. mannii Wick- ham (Wawawai, Whitman County, Washington). Fig. 11. S. regularis (LeConte) (Moscow Mountain, Latah County, Idaho). Fig. 12. S. relictus (G. Horn) (Mount Spokane, Spokane County, Washington). Scale lines = 1.0 mm." pageId="12" pageNumber="393">Fig. 12.</figureCitation>
</paragraph>
<paragraph id="8BA68273F917615BFF7CF990B7E9F96F" blockId="12.[123,1174,1154,1726]" pageId="12" pageNumber="393">
<emphasis id="B96D5E61F917615BFF7CF990B7EFF9A9" bold="true" box="[171,495,1637,1661]" pageId="12" pageNumber="393">
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<smallCapsWord id="8D4014AFF917615BFE8CF99DB7EFF9A9" baselines="1654" box="[347,495,1640,1659]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="12" pageNumber="393">DISTRIBUTION</smallCapsWord>
</emphasis>
.— This species is known from southwestern
<collectingRegion id="49DD4C91F917615BFC07F990B220F9AF" box="[976,1056,1637,1661]" country="Canada" name="Alberta" pageId="12" pageNumber="393">Alberta</collectingRegion>
and southeastern
<collectingRegion id="49DD4C91F917615BFF0FF970B79CF94F" box="[216,412,1669,1693]" country="Canada" name="British Columbia" pageId="12" pageNumber="393">British Columbia</collectingRegion>
, northwestern
<collectingRegion id="49DD4C91F917615BFD86F973B4B0F94F" box="[593,688,1670,1693]" country="United States of America" name="Montana" pageId="12" pageNumber="393">Montana</collectingRegion>
, northern
<collectingRegion id="49DD4C91F917615BFCE3F970B571F94F" box="[820,881,1669,1693]" country="United States of America" name="Idaho" pageId="12" pageNumber="393">Idaho</collectingRegion>
and eastern
<collectingRegion id="49DD4C91F917615BFBC1F970B296F94F" box="[1046,1174,1669,1693]" country="United States of America" name="Washington" pageId="12" pageNumber="393">Washington</collectingRegion>
(
<bibRefCitation id="EF88FF82F917615BFF53F950B72CF96C" author="GIDASPOW, T." box="[132,300,1701,1726]" pageId="12" pageNumber="393" pagination="53 - 98" refId="ref7559" refString="GIDASPOW, T. 1973. Revision of ground beetles of American genus Cychrus and four subgenera of genus Scaphinotus (Coleoptera, Carabidae). Bulletin of the American Museum of Natural History 152: 53 - 98." type="journal article" year="1973">Gidaspow 1973</bibRefCitation>
;
<bibRefCitation id="EF88FF82F917615BFEECF950B7D9F96C" author="BERGDAHL, J. C." box="[315,473,1701,1726]" pageId="12" pageNumber="393" pagination="1 - 12" refId="ref7343" refString="BERGDAHL, J. C. 2014. Carabid beetles (Coleoptera, Carabidae) of the Pacific Northwest, USA: Scaphinotus (Pseudonomaretus) manni Wickham (Cychrini). Bulletin of the Oregon Entomological Society, Winter 2014 / 2015: 1 - 12." type="journal article" year="2014">Bergdahl 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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