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<document ID-DOI="http://doi.org/10.5281/zenodo.3792320" ID-GBIF-Dataset="380557b0-b78e-4dd3-a582-75f6fa6d04b4" ID-GBIF-Taxon="163756041" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792320" ID-ZooBank="3B75A32E-5332-4F74-A3BA-974ADDF626C4" LSID="urn:lsid:zoobank.org:act:98429FDA-3B9D-44BA-8259-6D83D11CFFBB" checkinTime="1588008374112" checkinUser="plazi" docAuthor="Schmidt, Christian" docDate="2009" docId="03809242FFFF4C2830A5FCB0EAEFF915" docLanguage="en" docName="ZK_article_1989.pdf" docOrigin="ZooKeys 9 (9)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Leichosila wagneri Schmidt 2009, sp. n." docType="treatment" docUuid="98429FDA-3B9D-44BA-8259-6D83D11CFFBB" docUuidSource="ZooBank" docVersion="8" lastPageNumber="95" masterDocId="FFB9EA3AFFF94C2E3029FFF3EB5FFF92" masterDocTitle="A new genus and two new species of arctiine tiger moth (Noctuidae, Arctiinae, Arctiini) from Costa Rica" masterLastPageNumber="96" masterPageNumber="89" pageId="6" pageNumber="95" updateTime="1643565622888" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new genus and two new species of arctiine tiger moth (Noctuidae, Arctiinae, Arctiini) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmidt, Christian</mods:namePart>
<mods:affiliation>Canadian Food Inspection Agency, Ottawa ,, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-05-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>89</mods:start>
<mods:end>96</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.9.151</mods:identifier>
<mods:identifier type="GBIF-Dataset">380557b0-b78e-4dd3-a582-75f6fa6d04b4</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576455</mods:identifier>
<mods:identifier type="ZooBank">3B75A32E-5332-4F74-A3BA-974ADDF626C4</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792320" ID-GBIF-Taxon="163756041" ID-Zenodo-Dep="3792320" LSID="urn:lsid:zoobank.org:act:98429FDA-3B9D-44BA-8259-6D83D11CFFBB" httpUri="http://treatment.plazi.org/id/03809242FFFF4C2830A5FCB0EAEFF915" lastPageNumber="95" pageId="6" pageNumber="95">
<subSubSection box="[140,549,835,862]" pageId="6" pageNumber="95" type="nomenclature">
<paragraph blockId="6.[140,989,835,932]" box="[140,549,835,862]" pageId="6" pageNumber="95">
<heading box="[140,549,835,862]" fontSize="11" level="2" pageId="6" pageNumber="95" reason="3">
<emphasis bold="true" box="[140,549,835,862]" pageId="6" pageNumber="95">
<taxonomicName authority="Schmidt" authorityName="Schmidt" authorityYear="2009" box="[140,472,835,862]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="wagneri" status="sp. n.">
<emphasis bold="true" box="[140,362,835,862]" italics="true" pageId="6" pageNumber="95">
<emphasis box="[140,259,835,862]" inLineHeading="true" pageId="6" pageNumber="95" reason="1">Leichosila</emphasis>
wagneri
</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel box="[484,549,835,861]" pageId="6" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="description">
<paragraph blockId="6.[140,989,835,932]" box="[140,989,870,897]" pageId="6" pageNumber="95">
<uri box="[140,989,870,897]" pageId="6" pageNumber="95">
urn:lsid:zoobank.org:act:
<uuid box="[420,989,870,897]" pageId="6" pageNumber="95">98429FDA-3B9D-44BA-8259-6D83D11CFFBB</uuid>
</uri>
</paragraph>
<paragraph blockId="6.[140,989,835,932]" box="[140,291,906,932]" pageId="6" pageNumber="95">
(
<figureCitation box="[148,226,906,932]" captionStart="Figures 1-2" captionStartId="2.[140,222,1622,1644]" captionTargetBox="[140,1108,1276,1602]" captionTargetId="figure@2.[140,1108,1276,1602]" captionTargetPageId="2" captionText="Figures 1-2. Adult habitus of Leichosila gen. n. 1. Leichosila talamanca sp. n., male holotype. 2. Leichosila wagneri sp. n., male holotype." figureDoi="http://doi.org/10.5281/zenodo.3810733" httpUri="https://zenodo.org/record/3810733/files/figure.png" pageId="6" pageNumber="95">Figs. 2</figureCitation>
,
<figureCitation box="[238,255,906,932]" captionStart="Figures 3-5" captionStartId="4.[140,222,1526,1548]" captionTargetBox="[162,1087,174,1500]" captionTargetId="figure@4.[153,1094,165,1507]" captionTargetPageId="4" captionText="Figures 3-5. Male genitalia of Leichosila. 3a. L. talamanca (paratype), genital capsule (ventral view); 3b. phallus (left lateral view), with inflated vesica (ventrobasal diverticulum highlighted); 4a. L. wagneri (holotype), genital capsule (ventral view); 4b. phallus (left lateral view) with inflated vesica (ventrobasal diverticulum highlighted); 5. L. talamanca, 8th abdominal sternite, showing medioventral sclerite and lateral lobes." figureDoi="http://doi.org/10.5281/zenodo.3810737" httpUri="https://zenodo.org/record/3810737/files/figure.png" pageId="6" pageNumber="95">4</figureCitation>
,
<figureCitation box="[267,282,906,932]" captionStart="Figures 6-9" captionStartId="5.[140,222,1590,1612]" captionTargetBox="[190,1070,820,1565]" captionTargetId="figure@5.[140,1108,799,1573]" captionTargetPageId="5" captionText="Figures 6-9. Structural features of Leichosila. 6. L. talamanca, uncus (dorsal view); 7. L. wagneri, uncus (dorsal view); 8. L. talamanca, juxta; 9. L. talamanca, antenna (segments 14 – 18), ventral view (setae and cilia omitted except on segment 14)." figureDoi="http://doi.org/10.5281/zenodo.3810741" httpUri="https://zenodo.org/record/3810741/files/figure.png" pageId="6" pageNumber="95">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="materials_examined">
<paragraph blockId="6.[140,1108,976,1672]" pageId="6" pageNumber="95">
<emphasis bold="true" box="[140,310,976,1002]" pageId="6" pageNumber="95">
<emphasis box="[140,198,976,1002]" inLineHeading="true" pageId="6" pageNumber="95" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608997641" collectingDate="2002-02-15" collectionCode="CNC, INBIO" collectorName="David L. Wagner" elevation="2000" latitude="10.183333" location="Vara Blanca" longLatPrecision="1298" longitude="-84.11667" pageId="6" pageNumber="95" specimenCount="2" stateProvince="Heredia" typeStatus="holotype">
<emphasis bold="true" box="[316,429,976,1002]" pageId="6" pageNumber="95">
<typeStatus box="[316,429,976,1002]" pageId="6" pageNumber="95">Holotype</typeStatus>
</emphasis>
male – “C[osta]. RICA:
<collectingRegion box="[706,800,976,1002]" country="Costa Rica" name="Heredia" pageId="6" pageNumber="95">Heredia</collectingRegion>
: /
<quantity box="[825,883,976,1002]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="95" unit="km" value="6.0">6 km</quantity>
ENE
<location LSID="urn:lsid:plazi:treatment:03809242FFFF4C2830A5FCB0EAEFF915:8EF6758FFFFF4C283391FC22EF1FFC78" box="[952,1088,976,1003]" latitude="10.183333" longLatPrecision="1298" longitude="-84.11667" name="Vara Blanca" pageId="6" pageNumber="95" stateProvince="Heredia">Vara Blanca</location>
, /
<quantity box="[140,292,1011,1038]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="6" pageNumber="95" unit="m" value="2000.0" valueMax="2050.0" valueMin="1950.0">
<elevation box="[140,292,1011,1038]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="6" pageNumber="95" unit="m" value="2000.0" valueMax="2050.0" valueMin="1950.0">1950-2050m</elevation>
</quantity>
,
<geoCoordinate box="[304,401,1012,1038]" degrees="10" direction="north" minutes="11" orientation="latitude" pageId="6" pageNumber="95" precision="925" value="10.183333">10°11’N</geoCoordinate>
/
<geoCoordinate box="[425,527,1012,1038]" degrees="84" direction="west" minutes="07" orientation="longitude" pageId="6" pageNumber="95" precision="925" value="-84.11667">84 07’W</geoCoordinate>
;
<date box="[539,656,1011,1038]" pageId="6" pageNumber="95" value="2002-02-15">
<collectingDate box="[539,656,1011,1038]" pageId="6" pageNumber="95" value="2002-02-15">15 ii 2002</collectingDate>
</date>
/
<collectorName box="[680,874,1011,1038]" pageId="6" pageNumber="95">David L. Wagner</collectorName>
coll”; “INBio-OET- ALAS / transect / 20/L/01/011”; “Project / ALAS / INB0003216237”; “
<typeStatus box="[950,1107,1047,1073]" pageId="6" pageNumber="95">HOLOTYPE</typeStatus>
/
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[158,264,1082,1108]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis box="[158,264,1082,1108]" italics="true" pageId="6" pageNumber="95">Leichosila</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[292,379,1083,1108]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="wagneri">
<emphasis box="[292,379,1083,1108]" italics="true" pageId="6" pageNumber="95">wagneri</emphasis>
</taxonomicName>
/ Schmidt” [red label]; “Genitalia /
<collectionCode box="[796,860,1082,1108]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="6" pageNumber="95">CNC</collectionCode>
slide # / 14541”; deposited in
<collectionCode box="[259,347,1117,1143]" pageId="6" pageNumber="95">INBIO</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="diagnosis">
<paragraph blockId="6.[140,1108,976,1672]" pageId="6" pageNumber="95">
<emphasis bold="true" box="[185,312,1152,1178]" pageId="6" pageNumber="95">
<emphasis box="[185,307,1152,1178]" inLineHeading="true" pageId="6" pageNumber="95" reason="1">Diagnosis</emphasis>
.
</emphasis>
Distinguished from
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[554,701,1152,1179]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis box="[554,570,1153,1178]" italics="true" pageId="6" pageNumber="95">L</emphasis>
.
<emphasis box="[587,701,1152,1178]" italics="true" pageId="6" pageNumber="95">talamanca</emphasis>
</taxonomicName>
by the smaller size and darker colouration of
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[278,398,1188,1214]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="wagneri">
<emphasis box="[278,294,1188,1213]" italics="true" pageId="6" pageNumber="95">L</emphasis>
.
<emphasis box="[311,398,1188,1213]" italics="true" pageId="6" pageNumber="95">wagneri</emphasis>
</taxonomicName>
, and by the genitalic characters given under the diagnosis for
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[140,285,1223,1249]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis box="[140,156,1223,1248]" italics="true" pageId="6" pageNumber="95">L</emphasis>
.
<emphasis box="[171,285,1223,1249]" italics="true" pageId="6" pageNumber="95">talamanca</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="description">
<paragraph blockId="6.[140,1108,976,1672]" pageId="6" pageNumber="95">
<emphasis bold="true" box="[185,332,1257,1283]" pageId="6" pageNumber="95">
<emphasis box="[185,326,1257,1283]" inLineHeading="true" pageId="6" pageNumber="95" reason="1">Description</emphasis>
.
</emphasis>
A detailed description is given above in the genus description. Characters specific to
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[298,411,1294,1320]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="wagneri">
<emphasis box="[298,314,1294,1319]" italics="true" pageId="6" pageNumber="95">L</emphasis>
.
<emphasis box="[327,411,1294,1319]" italics="true" pageId="6" pageNumber="95">wagneri</emphasis>
</taxonomicName>
are as follows: Male (female unknown).
<emphasis bold="true" box="[847,956,1293,1319]" pageId="6" pageNumber="95">Forewing</emphasis>
<emphasis box="[962,976,1294,1319]" italics="true" pageId="6" pageNumber="95">–</emphasis>
length 13.4 (
<emphasis box="[149,164,1329,1354]" italics="true" pageId="6" pageNumber="95">n</emphasis>
= 1); ground colour dark grey.
<emphasis bold="true" box="[510,630,1328,1354]" pageId="6" pageNumber="95">Hindwing</emphasis>
<emphasis box="[637,651,1329,1354]" italics="true" pageId="6" pageNumber="95">–</emphasis>
ground colour dark grey, with indistinct, irregular dark grey subterminal spots and medial spot.
<emphasis bold="true" box="[745,911,1363,1389]" pageId="6" pageNumber="95">Male genitalia</emphasis>
– Uncus parallelsided basally, then tapering apically,
<emphasis box="[553,561,1399,1424]" italics="true" pageId="6" pageNumber="95">i</emphasis>
.
<emphasis box="[567,577,1399,1424]" italics="true" pageId="6" pageNumber="95">e</emphasis>
. bottle-shaped in dorsal profile (
<figureCitation box="[951,1020,1399,1425]" captionStart="Figures 6-9" captionStartId="5.[140,222,1590,1612]" captionTargetBox="[190,1070,820,1565]" captionTargetId="figure@5.[140,1108,799,1573]" captionTargetPageId="5" captionText="Figures 6-9. Structural features of Leichosila. 6. L. talamanca, uncus (dorsal view); 7. L. wagneri, uncus (dorsal view); 8. L. talamanca, juxta; 9. L. talamanca, antenna (segments 14 – 18), ventral view (setae and cilia omitted except on segment 14)." figureDoi="http://doi.org/10.5281/zenodo.3810741" httpUri="https://zenodo.org/record/3810741/files/figure.png" pageId="6" pageNumber="95">Fig. 7</figureCitation>
); vesica with left diverticulum of two ventro-basal diverticula conical and pointed (
<figureCitation box="[960,1042,1434,1461]" captionStart="Figures 3-5" captionStartId="4.[140,222,1526,1548]" captionTargetBox="[162,1087,174,1500]" captionTargetId="figure@4.[153,1094,165,1507]" captionTargetPageId="4" captionText="Figures 3-5. Male genitalia of Leichosila. 3a. L. talamanca (paratype), genital capsule (ventral view); 3b. phallus (left lateral view), with inflated vesica (ventrobasal diverticulum highlighted); 4a. L. wagneri (holotype), genital capsule (ventral view); 4b. phallus (left lateral view) with inflated vesica (ventrobasal diverticulum highlighted); 5. L. talamanca, 8th abdominal sternite, showing medioventral sclerite and lateral lobes." figureDoi="http://doi.org/10.5281/zenodo.3810737" httpUri="https://zenodo.org/record/3810737/files/figure.png" pageId="6" pageNumber="95">Fig. 4b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="etymology">
<paragraph blockId="6.[140,1108,976,1672]" pageId="6" pageNumber="95">
<emphasis bold="true" box="[185,320,1469,1495]" pageId="6" pageNumber="95">
<emphasis box="[185,315,1469,1495]" inLineHeading="true" pageId="6" pageNumber="95" reason="1">Etymology</emphasis>
.
</emphasis>
This species is named after David L. Wagner, who collected the
<typeStatus box="[1059,1107,1470,1496]" pageId="6" pageNumber="95">type</typeStatus>
specimen, and whose studies continue to vastly improve our knowledge of the larval biology of North American
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[456,597,1539,1565]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="95" type="distribution">
<paragraph blockId="6.[140,1108,976,1672]" pageId="6" pageNumber="95">
<emphasis bold="true" box="[185,492,1574,1601]" pageId="6" pageNumber="95">
<emphasis box="[185,336,1574,1600]" inLineHeading="true" pageId="6" pageNumber="95" reason="1">Distribution</emphasis>
and biology.
</emphasis>
<taxonomicName authorityName="Schmidt" authorityYear="2009" box="[500,702,1575,1601]" class="Insecta" family="Erebidae" genus="Leichosila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="wagneri">
<emphasis box="[500,702,1575,1601]" italics="true" pageId="6" pageNumber="95">Leichosila wagneri</emphasis>
</taxonomicName>
is known only from the
<typeStatus box="[986,1034,1575,1601]" pageId="6" pageNumber="95">type</typeStatus>
specimen, collected in high elevation forest (
<quantity box="[602,765,1610,1637]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.05" metricValueMin="1.95" pageId="6" pageNumber="95" unit="m" value="2000.0" valueMax="2050.0" valueMin="1950.0">1950-2050 m</quantity>
) on Volcán Barva (Cordillera Central) in mid February.
</paragraph>
</subSubSection>
</treatment>
</document>
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