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<document id="7B3D370C6089DA679B5811091E491A36" ID-CLB-Dataset="30140" ID-DOI="10.11646/zootaxa.4415.3.6" ID-GBIF-Dataset="f7dfd7e8-238c-432a-9eec-0929f785f747" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242185" ID-ZooBank="30CF1B69-020C-43E8-B8F0-C30B41E04FE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525600684827" checkinUser="plazi" docAuthor="Coelho, Beatriz W. T. &amp; Engel, Michael S." docDate="2018" docId="C55BAA03FFA57B45FF47FBEC3592F90F" docLanguage="en" docName="zootaxa.4415.3.6.imf" docOrigin="Zootaxa 4415 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ruizantheda gaullei Coelho &amp; Engel 2018, comb. n." docType="treatment" docVersion="10" lastPageNumber="529" masterDocId="3962D27BFFA87B55FFD0FFBB3400FFB9" masterDocTitle="A new group of species within the bee genus Ruizantheda, with a revised key to the males of the genus (Hymenoptera: Halictidae: Caenohalictini)" masterLastPageNumber="532" masterPageNumber="513" originalUpdateDomain="Boston" originalUpdateTime="1545217612389" originalUpdateUser="gglab" pageNumber="526" updateTime="1698541935543" updateUser="ExternalLinkService">
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<mods:title id="866FA04E9C26B713B4C9BC3A0FEFF685">A new group of species within the bee genus Ruizantheda, with a revised key to the males of the genus (Hymenoptera: Halictidae: Caenohalictini)</mods:title>
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<mods:namePart id="4BFEDD29CF88CA6D69C14E73CB92F031">Coelho, Beatriz W. T.</mods:namePart>
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<mods:namePart id="75BA6C5CC509A903551EAE58535F02BF">Engel, Michael S.</mods:namePart>
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<mods:title id="112F645ABD241180151CF98A0E7DE578">Zootaxa</mods:title>
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<treatment id="C55BAA03FFA57B45FF47FBEC3592F90F" ID-DOI="http://doi.org/10.5281/zenodo.6490344" ID-GBIF-Taxon="150641818" ID-Zenodo-Dep="6490344" LSID="urn:lsid:plazi:treatment:C55BAA03FFA57B45FF47FBEC3592F90F" httpUri="http://treatment.plazi.org/id/C55BAA03FFA57B45FF47FBEC3592F90F" lastPageId="16" lastPageNumber="529" pageId="13" pageNumber="526">
<subSubSection id="05E8489EFFA57B58FF47FBEC3663FB2A" pageId="13" pageNumber="526" type="nomenclature">
<paragraph id="4D4D1B15FFA57B58FF47FBEC36BCFBC8" blockId="13.[151,700,1111,1171]" box="[151,700,1111,1137]" pageId="13" pageNumber="526">
<heading id="1605AC79FFA57B58FF47FBEC36BCFBC8" bold="true" box="[151,700,1111,1137]" fontSize="11" level="1" pageId="13" pageNumber="526" reason="1">
<emphasis id="7F86C707FFA57B58FF47FBEC36BCFBC8" bold="true" box="[151,700,1111,1137]" pageId="13" pageNumber="526">
<taxonomicName id="8AF26096FFA57B58FF47FBEC3645FBC8" authority="(Vachal, 1903)" authorityName="Coelho &amp; Engel" authorityYear="2018" baseAuthorityName="Vachal" baseAuthorityYear="1903" box="[151,581,1111,1137]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="gaullei" status="comb. nov.">
<emphasis id="7F86C707FFA57B58FF47FBEC358AFBC8" bold="true" box="[151,394,1111,1137]" italics="true" pageId="13" pageNumber="526">Ruizantheda gaullei</emphasis>
(
<bibRefCitation id="296366E4FFA57B58FE49FBEC363CFBC8" box="[409,572,1111,1137]" pageId="13" pageNumber="526" refId="ref13971" refString="Vachal, J. (1903) Etude sur les Halictus d'Amerique (Hym.). Miscellanea Entomologica, 11, 89 - 104." type="journal article">Vachal, 1903</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="64B57A7CFFA57B58FD82FBEC36BCFBC8" box="[594,700,1111,1137]" pageId="13" pageNumber="526">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4D4D1B15FFA57B58FF47FBC13663FB2A" blockId="13.[151,700,1111,1171]" box="[151,611,1146,1171]" pageId="13" pageNumber="526">
(
<figureCitation id="D5C90790FFA57B58FF4EFBC1350AFB2B" box="[158,266,1146,1171]" captionStart="FIGURES38" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES38. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="13" pageNumber="526">Figures 8</figureCitation>
,
<figureCitation id="D5C90790FFA57B58FEC7FBC13560FB2A" box="[279,352,1146,1171]" captionStart="FIGURES 2025" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 2025. Male metasomal sterna. 2021, S45: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S24, R. gaullei (Vachal). 2324, S36: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="13" pageNumber="526">22, 25</figureCitation>
, 29, 32, 3637, 4850)
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFA57B58FF47FB7936BAFB45" pageId="13" pageNumber="526" type="reference_group">
<paragraph id="4D4D1B15FFA57B58FF47FB7936BAFB45" blockId="13.[151,1398,1218,1276]" pageId="13" pageNumber="526">
<treatmentCitationGroup id="6DE23C3BFFA57B58FF47FB7936BAFB45" pageId="13" pageNumber="526">
<taxonomicName id="8AF26096FFA57B58FF47FB7936EAFB63" authority="Vachal, 1903: 97" authorityName="Vachal" authorityPageNumber="97" authorityYear="1903" box="[151,746,1218,1243]" class="Insecta" family="Halictidae" genus="Halictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="gaullei" subGenus="Paragapostemon">
<emphasis id="7F86C707FFA57B58FF47FB7934F6FB62" box="[151,246,1218,1243]" italics="true" pageId="13" pageNumber="526">Halictus</emphasis>
(
<emphasis id="7F86C707FFA57B58FED6FB7835C5FB63" box="[262,453,1219,1242]" italics="true" pageId="13" pageNumber="526">Paragapostemon</emphasis>
)
<emphasis id="7F86C707FFA57B58FE05FB793623FB62" box="[469,547,1218,1243]" italics="true" pageId="13" pageNumber="526">gaullei</emphasis>
<treatmentCitation id="CC533D04FFA57B58FDFAFB7936EAFB63" author="" box="[554,746,1218,1243]" page="97" pageId="13" pageNumber="526">
<bibRefCitation id="296366E4FFA57B58FDFAFB7936C1FB63" box="[554,705,1218,1243]" pageId="13" pageNumber="526" refId="ref13971" refString="Vachal, J. (1903) Etude sur les Halictus d'Amerique (Hym.). Miscellanea Entomologica, 11, 89 - 104." type="journal article">Vachal, 1903</bibRefCitation>
: 97
</treatmentCitation>
</taxonomicName>
. Type locality: Argentina, Tucumán (MNHP) (examined).
<taxonomicName id="8AF26096FFA57B58FF47FB5D35D5FB45" ID-CoL="P9TK" authority="(Vachal)" authorityName="Vachal" baseAuthorityName="Vachal" baseAuthorityYear="1903" box="[151,469,1254,1276]" class="Insecta" family="Halictidae" genus="Caenohalictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="gaullei">
<emphasis id="7F86C707FFA57B58FF47FB5D3575FB45" box="[151,373,1254,1276]" italics="true" pageId="13" pageNumber="526">Caenohalictus gaullei</emphasis>
(Vachal)
</taxonomicName>
;
<treatmentCitation id="CC533D04FFA57B58FE0FFB5D36B5FB45" author="" box="[479,693,1254,1276]" page="240" pageId="13" pageNumber="526">
<bibRefCitation id="296366E4FFA57B58FE0FFB5D3682FB45" box="[479,642,1254,1276]" pageId="13" pageNumber="526" refId="ref13937" refString="Schrottky, C. (1913) La distribucion geografica de los himenopteros argentinos (conclusion). Anales de la Sociedad Cientifica Argentina, 75 (5), 225 - 286." type="journal article">Schrottky, 1913</bibRefCitation>
: 240
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFA57B58FF47FA9330ABFA17" pageId="13" pageNumber="526" type="diagnosis">
<paragraph id="4D4D1B15FFA57B58FF47FA9330ABFA17" blockId="13.[151,1437,1320,1993]" pageId="13" pageNumber="526">
<emphasis id="7F86C707FFA57B58FF47FA93350AFAF8" bold="true" box="[151,266,1320,1345]" pageId="13" pageNumber="526">Diagnosis</emphasis>
. Apical margin of S4 with a small medial projection (
<figureCitation id="D5C90790FFA57B58FCB1FA9337B2FAF8" box="[865,946,1320,1345]" captionStart="FIGURES 2025" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 2025. Male metasomal sterna. 2021, S45: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S24, R. gaullei (Vachal). 2324, S36: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="13" pageNumber="526">Fig. 22</figureCitation>
) is a distinct autapomorphy for this species. Other diagnostic characters include: small body size, mesobasitarsus with a thin keel on the inner surface, moderate size and rounded U-shaped gradulus of S5, with a patch of setae more conspicuous in profile (
<figureCitation id="D5C90790FFA57B58FB6FFACB3113FA30" box="[1215,1299,1392,1417]" captionStart="FIGURES 2025" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 2025. Male metasomal sterna. 2021, S45: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S24, R. gaullei (Vachal). 2324, S36: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="13" pageNumber="526">Fig. 25</figureCitation>
). Regarding the male genitalia, no marked differences between
<taxonomicName id="8AF26096FFA57B58FD01FA2D373EFA17" baseAuthorityName="Vachal" baseAuthorityYear="1903" box="[721,830,1429,1454]" class="Insecta" family="Halictidae" genus="Caenohalictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="gaullei">
<emphasis id="7F86C707FFA57B58FD01FA2D36E2FA14" box="[721,738,1430,1453]" italics="true" pageId="13" pageNumber="526">R</emphasis>
.
<emphasis id="7F86C707FFA57B58FD20FA2E373EFA17" box="[752,830,1429,1454]" italics="true" pageId="13" pageNumber="526">gaullei</emphasis>
</taxonomicName>
and that of
<taxonomicName id="8AF26096FFA57B58FC14FA2D301EFA17" authorityName="Coelho &amp; Engel" authorityYear="2018" baseAuthorityName="Coelho &amp; Engel" baseAuthorityYear="2018" box="[964,1054,1429,1454]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="7F86C707FFA57B58FC14FA2D301EFA17" box="[964,1054,1429,1454]" italics="true" pageId="13" pageNumber="526">R. baeri</emphasis>
</taxonomicName>
were found.
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFA57B45FF17FA03316DFF6D" lastPageId="16" lastPageNumber="529" pageId="13" pageNumber="526" type="description">
<paragraph id="4D4D1B15FFA57B58FF17FA033539FA4C" blockId="13.[151,1437,1320,1993]" pageId="13" pageNumber="526">
<emphasis id="7F86C707FFA57B58FF17FA03356DFA68" bold="true" box="[199,365,1464,1489]" pageId="13" pageNumber="526">Redescription</emphasis>
. Male:
<emphasis id="7F86C707FFA57B58FE1CFA0235F9FA69" box="[460,505,1465,1488]" italics="true" pageId="13" pageNumber="526">Size</emphasis>
. Length approximately 7.0 mm. Forewing length
<quantity id="8A0AB6F0FFA57B58FB97FA0330A4FA69" box="[1095,1188,1464,1489]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="526" unit="mm" value="6.4">6.4 mm</quantity>
. Intertegular distance about
<quantity id="8A0AB6F0FFA57B58FF0CFA663535FA4C" box="[220,309,1501,1525]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="526" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
<paragraph id="4D4D1B15FFA57B58FF17F9BB3588F977" blockId="13.[151,1437,1320,1993]" pageId="13" pageNumber="526">
<emphasis id="7F86C707FFA57B58FF17F9BB3515F9A0" box="[199,277,1536,1561]" italics="true" pageId="13" pageNumber="526">Colour</emphasis>
. Head and mesosoma blackish with dark blue highlights, green reflections on clypeus, lower paraocular area, and sometimes along inner orbit. About distal half of clypeus with yellow transverse band. Mandible blackish with reddish apex. Labral basal area yellow, brown elsewhere. Antenna blackish, flagellum lighter (yellow) below than above. Legs blackish to dark brown, sometimes protibia with small yellowish spot. Metasoma blackish to dark brown, discs of T15 with weak green tint; posterior marginal zones with yellowish-white translucent edges. Pygidial plate brown.
</paragraph>
<paragraph id="4D4D1B15FFA57B58FF17F96330ECF870" blockId="13.[151,1437,1320,1993]" pageId="13" pageNumber="526">
<emphasis id="7F86C707FFA57B58FF17F963354BF948" box="[199,331,1752,1777]" italics="true" pageId="13" pageNumber="526">Pubescence</emphasis>
. Compound eyes with brownish setae, approximately as long as ocellar diameter. Face with long, plumose, whitish setae, less-branched setae on frons and vertex; lower paraocular area with broad strip of short, plumose, white setae along inner margin of compound eye. Mesosoma and legs with whitish setae except metatarsus with yellowish setae. Discs of T25 with whitish setae of variable size. Posterior marginal zones of T35 with short, simple, decumbent setae. Apicolateral lobes of S4 with tuft of branched setae and about 5 stout, simple, recurved setae on margin, a few plumose setae near tuft. S5 with patch of short, decumbent setae, more conspicuous in profile, arising from inside U-shaped gradulus and extending toward apical margin.
</paragraph>
<caption id="198D4B9DFFA67B5BFF47FAF530E7FADC" box="[151,1255,1358,1381]" httpUri="https://zenodo.org/record/1242201/files/figure.png" pageId="14" pageNumber="527" startId="14.[151,264,1358,1381]" targetBox="[222,1364,193,1337]" targetPageId="14">
<paragraph id="4D4D1B15FFA67B5BFF47FAF530E7FADC" blockId="14.[151,1255,1358,1381]" box="[151,1255,1358,1381]" pageId="14" pageNumber="527">
<emphasis id="7F86C707FFA67B5BFF47FAF5354CFADD" bold="true" box="[151,332,1358,1381]" pageId="14" pageNumber="527">FIGURES 3841</emphasis>
. Male genital capsule of
<taxonomicName id="8AF26096FFA67B5BFD9DFAF43710FADC" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[589,784,1359,1381]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="527" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="7F86C707FFA67B5BFD9DFAF43710FADC" box="[589,784,1359,1381]" italics="true" pageId="14" pageNumber="527">Ruizantheda pilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFA67B5BFCC6FAF4374FFADD" box="[790,847,1359,1380]" pageId="14" pageNumber="527" rank="species">
<emphasis id="7F86C707FFA67B5BFCC6FAF4374FFADD" bold="true" box="[790,847,1359,1380]" pageId="14" pageNumber="527">n. sp.</emphasis>
</taxonomicNameLabel>
38, 40 ventral view; 39, 41 dorsal view.
</paragraph>
</caption>
<paragraph id="4D4D1B15FFA67B5BFF17FA29374BF8C7" blockId="14.[151,1437,1425,2026]" pageId="14" pageNumber="527">
<emphasis id="7F86C707FFA67B5BFF17FA29352CFA10" box="[199,300,1426,1449]" italics="true" pageId="14" pageNumber="527">Structure</emphasis>
. Head wider than longer, ratio about 1.2 (
<figureCitation id="D5C90790FFA67B5BFCD5FA2A3749FA13" box="[773,841,1425,1450]" captionStart="FIGURES38" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES38. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="14" pageNumber="527">Fig. 8</figureCitation>
). Ratio of upper to lower interocular distances about 1.3. Malar area linear. Mandible simple. Labral distal process in form of a minute inverted triangle. Epistomal sulcus forming obtuse angle. Clypeus projecting approximately one-half of its total length below lower orbital tangent. Ocellocular area slightly concave. Vertex slightly expanded behind ocelli. Frons slightly depressed. Preoccipital ridge rounded. Antenna unmodified, F2 1.7× length of F1. Profemur swollen. Outer surface of mesotrochanter with ventral margin convex, forming a very weak keel; mesofemur greatly swollen, ventrally flattened; mesotibia slightly swollen with flat, minutely ridged ventral area and minute distal tooth on outer surface; mesobasitarsus nearly one-half length of mesotibia, with thin keel restricted to one margin of inner surface. Hind legs unmodified. S4 shortened medially, largely hidden under S3, with transverse median depression and apical margin shallowly emarginate, with small medial projection (
<figureCitation id="D5C90790FFA67B5BFC20F96E304BF957" box="[1008,1099,1749,1774]" captionStart="FIGURES 2025" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 2025. Male metasomal sterna. 2021, S45: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S24, R. gaullei (Vachal). 2324, S36: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="14" pageNumber="527">Fig. 22</figureCitation>
). Apical margin of S5 very shallowly emarginate, moderate size and rounded U-shaped gradulus (
<figureCitation id="D5C90790FFA67B5BFC35F9423040F8AB" box="[997,1088,1785,1810]" captionStart="FIGURES 2025" captionStartId="12.[151,264,1391,1414]" captionTargetBox="[229,1357,598,1350]" captionTargetId="figure@12.[221,1365,590,1370]" captionTargetPageId="12" captionText="FIGURES 2025. Male metasomal sterna. 2021, S45: 20, Ruizantheda nigra n. sp.; 21, R. venezuelana n. sp. 22, S24, R. gaullei (Vachal). 2324, S36: 23, R.baeri (Vachal); 24, R. pilosa n. sp. 25, S5, R. gaullei." httpUri="https://zenodo.org/record/1242197/files/figure.png" pageId="14" pageNumber="527">Fig. 25</figureCitation>
). Gradulus of S6 interrupted medially (Fig. 29). S7 with median apical process angulate, short and without setae. S8 with median apical process broad and trapezoidal shaped, apical margin bearing lateral membranous lobes, separated by a transverse, very thin surface, a round lobe medially (Fig. 32). Pygidial plate ovoid.
</paragraph>
<paragraph id="4D4D1B15FFA67B5AFF17F832366FFE4C" blockId="14.[151,1437,1425,2026]" lastBlockId="15.[151,1436,151,501]" lastPageId="15" lastPageNumber="528" pageId="14" pageNumber="527">
<emphasis id="7F86C707FFA67B5BFF17F8323531F81B" box="[199,305,1929,1954]" italics="true" pageId="14" pageNumber="527">Genitalia</emphasis>
(
<figureCitation id="D5C90790FFA67B5BFE94F83235D0F81B" box="[324,464,1929,1954]" captionStart="FIGURES 3437" captionStartId="15.[151,264,1703,1726]" captionTargetBox="[225,1363,542,1683]" captionTargetId="figure@15.[224,1363,541,1683]" captionTargetPageId="15" captionText="FIGURES 3437. Male genital capsule. 34, 36 ventral view; 35, 37 dorsal view. 3435, R. colombiana n. sp.; 3637, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242203/files/figure.png" pageId="14" pageNumber="527">Figs. 3637</figureCitation>
). Basal region of gonobase with W-shaped in ventral view. Gonocoxae with outer margins divergent and convex near apex; inner dorsal margins progressively divergent, at about midpoint becoming slightly curved and more divergent. Gonostylus with basal region with
<emphasis id="7F86C707FFA67B5BFBE6F86A3058F853" bold="true" box="[1078,1112,2001,2026]" pageId="14" pageNumber="527">rvl</emphasis>
sclerotized, long, more than one-half length of gonocoxite and rounded apically, with rows of simple setae at extremity, overlapping those of opposing gonostylus; with middle and broad projection formed by a sclerotized area that partially borders a translucent and inflated region, inner lobe present.
<emphasis id="7F86C707FFA77B5AFD04FF643705FF41" bold="true" box="[724,773,223,248]" pageId="15" pageNumber="528">Mgl</emphasis>
an crescent-shaped, with outer surface expanded medially, and with scattered short setae.
<emphasis id="7F86C707FFA77B5AFE3CFEBF361FFEA4" bold="true" box="[492,543,260,285]" pageId="15" pageNumber="528">Ogp</emphasis>
large, terminating before apex of
<emphasis id="7F86C707FFA77B5AFC4AFEBF37C7FEA4" bold="true" box="[922,967,260,285]" pageId="15" pageNumber="528">mgl</emphasis>
, with many long, plumose setae arising at juncture of
<emphasis id="7F86C707FFA77B5AFEF1FE9C354DFEF9" bold="true" box="[289,333,295,320]" pageId="15" pageNumber="528">ogp</emphasis>
with
<emphasis id="7F86C707FFA77B5AFE46FE9C35C3FEF9" bold="true" box="[406,451,295,320]" pageId="15" pageNumber="528">mgl</emphasis>
, outer margin less sclerotized, extending to ventral region and laterally forming an inflated area with cuticular wrinkles; clump of setae of different lengths at base of
<emphasis id="7F86C707FFA77B5AFBE0FEF7305CFEDC" bold="true" box="[1072,1116,332,357]" pageId="15" pageNumber="528">ogp</emphasis>
, most setae semi decumbent and turning toward dorsal surface. Penis valve strongly arched in lateral view; strong, main, central dorsal ridge; apex pointed; outer lateral expansion strongly bilobed, nearly right-angled between two sections and with posterior projection; prong very narrow, extending past volsella posteriorly. Volsella medio-apical margin strongly emarginate; basal region slightly rounded.
</paragraph>
<caption id="198D4B9DFFA77B5AFF47F91C3541F964" httpUri="https://zenodo.org/record/1242203/files/figure.png" pageId="15" pageNumber="528" startId="15.[151,264,1703,1726]" targetBox="[225,1363,542,1683]" targetPageId="15">
<paragraph id="4D4D1B15FFA77B5AFF47F91C3541F964" blockId="15.[151,1436,1703,1757]" pageId="15" pageNumber="528">
<emphasis id="7F86C707FFA77B5AFF47F91C3550F904" bold="true" box="[151,336,1703,1726]" pageId="15" pageNumber="528">FIGURES 3437</emphasis>
. Male genital capsule. 34, 36 ventral view; 35, 37 dorsal view. 3435,
<taxonomicName id="8AF26096FFA77B5AFB99F91230E0F907" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[1097,1248,1704,1726]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="528" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="7F86C707FFA77B5AFB99F91230E0F907" box="[1097,1248,1704,1726]" italics="true" pageId="15" pageNumber="528">R. colombiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFA77B5AFB3AF9133126F904" box="[1258,1318,1704,1725]" pageId="15" pageNumber="528" rank="species">
<emphasis id="7F86C707FFA77B5AFB3AF9133126F904" bold="true" box="[1258,1318,1704,1725]" pageId="15" pageNumber="528">n. sp.</emphasis>
</taxonomicNameLabel>
; 3637,
<taxonomicName id="8AF26096FFA77B5AFA56F912353DF964" authority="(Vachal)" authorityName="Vachal" baseAuthorityName="Vachal" baseAuthorityYear="1903" class="Insecta" family="Halictidae" genus="Caenohalictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="528" phylum="Arthropoda" rank="species" species="gaullei">
<emphasis id="7F86C707FFA77B5AFA56F91234DDF964" italics="true" pageId="15" pageNumber="528">R. gaullei</emphasis>
(Vachal)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="4D4D1B15FFA77B5AFF17F8B8319DF84D" blockId="15.[151,1437,1795,2036]" pageId="15" pageNumber="528">
<emphasis id="7F86C707FFA77B5AFF17F8B83531F8A5" box="[199,305,1795,1820]" italics="true" pageId="15" pageNumber="528">Sculpture</emphasis>
. Upper paraocular area and frons densely punctate, with fine, contiguous punctures and some scattered setal bases intermixed. Lower paraocular area with coarse punctures separated by 13 PD. Supraclypeal area with punctures separated by 14 PD. Clypeus with punctures separated by 13 PD laterally, becoming sparser medially. Face with surface between punctures microreticulate. Mesoscutum with setal bases separated by 48 PD centrally. Mesoscutellum with a few coarse setal bases centrally, becoming more densely punctured posteriorly and laterally (14 PD). Pre-episternum with setal bases separated by 24 PD. Mesepisternum with setal bases separated by 24 PD. Metapostnotum with triangular area defined by microareolate sculpture, usually without rugosity.
</paragraph>
<paragraph id="4D4D1B15FFB87B45FF47FF2C316DFF6D" blockId="16.[151,1436,151,213]" pageId="16" pageNumber="529">Surface between setal bases reticulate throughout mesosoma. Discs of T23 with strong punctures separated by about 4 PD and finer punctures intermixed, posterior marginal zones with fine punctures separated by 23 PD.</paragraph>
</subSubSection>
<caption id="198D4B9DFFB87B45FF47FA293676FA7F" httpUri="https://zenodo.org/record/1242205/files/figure.png" pageId="16" pageNumber="529" startId="16.[151,264,1426,1449]" targetBox="[241,1331,260,1411]" targetPageId="16">
<paragraph id="4D4D1B15FFB87B45FF47FA293676FA7F" blockId="16.[151,1436,1426,1479]" pageId="16" pageNumber="529">
<emphasis id="7F86C707FFB87B45FF47FA29354EFA1E" bold="true" box="[151,334,1426,1449]" pageId="16" pageNumber="529">FIGURES 4244</emphasis>
. Male. 4243, genital capsule, ventral view; 44, gonostylus, ventral view. 42,
<taxonomicName id="8AF26096FFB87B45FBA0FA29312DFA11" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[1136,1325,1426,1448]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="529" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
<emphasis id="7F86C707FFB87B45FBA0FA29312DFA11" box="[1136,1325,1426,1448]" italics="true" pageId="16" pageNumber="529">Ruizantheda nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFB87B45FAE4FA28316EFA11" box="[1332,1390,1427,1448]" pageId="16" pageNumber="529" rank="species">
<emphasis id="7F86C707FFB87B45FAE4FA28316EFA11" bold="true" box="[1332,1390,1427,1448]" pageId="16" pageNumber="529">n. sp.</emphasis>
</taxonomicNameLabel>
; 43,
<taxonomicName id="8AF26096FFB87B45FF47FA093530FA7E" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[151,304,1457,1479]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="529" phylum="Arthropoda" rank="species" species="venezuelana" status="sp. nov.">
<emphasis id="7F86C707FFB87B45FF47FA093530FA7E" box="[151,304,1457,1479]" italics="true" pageId="16" pageNumber="529">R. venezuelana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFB87B45FEE6FA0A3570FA7F" box="[310,368,1457,1478]" pageId="16" pageNumber="529" rank="species">
<emphasis id="7F86C707FFB87B45FEE6FA0A3570FA7F" bold="true" box="[310,368,1457,1478]" pageId="16" pageNumber="529">n. sp.</emphasis>
</taxonomicNameLabel>
; 44,
<taxonomicName id="8AF26096FFB87B45FE73FA093637FA7E" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[419,567,1457,1479]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="529" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="7F86C707FFB87B45FE73FA093637FA7E" box="[419,567,1457,1479]" italics="true" pageId="16" pageNumber="529">R. colombiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFB87B45FDECFA0A3676FA7F" box="[572,630,1457,1478]" pageId="16" pageNumber="529" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="05E8489EFFB87B45FF17F9B63592F90F" pageId="16" pageNumber="529" type="distribution">
<paragraph id="4D4D1B15FFB87B45FF17F9B63592F90F" blockId="16.[151,1436,1549,1718]" pageId="16" pageNumber="529">
<emphasis id="7F86C707FFB87B45FF17F9B63560F99F" bold="true" box="[199,352,1549,1574]" pageId="16" pageNumber="529">Distribution.</emphasis>
<taxonomicName id="8AF26096FFB87B45FEB7F9B63648F99F" authorityName="Coelho &amp; Engel" authorityYear="2018" baseAuthorityName="Coelho &amp; Engel" baseAuthorityYear="2018" box="[359,584,1549,1574]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="529" phylum="Arthropoda" rank="species" species="gaullei">
<emphasis id="7F86C707FFB87B45FEB7F9B63648F99F" box="[359,584,1549,1574]" italics="true" pageId="16" pageNumber="529">Ruizantheda gaullei</emphasis>
</taxonomicName>
is found in
<collectingCountry id="35E55B85FFB87B45FD1DF9B63741F99F" box="[717,833,1549,1574]" name="Argentina" pageId="16" pageNumber="529">Argentina</collectingCountry>
: provinces of
<collectingRegion id="8F36D5F7FFB87B45FC0EF9B63016F99F" box="[990,1046,1549,1574]" country="Argentina" name="Salta" pageId="16" pageNumber="529">Salta</collectingRegion>
and
<collectingRegion id="8F36D5F7FFB87B45FB9DF9B530B9F99C" box="[1101,1209,1550,1573]" country="Argentina" name="Tucuman" pageId="16" pageNumber="529">Tucuman</collectingRegion>
.
<emphasis id="7F86C707FFB87B45FF17F98A3575F9F3" bold="true" box="[199,373,1585,1610]" pageId="16" pageNumber="529">
<typeStatus id="9249A5B7FFB87B45FF17F98A3502F9F3" box="[199,258,1585,1610]" pageId="16" pageNumber="529">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="FD9A1148FFB87B45FEACF98A30E3F9F3" ID-GBIF-Occurrence="1948728429" box="[380,1251,1585,1610]" collectionCode="MNHP" country="Argentina" pageId="16" pageNumber="529" specimenCount="1" specimenCount-female="1" stateProvince="Tucuman" typeStatus="holotype">
<typeStatus id="9249A5B7FFB87B45FEACF98A35E4F9F3" box="[380,484,1585,1610]" pageId="16" pageNumber="529" type="holotype">Holotype</typeStatus>
female (
<collectionCode id="2BE383D0FFB87B45FD97F98A369CF9F3" box="[583,668,1585,1610]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="16" pageNumber="529">
<emphasis id="7F86C707FFB87B45FD97F98A369CF9F3" bold="true" box="[583,668,1585,1610]" pageId="16" pageNumber="529">MNHP</emphasis>
</collectionCode>
),
<collectingRegion id="8F36D5F7FFB87B45FD64F98A3720F9F0" box="[692,800,1585,1609]" country="Argentina" name="Tucuman" pageId="16" pageNumber="529">Tucumán</collectingRegion>
,
<collectingCountry id="35E55B85FFB87B45FCFBF98A379EF9F3" box="[811,926,1585,1610]" name="Argentina" pageId="16" pageNumber="529">Argentina</collectingCountry>
. Specimen in good condition
</materialsCitation>
(
<figureCitation id="D5C90790FFB87B45FB23F98A317DF9F3" box="[1267,1405,1585,1610]" captionStart="FIGURES 4550" captionStartId="17.[151,264,723,746]" captionTargetBox="[222,1364,193,702]" captionTargetId="figure@17.[222,1364,193,702]" captionTargetPageId="17" captionText="FIGURES 4550. Female holotype. 45, 48, dorsal view; 46, 49, head; 47, 50, mesosoma, dorsal view. 4547, R. baeri (Vachal); 4850, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242207/files/figure.png" pageId="16" pageNumber="529">Figs. 4850</figureCitation>
).
<emphasis id="7F86C707FFB87B45FF17F9EE35AEF9D7" bold="true" box="[199,430,1621,1646]" pageId="16" pageNumber="529">Material examined</emphasis>
.
<materialsCitation id="FD9A1148FFB87B45FE11F9ED358CF928" ID-GBIF-Occurrence="1948728574" collectingDate="1984-02" collectionCode="AMNH" collectorName="Fritz" country="Argentina" elevation="2500" location="Cuesta Obispo" pageId="16" pageNumber="529" specimenCount="1" specimenCount-male="1" stateProvince="Salta">
<collectingCountry id="35E55B85FFB87B45FE11F9ED3667F9D7" box="[449,615,1622,1646]" name="Argentina" pageId="16" pageNumber="529">ARGENTINA</collectingCountry>
,
<collectingRegion id="8F36D5F7FFB87B45FDA8F9EE36B4F9D7" box="[632,692,1621,1646]" country="Argentina" name="Salta" pageId="16" pageNumber="529">Salta</collectingRegion>
: one male (
<collectionCode id="2BE383D0FFB87B45FC92F9ED379EF9D7" box="[834,926,1622,1646]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="529">AMNH</collectionCode>
), &quot;
<collectionCode id="2BE383D0FFB87B45FC6EF9ED37F9F9D7" box="[958,1017,1622,1646]" country="Malta" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="16" pageNumber="529">ARG</collectionCode>
.—
<collectionCode id="2BE383D0FFB87B45FBC8F9ED3071F9D7" box="[1048,1137,1622,1646]" pageId="16" pageNumber="529">
<collectingRegion id="8F36D5F7FFB87B45FBC8F9ED3071F9D7" box="[1048,1137,1622,1646]" country="Argentina" name="Salta" pageId="16" pageNumber="529">SALTA</collectingRegion>
</collectionCode>
\ Cuesta Obispo\
<quantity id="8A0AB6F0FFB87B45FA92F9EE319CF9D4" box="[1346,1436,1621,1646]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="529" unit="m" value="2500.0">2500 m</quantity>
aprox.\ Fritzb- 2.84&quot;.
</materialsCitation>
<materialsCitation id="FD9A1148FFB87B45FE46F9C13585F90F" ID-GBIF-Occurrence="1948728572" collectingDate="1971-12-06" collectionCode="AMNH" collectorName="Charles Porter" country="Argentina" location="Tafi del Valle" pageId="16" pageNumber="529" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman">
<collectingRegion id="8F36D5F7FFB87B45FE46F9C13603F928" box="[406,515,1658,1681]" country="Argentina" name="Tucuman" pageId="16" pageNumber="529">Tucuman</collectingRegion>
: one male (
<collectionCode id="2BE383D0FFB87B45FD5FF9C236EBF928" box="[655,747,1657,1681]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="529">AMNH</collectionCode>
), &quot;
<collectingCountry id="35E55B85FFB87B45FCC8F9C237BEF928" box="[792,958,1657,1681]" name="Argentina" pageId="16" pageNumber="529">ARGENTINA</collectingCountry>
,
<collectingRegion id="8F36D5F7FFB87B45FC1EF9C23036F928" box="[974,1078,1657,1681]" country="Argentina" name="Tucuman" pageId="16" pageNumber="529">Tucumán</collectingRegion>
\ Tafi del. Valle,\
<date id="394C3DD5FFB87B45FAD9F9C234CEF90F" pageId="16" pageNumber="529" value="1971-12-06">December 6, 1971</date>
\ Charles Porter
</materialsCitation>
&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>