208 lines
30 KiB
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208 lines
30 KiB
XML
<document id="65DEC95EA1283494265484E142B7C05F" ID-CLB-Dataset="30140" ID-DOI="10.11646/zootaxa.4415.3.6" ID-GBIF-Dataset="f7dfd7e8-238c-432a-9eec-0929f785f747" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242185" ID-ZooBank="30CF1B69-020C-43E8-B8F0-C30B41E04FE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525600684827" checkinUser="plazi" docAuthor="Coelho, Beatriz W. T. & Engel, Michael S." docDate="2018" docId="C55BAA03FFAA7B51FF47F9AA35B3FEF9" docLanguage="en" docName="zootaxa.4415.3.6.imf" docOrigin="Zootaxa 4415 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ruizantheda nigra Coelho & Engel 2018, n. sp." docType="treatment" docVersion="9" lastPageNumber="517" masterDocId="3962D27BFFA87B55FFD0FFBB3400FFB9" masterDocTitle="A new group of species within the bee genus Ruizantheda, with a revised key to the males of the genus (Hymenoptera: Halictidae: Caenohalictini)" masterLastPageNumber="532" masterPageNumber="513" originalUpdateDomain="Boston" originalUpdateTime="1545217612389" originalUpdateUser="gglab" pageNumber="515" updateTime="1698541935543" updateUser="ExternalLinkService">
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<mods:mods id="6D9C389D7FE0370A9A66FB5F7D9CE41C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AB4564109BCC0457727DAB7AD44D5DF3">
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<mods:title id="DBD8542493FE7B13B0ACED00608F5E66">A new group of species within the bee genus Ruizantheda, with a revised key to the males of the genus (Hymenoptera: Halictidae: Caenohalictini)</mods:title>
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<mods:name id="C55FB8F96AEBF269CDB70E44D469785F" type="personal">
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<mods:roleTerm id="7AF96D6D5D0266F552EEFD74F27444E3">Author</mods:roleTerm>
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<mods:namePart id="26F614F5FE74C2579C62B05022E5D0D4">Coelho, Beatriz W. T.</mods:namePart>
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<mods:name id="DDBC69F247883BDD21A23B694B36DD0A" type="personal">
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<mods:roleTerm id="32C8948ED61B28F9B6A4B4BE292E4913">Author</mods:roleTerm>
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<mods:namePart id="587C21152555463219216172C9813B97">Engel, Michael S.</mods:namePart>
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<mods:typeOfResource id="6FB7F87D4544F6552C8897F7D5C62C94">text</mods:typeOfResource>
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<mods:title id="7DBDE6C1C56700776EE252C7922FD3E5">Zootaxa</mods:title>
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<mods:part id="4F9F871385BFC292FA9B93773FFB74F3">
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<mods:date id="B8DEAE28F81981BFF9AFED14FF096B85">2018</mods:date>
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<mods:number id="B9400BCBE77E0FC32BFC3F7EB8614178">4415</mods:number>
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<mods:detail id="6C7C0755D22D3561A993B65650D165B1" type="issue">
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<mods:number id="392B43C666BA184D86BB3A6DB059C2E9">3</mods:number>
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<mods:classification id="7C7AC17F09CA1BB1DEBAE05AF6B18A73">journal article</mods:classification>
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<mods:identifier id="C9B777FD8E30F171039C7B52ADFBF879" type="DOI">10.11646/zootaxa.4415.3.6</mods:identifier>
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<treatment id="C55BAA03FFAA7B51FF47F9AA35B3FEF9" ID-DOI="http://doi.org/10.5281/zenodo.6490342" ID-GBIF-Taxon="143811444" ID-Zenodo-Dep="6490342" LSID="urn:lsid:plazi:treatment:C55BAA03FFAA7B51FF47F9AA35B3FEF9" httpUri="http://treatment.plazi.org/id/C55BAA03FFAA7B51FF47F9AA35B3FEF9" lastPageId="4" lastPageNumber="517" pageId="2" pageNumber="515">
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<subSubSection id="05E8489EFFAA7B57FF47F9AA360FF9F4" pageId="2" pageNumber="515" type="nomenclature">
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<paragraph id="4D4D1B15FFAA7B57FF47F9AA35C2F992" blockId="2.[151,527,1553,1613]" box="[151,450,1553,1579]" pageId="2" pageNumber="515">
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<heading id="1605AC79FFAA7B57FF47F9AA35C2F992" bold="true" box="[151,450,1553,1579]" fontSize="11" level="1" pageId="2" pageNumber="515" reason="1">
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<emphasis id="7F86C707FFAA7B57FF47F9AA35C2F992" bold="true" box="[151,450,1553,1579]" pageId="2" pageNumber="515">
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<taxonomicName id="8AF26096FFAA7B57FF47F9AA3578F992" authority="Coelho & Engel, 2018" authorityName="Coelho & Engel" authorityYear="2018" box="[151,376,1553,1579]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="515" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
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<emphasis id="7F86C707FFAA7B57FF47F9AA3578F992" bold="true" box="[151,376,1553,1579]" italics="true" pageId="2" pageNumber="515">Ruizantheda nigra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="64B57A7CFFAA7B57FE50F9AA35C2F992" box="[384,450,1553,1579]" pageId="2" pageNumber="515" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4D4D1B15FFAA7B57FF47F98F360FF9F4" blockId="2.[151,527,1553,1613]" box="[151,527,1588,1613]" pageId="2" pageNumber="515">
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(
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<figureCitation id="D5C90790FFAA7B57FF4EF98F350AF9F4" box="[158,266,1588,1613]" captionStart="FIGURES3–8" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES3–8. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="2" pageNumber="515">Figures 3</figureCitation>
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,
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<figureCitation id="D5C90790FFAA7B57FEC7F98E3535F9F4" box="[279,309,1589,1613]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1052,1075]" captionTargetBox="[228,1355,273,1025]" captionTargetId="figure@10.[223,1363,267,1031]" captionTargetPageId="10" captionText="FIGURES 14–19. Male legs. 14, 17 mid leg.15, inner surface—16, outer surface, 18–19 mesobasitarsus. 14–16, Ruizantheda pilosa n. sp.; 17–18, R.baeri (Vachal); 19, R. nigra n. sp." httpUri="https://zenodo.org/record/1242195/files/figure.png" pageId="2" pageNumber="515">19</figureCitation>
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,
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<figureCitation id="D5C90790FFAA7B57FE91F98E3560F9F4" box="[321,352,1589,1613]" captionStart="FIGURES 20–25" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 20–25. Male metasomal sterna. 20–21, S4–5: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S2–4, R. gaullei (Vachal). 23–24, S3–6: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="2" pageNumber="515">20</figureCitation>
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,
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<figureCitation id="D5C90790FFAA7B57FEBBF98F3589F9F4" box="[363,393,1588,1613]" captionStart="FIGURES 26–27" captionStartId="12.[151,264,1992,2015]" captionTargetBox="[259,1316,1494,1970]" captionTargetId="figure@12.[259,1328,1494,1971]" captionTargetPageId="12" captionText="FIGURES 26–27. Male.26, habitus, lateral view, Ruizanthedanigra n. sp. 27, S5,R. colombianan. sp." httpUri="https://zenodo.org/record/1242199/files/figure.png" pageId="2" pageNumber="515">26</figureCitation>
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, 28, 31, 42)
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</paragraph>
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</subSubSection>
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<subSubSection id="05E8489EFFAA7B57FF47F9C73782F8F1" pageId="2" pageNumber="515" type="diagnosis">
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<paragraph id="4D4D1B15FFAA7B57FF47F9C73782F8F1" blockId="2.[151,1437,1660,2009]" pageId="2" pageNumber="515">
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<emphasis id="7F86C707FFAA7B57FF47F9C7350AF92C" bold="true" box="[151,266,1660,1685]" pageId="2" pageNumber="515">Diagnosis</emphasis>
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. Mesobasitarsus short, about one-third length of mesotibia, subquadrate and flattened, with small ventral keel at apex (
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<figureCitation id="D5C90790FFAA7B57FEE4F91B358AF901" box="[308,394,1696,1721]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1052,1075]" captionTargetBox="[228,1355,273,1025]" captionTargetId="figure@10.[223,1363,267,1031]" captionTargetPageId="10" captionText="FIGURES 14–19. Male legs. 14, 17 mid leg.15, inner surface—16, outer surface, 18–19 mesobasitarsus. 14–16, Ruizantheda pilosa n. sp.; 17–18, R.baeri (Vachal); 19, R. nigra n. sp." httpUri="https://zenodo.org/record/1242195/files/figure.png" pageId="2" pageNumber="515">Fig. 19</figureCitation>
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); apical margin of S4 deeply emarginate and bent ventrally, surrounded by many plumose setae (
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<figureCitation id="D5C90790FFAA7B57FF0EF97F3531F964" box="[222,305,1732,1757]" captionStart="FIGURES 20–25" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 20–25. Male metasomal sterna. 20–21, S4–5: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S2–4, R. gaullei (Vachal). 23–24, S3–6: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="2" pageNumber="515">Fig. 20</figureCitation>
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); S5 with extremely-large, U-shaped gradulus (
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<figureCitation id="D5C90790FFAA7B57FC90F97F3793F964" box="[832,915,1732,1757]" captionStart="FIGURES 20–25" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 20–25. Male metasomal sterna. 20–21, S4–5: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S2–4, R. gaullei (Vachal). 23–24, S3–6: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="2" pageNumber="515">Fig. 20</figureCitation>
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); lower lamella of bilobed penis valve forming lateral lobe curved upwards (
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<figureCitation id="D5C90790FFAA7B57FE36F953363CF8B9" box="[486,572,1768,1793]" captionStart="FIGURES 42–44" captionStartId="16.[151,264,1426,1449]" captionTargetBox="[241,1331,260,1411]" captionTargetId="figure@16.[240,1346,251,1423]" captionTargetPageId="16" captionText="FIGURES 42–44. Male. 42–43, genital capsule, ventral view; 44, gonostylus, ventral view. 42, Ruizantheda nigra n. sp.; 43, R.venezuelana n. sp.; 44, R. colombianan. sp." httpUri="https://zenodo.org/record/1242205/files/figure.png" pageId="2" pageNumber="515">Fig. 42</figureCitation>
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). These traits are distinct autapomorphies for this species. Other diagnostic characters include: head and mesosoma blackish with faint green highlights; mesoscutum with abundant dark setae; head as long as wide giving an elongate appearance (
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<figureCitation id="D5C90790FFAA7B57FCE0F88B3774F8F1" box="[816,884,1840,1865]" captionStart="FIGURES3–8" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES3–8. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="2" pageNumber="515">Fig. 3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="05E8489EFFAA7B56FF17F8EF36B0F82C" lastPageId="3" lastPageNumber="516" pageId="2" pageNumber="515" type="description">
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<paragraph id="4D4D1B15FFAA7B57FF17F8EF34CAF829" blockId="2.[151,1437,1660,2009]" pageId="2" pageNumber="515">
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<emphasis id="7F86C707FFAA7B57FF17F8EF3551F8D4" bold="true" box="[199,337,1876,1901]" pageId="2" pageNumber="515">Description</emphasis>
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. Male:
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<emphasis id="7F86C707FFAA7B57FE7BF8EE35D8F8D5" box="[427,472,1877,1900]" italics="true" pageId="2" pageNumber="515">Size</emphasis>
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. Length approximately 11.0 mm. Forewing length 9.0 mm. Intertegular distance
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<quantity id="8A0AB6F0FFAA7B57FAA9F8EF34C6F829" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="515" unit="mm" value="1.6">1.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="4D4D1B15FFAA7B57FF17F827319DF860" blockId="2.[151,1437,1660,2009]" pageId="2" pageNumber="515">
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<emphasis id="7F86C707FFAA7B57FF17F8273515F80C" box="[199,277,1948,1973]" italics="true" pageId="2" pageNumber="515">Colour</emphasis>
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. Head and mesosoma blackish with faint green highlights on supraclypeal area, mesoscutellum, metanotum, and posterior surface of propodeum. About apical one third of clypeus with yellow transverse band.
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</paragraph>
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<paragraph id="4D4D1B15FFAB7B56FF47FF2C306CFF41" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="516">Mandible brown with reddish apex. Labral basal area brownish-yellow. Antenna dark brown, flagellum lighter below than above. Legs dark brown, but yellow on extremities of profemur and basal region of probasitarsus. Metasoma dark brown, bases of T1–6 with faint green highlights. Pygidial plate brown.</paragraph>
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<paragraph id="4D4D1B15FFAB7B56FF17FEBF366CFDD9" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="516">
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<emphasis id="7F86C707FFAB7B56FF17FEBF354BFEA4" box="[199,331,260,285]" italics="true" pageId="3" pageNumber="516">Pubescence</emphasis>
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. Compound eyes with dark setae, longer than ocellar diameter (approximately 1.3×). Face with long, plumose, dark setae, white setae around torulus. Lower paraocular area with strip of short, plumose, white setae along inner margin of compound eye. Anterior half of upper gena with long, plumose, dark setae, such setae white elsewhere. Mesosoma with dark setae, except anterior margin of mesoscutum with white setae. Legs with whitish setae, except profemur with some dark setae, inner surface of tibiae and tarsus with yellowish setae. Discs of T2–5 with semi decumbent to decumbent setae of variable length. Posterior marginal zones of T3–5 with many short, simple, yellowish, appressed setae. Apicolateral lobes of S4 with tuft of branched setae and about 12 stout, simple, recurved setae on margin; concavity of S4 surrounded by many plumose setae. S5 with conspicuous patch of short, decumbent setae inside U-shaped gradulus; apical margin with band of plumose, semi-decumbent setae, slightly longer than those inside gradulus.
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</paragraph>
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<paragraph id="4D4D1B15FFAB7B56FF17FDD636C5FBE1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="516">
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<emphasis id="7F86C707FFAB7B56FF17FDD6352CFD3D" box="[199,300,621,644]" italics="true" pageId="3" pageNumber="516">Structure</emphasis>
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. Head approximately as long as wide (
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<figureCitation id="D5C90790FFAB7B56FD32FDD73726FD3D" box="[738,806,620,645]" captionStart="FIGURES3–8" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES3–8. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="3" pageNumber="516">Fig. 3</figureCitation>
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). Ratio of upper to lower interocular distance about 1.2. Malar area linear. Mandible simple. Labral distal process in form of a minute inverted triangle. Epistomal sulcus forming obtuse angle and slightly rounded. Clypeus projecting approximately two thirds of its length below lower orbital tangent. Ocellocular area concave. Vertex expanded behind ocelli. Frons slightly depressed. Preoccipital ridge rounded. Antenna unmodified, F2 about 2.0× length of F1. Profemur swollen. Mesofemur greatly swollen, ventrally flattened; mesotibia slightly swollen with flat, minutely ridged ventral area and small distal tooth on outer surface; mesobasitarsus similar in shape to other tarsi, short, about one-third length of mesotibia, subquadrate and flattened, with small, distal, and ventral keel (
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<figureCitation id="D5C90790FFAB7B56FD66FCDC370EFC39" box="[694,782,871,896]" captionStart="FIGURES 14–19" captionStartId="10.[151,264,1052,1075]" captionTargetBox="[228,1355,273,1025]" captionTargetId="figure@10.[223,1363,267,1031]" captionTargetPageId="10" captionText="FIGURES 14–19. Male legs. 14, 17 mid leg.15, inner surface—16, outer surface, 18–19 mesobasitarsus. 14–16, Ruizantheda pilosa n. sp.; 17–18, R.baeri (Vachal); 19, R. nigra n. sp." httpUri="https://zenodo.org/record/1242195/files/figure.png" pageId="3" pageNumber="516">Fig. 19</figureCitation>
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). Hind leg unmodified. S4 shortened medially, partially hidden under S3 and bent ventrally (
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<figureCitation id="D5C90790FFAB7B56FD86FC3736B0FC1C" box="[598,688,908,933]" captionStart="FIGURES 26–27" captionStartId="12.[151,264,1992,2015]" captionTargetBox="[259,1316,1494,1970]" captionTargetId="figure@12.[259,1328,1494,1971]" captionTargetPageId="12" captionText="FIGURES 26–27. Male.26, habitus, lateral view, Ruizanthedanigra n. sp. 27, S5,R. colombianan. sp." httpUri="https://zenodo.org/record/1242199/files/figure.png" pageId="3" pageNumber="516">Fig. 26</figureCitation>
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), with transverse median depression and apical margin deeply emarginated (
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<figureCitation id="D5C90790FFAB7B56FEE2FC143588FC71" box="[306,392,943,968]" captionStart="FIGURES 20–25" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 20–25. Male metasomal sterna. 20–21, S4–5: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S2–4, R. gaullei (Vachal). 23–24, S3–6: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="3" pageNumber="516">Fig. 20</figureCitation>
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). Apical margin of S5 very shallowly emarginate, with extremely-large, U-shaped gradulus almost reaching apical margin (
|
||
<figureCitation id="D5C90790FFAB7B56FE26FC6F364AFC55" box="[502,586,980,1005]" captionStart="FIGURES 20–25" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 20–25. Male metasomal sterna. 20–21, S4–5: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S2–4, R. gaullei (Vachal). 23–24, S3–6: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="3" pageNumber="516">Fig. 20</figureCitation>
|
||
). Gradulus of S6 nearly interrupted medially, present only as a very thin line (Fig. 28). S7 with median apical process short, without setae. S8 with median apical process broad, apical margin bearing lateral lobes membranous, separated by a short, thin, sclerotized surface and, rounded lobe medially (Fig. 31). Pygidial plate oval, slightly truncate apically.
|
||
</paragraph>
|
||
<paragraph id="4D4D1B15FFAB7B56FF17FBDF35E4F994" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="516">
|
||
<emphasis id="7F86C707FFAB7B56FF17FBDF3531FBC4" box="[199,305,1124,1149]" italics="true" pageId="3" pageNumber="516">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="D5C90790FFAB7B56FE91FBDF3591FBC5" box="[321,401,1124,1149]" captionStart="FIGURES 42–44" captionStartId="16.[151,264,1426,1449]" captionTargetBox="[241,1331,260,1411]" captionTargetId="figure@16.[240,1346,251,1423]" captionTargetPageId="16" captionText="FIGURES 42–44. Male. 42–43, genital capsule, ventral view; 44, gonostylus, ventral view. 42, Ruizantheda nigra n. sp.; 43, R.venezuelana n. sp.; 44, R. colombianan. sp." httpUri="https://zenodo.org/record/1242205/files/figure.png" pageId="3" pageNumber="516">Fig 42</figureCitation>
|
||
). Basal region of gonobase with W-shaped in ventral view. Gonocoxae with outer margins divergent, convex near apex; inner dorsal margins progressively divergent, at about midpoint becoming slightly curved and much more divergent. Basal region of gonostylus with
|
||
<emphasis id="7F86C707FFAB7B56FC56FB1737A8FB7C" bold="true" box="[902,936,1196,1221]" pageId="3" pageNumber="516">rvl</emphasis>
|
||
sclerotized, long, more than one-half length of gonocoxite, rounded apically, with many rows of simple setae slightly curved upwards at extremity, not overlapping those of opposing gonostylus, and setae on half of inner lateral margin; with broad median projection formed by a sclerotized area that partially borders a translucent and inflated region; inner lobe present.
|
||
<emphasis id="7F86C707FFAB7B56FABCFAAC319DFA89" bold="true" box="[1388,1437,1303,1328]" pageId="3" pageNumber="516">Mgl</emphasis>
|
||
crescent-shaped with outer surface expanded medially, and with scattered short setae.
|
||
<emphasis id="7F86C707FFAB7B56FBA9FA8730ACFAEC" bold="true" box="[1145,1196,1340,1365]" pageId="3" pageNumber="516">Ogp</emphasis>
|
||
large, ending before apex of
|
||
<emphasis id="7F86C707FFAB7B56FF21FAE4351EFAC1" bold="true" box="[241,286,1375,1400]" pageId="3" pageNumber="516">mgl</emphasis>
|
||
, with many erect setae arising at juncture of
|
||
<emphasis id="7F86C707FFAB7B56FCC8FAE43743FAC1" bold="true" box="[792,835,1375,1400]" pageId="3" pageNumber="516">ogp</emphasis>
|
||
with
|
||
<emphasis id="7F86C707FFAB7B56FC54FAE437B1FAC1" bold="true" box="[900,945,1375,1400]" pageId="3" pageNumber="516">mgl</emphasis>
|
||
; outer margin less sclerotized, extending to ventral region and laterally forming an inflated, circular area with cuticular wrinkles; clump of setae at base of
|
||
<emphasis id="7F86C707FFAB7B56FAA0FA3F319BFA24" bold="true" box="[1392,1435,1412,1437]" pageId="3" pageNumber="516">ogp</emphasis>
|
||
absent. Penis valve strongly arched in lateral view; with strong, main, central dorsal ridge; apex pointed and long; outer lateral expansion bilobed with lower lamella forming lateral lobe curved upwards and without projection posteriorly; prong narrow, extending past volsella posteriorly. Volsella medio-apical margin strongly emarginate; basal region slightly rounded.
|
||
</paragraph>
|
||
<paragraph id="4D4D1B15FFAB7B56FF17F98C36B0F82C" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="516">
|
||
<emphasis id="7F86C707FFAB7B56FF17F98C3531F9E9" box="[199,305,1591,1616]" italics="true" pageId="3" pageNumber="516">Sculpture</emphasis>
|
||
. Upper paraocular area and frons densely punctate, with fine, contiguous punctures and setal bases intermixed, separated by 4–6 PD, surface finely roughened near median ocellus. Lower paraocular area with coarse punctures separated by 1–3 PD. Supraclypeal area with punctures separated by 1–2 PD laterally, becoming sparser centrally in lower half. Disc of clypeus with coarse punctures separated by 2–4 PD. Face with surface between punctures microreticulate. Mesoscutum with setal bases separated by 4–6 PD in central area. Mesoscutellum with coarse setal bases denser at sides and posterior margin (1–2 PD), becoming sparser centrally. Pre-episternum with setal bases separated by 3–4 PD. Mesepisternum with setal bases separated by 3–6 PD. Metapostnotum with triangular area defined by microareolate sculpture. Surface between setal bases reticulate throughout mesosoma. Discs of T2–3 with fine punctures separated by about 2–3 PD, punctation reaching apical margin, surface reticulate, slightly finer than remainder of body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05E8489EFFAB7B56FF17F81B302EF800" box="[199,1070,1952,1977]" pageId="3" pageNumber="516" type="distribution">
|
||
<paragraph id="4D4D1B15FFAB7B56FF17F81B302EF800" blockId="3.[151,1437,151,2013]" box="[199,1070,1952,1977]" pageId="3" pageNumber="516">
|
||
<emphasis id="7F86C707FFAB7B56FF17F81B3559F800" bold="true" box="[199,345,1952,1977]" pageId="3" pageNumber="516">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="8AF26096FFAB7B56FEB7F81B3637F801" authority="Coelho & Engel, 2018" authorityName="Coelho & Engel" authorityYear="2018" box="[359,567,1952,1977]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
|
||
<emphasis id="7F86C707FFAB7B56FEB7F81B3637F801" box="[359,567,1952,1977]" italics="true" pageId="3" pageNumber="516">Ruizantheda nigra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64B57A7CFFAB7B56FDEEF81B367CF800" box="[574,636,1952,1977]" pageId="3" pageNumber="516" rank="species">
|
||
<emphasis id="7F86C707FFAB7B56FDEEF81B367CF800" bold="true" box="[574,636,1952,1977]" pageId="3" pageNumber="516">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is presently known only from
|
||
<collectingCountry id="35E55B85FFAB7B56FC05F81B302BF800" box="[981,1067,1952,1977]" name="Bolivia" pageId="3" pageNumber="516">Bolivia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05E8489EFFAB7B51FF17F87F366EFF6C" lastPageId="4" lastPageNumber="517" pageId="3" pageNumber="516" type="materials_examined">
|
||
<paragraph id="4D4D1B15FFAB7B56FF17F87F319CF865" blockId="3.[151,1437,151,2013]" box="[199,1436,1988,2013]" pageId="3" pageNumber="516">
|
||
<materialsCitation id="FD9A1148FFAB7B56FF17F87F319CF865" ID-GBIF-Occurrence="1949259351" box="[199,1436,1988,2013]" collectingDate="1966-10-15" collectingDateMax="1966-10-16" collectingDateMin="1966-10-15" collectionCode="USNM" collectorName="B&K Burks" country="Bolivia" location="Br. Ichilo R." pageId="3" pageNumber="516" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="7F86C707FFAB7B56FF17F87F3572F864" bold="true" box="[199,370,1988,2013]" pageId="3" pageNumber="516">
|
||
<typeStatus id="9249A5B7FFAB7B56FF17F87F3502F864" box="[199,258,1988,2013]" pageId="3" pageNumber="516">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="9249A5B7FFAB7B56FE54F87F35ECF864" box="[388,492,1988,2013]" pageId="3" pageNumber="516" type="holotype">Holotype</typeStatus>
|
||
male (
|
||
<collectionCode id="2BE383D0FFAB7B56FD93F87F3695F864" box="[579,661,1988,2013]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="516">
|
||
<emphasis id="7F86C707FFAB7B56FD93F87F3695F864" bold="true" box="[579,661,1988,2013]" pageId="3" pageNumber="516">USNM</emphasis>
|
||
</collectionCode>
|
||
), “
|
||
<collectingCountry id="35E55B85FFAB7B56FD6BF87F3730F865" box="[699,816,1988,2012]" name="Bolivia" pageId="3" pageNumber="516">BOLIVIA</collectingCountry>
|
||
\
|
||
<collectorName id="E0077EC3FFAB7B56FC94F87E37D6F865" box="[836,982,1988,2013]" pageId="3" pageNumber="516">Br. Ichilo R.</collectorName>
|
||
\
|
||
<location id="482D4DCEFFAB7B56FC3BF87F3071F865" LSID="urn:lsid:plazi:treatment:C55BAA03FFAA7B51FF47F9AA35B3FEF9:482D4DCEFFAB7B56FC3BF87F3071F865" box="[1003,1137,1988,2013]" country="Bolivia" name="Rain Forest" pageId="3" pageNumber="516">Rain Forest</location>
|
||
\
|
||
<collectingDate id="2908C43DFFAB7B56FB54F87F30D4F864" box="[1156,1236,1988,2013]" pageId="3" pageNumber="516" value="2015-10">
|
||
<date id="394C3DD5FFAB7B56FB54F87F30D4F864" box="[1156,1236,1988,2013]" pageId="3" pageNumber="516" value="2015-10">Oct.15</date>
|
||
</collectingDate>
|
||
:16, 1966\ B.&K.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4D4D1B15FFAC7B51FF47FF2C366EFF6C" blockId="4.[151,1436,151,320]" pageId="4" pageNumber="517">
|
||
Burks” and “
|
||
<typeStatus id="9249A5B7FFAC7B51FEE0FF2335C6FF09" box="[304,454,152,176]" pageId="4" pageNumber="517" type="holotype">HOLOTYPE</typeStatus>
|
||
\
|
||
<taxonomicName id="8AF26096FFAC7B51FE36FF2C36B7FF09" authorityName="Coelho & Engel" authorityYear="2018" box="[486,695,151,176]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="517" phylum="Arthropoda" rank="species" species="nigra">Ruizantheda nigra</taxonomicName>
|
||
\Coelho & Engel” [red label]. Head, antennae, left mesofemur, mesotibia, and mesotarsus glued to board.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05E8489EFFAC7B51FF17FF6435B3FEF9" pageId="4" pageNumber="517" type="discussion">
|
||
<paragraph id="4D4D1B15FFAC7B51FF17FF6435B3FEF9" blockId="4.[151,1436,151,320]" pageId="4" pageNumber="517">
|
||
<emphasis id="7F86C707FFAC7B51FF17FF643548FF41" bold="true" box="[199,328,223,248]" pageId="4" pageNumber="517">Comments</emphasis>
|
||
. Among the six species studied,
|
||
<taxonomicName id="8AF26096FFAC7B51FD68FF5A3714FF41" authorityName="Coelho & Engel" authorityYear="2018" box="[696,788,225,248]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="517" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="7F86C707FFAC7B51FD68FF5A3714FF41" box="[696,788,225,248]" italics="true" pageId="4" pageNumber="517">R. nigra</emphasis>
|
||
</taxonomicName>
|
||
has the longest apex to the penis valve.
|
||
<emphasis id="7F86C707FFAC7B51FF17FEBF3546FEA4" bold="true" box="[199,326,260,285]" pageId="4" pageNumber="517">Etymology</emphasis>
|
||
. This specific epithet is taken from the Latin
|
||
<taxonomicName id="8AF26096FFAC7B51FC94FEBE3780FEA5" authorityName="Coelho & Engel" authorityYear="2018" box="[836,896,261,284]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="517" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="7F86C707FFAC7B51FC94FEBE3780FEA5" box="[836,896,261,284]" italics="true" pageId="4" pageNumber="517">nigra</emphasis>
|
||
</taxonomicName>
|
||
, meaning “black” and a reference to the general colour of the integument.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |