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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. &amp; Titarenko, Andrey Y." docDate="2014" docId="6AEE4401E25D512FBD635555311404E6" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus Kurnakov 1972, stat. n." docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="25" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="24" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
<mods:affiliation>Publicly Traded Company &quot; Morpho Absoloni &quot;, Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-12-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>463</mods:number>
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<mods:extent unit="page">
<mods:start>21</mods:start>
<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
<mods:identifier type="ZooBank">71C1AE92B0CE4D26B1E5076E480180AD</mods:identifier>
<mods:identifier type="Pensoft-UUID">3E5CFFC6FFC91628FFF6E355951DFFF8</mods:identifier>
<mods:identifier type="Zenodo-Dep">578654</mods:identifier>
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<treatment ID-GBIF-Taxon="182234817" LSID="urn:lsid:plazi:treatment:6AEE4401E25D512FBD635555311404E6" httpUri="http://treatment.plazi.org/id/6AEE4401E25D512FBD635555311404E6" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">
<subSubSection pageId="3" pageNumber="24" type="nomenclature">
<paragraph pageId="3" pageNumber="24">
<taxonomicName LSID="6AEE4401-E25D-512F-BD63-5555311404E6" authority="Kurnakov, 1972" authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="besleticus" status="stat. n." subGenus="Archiplectes">
<pageBreakToken pageId="3" pageNumber="24" start="start">Carabus</pageBreakToken>
(Archiplectes) besleticus Kurnakov, 1972
</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="24">stat. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="3" pageNumber="24">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="3" pageNumber="24">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="3" pageNumber="24">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="3" pageNumber="24">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="3" pageNumber="24">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus resheviensis (River Reshevie), Carabus (Archiplectes) pseudopshuensis (River Reshevie), and Carabus (Archiplectes) juenthneri juenthneri (Village Pskhu env.) constructed using discriminant analysis based on 8 parameters." pageId="3" pageNumber="24">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="3" pageNumber="24">, 7</figureCitation>
<figureCitation captionStart="Figures 16–21" captionStartId="F9" captionText="Figures 16 - 21. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus besleticus and Carabus (Archiplectes) besleticus mtsaranus. 16 - 18 Carabus (Archiplectes) besleticus besleticus 19 - 21 Carabus (Archiplectes) besleticus mtsaranus." pageId="3" pageNumber="24">, 16-21</figureCitation>
<figureCitation captionStart="Figures 22–27" captionStartId="F10" captionText="Figures 22 - 27. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus duripshensis and Carabus (Archiplectes) besleticus napraensis. 22 - 24 Carabus (Archiplectes) besleticus duripshensis 25 - 27 Carabus (Archiplectes) besleticus napraensis." pageId="3" pageNumber="24">, 22-27</figureCitation>
<figureCitation captionStart="Figures 28–33" captionStartId="F11" captionText="Figures 28 - 33. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus dsychvensis and Carabus (Archiplectes) besleticus resheviensis 28 - 30 Carabus (Archiplectes) besleticus dsychvensis 31 - 33 Carabus (Archiplectes) besleticus resheviensis." pageId="3" pageNumber="24">, 28-33</figureCitation>
<figureCitation captionStart="Figures 34–41" captionStartId="F12" captionText="Figures 34 - 41. Dorsal habitus, Carabus (Archiplectes) satyrus, Abkhazia, Village Merkheul env." pageId="3" pageNumber="24">, 34-41</figureCitation>
<figureCitation captionStart="Figures 42–49" captionStartId="F13" captionText="Figures 42 - 49. Dorsal habitus, Carabus (Archiplectes) besleticus besleticus, Abkhazia, Mt. Birtzkha." pageId="3" pageNumber="24">, 42-49</figureCitation>
<figureCitation captionStart="Figures 50–57" captionStartId="F14" captionText="Figures 50 - 57. Dorsal habitus, Carabus (Archiplectes) besleticus mtsaranus, Abkhazia, Zashirbara Mt. Range, valley of River Mtsara." pageId="3" pageNumber="24">, 50-57</figureCitation>
<figureCitation captionStart="Figures 58–61" captionStartId="F15" captionText="Figures 58 - 61. Dorsal habitus, Carabus (Archiplectes) besleticus duripshensis. 58 - 59 Abkhazia, S slope of Bsybian Mt. Range, valley of River Khipsta 60 - 61 Abkhazia, Bsybian Mt. Range, Duriph env. (after Retezar 2008)." pageId="3" pageNumber="24">, 58-61</figureCitation>
<figureCitation captionStart="Figures 62–65" captionStartId="F16" captionText="Figures 62 - 65. Dorsal habitus, Carabus (Archiplectes) besleticus napraensis. 62 - 64 Abkhazia, Bzybian Mt. Range, Mt. Chibzharga (paratypes) 65 same locality (paratype) (after Retezar 2008)." pageId="3" pageNumber="24">, 62-65</figureCitation>
<figureCitation captionStart="Figures 66–73" captionStartId="F17" captionText="Figures 66 - 73. Dorsal habitus, Carabus (Archiplectes) besleticus resheviensis. 66 Abkhazia, N slopes of Bzybian Mt. Range, right bank of River Reshevie (holotype) 67 - 73 same locality (paratypes)." pageId="3" pageNumber="24">, 66-73</figureCitation>
<figureCitation captionStart="Figures 74–77" captionStartId="F18" captionText="Figures 74 - 77. Dorsal habitus, Carabus (Archiplectes) besleticus dsychvensis, Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva." pageId="3" pageNumber="24">, 74-77</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="25" type="remarks">
<paragraph pageId="4" pageNumber="25">
<pageBreakToken pageId="4" pageNumber="25" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="25">
The structure of the endophallus testifies to the attribution of this form to the
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="reitteri" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis italics="true" pageId="4" pageNumber="25">reitteri</emphasis>
</taxonomicName>
group. Saccellus large, conical, smoothly rounded apically, with faintly developed or missing S-shaped fold on the right, paracellar lobe very faintly developed, endophallus with rather symmetric position of inner structures (when viewed frontally). Ostium lobe large, of dulled conical shape. Lamella of aedeagus elongated, with prominent hollow on the right before apex. Aggonoporius large, strongly sclerotized. This type of endophallus is characteristic for all subspecies described subsequently, earlier attributed to
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
. Based on such pronounced differences in the structure of the endophallus from that of
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
, it is proposed to raise the status of
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
from subspecies to species.
</paragraph>
<caption pageId="4" pageNumber="25" start="Figure 3" startId="F3">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 3.</emphasis>
Distribution of morphometric characters in males of
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
subspecies constructed using discriminant analysis based on 8 parameters.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figure 4" startId="F4">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 4.</emphasis>
Distribution of morphometric characters in females of
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
subspecies constructed using discriminant analysis based on 8 parameters.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figure 5" startId="F5">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 5.</emphasis>
Distribution of morphometric characters in males of
<taxonomicName authorityName="Solodovnikov, Zamotajlov &amp; Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
(River Reshevie),
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
(River Reshevie), and
<taxonomicName authority="juenthneri" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. juenthneri" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="juenthneri">Carabus (Archiplectes) juenthneri juenthneri</taxonomicName>
(Village Pskhu env.) constructed using discriminant analysis based on 8 parameters.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figure 6" startId="F6">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 6.</emphasis>
Distribution of morphometric characters in females of
<taxonomicName authorityName="Solodovnikov, Zamotajlov &amp; Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
(River Reshevie),
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
(River Reshevie), and
<taxonomicName authority="juenthneri" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. juenthneri" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="juenthneri">Carabus (Archiplectes) juenthneri juenthneri</taxonomicName>
(Village Pskhu env.) constructed using discriminant analysis based on 8 parameters.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figure 7" startId="F7">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 7.</emphasis>
Distribution map of
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
,
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
, and
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
in Abkhazia.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 8–15" startId="F8">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 8-15.</emphasis>
Fully inflated endophallus preparation and apical lamella of aedeagus of
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
and
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
.
<emphasis bold="true" pageId="4" pageNumber="25">8-11</emphasis>
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">12-15</emphasis>
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 16–21" startId="F9">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 16-21.</emphasis>
Fully inflated endophallus preparation and apical lamella of aedeagus of
<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus</taxonomicName>
and
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
.
<emphasis bold="true" pageId="4" pageNumber="25">16-18</emphasis>
<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">19-21</emphasis>
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 22–27" startId="F10">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 22-27.</emphasis>
Fully inflated endophallus preparation and apical lamella of aedeagus of
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. duripshensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) besleticus duripshensis</taxonomicName>
and
<taxonomicName authorityName="Belousov &amp; Zamotajlov" authorityYear="1993" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. napraensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="napraensis">Carabus (Archiplectes) besleticus napraensis</taxonomicName>
.
<emphasis bold="true" pageId="4" pageNumber="25">22-24</emphasis>
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. duripshensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) besleticus duripshensis</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">25-27</emphasis>
<taxonomicName authorityName="Belousov &amp; Zamotajlov" authorityYear="1993" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. napraensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="napraensis">Carabus (Archiplectes) besleticus napraensis</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 28–33" startId="F11">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 28-33.</emphasis>
Fully inflated endophallus preparation and apical lamella of aedeagus of
<taxonomicName authorityName="Gottwald" authorityYear="1985" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. dsychvensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) besleticus dsychvensis</taxonomicName>
and
<taxonomicName authorityName="Solodovnikov, Zamotajlov &amp; Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">28-30</emphasis>
<taxonomicName authorityName="Gottwald" authorityYear="1985" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. dsychvensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) besleticus dsychvensis</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">31-33</emphasis>
<taxonomicName authorityName="Solodovnikov, Zamotajlov &amp; Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 34–41" startId="F12">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 34-41.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
, Abkhazia, Village Merkheul env.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 42–49" startId="F13">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 42-49.</emphasis>
Dorsal habitus,
<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">Carabus (Archiplectes) besleticus besleticus</taxonomicName>
, Abkhazia, Mt. Birtzkha.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 50–57" startId="F14">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 50-57.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
, Abkhazia, Zashirbara Mt. Range, valley of River Mtsara.
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 58–61" startId="F15">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 58-61.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. duripshensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) besleticus duripshensis</taxonomicName>
.
<emphasis bold="true" pageId="4" pageNumber="25">58-59</emphasis>
Abkhazia, S slope of Bsybian Mt. Range, valley of River Khipsta
<emphasis bold="true" pageId="4" pageNumber="25">60-61</emphasis>
Abkhazia, Bsybian Mt. Range, Duriph env. (after
<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
2008
</bibRefCitation>
).
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 62–65" startId="F16">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 62-65.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="Belousov &amp; Zamotajlov" authorityYear="1993" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. napraensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="napraensis">Carabus (Archiplectes) besleticus napraensis</taxonomicName>
.
<emphasis bold="true" pageId="4" pageNumber="25">62-64</emphasis>
Abkhazia, Bzybian Mt. Range, Mt. Chibzharga (paratypes)
<emphasis bold="true" pageId="4" pageNumber="25">65</emphasis>
same locality (paratype) (after
<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
2008
</bibRefCitation>
).
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 66–73" startId="F17">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 66-73.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="Solodovnikov, Zamotajlov &amp; Fominykh" authorityYear="2014" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. resheviensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="resheviensis">Carabus (Archiplectes) besleticus resheviensis</taxonomicName>
.
<emphasis bold="true" pageId="4" pageNumber="25">66</emphasis>
Abkhazia, N slopes of Bzybian Mt. Range, right bank of River Reshevie (holotype)
<emphasis bold="true" pageId="4" pageNumber="25">67-73</emphasis>
same locality (paratypes).
</paragraph>
</caption>
<caption pageId="4" pageNumber="25" start="Figures 74–77" startId="F18">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figures 74-77.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="Gottwald" authorityYear="1985" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. dsychvensis" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) besleticus dsychvensis</taxonomicName>
, Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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