154 lines
16 KiB
XML
154 lines
16 KiB
XML
<document id="C2FF4DCD032EBA743638A323E5D22A6C" ID-CLB-Dataset="5843" ID-DOI="10.1206/3908.1" ID-GBIF-Dataset="2fc56743-3f64-4d91-9433-d5fe41fd7d1d" ID-ISSN="0003-0082" ID-Zenodo-Dep="4598569" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615482533448" checkinUser="felipe" docAuthor="Grimaldi, David A., Sunderlin, David, Aaroe, Georgene A., Dempsky, Michelle R., Parker, Nancy E., Tillery, George Q., White, Jaclyn G., Barden, Phillip, Nascimbene, Paul C. & Williams, Christopher J." docDate="2018" docId="03A187BFFF982F33AB9B425950C5FBF9" docLanguage="en" docName="AmMusNovit.2018.3908.1-37.pdf.imf" docOrigin="American Museum Novitates 2018 (3908)" docSource="http://www.bioone.org/doi/10.1206/3908.1" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Blattodea" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="FF98FFC7FF8B2F20AA7C43405143FF90" masterDocTitle="Biological Inclusions in Amber from the Paleogene Chickaloon Formation of Alaska" masterLastPageNumber="37" masterPageNumber="1" pageNumber="20" updateTime="1698933204900" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F3C06F8FE4F53030B48FFE76C488D4B3">Biological Inclusions in Amber from the Paleogene Chickaloon Formation of Alaska</mods:title>
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<mods:namePart id="CD63A53CDAB83C32A8FEC2AC07A89F50">Grimaldi, David A.</mods:namePart>
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<mods:affiliation id="C730851A91D97B2F025CE1EBC8EB2329">Division of Invertebrate Zoology, American Museum of Natural History, New York.</mods:affiliation>
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<mods:namePart id="ACF42E3B93BD55407AAD33CDE9470C08">Sunderlin, David</mods:namePart>
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<mods:affiliation id="F723478BF1D4BBFB70A08512D74A4C20">Department of Geology & Environmental Geosciences, Lafayette College, Easton, Pennsylvania.</mods:affiliation>
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<mods:namePart id="2BE999B7207C1472593BD2DACB465BF7">Aaroe, Georgene A.</mods:namePart>
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<mods:affiliation id="AC5AE8BE50D318754D317116530E97DE">Department of Geology & Environmental Geosciences, Lafayette College, Easton, Pennsylvania.</mods:affiliation>
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<mods:namePart id="4A55CA91CF98341A3F2CA4B79FC79674">Dempsky, Michelle R.</mods:namePart>
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<mods:affiliation id="5F042F0800253F21F3120D5F861A22B5">Department of Geology & Environmental Geosciences, Lafayette College, Easton, Pennsylvania.</mods:affiliation>
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<mods:namePart id="464D1C9C761C570D450B2FFC7CD5FBC1">Parker, Nancy E.</mods:namePart>
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<mods:affiliation id="B68FB540FEFD091B6CD21E4BF7C6F258">Department of Geology & Environmental Geosciences, Lafayette College, Easton, Pennsylvania.</mods:affiliation>
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<mods:namePart id="EE4C0599B7B09B90671FB80ABB6D15F6">Tillery, George Q.</mods:namePart>
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<mods:affiliation id="D916605AB94CAA621D5FA95362F306CD">Department of Geology & Environmental Geosciences, Lafayette College, Easton, Pennsylvania.</mods:affiliation>
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<mods:namePart id="DE1CB083DE3B42A892FD26DE994AA407">White, Jaclyn G.</mods:namePart>
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<mods:affiliation id="CF3E7D2C0468758B5E1C00DF19E354B8">Department of Geology & Environmental Geosciences, Lafayette College, Easton, Pennsylvania.</mods:affiliation>
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<mods:namePart id="D40D5ACB998E4DDB04BC0CCB6D1DF9B2">Barden, Phillip</mods:namePart>
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<mods:affiliation id="606BE73372697066745F4C3E6BD41317">Department of Biology, New Jersey Institute of Technology, Newark.</mods:affiliation>
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<mods:namePart id="AB78496690DE4DEB0F02CFAEE2280B65">Nascimbene, Paul C.</mods:namePart>
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<mods:affiliation id="D2A42F748E4817217BBF9A746D81AAAD">Division of Invertebrate Zoology, American Museum of Natural History, New York.</mods:affiliation>
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<mods:namePart id="30D2E1B8C92FDD15444BFD3D1D716159">Williams, Christopher J.</mods:namePart>
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<mods:affiliation id="48A1F8DB940AB3E8E5D070EB5B02E1B0">Department of Earth & Environment, Franklin & Marshall College, Lancaster, Pennsylvania.</mods:affiliation>
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<mods:date id="6BD6CDFBF4FCDFCAB7D18909B3127880">2018</mods:date>
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<mods:number id="31C6415A5DC0B5CC6BD1A858041F6E3C">2018-09-28</mods:number>
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<mods:number id="0EE7FFA56DDFA4C45CF1E333AE70585C">2018</mods:number>
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<mods:number id="D3FFEEC716953D97DB40FCD71228C2A2">3908</mods:number>
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<treatment id="03A187BFFF982F33AB9B425950C5FBF9" ID-DOI="http://doi.org/10.5281/zenodo.4598229" ID-GBIF-Taxon="183232801" ID-Zenodo-Dep="4598229" LSID="urn:lsid:plazi:treatment:03A187BFFF982F33AB9B425950C5FBF9" httpUri="http://treatment.plazi.org/id/03A187BFFF982F33AB9B425950C5FBF9" lastPageNumber="20" pageId="19" pageNumber="20">
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<subSubSection id="C3126522FF982F33AB9B42595202FEA3" box="[487,833,281,307]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8BB736A9FF982F33AB9B42595202FEA3" blockId="19.[487,833,281,307]" box="[487,833,281,307]" pageId="19" pageNumber="20">
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<heading id="D0FF81C5FF982F33AB9B42595202FEA3" allCaps="true" box="[487,833,281,307]" centered="true" fontSize="36" level="1" pageId="19" pageNumber="20" reason="1">
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<taxonomicName id="4C084D2AFF982F33AB9B42595202FEA3" ID-CoL="TM" ID-ENA="85823" authority="(COCKROACHES)" baseAuthorityName="COCKROACHES" box="[487,833,281,307]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">BLATTODEA (COCKROACHES)</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126522FF982F33A825421150C5FBF9" pageId="19" pageNumber="20" type="description">
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<paragraph id="8BB736A9FF982F33A8254211538CFEFC" blockId="19.[601,719,337,364]" box="[601,719,337,364]" pageId="19" pageNumber="20">
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<heading id="D0FF81C5FF982F33A8254211538CFEFC" box="[601,719,337,364]" centered="true" fontSize="36" level="3" pageId="19" pageNumber="20" reason="7">
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<figureCitation id="13332A2CFF982F33A8254211538CFEFC" box="[601,719,337,364]" captionStart="FIG" captionStartId="10.[108,150,1224,1245]" captionTargetBox="[108,1212,224,1198]" captionTargetId="figure-10@10.[108,1212,224,1198]" captionTargetPageId="10" captionText="FIG. 7. Various insect inclusions. A. AMNH WH-12, with legs from a roach (Blattodea), and the legs and distal portions of two wings from a caddisfly (Trichoptera). B. AMNH LC-A4, nymph of Heteroptera (Hemiptera). C, D. AMNH LC-B6, nymphal thrips (Thysanoptera), habitus (C) and detail of antennae (D). E. AMNH WH-3, with inclusion of entire male Chironomidae: Tanypodinae, and juvenile spider." figureDoi="http://doi.org/10.5281/zenodo.4598585" httpUri="https://zenodo.org/record/4598585/files/figure.png" pageId="19" pageNumber="20">Figure 7A</figureCitation>
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</heading>
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</paragraph>
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<paragraph id="8BB736A9FF982F33AAE042CA5577FCAE" blockId="19.[108,1214,393,1129]" pageId="19" pageNumber="20">
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Two pieces of amber, AMNH WH-12 and AMNH WH-11, contain fragmentary remains of small roaches, probably nymphs. WH-12 contains a portion of one leg: apical portion of the femur and the entire tibia and tarsus. The tibia has 18 spinelike setae (including a pair of apical tibial spurs), in roughly four longitudinal rows. WH-11 contains portions of three legs and an antenna, all very dark and surrounded by a dark reddish “halo” of pyritization/oxidation. One leg is preserved as a portion of a femur; the tarsi and portions of the tibia are preserved for the other two legs. Approximately 30 filiform flagellomeres are preserved; the basal ones short (lengths about twice the width), gradually increasing in length distad to about five times the width. Leg segments of WH-11 are shorter than in WH-12, and tarsomere four has a long ventral lobe. Thus, there appears to be two taxa of roaches. Though these specimens cannot be identified to family or superfamily, they are clearly
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<taxonomicName id="4C084D2AFF982F33A8B141BF527FFC89" box="[717,828,767,793]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
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based on the antennal structure and the tibial spines, which further have minute serrations on the ventral margin.
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</paragraph>
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<paragraph id="8BB736A9FF982F33AAE0400950C5FBF9" blockId="19.[108,1214,393,1129]" pageId="19" pageNumber="20">
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The fossil record and natural history of roaches are reviewed by
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<bibRefCitation id="EF994B58FF982F33A902400955FBFCF4" author="Grimaldi, D. A. & M. S. Engel" box="[894,1208,841,868]" pageId="19" pageNumber="20" refId="ref12927" refString="Grimaldi, D. A., and M. S. Engel. 2005. Evolution of the insects. New York: Cambridge University Press." type="book" year="2005">Grimaldi and Engel (2005)</bibRefCitation>
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. The natural distribution of
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<taxonomicName id="4C084D2AFF982F33ABD7402F5359FC19" box="[427,538,879,905]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
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worldwide is almost entirely tropical to warm temperate; pest species reach higher latitudes in association with human habitations. Of the 69 species of roaches in North America, 24 are introduced from other regions, and only three native species (
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<taxonomicName id="4C084D2AFF982F33AA0A409F50DAFC69" baseAuthorityName="De Geer" baseAuthorityYear="1773" box="[118,409,991,1017]" class="Insecta" family="Ectobiidae" genus="Parcoblatta" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pensylvanica">
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<emphasis id="B97CEABBFF982F33AA0A409F50DAFC69" box="[118,409,991,1017]" italics="true" pageId="19" pageNumber="20">Parcoblatta pensylvanica</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C084D2AFF982F33ABD6409F537AFC69" baseAuthorityName="Saussure" baseAuthorityYear="1862" box="[426,569,991,1017]" class="Insecta" family="Ectobiidae" genus="Parcoblatta" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="uhleriana">
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<emphasis id="B97CEABBFF982F33ABD6409F537AFC69" box="[426,569,991,1017]" italics="true" pageId="19" pageNumber="20">P. uhleriana</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C084D2AFF982F33A8FC409F5240FC68" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1865" box="[640,771,991,1016]" class="Insecta" family="Ectobiidae" genus="Parcoblatta" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="virginica">
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<emphasis id="B97CEABBFF982F33A8FC409F5240FC68" box="[640,771,991,1016]" italics="true" pageId="19" pageNumber="20">P. virginica</emphasis>
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</taxonomicName>
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) have distributions that extend into southernmost
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<collectingRegion id="49CCF84BFF982F33AB6447445036FB8E" box="[280,373,1028,1054]" country="Canada" name="Ontario" pageId="19" pageNumber="20">Ontario</collectingRegion>
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and
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<collectingRegion id="49CCF84BFF982F33ABCF4744534EFB8E" box="[435,525,1028,1054]" country="Canada" name="Quebec" pageId="19" pageNumber="20">Québec</collectingRegion>
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(Vickery and McKevan, 1985;
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<bibRefCitation id="EF994B58FF982F33A9FC47445538FB8E" author="Atkinson, T. H. & P. G. Koehler & R. S. Patterson" box="[896,1147,1028,1055]" pageId="19" pageNumber="20" pagination="1 - 86" refId="ref12003" refString="Atkinson, T. H., P. G. Koehler, and R. S. Patterson. 1991. Catalog and atlas of the cockroaches (Dictyoptera) of North America north of Mexico. Miscellaneous Publications of the Entomological Society of America 78: 1 - 86." type="journal article" year="1991">Atkinson et al., 1991</bibRefCitation>
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). The roaches are the most obvious example in Chickaloon amber of a taxon that has retreated from high northern latitudes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |