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<document id="36DEEE24797820609F164C3464401A27" ID-CLB-Dataset="31822" ID-DOI="10.11646/zootaxa.4337.2.1" ID-GBIF-Dataset="bca0e227-e4fb-4f93-b99b-c8794b19de8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1018631" ID-ZooBank="A712D90D-FCD1-49C9-928F-21C95814815B" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508314185452" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M." docDate="2017" docId="9D02C578FF81FF87FF12CE1DFA87FA41" docLanguage="en" docName="zootaxa.4337.2.1.pdf" docOrigin="Zootaxa 4337 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tegoribates walteri Behan-Pelletier 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="187" masterDocId="613BBD00FF9EFFA3FF85CC7CFF9CFFD0" masterDocTitle="Tegoribatidae of North America, with proposal of Protectoribates gen. nov., and new species (Acari, Oribatida, Tegoribatidae)" pageNumber="182" updateTime="1698469699374" updateUser="ExternalLinkService">
<mods:mods id="C227836B0D7869F7AF294381F6DDC14A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D493845725C24AC30F6AC24E3A8D3F71">
<mods:title id="423372AB7413DBA5AA2E4CEE90FCDEF3">Tegoribatidae of North America, with proposal of Protectoribates gen. nov., and new species (Acari, Oribatida, Tegoribatidae)</mods:title>
</mods:titleInfo>
<mods:name id="D60F44DB6701AF186FA548AE76424B13" type="personal">
<mods:role id="7F8FB2F3BD36CFA98E56D97323FECD98">
<mods:roleTerm id="6D216A1B9A86FD6FEA4B3F12BE40AA91">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF56D34AB07A4779AE639428FF921E39">Behan-Pelletier, Valerie M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="91FA72C2D231AAD28A6982B935FD6629">text</mods:typeOfResource>
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<mods:titleInfo id="B171CD60016910946E99D3A1C4313FF2">
<mods:title id="2A226157013401BFCA56539CD71283DE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6D3A5B6AFF837DD7EB2A5DC557D96326">
<mods:date id="BAB8B7159DD6A95C2E4CA38B4B6B64B7">2017</mods:date>
<mods:detail id="192106FE38D6FCE9B57D4C50A214B077" type="pubDate">
<mods:number id="15922DADC14C110DEB4D37ABED2EF32A">2017-10-18</mods:number>
</mods:detail>
<mods:detail id="3E47804EFF646978F1B4CBBF45E21485" type="volume">
<mods:number id="434F9A4BE0412C70AFC4F8FCC3135FE5">4337</mods:number>
</mods:detail>
<mods:detail id="AD6C77C5AC38B4923B992000BED39558" type="issue">
<mods:number id="673B643CCA1BD4472DC5B184161A95FD">2</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="E930E9E4025F738303D5B3777F8903F8">journal volume</mods:classification>
<mods:identifier id="B4FB35810B2CE99E17C2D1B510643B67" type="CLB-Dataset">31822</mods:identifier>
<mods:identifier id="F16451C78481167DDA5D4407092D7F1B" type="DOI">10.11646/zootaxa.4337.2.1</mods:identifier>
<mods:identifier id="63B934907202C73CEF029E4C0D731110" type="GBIF-Dataset">bca0e227-e4fb-4f93-b99b-c8794b19de8d</mods:identifier>
<mods:identifier id="B854D76CAE28641C7F319304FCC233DB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="07D77C56C8D83E729CD7CB9ABCD2BF83" type="Zenodo-Dep">1018631</mods:identifier>
<mods:identifier id="CB20CEA241A416BCD2553D1C6759098F" type="ZooBank">A712D90D-FCD1-49C9-928F-21C95814815B</mods:identifier>
</mods:mods>
<treatment id="9D02C578FF81FF87FF12CE1DFA87FA41" ID-DOI="http://doi.org/10.5281/zenodo.5664409" ID-GBIF-Taxon="135450340" ID-Zenodo-Dep="5664409" LSID="urn:lsid:plazi:treatment:9D02C578FF81FF87FF12CE1DFA87FA41" httpUri="http://treatment.plazi.org/id/9D02C578FF81FF87FF12CE1DFA87FA41" lastPageId="36" lastPageNumber="187" pageId="31" pageNumber="182">
<subSubSection id="5DB127E5FF81FFBCFF12CE1DFE24FD4C" pageId="31" pageNumber="182" type="nomenclature">
<paragraph id="1514746EFF81FFBCFF12CE1DFE7DFDAB" blockId="31.[151,481,609,669]" box="[151,481,609,635]" pageId="31" pageNumber="182">
<heading id="4E5CC302FF81FFBCFF12CE1DFE7DFDAB" bold="true" box="[151,481,609,635]" fontSize="11" level="1" pageId="31" pageNumber="182" reason="1">
<emphasis id="27DFA87CFF81FFBCFF12CE1DFE7DFDAB" bold="true" box="[151,481,609,635]" pageId="31" pageNumber="182">
<taxonomicName id="D2AB0FEDFF81FFBCFF12CE1DFEE0FDAB" ID-CoL="8TDNQ" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[151,380,609,635]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFF81FFBCFF12CE1DFEE0FDAB" bold="true" box="[151,380,609,635]" italics="true" pageId="31" pageNumber="182">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FF81FFBCFE06CE1DFE7DFDAB" box="[387,481,609,635]" pageId="31" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1514746EFF81FFBCFF12CEF8FE24FD4C" blockId="31.[151,481,609,669]" box="[151,440,644,669]" pageId="31" pageNumber="182">
(
<figureCitation id="8D9068EBFF81FFBCFF1ACEF8FF60FD4C" box="[159,252,644,669]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="31" pageNumber="182">Figs. 20</figureCitation>
,
<figureCitation id="8D9068EBFF81FFBCFE8DCEF8FEBAFD4C" box="[264,294,644,668]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">21</figureCitation>
,
<figureCitation id="8D9068EBFF81FFBCFEB7CEF8FE1BFD4C" box="[306,391,644,668]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="31" pageNumber="182">22A–D</figureCitation>
,
<figureCitation id="8D9068EBFF81FFBCFE16CEF8FE2CFD4C" box="[403,432,644,668]" captionStart="FIGURE 23" captionStartId="35.[151,250,1921,1943]" captionTargetBox="[210,1367,193,1875]" captionTargetId="figure@35.[210,1367,193,1894]" captionTargetPageId="35" captionText="FIGURE 23. Tegoribates walteri sp. nov., differential interference contrast microscope images of immatures; A, larval scalp (3 layers); B, protonymph (3 layers), dorsal aspect, larval scalp removed; C, larval and protonymphal scalp, with arrow to tubercle in region of setae dp; D, deutonymph, dorsal aspect, with scalps removed. Scale bars: A – C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/1018679/files/figure.png" pageId="31" pageNumber="182">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FF81FFBCFF12CEBAFEFAFC9B" pageId="31" pageNumber="182" type="materials_examined">
<paragraph id="1514746EFF81FFBCFF12CEBAFEFAFC9B" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<materialsCitation id="A5C37E33FF81FFBCFF12CEBAFEDDFCF7" ID-GBIF-Occurrence="1675310254" collectingDate="2009-03-06" collectionCode="CNC" collectorName="M. Behan" country="United States of America" county="Sonoma Co." location="Salt Point State Park" municipality="Material" pageId="31" pageNumber="182" specimenCount="2" stateProvince="California" typeStatus="holotype">
<emphasis id="27DFA87CFF81FFBCFF12CEBAFE1DFD0F" bold="true" box="[151,385,710,735]" pageId="31" pageNumber="182">
<collectingMunicipality id="F570EE14FF81FFBCFF12CEBAFF63FD0F" box="[151,255,710,735]" pageId="31" pageNumber="182">Material</collectingMunicipality>
examined
</emphasis>
:
<typeStatus id="CA10CACCFF81FFBCFE12CEBAFE62FD0F" box="[407,510,710,735]" pageId="31" pageNumber="182" type="holotype">
<emphasis id="27DFA87CFF81FFBCFE12CEBAFE62FD0F" box="[407,510,710,735]" italics="true" pageId="31" pageNumber="182">Holotype</emphasis>
</typeStatus>
: adult male,
<collectingCountry id="6DBC34FEFF81FFBCFD22CEBBFD7EFD0F" box="[679,738,711,735]" name="United States of America" pageId="31" pageNumber="182">USA</collectingCountry>
,
<collectingRegion id="D76FBA8CFF81FFBCFD70CEBAFCF6FD0F" box="[757,874,710,735]" country="United States of America" name="California" pageId="31" pageNumber="182">California</collectingRegion>
,
<collectingCounty id="FC750CE2FF81FFBCFCF9CEBBFB8CFD0F" box="[892,1040,711,735]" pageId="31" pageNumber="182">Sonoma Co.</collectingCounty>
,
<location id="107422B5FF81FFBCFBA1CEBAFAB8FD0F" LSID="urn:lsid:plazi:treatment:9D02C578FF81FF87FF12CE1DFA87FA41:107422B5FF81FFBCFBA1CEBAFAB8FD0F" box="[1060,1316,710,735]" country="United States of America" county="Sonoma Co." municipality="Material" name="Salt Point State Park" pageId="31" pageNumber="182" stateProvince="California">Salt Point State Park</location>
, 38°567N 123°333W,
<location id="107422B5FF81FFBCFE9BCE97FE59FCD3" LSID="urn:lsid:plazi:treatment:9D02C578FF81FF87FF12CE1DFA87FA41:107422B5FF81FFBCFE9BCE97FE59FCD3" box="[286,453,746,771]" country="United States of America" county="Sonoma Co." municipality="Material" name="Sea Cliff Trail" pageId="31" pageNumber="182" stateProvince="California">Sea Cliff Trail</location>
,
<collectingDate id="7151AB46FF81FFBCFE56CE96FDA2FCD3" box="[467,574,746,771]" pageId="31" pageNumber="182" value="2009-03-06">
<date id="611552AEFF81FFBCFE56CE96FDA2FCD3" box="[467,574,746,771]" pageId="31" pageNumber="182" value="2009-03-06">6.iii.2009</date>
</collectingDate>
(VBP and
<collectorName id="B85E11B8FF81FFBCFD46CE97FCA9FCD3" box="[707,821,746,771]" pageId="31" pageNumber="182">M. Behan</collectorName>
) from litter under lupins; deposited in the
<collectionCode id="73BAECABFF81FFBCFAA7CE97FAC2FCD3" box="[1314,1374,747,771]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="31" pageNumber="182">CNC</collectionCode>
,
<typeStatus id="CA10CACCFF81FFBCFAE9CE97FA00FCD3" box="[1388,1436,747,771]" pageId="31" pageNumber="182">type</typeStatus>
number 24350
</materialsCitation>
.
<materialsCitation id="A5C37E33FF81FFBCFED5CF73FEFAFC9B" ID-GBIF-Occurrence="1675310262" collectingDate="2009-03-06" collectionCode="CNC, USNM, RNC" collectorName="M. Behan" country="United States of America" county="Sonoma Co." location="Salt Point State Park" municipality="Material" pageId="31" pageNumber="182" specimenCount="12" specimenCount-female="2" specimenCount-male="10" stateProvince="California" typeStatus="paratype">
<typeStatus id="CA10CACCFF81FFBCFED5CF73FE5DFCF6" box="[336,449,783,806]" pageId="31" pageNumber="182" type="paratype">
<emphasis id="27DFA87CFF81FFBCFED5CF73FE5DFCF6" box="[336,449,783,806]" italics="true" pageId="31" pageNumber="182">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="03ADBFE7FF81FFBCFE51CF73FDDBFCF7" box="[468,583,782,807]" pageId="31" pageNumber="182" type="female">2 females</specimenCount>
,
<specimenCount id="03ADBFE7FF81FFBCFDD2CF73FD23FCF7" box="[599,703,782,807]" pageId="31" pageNumber="182" type="male">10 males</specimenCount>
with same data as holotype. Paratypes deposited in the
<collectionCode id="73BAECABFF81FFBCFADECF73FA04FCF7" box="[1371,1432,783,807]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="31" pageNumber="182">CNC</collectionCode>
,
<collectionCode id="73BAECABFF81FFBCFF12CF4FFF77FC9B" box="[151,235,819,843]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="31" pageNumber="182">USNM</collectionCode>
, and
<collectionCode id="73BAECABFF81FFBCFEA0CF4FFEFEFC9B" box="[293,354,819,843]" collectionName="RNC" pageId="31" pageNumber="182">RNC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FF81FFBCFF42CF2AFBAFFBF3" pageId="31" pageNumber="182" type="diagnosis">
<paragraph id="1514746EFF81FFBCFF42CF2AFBAFFBF3" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42CF2AFEA6FCBF" bold="true" box="[199,314,854,879]" pageId="31" pageNumber="182">Diagnosis</emphasis>
. Total length 350–400. Lamella 104 long, medially with V-shaped indentation, about 3 long. Lamellar seta arising anteriorly on fused lamella, 10–17 long. Interlamellar setae 7–11. Bothridial seta fusiform, 77–90 long. Pedotectum I pointed distally. Muscle sigillae strongly developed on notogaster (clearly visible with incident illumination). Octotaxic system as 4 pairs porose areas,
<emphasis id="27DFA87CFF81FFBCFCE0CFBFFC18FC0A" box="[869,900,963,986]" italics="true" pageId="31" pageNumber="182">Aa</emphasis>
about
<quantity id="D253D98BFF81FFBCFC55CFBFFB9BFC0B" box="[976,1031,962,987]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="31" pageNumber="182" unit="in" value="11.0">11 in</quantity>
diameter. Notogastral setae smooth, 6–8, setae subequal in length;
<emphasis id="27DFA87CFF81FFBCFE62CF9AFD9FFC2F" box="[487,515,998,1023]" italics="true" pageId="31" pageNumber="182">lm</emphasis>
posteromedial of
<emphasis id="27DFA87CFF81FFBCFD49CF9BFD77FC2E" box="[716,747,999,1022]" italics="true" pageId="31" pageNumber="182">Aa</emphasis>
and
<emphasis id="27DFA87CFF81FFBCFCA4CF9AFCABFC2F" box="[801,823,998,1023]" italics="true" pageId="31" pageNumber="182">lp</emphasis>
dorsocentrally positioned, medial of
<emphasis id="27DFA87CFF81FFBCFB53CF9BFB69FC2F" box="[1238,1269,999,1023]" italics="true" pageId="31" pageNumber="182">A1</emphasis>
. Tutorium 72– 80 long, of which cusp 17–21. Custodium pointed distally, 3–5 long. Tridactylous.
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FF81FF87FF42C852FA89FBDC" lastPageId="36" lastPageNumber="187" pageId="31" pageNumber="182" type="description">
<paragraph id="1514746EFF81FFBCFF42C852FC69FBBB" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42C852FEC5FB97" bold="true" box="[199,345,1070,1095]" pageId="31" pageNumber="182">Description.</emphasis>
Adults.
<emphasis id="27DFA87CFF81FFBCFE32C853FDD9FB96" box="[439,581,1071,1094]" italics="true" pageId="31" pageNumber="182">Dimensions:</emphasis>
Total length: females (
<emphasis id="27DFA87CFF81FFBCFCC2C853FCC9FB96" box="[839,853,1071,1094]" italics="true" pageId="31" pageNumber="182">n</emphasis>
= 2) 370, 400; males (
<emphasis id="27DFA87CFF81FFBCFBD6C853FBFDFB96" box="[1107,1121,1071,1094]" italics="true" pageId="31" pageNumber="182">n</emphasis>
= 10) 359 (range 350–374). Notogastral width: females (
<emphasis id="27DFA87CFF81FFBCFE53C828FE78FBBB" box="[470,484,1108,1131]" italics="true" pageId="31" pageNumber="182">n</emphasis>
= 1) 240; males (
<emphasis id="27DFA87CFF81FFBCFD2EC828FD25FBBB" box="[683,697,1108,1131]" italics="true" pageId="31" pageNumber="182">n</emphasis>
= 10) 230 (range 221–240).
</paragraph>
<paragraph id="1514746EFF81FFBCFF42C80BFAAFFB63" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42C80BFED2FB5E" box="[199,334,1143,1166]" italics="true" pageId="31" pageNumber="182">Integument:</emphasis>
Smooth; longitudinal shallow grooves ventrally extending from level of custodium to posterior of anal plate (
<figureCitation id="8D9068EBFF81FFBCFE97C8E6FEE5FB63" box="[274,377,1178,1203]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">Fig. 21B</figureCitation>
). Pteromorph, pedotectum I,
<taxonomicName id="D2AB0FEDFF81FFBCFD45C8E6FCD6FB63" box="[704,842,1178,1203]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="31" pageNumber="190" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
, epimeres microtuberculate (
<figureCitation id="8D9068EBFF81FFBCFB0BC8E6FABBFB63" box="[1166,1319,1178,1203]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">Figs. 21 C–E</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF81FFBCFF42C8C2FBE8FAB7" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42C8C2FED1FB07" box="[199,333,1214,1239]" italics="true" pageId="31" pageNumber="182">Prodorsum:</emphasis>
Seta
<emphasis id="27DFA87CFF81FFBCFE15C8C3FE3AFB06" box="[400,422,1215,1238]" italics="true" pageId="31" pageNumber="182">ro</emphasis>
about 25 long, strongly barbed along length, acuminate, curved anteromedially; mutual distance about 50 (
<figureCitation id="8D9068EBFF81FFBCFEEFC89EFE4DFB2B" box="[362,465,1250,1275]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">Fig. 21C</figureCitation>
). Lamella about 104 long, medially with V-shaped indentation; about 3 long. Lamellar seta arising anteriorly on fused lamella, 10–17 long; mutual distance 26–28 (
<figureCitation id="8D9068EBFF81FFBCFC69C97AFBD5FACF" box="[1004,1097,1286,1311]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="31" pageNumber="182">Figs. 20</figureCitation>
,
<figureCitation id="8D9068EBFF81FFBCFBD1C97BFB1BFACF" box="[1108,1159,1287,1311]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">21C</figureCitation>
). Seta
<emphasis id="27DFA87CFF81FFBCFB55C97BFB7AFACE" box="[1232,1254,1287,1310]" italics="true" pageId="31" pageNumber="182">in</emphasis>
7–11 long, thin, smooth, acuminate; mutual distance 53–56 (
<figureCitation id="8D9068EBFF81FFBCFD18C956FD63FA93" box="[669,767,1322,1347]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="31" pageNumber="182">Figs. 20</figureCitation>
,
<figureCitation id="8D9068EBFF81FFBCFC8AC957FCD8FA93" box="[783,836,1323,1347]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="31" pageNumber="182">22A</figureCitation>
). Bothridial seta fusiform, 77–90 long. Seta
<emphasis id="27DFA87CFF81FFBCFAD1C950FAF0FA93" box="[1364,1388,1324,1347]" italics="true" pageId="31" pageNumber="182">ex</emphasis>
not evident, alveolus present. Dorsophragmata fused, elongated, length about 22 (
<figureCitation id="8D9068EBFF81FFBCFC7AC932FBF4FAB7" box="[1023,1128,1358,1383]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="31" pageNumber="182">Fig. 22A</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF81FFBCFF42C90EFCA9FA03" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42C90EFD8FFA5B" box="[199,531,1394,1419]" italics="true" pageId="31" pageNumber="182">Lateral region of podosoma:</emphasis>
Taenidium extending from rostrum to acetabulum I (
<figureCitation id="8D9068EBFF81FFBCFB01C90EFB71FA5B" box="[1156,1261,1394,1419]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">Fig. 21E</figureCitation>
). Pedotectum I pointed distally. Tutorium narrow tectum, about 74 long, with tooth distally, about 3 long, covering insertion of seta
<emphasis id="27DFA87CFF81FFBCFF12C9C0FF32FA03" box="[151,174,1468,1491]" italics="true" pageId="31" pageNumber="182">ro</emphasis>
. Custodium tapered to point distally 3–5 long (
<figureCitation id="8D9068EBFF81FFBCFD44C9C6FCB4FA03" box="[705,808,1466,1491]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="31" pageNumber="182">Fig. 22C</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF81FFBCFF42C9A3FE55F97B" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42C9A3FED0FA26" box="[199,332,1503,1526]" italics="true" pageId="31" pageNumber="182">Notogaster:</emphasis>
Slightly longer than wide: 1.2:1; anterior margin straight. Lenticulus absent. Muscle sigillae clearly evident on anterior of notogaster with incident or transmitted illumination (
<figureCitation id="8D9068EBFF81FFBCFBD0CA7EFB5EF9CA" box="[1109,1218,1538,1563]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="31" pageNumber="182">Fig. 22D</figureCitation>
). Octotaxic system expressed as porose areas,
<emphasis id="27DFA87CFF81FFBCFE4ACA5AFE72F9ED" box="[463,494,1574,1597]" italics="true" pageId="31" pageNumber="182">Aa</emphasis>
about
<quantity id="D253D98BFF81FFBCFDC4CA5AFDE7F9EE" box="[577,635,1573,1598]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="31" pageNumber="182" unit="in" value="11.0">11 in</quantity>
diameter,
<emphasis id="27DFA87CFF81FFBCFD72CA5AFCDEF9EE" box="[759,834,1574,1598]" italics="true" pageId="31" pageNumber="182">A1–A3</emphasis>
about 9 at widest; porose area
<emphasis id="27DFA87CFF81FFBCFB36CA5AFB4EF9EE" box="[1203,1234,1574,1598]" italics="true" pageId="31" pageNumber="182">A3</emphasis>
variable, may be absent bilaterally (
<specimenCount id="03ADBFE7FF81FFBCFEEBCA35FE6CF9B2" box="[366,496,1609,1634]" pageId="31" pageNumber="182" type="generic">1 specimen</specimenCount>
) or unilaterally (
<specimenCount id="03ADBFE7FF81FFBCFD33CA35FCABF9B2" box="[694,823,1609,1634]" pageId="31" pageNumber="182" type="generic">1 specimen</specimenCount>
) (
<figureCitation id="8D9068EBFF81FFBCFCD5CA35FC3BF9B1" box="[848,935,1609,1634]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="31" pageNumber="182">Fig. 20</figureCitation>
). Notogastral setae smooth, 6–8 long, setae subequal in length;
<emphasis id="27DFA87CFF81FFBCFEF7CA10FE12F955" box="[370,398,1644,1669]" italics="true" pageId="31" pageNumber="182">lm</emphasis>
posteromedial of
<emphasis id="27DFA87CFF81FFBCFDDDCA11FDE4F954" box="[600,632,1645,1668]" italics="true" pageId="31" pageNumber="182">Aa</emphasis>
and
<emphasis id="27DFA87CFF81FFBCFD28CA10FD5FF955" box="[685,707,1644,1669]" italics="true" pageId="31" pageNumber="182">lp</emphasis>
dorsocentrally positioned, medial of
<emphasis id="27DFA87CFF81FFBCFBE1CA11FB1FF955" box="[1124,1155,1645,1669]" italics="true" pageId="31" pageNumber="182">A1</emphasis>
; mutual distance of
<emphasis id="27DFA87CFF81FFBCFAE2CA10FA00F959" box="[1383,1436,1644,1673]" italics="true" pageId="31" pageNumber="182">h1-h1</emphasis>
about 90, of
<emphasis id="27DFA87CFF81FFBCFEA1CAE8FEC5F97F" box="[292,345,1684,1711]" italics="true" pageId="31" pageNumber="182">p1-p1</emphasis>
about 59.
</paragraph>
<paragraph id="1514746EFF81FFBCFF42CAC5FE26F8EF" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42CAC5FEE7F901" box="[199,379,1721,1746]" italics="true" pageId="31" pageNumber="182">Ventral Region:</emphasis>
Epimeral setae mostly about 7–18 long, thin, smooth, acuminate, with
<emphasis id="27DFA87CFF81FFBCFB41CAC5FB43F901" box="[1220,1247,1721,1745]" italics="true" pageId="31" pageNumber="182">1c</emphasis>
slightly barbed. Aggenital, anal and adanal setae smooth; genital setae about 14, others about 9. Genital setae six pairs,
<emphasis id="27DFA87CFF81FFBCFAA7CAA2FACAF929" box="[1314,1366,1758,1785]" italics="true" pageId="31" pageNumber="182">g1-g3</emphasis>
about 14 long, posterior 3 pairs about 9 long. Lyrifissure
<emphasis id="27DFA87CFF81FFBCFD69CB7EFC8CF8CB" box="[748,784,1794,1819]" italics="true" pageId="31" pageNumber="182">iad</emphasis>
at anterolateral edge of anal plate. Postanal porose area about 33 long and 4 wide.
</paragraph>
<paragraph id="1514746EFF81FFBCFF42CB35FB2BF8B1" blockId="31.[151,1437,710,2032]" box="[199,1207,1865,1890]" pageId="31" pageNumber="182">
<taxonomicName id="D2AB0FEDFF81FFBCFF42CB35FE34F8B2" authority=": Small" authorityName="Small" box="[199,424,1865,1890]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="31" pageNumber="182" phylum="Nematoda" rank="genus">
<emphasis id="27DFA87CFF81FFBCFF42CB35FEC3F8B2" box="[199,351,1865,1890]" italics="true" pageId="31" pageNumber="182">Gnathosoma:</emphasis>
Small
</taxonomicName>
axillary saccule present at base of palp,
<date id="611552AEFF81FFBCFCE8CB36FC0CF8B1" box="[877,912,1866,1890]" pageId="31" pageNumber="182">4 x</date>
2. Length of seta
<emphasis id="27DFA87CFF81FFBCFBDFCB35FB2CF8B2" box="[1114,1200,1865,1890]" italics="true" pageId="31" pageNumber="182">m&gt;a&gt;h</emphasis>
.
</paragraph>
<paragraph id="1514746EFF81FFBCFF42CB12FE47F879" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42CB12FE9AF855" box="[199,262,1902,1925]" italics="true" pageId="31" pageNumber="182">Legs:</emphasis>
Setation as for
<taxonomicName id="D2AB0FEDFF81FFBCFE3BCB11FD52F855" authorityName="Hammer. Setae (Roman" authorityYear="1958" box="[446,718,1901,1926]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="182" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFF81FFBCFE3BCB11FD52F855" box="[446,718,1901,1926]" italics="true" pageId="31" pageNumber="182">Tegoribates americanus</emphasis>
</taxonomicName>
(
<figureCitation id="8D9068EBFF81FFBCFD5ACB11FCE8F856" box="[735,884,1901,1926]" captionStart="FIGURE 12" captionStartId="22.[151,250,1970,1992]" captionTargetBox="[236,1371,232,1949]" captionTargetId="figure@22.[204,1382,193,1949]" captionTargetPageId="22" captionText="FIGURE 12 A – C. Tegoribates americanus Hammer, adult female, legs, all antiaxial aspect (trochanter not shown in A, B): A, leg I, with arrow to bend in femur; B, leg II (femur to tibia), C, leg III. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018655/files/figure.png" pageId="31" pageNumber="182">Figs. 12A–C</figureCitation>
,
<tableCitation id="582941D5FF81FFBCFC07CB11FC44F856" box="[898,984,1901,1926]" captionStart="TABLE 3" captionStartId="27.[151,239,694,717]" captionText="TABLE 3. Setiform organs in immature and adult Tegoribates americanus Hammer. Setae (Roman) and solenidia (Greek) are indicated; hyphen indicates no addition; parentheses indicate pseudosymmetrical pair of setae; brackets indicate that this seta is lost in the adult. Note: setation is cumulative, i. e., it indicates when the structure was added." pageId="31" pageNumber="182">Table 3</tableCitation>
). Seta
<emphasis id="27DFA87CFF81FFBCFBAECB11FBDEF856" box="[1067,1090,1901,1926]" italics="true" pageId="31" pageNumber="182">l”</emphasis>
of genu II strongly spine-like (
<figureCitation id="8D9068EBFF81FFBCFF1ACBECFEAFF879" box="[159,307,1936,1961]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="31" pageNumber="182">Figs. 21A, C</figureCitation>
). Tridactylous.
</paragraph>
<paragraph id="1514746EFF81FFBCFF42CBC9FBFDF820" blockId="31.[151,1437,710,2032]" pageId="31" pageNumber="182">
<emphasis id="27DFA87CFF81FFBCFF42CBC9FE78F81C" box="[199,484,1973,1996]" italics="true" pageId="31" pageNumber="182">Immatures. Dimensions:</emphasis>
Mean length (based on slide mounted specimens): protonymph (n=1) 260; deutonymph (
<emphasis id="27DFA87CFF81FFBCFEB6CBA4FEDDF83F" box="[307,321,2008,2031]" italics="true" pageId="31" pageNumber="182">n</emphasis>
= 1) 360. Length of prodorsal and hysterosomal setae given in
<tableCitation id="582941D5FF81FFBCFB8DCBABFBC7F820" box="[1032,1115,2007,2032]" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionText="TABLE 1. Prodorsal and hysterosomal setal lengths (in micrometers) of known immatures of Protectoribates occcidentalis sp. nov., Tegoribates americanus Hammer, and Tegoribates walteri sp. nov. NA indicates data not available, empty cell indicates character not present." pageId="31" pageNumber="182">Table 1</tableCitation>
.
</paragraph>
<caption id="41D424E6FFBEFF83FF12CBC4FB76F81E" box="[151,1258,1976,1998]" httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="32" pageNumber="183" targetBox="[244,1353,224,1923]" targetPageId="32">
<paragraph id="1514746EFFBEFF83FF12CBC4FB76F81E" blockId="32.[151,1258,1976,1998]" box="[151,1258,1976,1998]" pageId="32" pageNumber="183">
<emphasis id="27DFA87CFFBEFF83FF12CBC4FE83F81E" bold="true" box="[151,287,1976,1998]" pageId="32" pageNumber="183">FIGURE 20.</emphasis>
<taxonomicName id="D2AB0FEDFFBEFF83FEA0CBC4FE77F81E" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[293,491,1976,1998]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="183" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBEFF83FEA0CBC4FE77F81E" box="[293,491,1976,1998]" italics="true" pageId="32" pageNumber="183">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBEFF83FE75CBC4FDDDF81E" box="[496,577,1976,1998]" pageId="32" pageNumber="183" rank="species">
<emphasis id="27DFA87CFFBEFF83FE75CBC4FDDDF81E" bold="true" box="[496,577,1976,1998]" pageId="32" pageNumber="183">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm.
</paragraph>
</caption>
<caption id="41D424E6FFBFFF82FF12CB22FB43F800" httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="33" pageNumber="184" targetBox="[225,1362,193,1858]" targetPageId="33">
<paragraph id="1514746EFFBFFF82FF12CB22FB43F800" blockId="33.[151,1436,1886,2000]" pageId="33" pageNumber="184">
<emphasis id="27DFA87CFFBFFF82FF12CB22FEBDF8A4" bold="true" box="[151,289,1886,1908]" pageId="33" pageNumber="184">FIGURE 21.</emphasis>
<taxonomicName id="D2AB0FEDFFBFFF82FEACCB22FE73F8A4" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[297,495,1886,1908]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="184" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBFFF82FEACCB22FE73F8A4" box="[297,495,1886,1908]" italics="true" pageId="33" pageNumber="184">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBFFF82FE72CB22FDD5F8A4" box="[503,585,1886,1908]" pageId="33" pageNumber="184" rank="species">
<emphasis id="27DFA87CFFBFFF82FE72CB22FDD5F8A4" bold="true" box="[503,585,1886,1908]" pageId="33" pageNumber="184">sp. nov.</emphasis>
</taxonomicNameLabel>
, scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta
<emphasis id="27DFA87CFFBFFF82FD88CB01FDBEF843" box="[525,546,1917,1939]" italics="true" pageId="33" pageNumber="184">l”</emphasis>
of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae
<emphasis id="27DFA87CFFBFFF82FE21CBE0FE26F862" box="[420,442,1948,1970]" italics="true" pageId="33" pageNumber="184">l”</emphasis>
of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta
<emphasis id="27DFA87CFFBFFF82FF41CBC6FF46F800" box="[196,218,1978,2000]" italics="true" pageId="33" pageNumber="184">l”</emphasis>
of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm.
</paragraph>
</caption>
<caption id="41D424E6FFBCFF81FF12CB5FFEB6F821" httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="34" pageNumber="185" targetBox="[222,1338,193,1774]" targetPageId="34">
<paragraph id="1514746EFFBCFF81FF12CB5FFEB6F821" blockId="34.[151,1437,1827,2033]" pageId="34" pageNumber="185">
<emphasis id="27DFA87CFFBCFF81FF12CB5FFEBEF8E9" bold="true" box="[151,290,1827,1849]" pageId="34" pageNumber="185">FIGURE 22.</emphasis>
Differential interference contrast microscope images of adults; A–D,
<taxonomicName id="D2AB0FEDFFBCFF81FB85CB5FFB54F8E9" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[1024,1224,1827,1849]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBCFF81FB85CB5FFB54F8E9" box="[1024,1224,1827,1849]" italics="true" pageId="34" pageNumber="185">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBCFF81FB54CB5FFAB8F8E9" box="[1233,1316,1827,1849]" pageId="34" pageNumber="185" rank="species">
<emphasis id="27DFA87CFFBCFF81FB54CB5FFAB8F8E9" bold="true" box="[1233,1316,1827,1849]" pageId="34" pageNumber="185">sp. nov.</emphasis>
</taxonomicNameLabel>
, A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area
<emphasis id="27DFA87CFFBCFF81FA05CB1DFA01F8A6" box="[1408,1437,1889,1910]" italics="true" pageId="34" pageNumber="185">Aa</emphasis>
and surrounding muscle sigillae (5 layers). E,
<taxonomicName id="D2AB0FEDFFBCFF81FDE3CB03FC1AF845" authority="Ewing" authorityName="Ewing" authorityYear="1917" box="[614,902,1919,1941]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="subniger">
<emphasis id="27DFA87CFFBCFF81FDE3CB03FCA7F845" box="[614,827,1919,1941]" italics="true" pageId="34" pageNumber="185">Tegoribates subniger</emphasis>
Ewing
</taxonomicName>
, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G,
<taxonomicName id="D2AB0FEDFFBCFF81FE4DCBE1FD74F863" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[456,744,1949,1971]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="27DFA87CFFBCFF81FE4DCBE1FD74F863" box="[456,744,1949,1971]" italics="true" pageId="34" pageNumber="185">Protectoribates occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBCFF81FD74CBE1FCD9F863" box="[753,837,1949,1971]" pageId="34" pageNumber="185" rank="species">
<emphasis id="27DFA87CFFBCFF81FD74CBE1FCD9F863" bold="true" box="[753,837,1949,1971]" pageId="34" pageNumber="185">sp. nov.</emphasis>
</taxonomicNameLabel>
, detail of preanal organ; H,
<taxonomicName id="D2AB0FEDFFBCFF81FBF1CBE3FA04F863" authority="Aoki" authorityName="Aoki" authorityYear="1964" box="[1140,1432,1949,1972]" class="Arachnida" family="Austrachipteriidae" genus="Hypozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="laysanensis">
<emphasis id="27DFA87CFFBCFF81FBF1CBE3FAC6F863" box="[1140,1370,1949,1972]" italics="true" pageId="34" pageNumber="185">Hypozetes laysanensis</emphasis>
Aoki
</taxonomicName>
, detail of preanal organ; I,
<taxonomicName id="D2AB0FEDFFBCFF81FE1BCBC0FCE7F802" authority="Behan-Pelletier &amp; Walter" authorityName="Behan-Pelletier &amp; Walter" authorityYear="2013" box="[414,891,1980,2002]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="27DFA87CFFBCFF81FE1BCBC0FDEEF802" box="[414,626,1980,2002]" italics="true" pageId="34" pageNumber="185">Tectoribates borealis</emphasis>
Behan-Pelletier &amp; Walter
</taxonomicName>
, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm.
</paragraph>
</caption>
<caption id="41D424E6FFBDFF80FF12CBFDFA93F804" httpUri="https://zenodo.org/record/1018679/files/figure.png" pageId="35" pageNumber="186" targetBox="[210,1367,193,1875]" targetPageId="35">
<paragraph id="1514746EFFBDFF80FF12CBFDFA93F804" blockId="35.[151,1436,1921,2004]" pageId="35" pageNumber="186">
<emphasis id="27DFA87CFFBDFF80FF12CBFDFE83F847" bold="true" box="[151,287,1921,1943]" pageId="35" pageNumber="186">FIGURE 23.</emphasis>
<taxonomicName id="D2AB0FEDFFBDFF80FEA1CBFDFE74F847" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[292,488,1921,1943]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBDFF80FEA1CBFDFE74F847" box="[292,488,1921,1943]" italics="true" pageId="35" pageNumber="186">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBDFF80FE6BCBFDFDA2F847" box="[494,574,1921,1943]" pageId="35" pageNumber="186" rank="species">
<emphasis id="27DFA87CFFBDFF80FE6BCBFDFDA2F847" bold="true" box="[494,574,1921,1943]" pageId="35" pageNumber="186">sp. nov.</emphasis>
</taxonomicNameLabel>
, differential interference contrast microscope images of immatures; A, larval scalp (3 layers); B, protonymph (3 layers), dorsal aspect, larval scalp removed; C, larval and protonymphal scalp, with arrow to tubercle in region of setae
<emphasis id="27DFA87CFFBDFF80FEC9CBC2FEF9F804" box="[332,357,1982,2004]" italics="true" pageId="35" pageNumber="186">dp</emphasis>
; D, deutonymph, dorsal aspect, with scalps removed. Scale bars: A–C = 20 µm; D = 40 µm.
</paragraph>
</caption>
<paragraph id="1514746EFFBAFF87FF42CCEBFCE9FF04" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CCEBFE8DFF60" bold="true" box="[199,273,151,176]" pageId="36" pageNumber="187">Larva</emphasis>
(
<figureCitation id="8D9068EBFFBAFF87FEA4CCEBFE14FF60" box="[289,392,151,176]" captionStart="FIGURE 23" captionStartId="35.[151,250,1921,1943]" captionTargetBox="[210,1367,193,1875]" captionTargetId="figure@35.[210,1367,193,1894]" captionTargetPageId="35" captionText="FIGURE 23. Tegoribates walteri sp. nov., differential interference contrast microscope images of immatures; A, larval scalp (3 layers); B, protonymph (3 layers), dorsal aspect, larval scalp removed; C, larval and protonymphal scalp, with arrow to tubercle in region of setae dp; D, deutonymph, dorsal aspect, with scalps removed. Scale bars: A – C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/1018679/files/figure.png" pageId="36" pageNumber="187">Fig. 23A</figureCitation>
). Only scalp available. Length scalp 159 (
<emphasis id="27DFA87CFFBAFF87FCE9CCE5FCE6FF60" box="[876,890,153,176]" italics="true" pageId="36" pageNumber="187">n</emphasis>
=1). Setae
<emphasis id="27DFA87CFFBAFF87FC73CCE5FB9EFF60" box="[1014,1026,153,176]" italics="true" pageId="36" pageNumber="187">c</emphasis>
,
<emphasis id="27DFA87CFFBAFF87FB97CCEBFB17FF60" box="[1042,1163,151,176]" italics="true" pageId="36" pageNumber="187">da, dm, dp</emphasis>
small, smooth, tapered; mutual distance
<emphasis id="27DFA87CFFBAFF87FECBCCC7FDA7FF04" box="[334,571,187,212]" italics="true" pageId="36" pageNumber="187">da-da, dm-dm, dp-dp</emphasis>
12, 16 and 10, respectively.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CCA2FA8FFECB" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CCA2FEC1FF27" bold="true" box="[199,349,222,247]" pageId="36" pageNumber="187">Protonymph</emphasis>
(
<figureCitation id="8D9068EBFFBAFF87FEF1CCA2FE7FFF27" box="[372,483,222,247]" captionStart="FIGURE 23" captionStartId="35.[151,250,1921,1943]" captionTargetBox="[210,1367,193,1875]" captionTargetId="figure@35.[210,1367,193,1894]" captionTargetPageId="35" captionText="FIGURE 23. Tegoribates walteri sp. nov., differential interference contrast microscope images of immatures; A, larval scalp (3 layers); B, protonymph (3 layers), dorsal aspect, larval scalp removed; C, larval and protonymphal scalp, with arrow to tubercle in region of setae dp; D, deutonymph, dorsal aspect, with scalps removed. Scale bars: A – C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/1018679/files/figure.png" pageId="36" pageNumber="187">Fig. 23B</figureCitation>
).
<emphasis id="27DFA87CFFBAFF87FE7BCC9CFDE0FF27" box="[510,636,224,247]" italics="true" pageId="36" pageNumber="187">Integument</emphasis>
: Sclerotized with light brown coloration. All of body, other than centrodorsal region of hysterosoma covered by layer of cerotegument as globules, about
<quantity id="D253D98BFFBAFF87FBFCCD7FFB38FECB" box="[1145,1188,258,283]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="187" unit="in" value="1.0">1 in</quantity>
diameter.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CD59FEB3FE55" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CD59FED1FEEE" box="[199,333,293,318]" italics="true" pageId="36" pageNumber="187">Prodorsum:</emphasis>
Porose region on aspis not evident. Seta
<emphasis id="27DFA87CFFBAFF87FCA4CD5BFCA5FEEE" box="[801,825,295,318]" italics="true" pageId="36" pageNumber="187">ro</emphasis>
finely barbed along length, tapered. Seta
<emphasis id="27DFA87CFFBAFF87FA97CD59FAD7FEEE" box="[1298,1355,293,318]" italics="true" pageId="36" pageNumber="187">le, in</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FA06CD5BFA00FEEE" box="[1411,1436,295,318]" italics="true" pageId="36" pageNumber="187">ex</emphasis>
smooth, isodiametric; mutual distance of setae
<emphasis id="27DFA87CFFBAFF87FD2ACD36FCB6FEB2" box="[687,810,329,354]" italics="true" pageId="36" pageNumber="187">ro-ro, le-le</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FCE1CD36FC05FEB1" box="[868,921,330,353]" italics="true" pageId="36" pageNumber="187">in-in</emphasis>
about 19, 20 and 34, respectively. Bothridial seta setiform.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CDEDFD64FDC9" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CDEDFE3FFE78" box="[199,419,401,424]" italics="true" pageId="36" pageNumber="187">Gastronotic Region</emphasis>
: Integument without sclerotization; centrodorsal region normally covered by larval scalp, smooth; integument slightly plicate laterally. Gastronotal setae of
<emphasis id="27DFA87CFFBAFF87FC15CDC9FC46FE1C" box="[912,986,435,460]" italics="true" pageId="36" pageNumber="187">c, d, p</emphasis>
series and
<emphasis id="27DFA87CFFBAFF87FBE5CDCFFB6CFE00" box="[1120,1264,435,464]" italics="true" pageId="36" pageNumber="187">la, lm, h2, h1</emphasis>
short, smooth, isodiametric;
<emphasis id="27DFA87CFFBAFF87FEB4CDA6FEDBFE23" box="[305,327,474,499]" italics="true" pageId="36" pageNumber="187">lp</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FE04CDA6FE0AFE26" box="[385,406,474,502]" italics="true" pageId="36" pageNumber="187">h3</emphasis>
long, setiform, embedded in thick covering of cerotegument. Mutual distance of setae
<emphasis id="27DFA87CFFBAFF87FAF3CDA6FE93FDC9" italics="true" pageId="36" pageNumber="187">dada, dm-dm</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FEC1CE7CFE1AFDC9" box="[324,390,512,537]" italics="true" pageId="36" pageNumber="187">dp-dp</emphasis>
about 12, 21 and 9, respectively.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CE5FFC45FD53" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CE5FFEE8FDEC" box="[199,372,547,572]" italics="true" pageId="36" pageNumber="187">Ventral Region</emphasis>
: Medial margin of epimeral plates weakly defined. Epimere I with narrow mentotectum overlapping medially. Epimeral, genital, aggenital, anal and adanal setae smooth, isodiametric, about 9 long. Porose integument surrounding opening of opisthonotal gland not evident.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CEF2FE44FD1A" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<taxonomicName id="D2AB0FEDFFBAFF87FF42CEF2FE5BFD77" authority=": Axillary" authorityName="Axillary" box="[199,455,654,679]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="36" pageNumber="187" phylum="Nematoda" rank="genus">
<emphasis id="27DFA87CFFBAFF87FF42CEF2FECAFD77" box="[199,342,654,679]" italics="true" pageId="36" pageNumber="187">Gnathosoma</emphasis>
: Axillary
</taxonomicName>
saccule present. Palpal eupathidium
<emphasis id="27DFA87CFFBAFF87FC05CEF3FC32FD76" box="[896,942,655,678]" italics="true" pageId="36" pageNumber="187">acm</emphasis>
attached to solenidion along distal threequarters of solenidial length.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CEAAFDF4FCC1" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CEAAFF60FD3D" box="[199,252,726,749]" italics="true" pageId="36" pageNumber="187">Legs</emphasis>
: Genua and tibiae I-IV with seta
<emphasis id="27DFA87CFFBAFF87FDF1CEA9FD1EFD3E" box="[628,642,725,750]" italics="true" pageId="36" pageNumber="187">d</emphasis>
subequal in length to companion solenidion; solenidion on genua I-III and tibiae II, III with slightly bulbous tip.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CF60FB53FC72" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42CF60FEFFFCE5" bold="true" box="[199,355,796,821]" pageId="36" pageNumber="187">Deutonymph</emphasis>
(
<figureCitation id="8D9068EBFFBAFF87FEF1CF60FD8BFCE4" box="[372,535,796,821]" captionStart="FIGURE 23" captionStartId="35.[151,250,1921,1943]" captionTargetBox="[210,1367,193,1875]" captionTargetId="figure@35.[210,1367,193,1894]" captionTargetPageId="35" captionText="FIGURE 23. Tegoribates walteri sp. nov., differential interference contrast microscope images of immatures; A, larval scalp (3 layers); B, protonymph (3 layers), dorsal aspect, larval scalp removed; C, larval and protonymphal scalp, with arrow to tubercle in region of setae dp; D, deutonymph, dorsal aspect, with scalps removed. Scale bars: A – C = 20 µm; D = 40 µm." httpUri="https://zenodo.org/record/1018679/files/figure.png" pageId="36" pageNumber="187">Figs. 23 C, D</figureCitation>
). As for protonymph, except for overall size. Dorsal of hysterosoma with few plicae. Length of combined larval and protonymphal scalps about 220. Mutual distance of
<emphasis id="27DFA87CFFBAFF87FB35CF3CFAB7FC88" box="[1200,1323,831,856]" italics="true" pageId="36" pageNumber="187">ro-ro, le-le</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FAE2CF3CFA00FC87" box="[1383,1436,832,855]" italics="true" pageId="36" pageNumber="187">in-in</emphasis>
about 20, 17, and 42, respectively. Gastronotal setae
<emphasis id="27DFA87CFFBAFF87FD63CF18FC09FCAF" box="[742,917,867,895]" italics="true" pageId="36" pageNumber="187">c, d, l, p, and h1</emphasis>
short, isodiametric; setae
<emphasis id="27DFA87CFFBAFF87FB3FCF1FFB4CFCAF" box="[1210,1232,867,895]" italics="true" pageId="36" pageNumber="187">h3</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FA83CF1FFA80FCAF" box="[1286,1308,867,895]" italics="true" pageId="36" pageNumber="187">h2</emphasis>
brush-like, broad, long; mutual distance of setae
<emphasis id="27DFA87CFFBAFF87FDBFCFF5FD4BFC72" box="[570,727,905,930]" italics="true" pageId="36" pageNumber="187">da-da, dm-dm</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FC88CFF5FCD2FC72" box="[781,846,905,930]" italics="true" pageId="36" pageNumber="187">dp-dp</emphasis>
about 12, 25 and 10, respectively.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42CFD0FCC2FC39" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">Epimeral setae (protonymph and deutonymph): 3-1-2-1, 3-1-2-2. Genital, aggenital, anal and adanal setae (protonymph and deutonymph): 1-3, 1-1, 0-0, 0-2, respectively.</paragraph>
<paragraph id="1514746EFFBAFF87FF42CF88FA89FBDC" blockId="36.[151,1437,151,1426]" box="[199,1301,1011,1036]" pageId="36" pageNumber="187">
Leg setae and solenidia as for protonymph and deutonymph of
<taxonomicName id="D2AB0FEDFFBAFF87FC0FCF8FFB04FBDC" authorityName="Hammer. Setae (Roman" authorityYear="1958" box="[906,1176,1011,1036]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFFBAFF87FC0FCF8FFB04FBDC" box="[906,1176,1011,1036]" italics="true" pageId="36" pageNumber="187">Tegoribates americanus</emphasis>
</taxonomicName>
in
<tableCitation id="582941D5FFBAFF87FB3ECF8FFA8CFBDC" box="[1211,1296,1011,1036]" captionStart="TABLE 3" captionStartId="27.[151,239,694,717]" captionText="TABLE 3. Setiform organs in immature and adult Tegoribates americanus Hammer. Setae (Roman) and solenidia (Greek) are indicated; hyphen indicates no addition; parentheses indicate pseudosymmetrical pair of setae; brackets indicate that this seta is lost in the adult. Note: setation is cumulative, i. e., it indicates when the structure was added." pageId="36" pageNumber="187">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FFBAFF87FF42C86AFC5CFB83" pageId="36" pageNumber="187" type="etymology">
<paragraph id="1514746EFFBAFF87FF42C86AFC5CFB83" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42C86AFED2FBFF" bold="true" box="[199,334,1046,1071]" pageId="36" pageNumber="187">Etymology.</emphasis>
The specific epithet honours my colleague and friend Dr. David Evans Walter, who has contributed so significantly to acarology in North America and globally.
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FFBAFF87FF42C821FA87FA41" pageId="36" pageNumber="187" type="discussion">
<paragraph id="1514746EFFBAFF87FF42C821FCC8FB31" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
<emphasis id="27DFA87CFFBAFF87FF42C821FEA7FBA6" bold="true" box="[199,315,1117,1142]" pageId="36" pageNumber="187">Remarks.</emphasis>
Adults of
<taxonomicName id="D2AB0FEDFFBAFF87FE4DC821FD35FBA6" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[456,681,1117,1142]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBAFF87FE4DC821FD35FBA6" box="[456,681,1117,1142]" italics="true" pageId="36" pageNumber="187">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBAFF87FD3DC821FC85FBA6" box="[696,793,1117,1142]" pageId="36" pageNumber="187" rank="species">
<emphasis id="27DFA87CFFBAFF87FD3DC821FC85FBA6" bold="true" box="[696,793,1117,1142]" pageId="36" pageNumber="187">sp. nov.</emphasis>
</taxonomicNameLabel>
, are clearly distinguished from those of other North American species of
<taxonomicName id="D2AB0FEDFFBAFF87FE01C8FDFD9BFB4A" authorityName="Ewing" authorityYear="1917" box="[388,519,1153,1178]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFFBAFF87FE01C8FDFD9BFB4A" box="[388,519,1153,1178]" italics="true" pageId="36" pageNumber="187">Tegoribates</emphasis>
</taxonomicName>
by having the octotaxic system in the form of porose areas; porose areas are also found in
<taxonomicName id="D2AB0FEDFFBAFF87FF79C8DAFEE6FB6D" authorityName="Engelbrecht" authorityYear="1986" box="[252,378,1190,1213]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="27DFA87CFFBAFF87FF79C8DAFEE6FB6D" box="[252,378,1190,1213]" italics="true" pageId="36" pageNumber="187">T. montana</emphasis>
</taxonomicName>
and
<taxonomicName id="D2AB0FEDFFBAFF87FE34C8DAFDA3FB6E" authorityName="Engelbrecht" authorityYear="1986" box="[433,575,1189,1214]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="27DFA87CFFBAFF87FE34C8DAFDA3FB6E" box="[433,575,1189,1214]" italics="true" pageId="36" pageNumber="187">T. natalensis</emphasis>
</taxonomicName>
, known from
<collectingCountry id="6DBC34FEFFBAFF87FD5BC8D9FCF2FB6E" box="[734,878,1189,1214]" name="South Africa" pageId="36" pageNumber="187">South Africa</collectingCountry>
(
<bibRefCitation id="713A099FFFBAFF87FCF8C8D9FBD6FB6E" author="Engelbrecht, C. M." box="[893,1098,1189,1214]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="36" pageNumber="187" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht 1986</bibRefCitation>
). The 2 South African species are monodactylous, whereas
<taxonomicName id="D2AB0FEDFFBAFF87FE59C8B5FDDAFB31" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[476,582,1224,1249]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBAFF87FE59C8B5FDDAFB31" box="[476,582,1224,1249]" italics="true" pageId="36" pageNumber="187">T. walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBAFF87FDC8C8B4FD3BFB31" box="[589,679,1224,1249]" pageId="36" pageNumber="187" rank="species">
<emphasis id="27DFA87CFFBAFF87FDC8C8B4FD3BFB31" bold="true" box="[589,679,1224,1249]" pageId="36" pageNumber="187">sp. nov.</emphasis>
</taxonomicNameLabel>
is tridactylous.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42C890FD9FFABE" blockId="36.[151,1437,151,1426]" pageId="36" pageNumber="187">
Immatures of
<taxonomicName id="D2AB0FEDFFBAFF87FEE3C891FE4EFAD4" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[358,466,1259,1284]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBAFF87FEE3C891FE4EFAD4" box="[358,466,1259,1284]" italics="true" pageId="36" pageNumber="187">T. walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBAFF87FE5EC897FDAAFAD4" box="[475,566,1259,1284]" pageId="36" pageNumber="187" rank="species">
<emphasis id="27DFA87CFFBAFF87FE5EC897FDAAFAD4" bold="true" box="[475,566,1259,1284]" pageId="36" pageNumber="187">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="D2AB0FEDFFBAFF87FDEBC891FC92FAD4" authorityName="Hammer. Setae (Roman" authorityYear="1958" box="[622,782,1261,1284]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFFBAFF87FDEBC891FC92FAD4" box="[622,782,1261,1284]" italics="true" pageId="36" pageNumber="187">T. americanus</emphasis>
</taxonomicName>
are similar in the shape and length of setae
<emphasis id="27DFA87CFFBAFF87FA80C891FA8DFAD4" box="[1285,1297,1261,1284]" italics="true" pageId="36" pageNumber="187">c</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FACEC897FAC5FAD4" box="[1355,1369,1259,1284]" italics="true" pageId="36" pageNumber="187">d</emphasis>
, but
<emphasis id="27DFA87CFFBAFF87FA11C897FA00FAD4" box="[1428,1436,1259,1284]" italics="true" pageId="36" pageNumber="187">l</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FF43C973FF48FAF8" box="[198,212,1295,1320]" italics="true" pageId="36" pageNumber="187">h</emphasis>
series setae differ in shape and length (
<tableCitation id="582941D5FFBAFF87FD15C973FD78FAF7" box="[656,740,1295,1320]" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionText="TABLE 1. Prodorsal and hysterosomal setal lengths (in micrometers) of known immatures of Protectoribates occcidentalis sp. nov., Tegoribates americanus Hammer, and Tegoribates walteri sp. nov. NA indicates data not available, empty cell indicates character not present." pageId="36" pageNumber="187">Table 1</tableCitation>
). Comparisons with immatures of
<taxonomicName id="D2AB0FEDFFBAFF87FBE8C96CFB6AFAF8" authorityName="Koch. Jacot" authorityYear="1929" box="[1133,1270,1295,1320]" class="Arachnida" family="Galumnidae" genus="Zetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFFBAFF87FBE8C96CFB6AFAF8" box="[1133,1270,1295,1320]" italics="true" pageId="36" pageNumber="187">T. latirostris</emphasis>
</taxonomicName>
are limited, as only the illustrations of
<bibRefCitation id="713A099FFFBAFF87FE24C94FFDA4FA9B" author="Tuxen, S. L." box="[417,568,1331,1355]" journalOrPublisher="Entomologiske Meddelleilser" pageId="36" pageNumber="187" pagination="321 - 336" part="23" refString="Tuxen, S. L. (1943) Die zeitliche und raumliche Verteilung der Oribatiden-Fauna (Acar.) bei Maelifell, Nord-Island. Entomologiske Meddelleilser, 23, 321 - 336." title="Die zeitliche und raumliche Verteilung der Oribatiden-Fauna (Acar.) bei Maelifell, Nord-Island" type="journal article" year="1943">Tuxen (1943)</bibRefCitation>
are available. The larva he illustrated appears similar to that of
<taxonomicName id="D2AB0FEDFFBAFF87FB7BC948FA00FA9B" authorityName="Hammer. Setae (Roman" authorityYear="1958" box="[1278,1436,1332,1355]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFFBAFF87FB7BC948FA00FA9B" box="[1278,1436,1332,1355]" italics="true" pageId="36" pageNumber="187">T. americanus</emphasis>
</taxonomicName>
with long, thick setae
<emphasis id="27DFA87CFFBAFF87FE08C92AFE35FABF" box="[397,425,1366,1391]" italics="true" pageId="36" pageNumber="187">h3</emphasis>
and
<emphasis id="27DFA87CFFBAFF87FE65C92AFE60FABF" box="[480,508,1366,1391]" italics="true" pageId="36" pageNumber="187">h2</emphasis>
.
</paragraph>
<paragraph id="1514746EFFBAFF87FF42C906FA87FA41" blockId="36.[151,1437,151,1426]" box="[199,1307,1401,1426]" pageId="36" pageNumber="187">
Gut contents of immature and adult
<taxonomicName id="D2AB0FEDFFBAFF87FDDEC905FCA8FA42" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[603,820,1401,1426]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFFBAFF87FDDEC905FCA8FA42" box="[603,820,1401,1426]" italics="true" pageId="36" pageNumber="187">Tegoribates walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FFBAFF87FCB9C905FC09FA42" box="[828,917,1401,1426]" pageId="36" pageNumber="187" rank="species">
<emphasis id="27DFA87CFFBAFF87FCB9C905FC09FA42" bold="true" box="[828,917,1401,1426]" pageId="36" pageNumber="187">sp. nov.</emphasis>
</taxonomicNameLabel>
include fungal hyphae and spores.
</paragraph>
</subSubSection>
</treatment>
</document>
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