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<document id="910D42A07BB92D93AEB8D5324F92A416" ID-DOI="10.5281/zenodo.170449" ID-GBIF-Dataset="7053f917-ef1a-4864-98aa-ec3d5d4edec7" ID-ISSN="1175­5326" ID-Zenodo-Dep="170449" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459748772729" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A." docDate="2005" docId="03A187A2FF94FFB6FED0FD1AFD08FADF" docLanguage="en" docName="zt01084p064.pdf" docOrigin="Zootaxa 1084" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Arctodiamesa marinae Makarchenko, 2005, sp. n." docType="treatment" docVersion="9" lastPageNumber="63" masterDocId="FF98FFDAFF95FFB2FFD8FFBCFFA8FFAE" masterDocTitle="A new species of Arctodiamesa Makarchenko (Diptera: Chironomidae: Diamesinae) from the Russian Far East, with a key to known species of the genus" masterLastPageNumber="64" masterPageNumber="59" pageNumber="60" updateTime="1698211150242" updateUser="plazi">
<mods:mods id="02E8D9B1765226542D942367775D1975" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1D635039C35900A571FAF431B2C19092">
<mods:title id="F81AF66428998BE8B55E4B6EDD9D77D2">A new species of Arctodiamesa Makarchenko (Diptera: Chironomidae: Diamesinae) from the Russian Far East, with a key to known species of the genus</mods:title>
</mods:titleInfo>
<mods:name id="BD6C9960D5A95F61B1EA29C0B85569B3" type="personal">
<mods:role id="4356B0C7E241E28587EC70FCA57ACE61">
<mods:roleTerm id="585CB89FA8072C77BD997E9DE73D773C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90735FDB1850C3C8C8B88F35D9C66D8E">Makarchenko, Eugenyi A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3EE99B69117789E246D1AA087EA053F3">text</mods:typeOfResource>
<mods:relatedItem id="13C7DE17C2AA67BE25EEC8432CDD6836" type="host">
<mods:titleInfo id="841EADE7606FA3C98525278860A969A4">
<mods:title id="BE468FB0C72F2316015C3BB77465661C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6BFFC9FF2C8A3A46780F802E8486B64D">
<mods:date id="EDEFD0ADAB4E1CBE07037CF1DECC4003">2005</mods:date>
<mods:detail id="0D0142A6493B6B14352CB06124996B63" type="volume">
<mods:number id="578D43B6110EDBDBBE37B28BB76F1670">1084</mods:number>
</mods:detail>
<mods:extent id="8159A477FE687C1B6F7ED3281DDF384B" unit="page">
<mods:start id="26D3BAAAC48A8F30E250A7CDCE7DD405">59</mods:start>
<mods:end id="451B048AEC06A7E2A4D17BFF358B50E6">64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="91BFB1FB5EDD2EDE8FF35919D98B4C8D">journal article</mods:classification>
<mods:identifier id="9E30F6903D9E2CEC9E29682C39D3814E" type="DOI">10.5281/zenodo.170449</mods:identifier>
<mods:identifier id="5C6297CEC388D99193EFC46861DD69A4" type="GBIF-Dataset">7053f917-ef1a-4864-98aa-ec3d5d4edec7</mods:identifier>
<mods:identifier id="2B1E833312CA20ED3046709732F2DFB0" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="09EA57D9FD490E0968AC06C515601605" type="Zenodo-Dep">170449</mods:identifier>
</mods:mods>
<treatment id="03A187A2FF94FFB6FED0FD1AFD08FADF" ID-DOI="http://doi.org/10.5281/zenodo.6266220" ID-GBIF-Taxon="119337173" ID-Zenodo-Dep="6266220" LSID="urn:lsid:plazi:treatment:03A187A2FF94FFB6FED0FD1AFD08FADF" httpUri="http://treatment.plazi.org/id/03A187A2FF94FFB6FED0FD1AFD08FADF" lastPageId="4" lastPageNumber="63" pageId="1" pageNumber="60">
<subSubSection id="C312653FFF94FFB3FED0FD1AFE3FFD4D" pageId="1" pageNumber="60" type="nomenclature">
<paragraph id="8BB736B4FF94FFB3FED0FD1AFDCCFD6E" blockId="1.[264,612,678,739]" box="[264,612,678,704]" pageId="1" pageNumber="60">
<heading id="D0FF81D8FF94FFB3FED0FD1AFDCCFD6E" bold="true" box="[264,612,678,704]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
<emphasis id="B97CEAA6FF94FFB3FED0FD1AFDCCFD6E" bold="true" box="[264,612,678,704]" pageId="1" pageNumber="60">
<taxonomicName id="4C084D37FF94FFB3FED0FD1AFDB2FD6E" ID-CoL="67S2C" box="[264,538,678,704]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B97CEAA6FF94FFB3FED0FD1AFDB2FD6E" bold="true" box="[264,538,678,704]" italics="true" pageId="1" pageNumber="60">Arctodiamesa marinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57DDFF94FFB3FDFAFD1AFDCCFD6E" box="[546,612,678,704]" pageId="1" pageNumber="60" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736B4FF94FFB3FED0FD75FE3FFD4D" blockId="1.[264,612,678,739]" box="[264,407,713,739]" pageId="1" pageNumber="60">
(
<figureCitation id="13332A31FF94FFB3FEC9FD75FE26FD4D" box="[273,398,713,739]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 17" captionStartId-0="2.[264,382,1523,1547]" captionStartId-1="3.[264,382,1745,1769]" captionTargetBox-0="[302,1280,558,1479]" captionTargetBox-1="[311,1284,307,1717]" captionTargetId-0="figure@2.[302,1288,555,1489]" captionTargetId-1="figure@3.[297,1290,300,1717]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. Male of Arctodiamesa marinae sp. n. 1, hypopygium in dorsal view; 2, gonostylus deflected from gonocoxite; 3, gonostylus pressed to gonocoxite; 4, sternapodeme and medial aedeagal lobe. Scale bars = 50 µm." captionText-1="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri-0="https://zenodo.org/record/170450/files/figure.png" httpUri-1="https://zenodo.org/record/170451/files/figure.png" pageId="1" pageNumber="60">Figs. 1–17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C312653FFF94FFB3FED0FCA5FE19FC7D" pageId="1" pageNumber="60" type="materials_examined">
<paragraph id="8BB736B4FF94FFB3FED0FCA5FE19FC7D" blockId="1.[264,1323,793,1632]" pageId="1" pageNumber="60">
<emphasis id="B97CEAA6FF94FFB3FED0FCA5FED4FC9D" bold="true" box="[264,380,793,819]" pageId="1" pageNumber="60">Material.</emphasis>
<typeStatus id="54B38816FF94FFB3FE52FCA5FE54FC9D" box="[394,508,793,819]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
: male,
<collectingCountry id="F31F7624FF94FFB3FD83FCA5FD04FC9D" box="[603,684,793,819]" name="Russia" pageId="1" pageNumber="60">Russia</collectingCountry>
, Kedrovaya River, Kedrovaya Pad Nature Reserve, Khasansk District, Primorye Territory,
<geoCoordinate id="EE3C5073FF94FFB3FD07FCFDFCE6FCF5" box="[735,846,833,859]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="925" value="43.083332">N 43°05’</geoCoordinate>
874”,
<geoCoordinate id="EE3C5073FF94FFB3FC42FCFDFBBAFCF5" box="[922,1042,833,859]" direction="east" orientation="longitude" pageId="1" pageNumber="60" precision="925" value="131.55">E 131°33’</geoCoordinate>
412”,
<date id="FFB61074FF94FFB3FBB9FCFDFB4DFCF5" box="[1121,1253,833,859]" pageId="1" pageNumber="60" value="2005-06-04">4.VI. 2005</date>
, light trap, leg. E. Makarchenko.
<typeStatus id="54B38816FF94FFB3FD94FCD5FD6BFC2D" box="[588,707,873,899]" pageId="1" pageNumber="60" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0EFD3DFF94FFB3FD16FCD5FC81FC2D" box="[718,809,873,899]" pageId="1" pageNumber="60" type="male">3 males</specimenCount>
reared from pupae,
<specimenCount id="9D0EFD3DFF94FFB3FBC3FCD5FBD4FC2D" box="[1051,1148,873,899]" pageId="1" pageNumber="60" type="pupa">4 pupae</specimenCount>
,
<specimenCount id="9D0EFD3DFF94FFB3FB50FCD5FB70FC2D" box="[1160,1240,873,899]" pageId="1" pageNumber="60" type="pupa">1 pupa</specimenCount>
reared from larva, 5 fourth­instar larvae, same data as
<typeStatus id="54B38816FF94FFB3FC81FC2DFC17FC05" box="[857,959,913,939]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
except
<date id="FFB61074FF94FFB3FBFCFC2DFB6FFC05" box="[1060,1223,913,939]" pageId="1" pageNumber="60" value="2005-06-02" valueMax="2005-06-04" valueMin="2005-06-02">2–4.VI. 2005</date>
, leg. E. Makarchenko.
</paragraph>
</subSubSection>
<subSubSection id="C312653FFF94FFB6FEE0FC5DFC8EFD67" lastPageId="4" lastPageNumber="63" pageId="1" pageNumber="60" type="description">
<paragraph id="8BB736B4FF94FFB3FEE0FC5DFB3BFB8D" blockId="1.[264,1323,793,1632]" pageId="1" pageNumber="60">
<emphasis id="B97CEAA6FF94FFB3FEE0FC5DFE65FC55" bold="true" box="[312,461,993,1019]" pageId="1" pageNumber="60">Male imago</emphasis>
(
<emphasis id="B97CEAA6FF94FFB3FE39FC5DFE58FC54" box="[481,496,993,1018]" italics="true" pageId="1" pageNumber="60">n</emphasis>
= 4, except when otherwise stated). Total length
<quantity id="4CF09B51FF94FFB3FB91FC5DFB75FC55" box="[1097,1245,993,1019]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="1" pageNumber="60" unit="mm" value="4.1" valueMax="4.3" valueMin="3.9">3.9–4.3 mm</quantity>
. Wing length
<quantity id="4CF09B51FF94FFB3FE80FBB5FDAFFB8D" box="[344,519,1033,1059]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.28" metricValueMin="3.12" pageId="1" pageNumber="60" unit="mm" value="3.2" valueMax="3.28" valueMin="3.12">3.12–3.28 mm</quantity>
. Total length/wing length 1.25–1.31. Coloration brown.
</paragraph>
<paragraph id="8BB736B4FF94FFB3FEE0FB8DFE3CFABD" blockId="1.[264,1323,793,1632]" pageId="1" pageNumber="60">
Head. Eyes hairy. Temporal setae including only 14–16 postorbitals, verticals absent. Clypeus with 6 setae. Antenna with 13 flagellomeres and well­developed plume; pedicel with 2 or 3 setae. Length of subapical seta of terminal flagellomere
<quantity id="4CF09B51FF94FFB3FBECFB3DFB19FB35" box="[1076,1201,1153,1179]" metricMagnitude="1" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="1" pageNumber="60" unit="m" value="42.0" valueMax="44.0" valueMin="40.0">
40–
<specimenCount id="9D0EFD3DFF94FFB3FBB9FB3DFB19FB35" box="[1121,1201,1153,1179]" pageId="1" pageNumber="60" type="generic">44 m</specimenCount>
</quantity>
. AR 1.29. Lengths (m) of palpomeres 1­5: 44–48: 108–120: 124–160: 124–160: 150–172; second palpomere with sensilla capitata (diameter about
<quantity id="4CF09B51FF94FFB3FC87FB6DFC0BFB45" box="[863,931,1233,1259]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="60" unit="m" value="4.0">4 m</quantity>
) in distal part. Head width/palp length 1.34.
</paragraph>
<paragraph id="8BB736B4FF94FFB3FEE0FA9DFCC1FACD" blockId="1.[264,1323,793,1632]" pageId="1" pageNumber="60">Thorax. Antepronotum with 11–13 lateral setae. Acrostichals absent, dorsocentrals 16–21, prealars 13–16. Scutellum with 34–37 setae.</paragraph>
<paragraph id="8BB736B4FF94FFB3FEE0FACDFD57FA4F" blockId="1.[264,1323,793,1632]" pageId="1" pageNumber="60">Wing. Anal lobe well developed. Squama with 41–43 setae. R with 12–15 setae, R1 with 2 campaniform sensillae and 8 or 9 setae, R4+5 with 2 or 3 setae subapically. R2+3 faint with 2 campaniform sensillae in basal part.</paragraph>
<paragraph id="8BB736B4FF94FFB3FEE0FA53FB4AF9F2" blockId="1.[264,1323,793,1632]" pageId="1" pageNumber="60">
Legs. BR1 2.7, BR2 2.0, BR3 4.3. Spur of front tibia
<specimenCount id="9D0EFD3DFF94FFB3FC72FA53FC57F9A7" box="[938,1023,1519,1545]" pageId="1" pageNumber="60" type="generic">
<quantity id="4CF09B51FF94FFB3FC72FA53FC53F9A7" box="[938,1019,1519,1545]" metricMagnitude="1" metricUnit="m" metricValue="9.2" pageId="1" pageNumber="60" unit="m" value="92.0">92 m</quantity>
.
</specimenCount>
Spurs of middle tibia
<quantity id="4CF09B51FF94FFB3FAD5FA53FE98F99A" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="60" unit="m" value="48.0">48 m</quantity>
and
<quantity id="4CF09B51FF94FFB3FEB2F9A6FE1FF99A" box="[362,439,1562,1588]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="60" unit="m" value="40.0">40 m</quantity>
long, of hind tibia
<quantity id="4CF09B51FF94FFB3FD42F9A6FD4FF99A" box="[666,743,1562,1588]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="60" unit="m" value="76.0">76 m</quantity>
and
<quantity id="4CF09B51FF94FFB3FCF8F9A6FCC5F99A" box="[800,877,1562,1588]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="60" unit="m" value="44.0">44 m</quantity>
long.
<collectingCountry id="F31F7624FF94FFB3FC60F9A6FC5BF99A" box="[952,1011,1562,1588]" name="India" pageId="1" pageNumber="60">Hind</collectingCountry>
tibial comb with 10 setae. Middle ta1 with 2 pseudospurs
<quantity id="4CF09B51FF94FFB3FDAAF9FEFD17F9F2" box="[626,703,1602,1628]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="60" unit="m" value="28.0">28 m</quantity>
long, hind ta1 with 2 pseudospurs
<quantity id="4CF09B51FF94FFB3FB8AF9FEFB37F9F2" box="[1106,1183,1602,1628]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="60" unit="m" value="36.0">36 m</quantity>
long.
</paragraph>
<paragraph id="8BB736B4FF94FFB0FED0F936FE1DFE50" blockId="1.[264,799,1674,1700]" lastBlockId="2.[264,1323,284,510]" lastPageId="2" lastPageNumber="61" pageId="1" pageNumber="60">
Lengths (m) and proportions of legs (
<emphasis id="B97CEAA6FF94FFB3FD08F937FD77F90A" box="[720,735,1675,1700]" italics="true" pageId="1" pageNumber="60">n</emphasis>
= 1): Hypopygium (
<figureCitation id="13332A31FF97FFB0FE3BFEA0FDFCFE98" box="[483,596,284,310]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1523,1547]" captionTargetBox="[302,1280,558,1479]" captionTargetId="figure@2.[302,1288,555,1489]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male of Arctodiamesa marinae sp. n. 1, hypopygium in dorsal view; 2, gonostylus deflected from gonocoxite; 3, gonostylus pressed to gonocoxite; 4, sternapodeme and medial aedeagal lobe. Scale bars = 50 µm." httpUri="https://zenodo.org/record/170450/files/figure.png" pageId="2" pageNumber="61">Figs. 1–4</figureCitation>
). Tergite IX with 30 setae and narrow, anal point
<quantity id="4CF09B51FF97FFB0FB43FEA0FB40FE98" box="[1179,1256,284,310]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="61" unit="m" value="68.0">68 m</quantity>
long. Laterosternite IX with 12 setae. Transverse sternapodeme
<quantity id="4CF09B51FF97FFB0FC61FEF8FBBFFEF0" box="[953,1047,324,350]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="2" pageNumber="61" unit="m" value="144.0">144 m</quantity>
wide. Medial aedeagal lobe weakly sclerotized, with strong apical spines (
<figureCitation id="13332A31FF97FFB0FCBAFED0FC03FE28" box="[866,939,364,390]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1523,1547]" captionTargetBox="[302,1280,558,1479]" captionTargetId="figure@2.[302,1288,555,1489]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male of Arctodiamesa marinae sp. n. 1, hypopygium in dorsal view; 2, gonostylus deflected from gonocoxite; 3, gonostylus pressed to gonocoxite; 4, sternapodeme and medial aedeagal lobe. Scale bars = 50 µm." httpUri="https://zenodo.org/record/170450/files/figure.png" pageId="2" pageNumber="61">Fig. 4</figureCitation>
). Gonocoxite
<quantity id="4CF09B51FF97FFB0FB8AFED0FB4FFE28" box="[1106,1255,364,390]" metricMagnitude="2" metricUnit="m" metricValue="2.86" metricValueMax="2.88" metricValueMin="2.84" pageId="2" pageNumber="61" unit="m" value="286.0" valueMax="288.0" valueMin="284.0">284–288 m</quantity>
long, superior and inferior volsellae reduced. Gonostylus
<quantity id="4CF09B51FF97FFB0FC50FE28FC40FE00" box="[904,1000,404,430]" metricMagnitude="2" metricUnit="m" metricValue="1.56" pageId="2" pageNumber="61" unit="m" value="156.0">156 m</quantity>
long, with rounded, wide (
<quantity id="4CF09B51FF97FFB0FEC8FE00FE23FE78" box="[272,395,444,470]" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="2" pageNumber="61" unit="m" value="38.0" valueMax="40.0" valueMin="36.0">36–40 m</quantity>
) lobe in basal half and short projection in subapical part; megaseta
<quantity id="4CF09B51FF97FFB0FB45FE00FB40FE78" box="[1181,1256,444,470]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="61" unit="m" value="20.0">20 m</quantity>
long. HR 1.82–1.85.
</paragraph>
<paragraph id="8BB736B4FF94FFB3FE92F971FC66F8C6" pageId="1" pageNumber="60">
<table id="F908C414FF94004DFED0F971FC66F8C2" box="[264,974,1741,1900]" gridcols="10" gridrows="4" pageId="1" pageNumber="60">
<tr id="353834F6FF94004DFED0F971FC66F945" box="[264,974,1741,1771]" gridrow="0" pageId="1" pageNumber="60">
<th id="76E95D8AFF94004DFED0F971FECFF945" box="[264,359,1741,1771]" gridcol="0" gridrow="0" pageId="1" pageNumber="60">fe</th>
<th id="76E95D8AFF94004DFEA2F971FE1FF945" box="[378,439,1741,1771]" gridcol="1" gridrow="0" pageId="1" pageNumber="60">ti</th>
<th id="76E95D8AFF94004DFE13F971FE54F945" box="[459,508,1741,1771]" gridcol="2" gridrow="0" pageId="1" pageNumber="60">ta 1</th>
<th id="76E95D8AFF94004DFDD5F971FD93F945" box="[525,571,1741,1771]" gridcol="3" gridrow="0" pageId="1" pageNumber="60">ta2</th>
<th id="76E95D8AFF94004DFD97F971FDD5F945" box="[591,637,1741,1771]" gridcol="4" gridrow="0" pageId="1" pageNumber="60">ta3</th>
<th id="76E95D8AFF94004DFD56F971FD18F945" box="[654,688,1741,1771]" gridcol="5" gridrow="0" pageId="1" pageNumber="60">ta4</th>
<th id="76E95D8AFF94004DFD1CF971FD5AF945" box="[708,754,1741,1771]" gridcol="6" gridrow="0" pageId="1" pageNumber="60">ta5</th>
<th id="76E95D8AFF94004DFCDDF971FC93F945" box="[773,827,1741,1771]" gridcol="7" gridrow="0" pageId="1" pageNumber="60">LR</th>
<th id="76E95D8AFF94004DFC96F971FC2CF945" box="[846,900,1741,1771]" gridcol="8" gridrow="0" pageId="1" pageNumber="60">SV</th>
<th id="76E95D8AFF94004DFC4FF971FC66F945" box="[919,974,1741,1771]" gridcol="9" gridrow="0" pageId="1" pageNumber="60">BV</th>
</tr>
<tr id="353834F6FF94004DFED0F944FC66F8B8" box="[264,974,1784,1814]" gridrow="1" pageId="1" pageNumber="60">
<th id="76E95D8AFF94004DFED0F944FECFF8B8" box="[264,359,1784,1814]" gridcol="0" gridrow="1" pageId="1" pageNumber="60">P1 816</th>
<td id="76E95D8AFF94004DFEA2F944FE1FF8B8" box="[378,439,1784,1814]" gridcol="1" gridrow="1" pageId="1" pageNumber="60">1216</td>
<td id="76E95D8AFF94004DFE13F944FE54F8B8" box="[459,508,1784,1814]" gridcol="2" gridrow="1" pageId="1" pageNumber="60">944</td>
<td id="76E95D8AFF94004DFDD5F944FD93F8B8" box="[525,571,1784,1814]" gridcol="3" gridrow="1" pageId="1" pageNumber="60">448</td>
<td id="76E95D8AFF94004DFD97F944FDD5F8B8" box="[591,637,1784,1814]" gridcol="4" gridrow="1" pageId="1" pageNumber="60">272</td>
<td id="76E95D8AFF94004DFD56F944FD18F8B8" box="[654,688,1784,1814]" gridcol="5" gridrow="1" pageId="1" pageNumber="60">88</td>
<td id="76E95D8AFF94004DFD1CF944FD5AF8B8" box="[708,754,1784,1814]" gridcol="6" gridrow="1" pageId="1" pageNumber="60">112</td>
<td id="76E95D8AFF94004DFCDDF944FC93F8B8" box="[773,827,1784,1814]" gridcol="7" gridrow="1" pageId="1" pageNumber="60">0.78</td>
<td id="76E95D8AFF94004DFC96F944FC2CF8B8" box="[846,900,1784,1814]" gridcol="8" gridrow="1" pageId="1" pageNumber="60">2.15</td>
<td id="76E95D8AFF94004DFC4FF944FC66F8B8" box="[919,974,1784,1814]" gridcol="9" gridrow="1" pageId="1" pageNumber="60">3.23</td>
</tr>
<tr id="353834F6FF94004DFED0F89FFC66F8EF" box="[264,974,1827,1857]" gridrow="2" pageId="1" pageNumber="60">
<th id="76E95D8AFF94004DFED0F89FFECFF8EF" box="[264,359,1827,1857]" gridcol="0" gridrow="2" pageId="1" pageNumber="60">P2 992</th>
<td id="76E95D8AFF94004DFEA2F89FFE1FF8EF" box="[378,439,1827,1857]" gridcol="1" gridrow="2" pageId="1" pageNumber="60">1064</td>
<td id="76E95D8AFF94004DFE13F89FFE54F8EF" box="[459,508,1827,1857]" gridcol="2" gridrow="2" pageId="1" pageNumber="60">608</td>
<td id="76E95D8AFF94004DFDD5F89FFD93F8EF" box="[525,571,1827,1857]" gridcol="3" gridrow="2" pageId="1" pageNumber="60">320</td>
<td id="76E95D8AFF94004DFD97F89FFDD5F8EF" box="[591,637,1827,1857]" gridcol="4" gridrow="2" pageId="1" pageNumber="60">184</td>
<td id="76E95D8AFF94004DFD56F89FFD18F8EF" box="[654,688,1827,1857]" gridcol="5" gridrow="2" pageId="1" pageNumber="60">80</td>
<td id="76E95D8AFF94004DFD1CF89FFD5AF8EF" box="[708,754,1827,1857]" gridcol="6" gridrow="2" pageId="1" pageNumber="60">96</td>
<td id="76E95D8AFF94004DFCDDF89FFC93F8EF" box="[773,827,1827,1857]" gridcol="7" gridrow="2" pageId="1" pageNumber="60">0.57</td>
<td id="76E95D8AFF94004DFC96F89FFC2CF8EF" box="[846,900,1827,1857]" gridcol="8" gridrow="2" pageId="1" pageNumber="60">2.38</td>
<td id="76E95D8AFF94004DFC4FF89FFC66F8EF" box="[919,974,1827,1857]" gridcol="9" gridrow="2" pageId="1" pageNumber="60">3.92</td>
</tr>
<tr id="353834F6FF94004DFED0F8F2FC66F8C2" box="[264,974,1870,1900]" gridrow="3" pageId="1" pageNumber="60">
<th id="76E95D8AFF94004DFED0F8F2FECFF8C2" box="[264,359,1870,1900]" gridcol="0" gridrow="3" pageId="1" pageNumber="60">P3 1088</th>
<td id="76E95D8AFF94004DFEA2F8F2FE1FF8C2" box="[378,439,1870,1900]" gridcol="1" gridrow="3" pageId="1" pageNumber="60">1248</td>
<td id="76E95D8AFF94004DFE13F8F2FE54F8C2" box="[459,508,1870,1900]" gridcol="2" gridrow="3" pageId="1" pageNumber="60">768</td>
<td id="76E95D8AFF94004DFDD5F8F2FD93F8C2" box="[525,571,1870,1900]" gridcol="3" gridrow="3" pageId="1" pageNumber="60">432</td>
<td id="76E95D8AFF94004DFD97F8F2FDD5F8C2" box="[591,637,1870,1900]" gridcol="4" gridrow="3" pageId="1" pageNumber="60">256</td>
<td id="76E95D8AFF94004DFD56F8F2FD18F8C2" box="[654,688,1870,1900]" gridcol="5" gridrow="3" pageId="1" pageNumber="60">96</td>
<td id="76E95D8AFF94004DFD1CF8F2FD5AF8C2" box="[708,754,1870,1900]" gridcol="6" gridrow="3" pageId="1" pageNumber="60">104</td>
<td id="76E95D8AFF94004DFCDDF8F2FC93F8C2" box="[773,827,1870,1900]" gridcol="7" gridrow="3" pageId="1" pageNumber="60">0.62</td>
<td id="76E95D8AFF94004DFC96F8F2FC2CF8C2" box="[846,900,1870,1900]" gridcol="8" gridrow="3" pageId="1" pageNumber="60">3.04</td>
<td id="76E95D8AFF94004DFC4FF8F2FC66F8C2" box="[919,974,1870,1900]" gridcol="9" gridrow="3" pageId="1" pageNumber="60">3.50</td>
</tr>
</table>
</paragraph>
<caption id="DF77663CFF97FFB0FED0FA4FFDD5F9E1" httpUri="https://zenodo.org/record/170450/files/figure.png" pageId="2" pageNumber="61" targetBox="[302,1280,558,1479]" targetPageId="2">
<paragraph id="8BB736B4FF97FFB0FED0FA4FFDD5F9E1" blockId="2.[264,1323,1523,1615]" pageId="2" pageNumber="61">
<emphasis id="B97CEAA6FF97FFB0FED0FA4FFE1CF9A5" bold="true" box="[264,436,1523,1547]" pageId="2" pageNumber="61">FIGURES 1–4.</emphasis>
Male of
<taxonomicName id="4C084D37FF97FFB0FDCDFA4FFCA0F9A4" box="[533,776,1523,1546]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B97CEAA6FF97FFB0FDCDFA4FFCA0F9A4" box="[533,776,1523,1546]" italics="true" pageId="2" pageNumber="61">Arctodiamesa marinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57DDFF97FFB0FCD7FA4FFCE0F9A4" box="[783,840,1523,1546]" pageId="2" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="B97CEAA6FF97FFB0FC96FA4FFCF3F9A5" bold="true" box="[846,859,1523,1547]" pageId="2" pageNumber="61">1</emphasis>
, hypopygium in dorsal view;
<emphasis id="B97CEAA6FF97FFB0FB44FA4FFB01F9A5" bold="true" box="[1180,1193,1523,1547]" pageId="2" pageNumber="61">2</emphasis>
, gonostylus deflected from gonocoxite;
<emphasis id="B97CEAA6FF97FFB0FD9AF9A9FDE7F983" bold="true" box="[578,591,1557,1581]" pageId="2" pageNumber="61">3</emphasis>
, gonostylus pressed to gonocoxite;
<emphasis id="B97CEAA6FF97FFB0FC35F9A9FC52F983" bold="true" box="[1005,1018,1557,1581]" pageId="2" pageNumber="61">4</emphasis>
, sternapodeme and medial aedeagal lobe. Scale bars = 50 µm.
</paragraph>
</caption>
<paragraph id="8BB736B4FF97FFB0FEE0F919FE1EF946" blockId="2.[264,1323,1701,1891]" pageId="2" pageNumber="61">
<emphasis id="B97CEAA6FF97FFB0FEE0F919FED1F911" bold="true" box="[312,377,1701,1727]" pageId="2" pageNumber="61">Pupa</emphasis>
(
<emphasis id="B97CEAA6FF97FFB0FE4AF91AFE09F911" box="[402,417,1702,1727]" italics="true" pageId="2" pageNumber="61">n</emphasis>
= 4). Total length
<quantity id="4CF09B51FF97FFB0FD7BF91AFC93F96E" box="[675,827,1702,1728]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" metricValueMax="5.5" metricValueMin="4.9" pageId="2" pageNumber="61" unit="mm" value="5.2" valueMax="5.5" valueMin="4.9">4.9–5.5 mm</quantity>
. Coloration brownish. Exuviae grey or brownish grey.
</paragraph>
<paragraph id="8BB736B4FF97FFB0FEE0F94AFB09F8CD" blockId="2.[264,1323,1701,1891]" pageId="2" pageNumber="61">
Cephalothorax. Frontal tubercles low, with
<quantity id="4CF09B51FF97FFB0FC9EF94AFC77F8BE" box="[838,991,1782,1808]" metricMagnitude="2" metricUnit="m" metricValue="2.54" metricValueMax="2.56" metricValueMin="2.52" pageId="2" pageNumber="61" unit="m" value="254.0" valueMax="256.0" valueMin="252.0">252–256 m</quantity>
long setae. Thorax smooth and granulated. Pc­setae lengths (m): Pc
<date id="FFB61074FF97FFB0FD2BF892FCC9F896" box="[755,865,1822,1852]" pageId="2" pageNumber="61" value="1911-01-02">1 — 11–2</date>
–116, Pc2 — 305–310, Pc3 — 124–128 (
<figureCitation id="13332A31FF97FFB0FEC9F8F5FEF0F8CD" box="[273,344,1865,1891]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="2" pageNumber="61">Fig. 5</figureCitation>
). Dorsocentrals divided into 5–12 branches
<quantity id="4CF09B51FF97FFB0FC86F8F5FC7FF8CD" box="[862,983,1865,1891]" metricMagnitude="1" metricUnit="m" metricValue="2.6" metricValueMax="3.6" metricValueMin="1.6" pageId="2" pageNumber="61" unit="m" value="26.0" valueMax="36.0" valueMin="16.0">16–36 m</quantity>
long (
<figureCitation id="13332A31FF97FFB0FBFAF8F5FB3CF8CD" box="[1058,1172,1865,1891]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="2" pageNumber="61">Figs. 6–8</figureCitation>
).
</paragraph>
<caption id="DF77663CFF96FFB1FED0F96DFB51F8F5" httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="3" pageNumber="62" targetBox="[311,1284,307,1717]" targetPageId="3">
<paragraph id="8BB736B4FF96FFB1FED0F96DFB51F8F5" blockId="3.[264,1323,1745,1883]" pageId="3" pageNumber="62">
<emphasis id="B97CEAA6FF96FFB1FED0F96DFE6CF947" bold="true" box="[264,452,1745,1769]" pageId="3" pageNumber="62">FIGURES 5–17.</emphasis>
Pupa (5–11) and larva (12–17) of
<taxonomicName id="4C084D37FF96FFB1FC9FF96EFB94F947" box="[839,1084,1746,1769]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B97CEAA6FF96FFB1FC9FF96EFB94F947" box="[839,1084,1746,1769]" italics="true" pageId="3" pageNumber="62">Arctodiamesa marinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57DDFF96FFB1FB9EF96EFB2AF947" box="[1094,1154,1746,1769]" pageId="3" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="B97CEAA6FF96FFB1FB53F96DFB30F947" bold="true" box="[1163,1176,1745,1769]" pageId="3" pageNumber="62">5</emphasis>
, precorneals;
<emphasis id="B97CEAA6FF96FFB1FED0F94BFE98F8A1" bold="true" box="[264,304,1783,1807]" pageId="3" pageNumber="62">6–8</emphasis>
, dorsocentrals of thorax;
<emphasis id="B97CEAA6FF96FFB1FD90F94BFDFDF8A1" bold="true" box="[584,597,1783,1807]" pageId="3" pageNumber="62">9</emphasis>
, tergite IV;
<emphasis id="B97CEAA6FF96FFB1FD02F94BFD5DF8A1" bold="true" box="[730,757,1783,1807]" pageId="3" pageNumber="62">10</emphasis>
, netlike surface of tergites;
<emphasis id="B97CEAA6FF96FFB1FBFDF94BFB95F8A1" bold="true" box="[1061,1085,1783,1807]" pageId="3" pageNumber="62">11</emphasis>
, tergite VIII and anal segment;
<emphasis id="B97CEAA6FF96FFB1FEADF8A1FE38F89B" bold="true" box="[373,400,1821,1845]" pageId="3" pageNumber="62">12</emphasis>
, part of labrum;
<emphasis id="B97CEAA6FF96FFB1FD84F8A1FDDFF89B" bold="true" box="[604,631,1821,1845]" pageId="3" pageNumber="62">13</emphasis>
, premandible;
<emphasis id="B97CEAA6FF96FFB1FCFCF8A1FC97F89B" bold="true" box="[804,831,1821,1845]" pageId="3" pageNumber="62">14</emphasis>
, mandible;
<emphasis id="B97CEAA6FF96FFB1FC12F8A1FC4CF89B" bold="true" box="[970,996,1821,1845]" pageId="3" pageNumber="62">15</emphasis>
, procercus;
<emphasis id="B97CEAA6FF96FFB1FBACF8A1FB27F89B" bold="true" box="[1140,1167,1821,1845]" pageId="3" pageNumber="62">16</emphasis>
, antenna;
<emphasis id="B97CEAA6FF96FFB1FAD1F8A1FA8BF89B" bold="true" box="[1289,1315,1821,1845]" pageId="3" pageNumber="62">17</emphasis>
, mentum. Scale bars = 50 m for Figs. 5–8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12–17.
</paragraph>
</caption>
<paragraph id="8BB736B4FF91FFB6FEE0FEA7FB18FE98" blockId="4.[264,1325,283,1393]" box="[312,1200,283,310]" pageId="4" pageNumber="63">
<emphasis id="B97CEAA6FF91FFB6FEE0FEA7FD83FE9B" bold="true" box="[312,555,283,309]" pageId="4" pageNumber="63">Fourth­instar larva</emphasis>
(
<emphasis id="B97CEAA6FF91FFB6FDE4FEA0FDE3FE9B" box="[572,587,284,309]" italics="true" pageId="4" pageNumber="63">n</emphasis>
= 5). Coloration greenish. Total length
<quantity id="4CF09B51FF91FFB6FBC3FEA0FB04FE98" box="[1051,1196,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.4" metricValueMin="6.8" pageId="4" pageNumber="63" unit="mm" value="7.1" valueMax="7.4" valueMin="6.8">6.8–7.4 mm</quantity>
.
</paragraph>
<paragraph id="8BB736B4FF91FFB6FEE0FEF8FBCCFDD7" blockId="4.[264,1325,283,1393]" pageId="4" pageNumber="63">
Head. Head capsule
<quantity id="4CF09B51FF91FFB6FDE9FEF8FD76FEF0" box="[561,734,324,350]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="4" pageNumber="63" unit="mm" value="0.43" valueMax="0.46" valueMin="0.4">0.40–0.46 mm</quantity>
long,
<quantity id="4CF09B51FF91FFB6FCF5FEF8FC73FEF0" box="[813,987,324,350]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="4" pageNumber="63" unit="mm" value="0.32" valueMax="0.34" valueMin="0.3">0.30–0.34 mm</quantity>
wide, light yellow. Labrum with simple S I–S IV setae and large lamellae (
<figureCitation id="13332A31FF91FFB6FCC2FED0FCDBFE28" box="[794,883,364,390]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="4" pageNumber="63">Fig. 12</figureCitation>
). Premandible brown to light brown, distally with 6 or 7 teeth (
<figureCitation id="13332A31FF91FFB6FD89FE2BFD04FE1F" box="[593,684,407,433]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="4" pageNumber="63">Fig. 13</figureCitation>
). Antenna yellow, large ring organ situated in basal quarter of first segment; antennal blade ending near base of fourth segment (
<figureCitation id="13332A31FF91FFB6FB4BFE03FB42FE77" box="[1171,1258,447,473]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="4" pageNumber="63">Fig. 16</figureCitation>
); AR 1.56–1.61. Lengths (m) of antennal segments 1–5: 65.1: 15.0: 13.4: 4.2: 3.3. Mandible dark brown or black in distal part; apical tooth longest; seta interna with 15–18 branches, seta subdentalis short and spinelike (
<figureCitation id="13332A31FF91FFB6FD6DFD8BFCA4FDFF" box="[693,780,567,593]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="4" pageNumber="63">Fig. 14</figureCitation>
). Mentum with 1 median and 7 lateral pairs of teeth; median tooth 4.7–5.0 times broader than first lateral tooth (
<figureCitation id="13332A31FF91FFB6FBDEFDE3FBFFFDD7" box="[1030,1111,607,633]" captionStart="FIGURES 5 – 17" captionStartId="3.[264,382,1745,1769]" captionTargetBox="[311,1284,307,1717]" captionTargetId="figure@3.[297,1290,300,1717]" captionTargetPageId="3" captionText="FIGURES 5 – 17. Pupa (5 – 11) and larva (12 – 17) of Arctodiamesa marinae sp. n. 5, precorneals; 6 – 8, dorsocentrals of thorax; 9, tergite IV; 10, netlike surface of tergites; 11, tergite VIII and anal segment; 12, part of labrum; 13, premandible; 14, mandible; 15, procercus; 16, antenna; 17, mentum. Scale bars = 50 m for Figs. 5 – 8, 10; 200 m for Figs. 9, 11; 20 m for Figs. 12 – 17." httpUri="https://zenodo.org/record/170451/files/figure.png" pageId="4" pageNumber="63">Fig.17</figureCitation>
).
</paragraph>
<paragraph id="8BB736B4FF91FFB6FEE0FD3BFC8EFD67" blockId="4.[264,1325,283,1393]" pageId="4" pageNumber="63">
Abdomen. Procercus dark brown, slightly longer than wide, with 7 anal setae. Two dark brown subapical setae
<quantity id="4CF09B51FF91FFB6FD96FD13FD4BFD67" box="[590,739,687,713]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.32" metricValueMin="2.08" pageId="4" pageNumber="63" unit="m" value="220.0" valueMax="232.0" valueMin="208.0">208–232 m</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="C312653FFF91FFB6FEE0FD6BFAA9FB9F" pageId="4" pageNumber="63" type="discussion">
<paragraph id="8BB736B4FF91FFB6FEE0FD6BFAA9FB9F" blockId="4.[264,1325,283,1393]" pageId="4" pageNumber="63">
<emphasis id="B97CEAA6FF91FFB6FEE0FD6BFE19FD5F" bold="true" box="[312,433,727,753]" pageId="4" pageNumber="63">Remarks.</emphasis>
The male of
<taxonomicName id="4C084D37FF91FFB6FD87FD6BFD4FFD5E" box="[607,743,727,752]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B97CEAA6FF91FFB6FD87FD6BFD4FFD5E" box="[607,743,727,752]" italics="true" pageId="4" pageNumber="63">A. marinae</emphasis>
</taxonomicName>
can be separated from other known species of
<taxonomicName id="4C084D37FF91FFB6FED0FD43FE01FCB6" box="[264,425,767,792]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAA6FF91FFB6FED0FD43FE01FCB6" box="[264,425,767,792]" italics="true" pageId="4" pageNumber="63">Arctodiamesa</emphasis>
</taxonomicName>
by the shape of the gonostyli. The basally wide and rounded lobe, combined with the short, subapical projection, is characteristic. Larvae of
<taxonomicName id="4C084D37FF91FFB6FBCAFC9BFB30FCEE" box="[1042,1176,807,832]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B97CEAA6FF91FFB6FBCAFC9BFB30FCEE" box="[1042,1176,807,832]" italics="true" pageId="4" pageNumber="63">A. marinae</emphasis>
</taxonomicName>
have six or seven teeth on the premandible, whereas the other species have only three.
<taxonomicName id="4C084D37FF91FFB6FB51FCF3FEC2FC3E" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B97CEAA6FF91FFB6FB51FCF3FEC2FC3E" italics="true" pageId="4" pageNumber="63">Arctodiamesa marinae</emphasis>
</taxonomicName>
also differs from the other species by the wide median tooth of the mentum. The pupae are similar to those of the previously described but unnamed
<taxonomicName id="4C084D37FF91FFB6FB83FC23FB55FC16" box="[1115,1277,927,952]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAA6FF91FFB6FB83FC23FB55FC16" box="[1115,1277,927,952]" italics="true" pageId="4" pageNumber="63">Arctodiamesa</emphasis>
</taxonomicName>
sp. (
<bibRefCitation id="EF994B45FF91FFB6FEC8FC7BFE53FC4F" author="Makarchenko" box="[272,507,967,993]" pageId="4" pageNumber="64" refString="Makarchenko, E. A. (1995) New species of Lappodiamesa Serra-Tosio and Arctodiamesa Makartshenko (Diptera, Chironomidae) from the East Palaearctic. Aquatic Insects, 17, 83 - 93." type="journal article" year="1995">Makarchenko 1995</bibRefCitation>
), but can be distinguished by details of the tergite shagreen and the number of branches on the dorsocental setae of the thorax.
<taxonomicName id="4C084D37FF91FFB6FC31FC53FB50FBA6" box="[1001,1272,1007,1032]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B97CEAA6FF91FFB6FC31FC53FB50FBA6" box="[1001,1272,1007,1032]" italics="true" pageId="4" pageNumber="63">Arctodiamesa marinae</emphasis>
</taxonomicName>
has 5–12 branches on each dorsocentral, whereas
<taxonomicName id="4C084D37FF91FFB6FCFAFBABFC6CFB9E" box="[802,964,1047,1072]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAA6FF91FFB6FCFAFBABFC6CFB9E" box="[802,964,1047,1072]" italics="true" pageId="4" pageNumber="63">Arctodiamesa</emphasis>
</taxonomicName>
sp. has only 2–4 branches.
</paragraph>
</subSubSection>
<subSubSection id="C312653FFF91FFB6FEE0FB83FD78FB07" pageId="4" pageNumber="63" type="etymology">
<paragraph id="8BB736B4FF91FFB6FEE0FB83FD78FB07" blockId="4.[264,1325,283,1393]" pageId="4" pageNumber="63">
<emphasis id="B97CEAA6FF91FFB6FEE0FB83FE6CFBF7" bold="true" box="[312,452,1087,1113]" pageId="4" pageNumber="63">Etymology.</emphasis>
The new species is named in honour of my wife
<taxonomicName id="4C084D37FF91FFB6FBFCFB83FA8FFBF7" authority="Makarchenko" authorityName="Makarchenko" box="[1060,1319,1087,1113]" class="Magnoliopsida" family="Fabaceae" genus="Marina" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="63" phylum="Tracheophyta" rank="genus">Marina Makarchenko</taxonomicName>
, who studies the taxonomy of the Orthocladiinae of the Russian Far East. She is a driving force in my chironomid investigations.
</paragraph>
</subSubSection>
<subSubSection id="C312653FFF91FFB6FEE0FB0BFD08FADF" pageId="4" pageNumber="63" type="distribution">
<paragraph id="8BB736B4FF91FFB6FEE0FB0BFD08FADF" blockId="4.[264,1325,283,1393]" pageId="4" pageNumber="63">
<emphasis id="B97CEAA6FF91FFB6FEE0FB0BFD29FB7F" bold="true" box="[312,641,1207,1233]" pageId="4" pageNumber="63">Distribution and biology.</emphasis>
<taxonomicName id="4C084D37FF91FFB6FD49FB0BFC0DFB7E" box="[657,933,1207,1232]" class="Insecta" family="Chironomidae" genus="Arctodiamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B97CEAA6FF91FFB6FD49FB0BFC0DFB7E" box="[657,933,1207,1232]" italics="true" pageId="4" pageNumber="63">Arctodiamesa marinae</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54B38816FF91FFB6FB20FB0BFA83FB7F" box="[1272,1323,1207,1233]" pageId="4" pageNumber="63">type</typeStatus>
locality in Kedrovaya Pad Nature reserve in the southern part of Primorye Territory of the Russian Far East. Pupae and larvae were collected in the middle reaches of the Kedrovaya River on small and medium­sized stones covered with green algae. The water temperature was 14°C at the time of collection.
</paragraph>
</subSubSection>
</treatment>
</document>
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