402 lines
55 KiB
XML
402 lines
55 KiB
XML
<document id="A243262D5C5C6F16AC7212E8AFC89455" ID-CLB-Dataset="305522" ID-DOI="10.5852/cr-palevol2023v22a22" ID-GBIF-Dataset="8a406885-3429-4c59-b846-09b346ee9f5b" ID-ISSN="1777-571X" ID-Zenodo-Dep="14224492" ID-ZooBank="urn:lsid:zoobank.org:pub:B2995358-3451-4662-9C7E-FDF4A51838A7" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1732634267618" checkinUser="felipe" docAuthor="Branger, Patrick" docDate="2023" docId="201087CBFFD8BE69FF55F960FE5AFA36" docLanguage="en" docName="CRPalevol.22.22.479-490.pdf" docOrigin="Comptes Rendus Palevol 22 (22)" docSource="http://dx.doi.org/10.5852/cr-palevol2023v22a22" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Micronautilus evolutus Branger, 2023, n. gen., n. sp." docType="treatment" docVersion="3" lastPageNumber="484" masterDocId="DC29FFB3FFDDBE6EFFA9FFF4FFA0FFF8" masterDocTitle="Micronautilus n. gen., a new dwarf Bathonian (Middle Jurassic) nautilid from Western France" masterLastPageNumber="490" masterPageNumber="479" pageNumber="482" updateTime="1732751156522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C297EFEE5571F120C54C607F70B0D968">Micronautilus n. gen., a new dwarf Bathonian (Middle Jurassic) nautilid from Western France</mods:title>
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<mods:namePart id="265941F92CAB274E7192300A0E86376B">Branger, Patrick</mods:namePart>
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<mods:affiliation id="597C9EC5F7884915A2941FC8CB9D8C19">Franche, F- 79410 Cherveux (France)</mods:affiliation>
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<mods:nameIdentifier id="CFEAE86BE380DD65DEC0D52A8B17CA9D" type="email">patrick.branger@wanadoo.fr</mods:nameIdentifier>
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<mods:title id="002F7787C7B0CBD913D7F8DF7EA2EF43">Comptes Rendus Palevol</mods:title>
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<mods:part id="69CA190714A888012DD83EC26D848701">
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<mods:date id="F17BA778EA26FCCB3AC4D96F957B4B2C">2023</mods:date>
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<mods:number id="6BBD7822BC91490B4A46D3B1E3B63118">2023-07-04</mods:number>
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<treatment id="201087CBFFD8BE69FF55F960FE5AFA36" ID-DOI="http://doi.org/10.5281/zenodo.14232249" ID-GBIF-Taxon="247601402" ID-Zenodo-Dep="14232249" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36" httpUri="http://treatment.plazi.org/id/201087CBFFD8BE69FF55F960FE5AFA36" lastPageId="7" lastPageNumber="484" pageId="5" pageNumber="482" scope_class="Cephalopoda" scope_family="Paracenoceratidae" scope_order="Nautilida" scope_subClass="Nautilia" scope_superFamily="Nautiloidea">
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<subSubSection id="E0A36556FFD8BE6BFF55F960FD2FF955" box="[252,655,1683,1710]" pageId="5" pageNumber="482" type="nomenclature">
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<paragraph id="A80636DDFFD8BE6BFF55F960FD2FF955" blockId="5.[252,655,1683,1710]" box="[252,655,1683,1710]" pageId="5" pageNumber="482">
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<heading id="F34E81B1FFD8BE6BFF55F960FD2FF955" box="[252,655,1683,1710]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="482" reason="2">
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<taxonomicName id="6FB94D5EFFD8BE6BFF55F960FE55F956" authorityName="Branger" authorityYear="2023" box="[252,501,1684,1710]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
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<emphasis id="9ACDEACFFFD8BE6BFF55F960FE55F956" bold="true" box="[252,501,1684,1710]" italics="true" pageId="5" pageNumber="482">Micronautilus evolutus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="81FE57B4FFD8BE6BFE55F967FD2FF955" box="[508,655,1683,1709]" pageId="5" pageNumber="482" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E0A36556FFD8BE6BFE9FF947FCA7F8FE" pageId="5" pageNumber="482" type="description">
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<paragraph id="A80636DDFFD8BE6BFE9FF947FDF5F935" blockId="5.[310,597,1714,1741]" box="[310,597,1714,1741]" pageId="5" pageNumber="482">
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(
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<figureCitation id="30822A58FFD8BE6BFE96F946FE36F935" box="[319,406,1714,1741]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStartId-0="4.[132,143,875,892]" captionStartId-1="5.[520,531,1369,1386]" captionStartId-2="6.[132,143,999,1016]" captionStartId-3="6.[813,824,1682,1699]" captionStartId-4="7.[211,221,815,832]" captionTargetBox-0="[210,1377,214,833]" captionTargetBox-1="[157,1406,214,1322]" captionTargetBox-2="[416,1268,190,982]" captionTargetBox-3="[867,1426,1113,1641]" captionTargetBox-4="[155,1436,218,778]" captionTargetId-0="figure-232@4.[210,840,214,745]" captionTargetId-1="figure-336@5.[698,1267,762,1328]" captionTargetId-2="figure-434@6.[414,983,228,770]" captionTargetId-3="figure-471@6.[811,1456,1090,1587]" captionTargetId-4="graphics-781@7.[213,1413,225,697]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIG. 1. — Geographical map reporting the position of the localities that provided specimens of Micronautilus n. gen. Numbers: 1, Buffevent; 2, Saint-Maixentl’École; 3, Sainte-Éanne; 4, Trottebuie; 5, Saint-Gelais; 6, les Lucs; 7, Souché; 8, Jard-sur-Mer. TABLE 1. — Measurements in millimetres for the shells of Micronautilus evolutus n. gen., n. sp." captionText-1="FIG. 2. — Stratigraphic section at Buffevent quarry fromBranger (2009)." captionText-2="FIG. 3. — Suture lines of Micronautilus evolutus n. gen., n. sp. at the end of the phragmocone compared with Pictonautilus Branger, 2004 and Pseudaganides Spath, 1927: A-C, Micronautilus evolutus n. gen., n. sp.; D, Pictonautilus verciacensis (Lissajous, 1923), middle Bathonian, Bremeri Zone, Saint-Maixent-l’École (Deux-Sèvres); E, Pseudaganides n. sp. upper Bathonian, Retrocostatum Zone, Aiffres, Deux-Sèvres." captionText-3="FIG. 4. — Measurement of the septal spacing in the studied Micronautilus n. gen. Scale bar: 1 cm." captionText-4="FIG. 5. — Septum distances of Micronautilus evolutusn.gen.,n. sp.: A, 2021.BR.N.066;B, 2021.BR.N.091;C, 2021.BR.N.136;D, 2021.BR.N.138." pageId="5" pageNumber="482">Figs 1-5</figureCitation>
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;
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<figureCitation id="30822A58FFD8BE6BFE0DF946FD81F935" box="[420,545,1714,1741]" captionStart="APPENDIX" captionStartId="11.[132,144,316,333]" captionTargetId="figure-151@11.[748,1286,1040,1602]" captionTargetPageId="11" captionText="APPENDIX 1. — Micronautilus evolutus n. gen., n. sp.: A, holotype 2021.BR.N.103, Buffevent quarry, bed 8, upper Bathonian, Retrocostatum Zone,Hannoveranus horizon;B, paratype 2021.BR.N.091, Buffevent quarry,top of bed 8, upper Bathonian, Retrocostatum Zone, Hannoveranus horizon;C, paratype 2021.BR.N.138, Buffevent quarry,bed 8, upper Bathonian,Retrocostatum Zone,Hannoveranus horizon.The smallest adult with approximated septas;D, paratype 2021.BR.N.066, Saint-Maixent-l’École, upper Bathonian,base of the Retrocostatum Zone; E, paratype 2021.BR.N.136, Buffevent quarry, bed 3, middle Bathonian, Bremeri Zone, Fortecostatum horizon; F, Micronautilus minoti n. gen., n. sp. paratype 2021.BR.N.135, les Hauts-de-Rochefort (Sainte-Éanne quarry), middle Bathonian, Progracilis Zone, Orbignyi horizon. An asterisk (*) marks the beginning of the body chamber. Scale bar: 1 cm." pageId="5" pageNumber="482">Appendix 1</figureCitation>
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A-E)
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</paragraph>
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<paragraph id="A80636DDFFD8BE6BFF2DF901FCA7F8FE" blockId="5.[132,775,1781,1798]" box="[132,775,1781,1798]" pageId="5" pageNumber="482">
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<uri id="DC283ADFFFD8BE6BFF2DF901FCA7F8FE" box="[132,775,1781,1798]" pageId="5" pageNumber="482">
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urn:lsid:zoobank.org:act:
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<uuid id="DC1F0C08FFD8BE6BFECBF901FCA7F8FE" box="[354,775,1781,1798]" pageId="5" pageNumber="482">FA6CB531-2520-41D7-B6B9-758B95FB3984</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="E0A36556FFD8BE6BFF2DF8C8FE59F871" pageId="5" pageNumber="482" type="reference_group">
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<paragraph id="A80636DDFFD8BE6BFF2DF8C8FCA7F8AC" blockId="5.[132,775,1851,1876]" box="[132,775,1851,1876]" pageId="5" pageNumber="482">
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<treatmentCitationGroup id="88A911F3FFD8BE6BFF2DF8C8FCA7F8AC" box="[132,775,1851,1876]" pageId="5" pageNumber="482">
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?
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<taxonomicName id="6FB94D5EFFD8BE6BFF25F8CFFE70F8AB" authority="(Lissajous)" baseAuthorityName="Lissajous" baseAuthorityYear="1923" box="[140,464,1851,1875]" class="Cephalopoda" family="Cenoceratidae" genus="Pictonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="verciacensis">
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<emphasis id="9ACDEACFFFD8BE6BFF25F8CFFECAF8AB" box="[140,362,1851,1875]" italics="true" pageId="5" pageNumber="482">Pictonautilus verciacensis</emphasis>
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(Lissajous)
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</taxonomicName>
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–
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<treatmentCitation id="291810CCFFD8BE6BFE4CF8C8FD37F8AC" author="BRANGER P." box="[485,663,1852,1876]" page="143" pageId="5" pageNumber="482" year="2004">
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<bibRefCitation id="CC284B2CFFD8BE6BFE4CF8C8FD37F8AC" author="BRANGER P." box="[485,663,1852,1876]" pageId="5" pageNumber="482" pagination="142 - 149" refId="ref5498" refString="BRANGER P. 2004. - Middle Jurassic Nautiloidea from Western France. Rivista Italiana di Paleontologia e Stratigraphia (Special issue the 6 th Int. Symposium on the Jurassic System, 16 - 19 September 2002) 110 (1): 142 - 149, 3 pl." type="journal article" year="2004">Branger 2004: 143</bibRefCitation>
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</treatmentCitation>
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||
, pl. 1, fig. 3.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A80636DDFFD8BE6BFF2DF884FE59F871" blockId="5.[132,505,1904,1929]" box="[132,505,1904,1929]" pageId="5" pageNumber="482">
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<treatmentCitationGroup id="88A911F3FFD8BE6BFF2DF884FE59F871" box="[132,505,1904,1929]" pageId="5" pageNumber="482">
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<taxonomicName id="6FB94D5EFFD8BE6BFF2DF884FE80F871" box="[132,288,1904,1929]" class="Cephalopoda" family="Cenoceratidae" genus="Pictonautilus" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="9ACDEACFFFD8BE6BFF2DF884FF5FF870" box="[132,255,1904,1928]" italics="true" pageId="5" pageNumber="482">Pictonautilus</emphasis>
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sp.
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</taxonomicName>
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? –
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<treatmentCitation id="291810CCFFD8BE6BFEEBF884FE55F871" author="GRULKE W." box="[322,501,1904,1929]" page="141" pageId="5" pageNumber="482" year="2016">
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<bibRefCitation id="CC284B2CFFD8BE6BFEEBF884FE55F871" author="GRULKE W." box="[322,501,1904,1929]" pageId="5" pageNumber="482" refId="ref5854" refString="GRULKE W. 2016. - Nautilus, Beautiful Survivor: 500 Million Years of Evolutionary. @ One Edition, Sherborne, 224 p." type="book" year="2016">Grulke 2016: 141</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E0A36556FFD8BE6BFF2DF86BFBA0FA1E" pageId="5" pageNumber="482" type="diagnosis">
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<paragraph id="A80636DDFFD8BE6BFF2DF86BFBA0FA1E" blockId="5.[132,777,1950,2028]" lastBlockId="5.[814,1456,1459,1510]" pageId="5" pageNumber="482">
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. — Small sized nautilid (less than
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<quantity id="6F419B38FFD8BE6BFDE8F86AFD29F84E" box="[577,649,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="482" unit="mm" value="45.0">45 mm</quantity>
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in diameter) with a compressed and subrectangular section of the shell. The umbilical area is always open and quite wide, up to 20% of shell diameter. Septa are relatively closely spaced and sutures exhibit a shallow lateral lobe.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E0A36556FFD8BE6BFC84F9F7FB59F895" pageId="5" pageNumber="482" type="materials_examined">
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<paragraph id="A80636DDFFD8BE6BFC84F9F7FAFDF9CD" blockId="5.[812,1458,1538,1750]" pageId="5" pageNumber="482">
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<materialsCitation id="18D13C80FFD8BE6BFC84F9F7FAFDF9CD" ID-GBIF-Occurrence="4985331304" country="France" location="Buffevent" municipality="Deux-Sevres" pageId="5" pageNumber="482" specimenCode="N103" specimenCount="1" typeStatus="holotype">
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<smallCapsWord id="AEE0A001FFD8BE6BFC84F9F7FCFDF9E0" baselines="1556,1556" box="[813,861,1539,1562]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="5" pageNumber="482">
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<collectionCode id="CEA8AE18FFD8BE6BFC84F9F7FCFDF9E0" box="[813,861,1539,1562]" pageId="5" pageNumber="482">TYPE</collectionCode>
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</smallCapsWord>
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<smallCapsWord id="AEE0A001FFD8BE6BFCCDF9FCFC62F9E1" baselines="1557" box="[868,962,1544,1561]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="5" pageNumber="482">
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<collectionCode id="CEA8AE18FFD8BE6BFCCDF9FCFC62F9E1" box="[868,962,1544,1561]" pageId="5" pageNumber="482">MATERIAL</collectionCode>
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</smallCapsWord>
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.
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||
<emphasis id="9ACDEACFFFD8BE6BFC67F9F7FC47F9E2" box="[974,999,1539,1562]" italics="true" pageId="5" pageNumber="482">—</emphasis>
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<typeStatus id="7702887FFFD8BE6BFC45F9F6FBE9F9E2" box="[1004,1097,1538,1562]" pageId="5" pageNumber="482" type="holotype">Holotype</typeStatus>
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||
N103 from
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<location id="AD666006FFD8BE6BFB6DF9F6FA82F9E2" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36:AD666006FFD8BE6BFB6DF9F6FA82F9E2" box="[1220,1314,1538,1562]" country="France" county="It" municipality="Deux-Sevres" name="Buffevent" pageId="5" pageNumber="482">Buffevent</location>
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quarry,
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<location id="AD666006FFD8BE6BFADBF9F6FA0BF9E2" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36:AD666006FFD8BE6BFADBF9F6FA0BF9E2" box="[1394,1451,1538,1562]" country="France" county="It" municipality="Deux-Sevres" name="Niort" pageId="5" pageNumber="482">Niort</location>
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||
,
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||
<collectingMunicipality id="4862ACA7FFD8BE6BFC85F9E9FC08F9CD" box="[812,936,1565,1589]" pageId="5" pageNumber="482">Deux-Sèvres</collectingMunicipality>
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||
,
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||
<collectingCountry id="D0AE764DFFD8BE6BFC1BF9E9FC56F9CC" box="[946,1014,1565,1588]" name="France" pageId="5" pageNumber="482">France</collectingCountry>
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||
.
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||
<collectingCounty id="41674E51FFD8BE6BFC56F9E9FBAFF9CC" box="[1023,1039,1565,1588]" pageId="5" pageNumber="482">It</collectingCounty>
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||
is figured in
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||
<figureCitation id="30822A58FFD8BE6BFB39F9E9FB68F9CD" box="[1168,1224,1565,1589]" captionStart="FIG" captionStartId="4.[132,143,875,892]" captionTargetBox="[210,1377,214,833]" captionTargetId="figure-232@4.[210,840,214,745]" captionTargetPageId="4" captionText="FIG. 1. — Geographical map reporting the position of the localities that provided specimens of Micronautilus n. gen. Numbers: 1, Buffevent; 2, Saint-Maixentl’École; 3, Sainte-Éanne; 4, Trottebuie; 5, Saint-Gelais; 6, les Lucs; 7, Souché; 8, Jard-sur-Mer. TABLE 1. — Measurements in millimetres for the shells of Micronautilus evolutus n. gen., n. sp." pageId="5" pageNumber="482">Fig. 1</figureCitation>
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||
;
|
||
<figureCitation id="30822A58FFD8BE6BFB7CF9E9FAF9F9CD" box="[1237,1369,1565,1589]" captionStart="APPENDIX" captionStartId="11.[132,144,316,333]" captionTargetId="figure-151@11.[748,1286,1040,1602]" captionTargetPageId="11" captionText="APPENDIX 1. — Micronautilus evolutus n. gen., n. sp.: A, holotype 2021.BR.N.103, Buffevent quarry, bed 8, upper Bathonian, Retrocostatum Zone,Hannoveranus horizon;B, paratype 2021.BR.N.091, Buffevent quarry,top of bed 8, upper Bathonian, Retrocostatum Zone, Hannoveranus horizon;C, paratype 2021.BR.N.138, Buffevent quarry,bed 8, upper Bathonian,Retrocostatum Zone,Hannoveranus horizon.The smallest adult with approximated septas;D, paratype 2021.BR.N.066, Saint-Maixent-l’École, upper Bathonian,base of the Retrocostatum Zone; E, paratype 2021.BR.N.136, Buffevent quarry, bed 3, middle Bathonian, Bremeri Zone, Fortecostatum horizon; F, Micronautilus minoti n. gen., n. sp. paratype 2021.BR.N.135, les Hauts-de-Rochefort (Sainte-Éanne quarry), middle Bathonian, Progracilis Zone, Orbignyi horizon. An asterisk (*) marks the beginning of the body chamber. Scale bar: 1 cm." pageId="5" pageNumber="482">Appendix 1A</figureCitation>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="A80636DDFFD8BE6BFC84F9C3FBE1F92F" blockId="5.[812,1458,1538,1750]" pageId="5" pageNumber="482">
|
||
<materialsCitation id="18D13C80FFD8BE6BFC84F9C3FC3BF97C" ID-GBIF-Occurrence="4985331312" collectionCode="BR" location="Buffevent quarry near Niort" pageId="5" pageNumber="482" specimenCode="2021. BR.N.091" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="BEBFFD54FFD8BE6BFC84F9C3FB87F9B7" box="[813,1063,1591,1616]" count="8" pageId="5" pageNumber="482" type="generic">Eight paratype specimens</specimenCount>
|
||
(figured in
|
||
<figureCitation id="30822A58FFD8BE6BFB09F9C3FB78F9A8" box="[1184,1240,1591,1616]" captionStart="FIG" captionStartId="5.[520,531,1369,1386]" captionTargetBox="[157,1406,214,1322]" captionTargetId="figure-336@5.[698,1267,762,1328]" captionTargetPageId="5" captionText="FIG. 2. — Stratigraphic section at Buffevent quarry fromBranger (2009)." pageId="5" pageNumber="482">Fig. 2</figureCitation>
|
||
;
|
||
<figureCitation id="30822A58FFD8BE6BFB4CF9C3FAF9F9A8" box="[1253,1369,1591,1616]" captionStart="APPENDIX" captionStartId="11.[132,144,316,333]" captionTargetId="figure-151@11.[748,1286,1040,1602]" captionTargetPageId="11" captionText="APPENDIX 1. — Micronautilus evolutus n. gen., n. sp.: A, holotype 2021.BR.N.103, Buffevent quarry, bed 8, upper Bathonian, Retrocostatum Zone,Hannoveranus horizon;B, paratype 2021.BR.N.091, Buffevent quarry,top of bed 8, upper Bathonian, Retrocostatum Zone, Hannoveranus horizon;C, paratype 2021.BR.N.138, Buffevent quarry,bed 8, upper Bathonian,Retrocostatum Zone,Hannoveranus horizon.The smallest adult with approximated septas;D, paratype 2021.BR.N.066, Saint-Maixent-l’École, upper Bathonian,base of the Retrocostatum Zone; E, paratype 2021.BR.N.136, Buffevent quarry, bed 3, middle Bathonian, Bremeri Zone, Fortecostatum horizon; F, Micronautilus minoti n. gen., n. sp. paratype 2021.BR.N.135, les Hauts-de-Rochefort (Sainte-Éanne quarry), middle Bathonian, Progracilis Zone, Orbignyi horizon. An asterisk (*) marks the beginning of the body chamber. Scale bar: 1 cm." pageId="5" pageNumber="482">Appendix 1</figureCitation>
|
||
B-E) obtained from several localities (
|
||
<figureCitation id="30822A58FFD8BE6BFBE4F9A6FB24F992" box="[1101,1156,1618,1642]" captionStart="FIG" captionStartId="4.[132,143,875,892]" captionTargetBox="[210,1377,214,833]" captionTargetId="figure-232@4.[210,840,214,745]" captionTargetPageId="4" captionText="FIG. 1. — Geographical map reporting the position of the localities that provided specimens of Micronautilus n. gen. Numbers: 1, Buffevent; 2, Saint-Maixentl’École; 3, Sainte-Éanne; 4, Trottebuie; 5, Saint-Gelais; 6, les Lucs; 7, Souché; 8, Jard-sur-Mer. TABLE 1. — Measurements in millimetres for the shells of Micronautilus evolutus n. gen., n. sp." pageId="5" pageNumber="482">Fig. 1</figureCitation>
|
||
) have also been studied:
|
||
<specimenCode id="F81F9EA6FFD8BE6BFADFF9A6FC37F97C" pageId="5" pageNumber="482">2021. BR.N.091</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation id="18D13C80FFD8BE6BFC0DF998FBECF97C" ID-GBIF-Occurrence="4985331308" box="[932,1100,1644,1668]" collectionCode="BR" location="Buffevent quarry near Niort" pageId="5" pageNumber="482" specimenCode="2021.BR.N.103" specimenCount="1">
|
||
<specimenCode id="F81F9EA6FFD8BE6BFC0DF998FBE9F97C" box="[932,1097,1644,1668]" pageId="5" pageNumber="482">2021.BR.N.103</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation id="18D13C80FFD8BE6BFBFCF998FB5EF97C" ID-GBIF-Occurrence="4985331303" box="[1109,1278,1644,1668]" collectionCode="BR" location="Buffevent quarry near Niort" pageId="5" pageNumber="482" specimenCode="2021.BR.N.136" specimenCount="1">
|
||
<specimenCode id="F81F9EA6FFD8BE6BFBFCF998FB5AF97C" box="[1109,1274,1644,1668]" pageId="5" pageNumber="482">2021.BR.N.136</specimenCode>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation id="18D13C80FFD8BE6BFAAFF998FA0FF97C" ID-GBIF-Occurrence="4985331306" box="[1286,1455,1644,1668]" collectionCode="BR" location="Buffevent quarry near Niort" pageId="5" pageNumber="482" specimenCount="1">
|
||
2021.
|
||
<collectionCode id="CEA8AE18FFD8BE6BFAE8F998FAC7F97C" box="[1345,1383,1644,1668]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="5" pageNumber="482">BR</collectionCode>
|
||
.N.137,
|
||
</materialsCitation>
|
||
<materialsCitation id="18D13C80FFD8BE6BFC87F973FACDF967" ID-GBIF-Occurrence="4985331310" box="[814,1389,1671,1695]" collectionCode="BR" location="Buffevent quarry near Niort" pageId="5" pageNumber="482" specimenCount="1">
|
||
2021.
|
||
<collectionCode id="CEA8AE18FFD8BE6BFCC3F973FC31F967" box="[874,913,1671,1695]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="5" pageNumber="482">BR</collectionCode>
|
||
.N.138 are from
|
||
<location id="AD666006FFD8BE6BFBE9F973FAC9F967" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36:AD666006FFD8BE6BFBE9F973FAC9F967" box="[1088,1385,1671,1695]" name="Buffevent quarry near Niort" pageId="5" pageNumber="482">Buffevent quarry near Niort</location>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation id="18D13C80FFD8BE6BFADFF973FAA2F941" ID-GBIF-Occurrence="4985331305" location="Jard-sur-Mer (Vendee)" pageId="5" pageNumber="482" specimenCode="2021. BR.N.139" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="F81F9EA6FFD8BE6BFADFF973FC36F941" pageId="5" pageNumber="482">2021. BR.N.139</specimenCode>
|
||
is from
|
||
<location id="AD666006FFD8BE6BFC59F955FB6EF941" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36:AD666006FFD8BE6BFC59F955FB6EF941" box="[1008,1230,1697,1721]" name="Jard-sur-Mer (Vendee" pageId="5" pageNumber="482">Jard-sur-Mer (Vendée</location>
|
||
) and
|
||
</materialsCitation>
|
||
<materialsCitation id="18D13C80FFD8BE6BFAA2F955FBE1F92F" ID-GBIF-Occurrence="4985331316" location="Saint-Maixent-l'Ecole" pageId="5" pageNumber="482" specimenCode="2021.BR.N.066" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="F81F9EA6FFD8BE6BFAA2F955FA0FF941" box="[1291,1455,1697,1721]" pageId="5" pageNumber="482">2021.BR.N.066</specimenCode>
|
||
from
|
||
<location id="AD666006FFD8BE6BFCCCF949FB9DF92F" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36:AD666006FFD8BE6BFCCCF949FB9DF92F" box="[869,1085,1725,1751]" name="Saint-Maixent-l'Ecole" pageId="5" pageNumber="482">Saint-Maixent-l’École</location>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="A80636DDFFD8BE6BFC84F918FA10F8FB" blockId="5.[811,1456,1772,1902]" box="[813,1456,1771,1795]" pageId="5" pageNumber="482">
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|
||
<typeStatus id="7702887FFFD8BE6BFC84F918FCFDF8F9" box="[813,861,1772,1795]" pageId="5" pageNumber="482">
|
||
<collectionCode id="CEA8AE18FFD8BE6BFC84F918FCFDF8F9" box="[813,861,1772,1795]" pageId="5" pageNumber="482">TYPE</collectionCode>
|
||
</typeStatus>
|
||
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|
||
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|
||
<collectionCode id="CEA8AE18FFD8BE6BFCCDF905FC1FF8FA" box="[868,959,1777,1794]" pageId="5" pageNumber="482">LOCALITY</collectionCode>
|
||
</smallCapsWord>
|
||
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|
||
<collectionCode id="CEA8AE18FFD8BE6BFC6FF905FC50F8FA" box="[966,1008,1777,1794]" pageId="5" pageNumber="482">AND</collectionCode>
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</smallCapsWord>
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|
||
<collectionCode id="CEA8AE18FFD8BE6BFC5FF905FBF0F8FA" box="[1014,1104,1777,1794]" pageId="5" pageNumber="482">STRATUM</collectionCode>
|
||
</smallCapsWord>
|
||
<smallCapsWord id="AEE0A001FFD8BE6BFBFEF905FB0CF8FA" baselines="1790" box="[1111,1196,1777,1794]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="typicum" pageId="5" pageNumber="482">
|
||
<collectionCode id="CEA8AE18FFD8BE6BFBFEF905FB0CF8FA" box="[1111,1196,1777,1794]" pageId="5" pageNumber="482">TYPICUM</collectionCode>
|
||
</smallCapsWord>
|
||
. —
|
||
<taxonomicName id="6FB94D5EFFD8BE6BFB73F91FFA10F8FB" authorityName="Branger" authorityYear="2023" box="[1242,1456,1771,1795]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFD8BE6BFB73F91FFA10F8FB" box="[1242,1456,1771,1795]" italics="true" pageId="5" pageNumber="482">Micronautilus evolutus</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="A80636DDFFD8BE6BFC84F8F2FB59F895" blockId="5.[811,1456,1772,1902]" pageId="5" pageNumber="482">
|
||
n. gen., n. sp. is relatively common at the
|
||
<location id="AD666006FFD8BE6BFB17F8F2FC2EF8C0" LSID="urn:lsid:plazi:treatment:201087CBFFD8BE69FF55F960FE5AFA36:AD666006FFD8BE6BFB17F8F2FC2EF8C0" country="Western France" name="top of the Calcaire a silex formation" pageId="5" pageNumber="482">top of the Calcaire à silex formation</location>
|
||
(
|
||
<bibRefCitation id="CC284B2CFFD8BE6BFC34F8D4FBBCF8C0" author="WELSCH J." box="[925,1052,1824,1848]" pageId="5" pageNumber="482" pagination="797 - 881" refId="ref6409" refString="WELSCH J. 1903. - Etude des terrains du Poitou dans le Detroit poitevin et sur les bords du Massif ancien de la Gatine: 797 - 881." type="book chapter" year="1903">Welsch 1903</bibRefCitation>
|
||
) of
|
||
<geologicalTimeScale id="DA99B676FFD8BE6BFBEBF8D4FB70F8C0" box="[1090,1232,1824,1848]" pageId="5" pageNumber="482">late Bathonian</geologicalTimeScale>
|
||
(Retrocostatum Zone) age in
|
||
<collectingCountry id="D0AE764DFFD8BE6BFCC0F8CFFBA2F8AA" box="[873,1026,1851,1874]" pageId="5" pageNumber="482">Western France</collectingCountry>
|
||
, especially in the vicinity of Niort. The taxon has not yet been described from any other area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E0A36556FFD8BE6BFC84F871FBC6F864" box="[813,1126,1925,1949]" pageId="5" pageNumber="482" type="description">
|
||
<paragraph id="A80636DDFFD8BE6BFC84F871FBC6F864" blockId="5.[813,1126,1925,1949]" box="[813,1126,1925,1949]" pageId="5" pageNumber="482">
|
||
<smallCapsWord id="AEE0A001FFD8BE6BFC84F871FC6BF863" baselines="1942,1943" box="[813,971,1925,1948]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Measurements" pageId="5" pageNumber="482">MEASUREMENTS</smallCapsWord>
|
||
. — see Table 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E0A36556FFD8BE6BFC84F84EFB9AF814" pageId="5" pageNumber="482" type="etymology">
|
||
<paragraph id="A80636DDFFD8BE6BFC84F84EFB9AF814" blockId="5.[812,1455,1978,2028]" pageId="5" pageNumber="482">
|
||
<smallCapsWord id="AEE0A001FFD8BE6BFC84F84EFC06F828" baselines="1995,1996" box="[813,934,1978,2001]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="5" pageNumber="482">ETYMOLOGY</smallCapsWord>
|
||
. — The specific name
|
||
<taxonomicName id="6FB94D5EFFD8BE6BFB2CF84EFB70F82A" authorityName="Branger" authorityYear="2023" box="[1157,1232,1978,2002]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="evolutus">
|
||
<emphasis id="9ACDEACFFFD8BE6BFB2CF84EFB70F82A" box="[1157,1232,1978,2002]" italics="true" pageId="5" pageNumber="482">evolutus</emphasis>
|
||
</taxonomicName>
|
||
relates to the relatively wide umbilicus of the shell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FCC66655FFDBBE68FF2DFC13FC6FFBDE" pageId="6" pageNumber="483" startId="6.[132,143,999,1016]" targetBox="[416,1268,190,982]" targetPageId="6" targetType="figure">
|
||
<paragraph id="A80636DDFFDBBE68FF2DFC13FC6FFBDE" blockId="6.[132,1455,999,1062]" pageId="6" pageNumber="483">
|
||
<smallCapsWord id="AEE0A001FFDBBE68FF2DFC13FF3DFC00" baselines="1012,1013" box="[132,157,999,1016]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="483">FIG</smallCapsWord>
|
||
. 3. — Suture lines of
|
||
<taxonomicName id="6FB94D5EFFDBBE68FEFDFC13FDAFFC00" authorityName="Branger" authorityYear="2023" box="[340,527,999,1016]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFDBBE68FEFDFC13FDAFFC00" box="[340,527,999,1016]" italics="true" pageId="6" pageNumber="483">Micronautilus evolutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDBBE68FDBFFC13FD2BFC00" box="[534,651,999,1016]" pageId="6" pageNumber="483" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
at the end of the phragmocone compared with
|
||
<taxonomicName id="6FB94D5EFFDBBE68FBB4FC13FAA5FC00" authority="Branger, 2004" authorityName="Branger" authorityYear="2004" box="[1053,1285,999,1016]" class="Cephalopoda" family="Cenoceratidae" genus="Pictonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="genus">
|
||
<emphasis id="9ACDEACFFFDBBE68FBB4FC13FB28FC00" box="[1053,1160,999,1016]" italics="true" pageId="6" pageNumber="483">Pictonautilus</emphasis>
|
||
<bibRefCitation id="CC284B2CFFDBBE68FB27FC13FAA5FC00" author="BRANGER P." box="[1166,1285,999,1016]" pageId="6" pageNumber="483" pagination="142 - 149" refId="ref5498" refString="BRANGER P. 2004. - Middle Jurassic Nautiloidea from Western France. Rivista Italiana di Paleontologia e Stratigraphia (Special issue the 6 th Int. Symposium on the Jurassic System, 16 - 19 September 2002) 110 (1): 142 - 149, 3 pl." type="journal article" year="2004">Branger, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6FB94D5EFFDBBE68FA99FC13FF4DFBF7" authority="Spath, 1927" authorityName="Spath" authorityYear="1927" class="Cephalopoda" family="Pseudonautilidae" genus="Pseudaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="genus">
|
||
<emphasis id="9ACDEACFFFDBBE68FA99FC13FA0FFC00" box="[1328,1455,999,1016]" italics="true" pageId="6" pageNumber="483">Pseudaganides</emphasis>
|
||
Spath, 1927
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="9ACDEACFFFDBBE68FF5DFC0AFEB6FBF7" bold="true" box="[244,278,1022,1039]" pageId="6" pageNumber="483">A -C</emphasis>
|
||
,
|
||
<taxonomicName id="6FB94D5EFFDBBE68FE89FC0AFE7AFBF7" authorityName="Branger" authorityYear="2023" box="[288,474,1022,1039]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFDBBE68FE89FC0AFE7AFBF7" box="[288,474,1022,1039]" italics="true" pageId="6" pageNumber="483">Micronautilus evolutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDBBE68FE76FC0AFDF5FBF7" box="[479,597,1022,1039]" pageId="6" pageNumber="483" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="9ACDEACFFFDBBE68FDF5FC0AFDCAFBF7" bold="true" box="[604,618,1022,1039]" pageId="6" pageNumber="483">D</emphasis>
|
||
,
|
||
<taxonomicName id="6FB94D5EFFDBBE68FDDDFC0AFC7DFBF7" authority="(Lissajous, 1923)" baseAuthorityName="Lissajous" baseAuthorityYear="1923" box="[628,989,1022,1039]" class="Cephalopoda" family="Cenoceratidae" genus="Pictonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="verciacensis">
|
||
<emphasis id="9ACDEACFFFDBBE68FDDDFC0AFCEAFBF7" box="[628,842,1022,1039]" italics="true" pageId="6" pageNumber="483">Pictonautilus verciacensis</emphasis>
|
||
(Lissajous, 1923)
|
||
</taxonomicName>
|
||
, middle Bathonian, Bremeri Zone, Saint-Maixent-l’École (Deux-Sèvres);
|
||
<emphasis id="9ACDEACFFFDBBE68FEADFBE1FEB0FBDE" bold="true" box="[260,272,1045,1062]" pageId="6" pageNumber="483">E</emphasis>
|
||
,
|
||
<taxonomicName id="6FB94D5EFFDBBE68FEB3FBE1FE39FBDE" authorityName="Branger" authorityYear="2023" box="[282,409,1045,1062]" class="Cephalopoda" family="Pseudonautilidae" genus="Pseudaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="genus" status="n. sp.">
|
||
<emphasis id="9ACDEACFFFDBBE68FEB3FBE1FE39FBDE" box="[282,409,1045,1062]" italics="true" pageId="6" pageNumber="483">Pseudaganides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDBBE68FE36FBE1FE6FFBDE" box="[415,463,1045,1062]" pageId="6" pageNumber="483" rank="species">n. sp.</taxonomicNameLabel>
|
||
upper Bathonian, Retrocostatum Zone, Aiffres, Deux-Sèvres.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E0A36556FFDBBE68FF2DFBA1FEF7F813" pageId="6" pageNumber="483" type="description">
|
||
<paragraph id="A80636DDFFDBBE68FF2DFBA1FEF7F813" blockId="6.[130,777,1109,2028]" pageId="6" pageNumber="483">
|
||
<smallCapsWord id="AEE0A001FFDBBE68FF2DFBA1FEB5FB95" baselines="1128,1128" box="[132,277,1109,1135]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="6" pageNumber="483">DESCRIPTION</smallCapsWord>
|
||
<taxonomicName id="6FB94D5EFFDBBE68FF2DFB80FEDCFB76" authorityName="Branger" authorityYear="2023" box="[132,380,1140,1166]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFDBBE68FF2DFB80FEDCFB76" box="[132,380,1140,1166]" italics="true" pageId="6" pageNumber="483">Micronautilus evolutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDBBE68FE2EFB81FD85FB77" box="[391,549,1141,1167]" pageId="6" pageNumber="483" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
is a very distinctive species characterized by a very small size at adult stage, ranging from
|
||
<quantity id="6F419B38FFDBBE68FF42FB40FE9CFB37" box="[235,316,1204,1231]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="483" unit="mm" value="30.0">30 mm</quantity>
|
||
(estimated diameter) for 2021.BR.N.138 (
|
||
<figureCitation id="30822A58FFDBBE68FF24FB20FF6AFB16" box="[141,202,1236,1262]" captionStart="FIG" captionStartId="6.[132,143,999,1016]" captionTargetBox="[416,1268,190,982]" captionTargetId="figure-434@6.[414,983,228,770]" captionTargetPageId="6" captionText="FIG. 3. — Suture lines of Micronautilus evolutus n. gen., n. sp. at the end of the phragmocone compared with Pictonautilus Branger, 2004 and Pseudaganides Spath, 1927: A-C, Micronautilus evolutus n. gen., n. sp.; D, Pictonautilus verciacensis (Lissajous, 1923), middle Bathonian, Bremeri Zone, Saint-Maixent-l’École (Deux-Sèvres); E, Pseudaganides n. sp. upper Bathonian, Retrocostatum Zone, Aiffres, Deux-Sèvres." pageId="6" pageNumber="483">Fig. 3</figureCitation>
|
||
A-C;
|
||
<figureCitation id="30822A58FFDBBE68FEAFFB20FE38FB16" box="[262,408,1236,1262]" captionStart="APPENDIX" captionStartId="11.[132,144,316,333]" captionTargetId="figure-151@11.[748,1286,1040,1602]" captionTargetPageId="11" captionText="APPENDIX 1. — Micronautilus evolutus n. gen., n. sp.: A, holotype 2021.BR.N.103, Buffevent quarry, bed 8, upper Bathonian, Retrocostatum Zone,Hannoveranus horizon;B, paratype 2021.BR.N.091, Buffevent quarry,top of bed 8, upper Bathonian, Retrocostatum Zone, Hannoveranus horizon;C, paratype 2021.BR.N.138, Buffevent quarry,bed 8, upper Bathonian,Retrocostatum Zone,Hannoveranus horizon.The smallest adult with approximated septas;D, paratype 2021.BR.N.066, Saint-Maixent-l’École, upper Bathonian,base of the Retrocostatum Zone; E, paratype 2021.BR.N.136, Buffevent quarry, bed 3, middle Bathonian, Bremeri Zone, Fortecostatum horizon; F, Micronautilus minoti n. gen., n. sp. paratype 2021.BR.N.135, les Hauts-de-Rochefort (Sainte-Éanne quarry), middle Bathonian, Progracilis Zone, Orbignyi horizon. An asterisk (*) marks the beginning of the body chamber. Scale bar: 1 cm." pageId="6" pageNumber="483">Appendix 1C</figureCitation>
|
||
) up to
|
||
<quantity id="6F419B38FFDBBE68FE4DFB20FD93FB17" box="[484,563,1236,1263]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="483" unit="mm" value="44.0">44 mm</quantity>
|
||
in 2021.BR.N.136 (
|
||
<figureCitation id="30822A58FFDBBE68FF27FB00FEA6FAF7" box="[142,262,1268,1295]" captionStart="FIG" captionStartId="7.[211,221,815,832]" captionTargetBox="[155,1436,218,778]" captionTargetId="graphics-781@7.[213,1413,225,697]" captionTargetPageId="7" captionText="FIG. 5. — Septum distances of Micronautilus evolutusn.gen.,n. sp.: A, 2021.BR.N.066;B, 2021.BR.N.091;C, 2021.BR.N.136;D, 2021.BR.N.138." pageId="6" pageNumber="483">Fig. 5A, B</figureCitation>
|
||
;
|
||
<figureCitation id="30822A58FFDBBE68FEBFFB00FE0CFAF6" box="[278,428,1268,1294]" captionStart="APPENDIX" captionStartId="11.[132,144,316,333]" captionTargetId="figure-151@11.[748,1286,1040,1602]" captionTargetPageId="11" captionText="APPENDIX 1. — Micronautilus evolutus n. gen., n. sp.: A, holotype 2021.BR.N.103, Buffevent quarry, bed 8, upper Bathonian, Retrocostatum Zone,Hannoveranus horizon;B, paratype 2021.BR.N.091, Buffevent quarry,top of bed 8, upper Bathonian, Retrocostatum Zone, Hannoveranus horizon;C, paratype 2021.BR.N.138, Buffevent quarry,bed 8, upper Bathonian,Retrocostatum Zone,Hannoveranus horizon.The smallest adult with approximated septas;D, paratype 2021.BR.N.066, Saint-Maixent-l’École, upper Bathonian,base of the Retrocostatum Zone; E, paratype 2021.BR.N.136, Buffevent quarry, bed 3, middle Bathonian, Bremeri Zone, Fortecostatum horizon; F, Micronautilus minoti n. gen., n. sp. paratype 2021.BR.N.135, les Hauts-de-Rochefort (Sainte-Éanne quarry), middle Bathonian, Progracilis Zone, Orbignyi horizon. An asterisk (*) marks the beginning of the body chamber. Scale bar: 1 cm." pageId="6" pageNumber="483">Appendix 1E</figureCitation>
|
||
). The individuals are relatively evolute for Jurassic nautilids. The umbilicus is widely open since U/D can reach 20% of the diameter on full grown shells. At the very beginning of the embryonic shell, the whorl section is rounded and, until a diameter of about
|
||
<quantity id="6F419B38FFDBBE68FD7DFA80FCA9FA76" box="[724,777,1396,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="483" unit="cm" value="1.0">1 cm</quantity>
|
||
is reached, the shell displays reticulate ornamentation. This diameter seems to be approximately the hatching size of the animal according to the weak depression, corresponding to the nepionic constriction, observed on inner whorls of the
|
||
<typeStatus id="7702887FFFDBBE68FF2CF9E7FF42F9D5" box="[133,226,1555,1581]" pageId="6" pageNumber="483" type="holotype">holotype</typeStatus>
|
||
(2021.BR.N.103). Further adoral, the whorl section becomes sub-rectangular to sub-trapezoidal, compressed with a maximum width located close to the umbilicus. Flanks are flat, sub-parallel, boarded by angular but never sharp ventral edges; the wall of the umbilicus is sub-vertical. The venter is very slightly rounded on the inner whorls and flattens before the beginning of the body chamber, tending to be a little concave on stratigraphically younger specimens. The shell is narrowly camerate, with twelve to thirteen chambers occurring in each half of a whorl when adult. The suture lines are partly hidden on the
|
||
<typeStatus id="7702887FFFDBBE68FE73F8A6FD9BF894" box="[474,571,1874,1900]" pageId="6" pageNumber="483" type="holotype">holotype</typeStatus>
|
||
(2021.BR.N.103) but, on the
|
||
<typeStatus id="7702887FFFDBBE68FEA1F886FED3F874" box="[264,371,1906,1932]" pageId="6" pageNumber="483" type="paratype">paratypes</typeStatus>
|
||
, exhibit wide and shallow lateral and ventral lobes (
|
||
<figureCitation id="30822A58FFDBBE68FE88F866FEC1F854" box="[289,353,1938,1964]" captionStart="FIG" captionStartId="6.[132,143,999,1016]" captionTargetBox="[416,1268,190,982]" captionTargetId="figure-434@6.[414,983,228,770]" captionTargetPageId="6" captionText="FIG. 3. — Suture lines of Micronautilus evolutus n. gen., n. sp. at the end of the phragmocone compared with Pictonautilus Branger, 2004 and Pseudaganides Spath, 1927: A-C, Micronautilus evolutus n. gen., n. sp.; D, Pictonautilus verciacensis (Lissajous, 1923), middle Bathonian, Bremeri Zone, Saint-Maixent-l’École (Deux-Sèvres); E, Pseudaganides n. sp. upper Bathonian, Retrocostatum Zone, Aiffres, Deux-Sèvres." pageId="6" pageNumber="483">Fig. 3</figureCitation>
|
||
). The hyponomic sinus of the shell is quite deep and narrow. The position of the siphuncle has not been observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FCC66655FFDBBE68FC84F966FC54F941" pageId="6" pageNumber="483" startId="6.[813,824,1682,1699]" targetBox="[867,1426,1113,1641]" targetPageId="6" targetType="figure">
|
||
<paragraph id="A80636DDFFDBBE68FC84F966FC54F941" blockId="6.[813,1455,1682,1721]" pageId="6" pageNumber="483">
|
||
<smallCapsWord id="AEE0A001FFDBBE68FC84F966FCE5F95A" baselines="1695,1695" box="[813,837,1682,1699]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="483">FIG</smallCapsWord>
|
||
. 4. — Measurement of the septal spacing in the studied
|
||
<taxonomicName id="6FB94D5EFFDBBE68FA95F966FA0FF95B" authorityName="Branger" authorityYear="2023" box="[1340,1455,1682,1699]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="9ACDEACFFFDBBE68FA95F966FA0FF95B" box="[1340,1455,1682,1699]" italics="true" pageId="6" pageNumber="483">Micronautilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDBBE68FC84F95CFCC7F941" box="[813,871,1704,1721]" pageId="6" pageNumber="483" rank="genus">n. gen.</taxonomicNameLabel>
|
||
Scale bar: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E0A36556FFDBBE69FC84F907FE5AFA36" lastPageId="7" lastPageNumber="484" pageId="6" pageNumber="483" type="discussion">
|
||
<paragraph id="A80636DDFFDBBE68FC84F907FC31F8F3" blockId="6.[811,1457,1779,2027]" box="[813,913,1779,1805]" pageId="6" pageNumber="483">
|
||
<smallCapsWord id="AEE0A001FFDBBE68FC84F907FC31F8F3" baselines="1798,1798" box="[813,913,1779,1805]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="6" pageNumber="483">REMARKS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="A80636DDFFDBBE69FC84F8E6FE59FC28" blockId="6.[811,1457,1779,2027]" lastBlockId="7.[130,777,885,1486]" lastPageId="7" lastPageNumber="484" pageId="6" pageNumber="483">
|
||
As discussed above, apart from the
|
||
<specimenCount id="BEBFFD54FFDBBE68FB11F8E7FAF9F8D4" box="[1208,1369,1810,1837]" count="2" pageId="6" pageNumber="483" type="generic">two specimens</specimenCount>
|
||
figured under the name of?
|
||
<taxonomicName id="6FB94D5EFFDBBE68FC57F8C6FA10F8B4" authority="(Branger, 2004)" baseAuthorityName="Branger" baseAuthorityYear="2004" box="[1022,1456,1842,1868]" class="Cephalopoda" family="Cenoceratidae" genus="Pictonautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="verciacensis">
|
||
<emphasis id="9ACDEACFFFDBBE68FC57F8C6FAA0F8B4" box="[1022,1280,1842,1868]" italics="true" pageId="6" pageNumber="483">Pictonautilus verciacensis</emphasis>
|
||
(
|
||
<bibRefCitation id="CC284B2CFFDBBE68FAA4F8C6FA07F8B4" author="BRANGER P." box="[1293,1447,1842,1868]" pageId="6" pageNumber="483" pagination="142 - 149" refId="ref5498" refString="BRANGER P. 2004. - Middle Jurassic Nautiloidea from Western France. Rivista Italiana di Paleontologia e Stratigraphia (Special issue the 6 th Int. Symposium on the Jurassic System, 16 - 19 September 2002) 110 (1): 142 - 149, 3 pl." type="journal article" year="2004">Branger, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6FB94D5EFFDBBE68FCF2F8A6FBA7F895" box="[859,1031,1874,1901]" class="Cephalopoda" family="Cenoceratidae" genus="Pictonautilus" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="9ACDEACFFFDBBE68FCF2F8A6FC43F894" box="[859,995,1874,1900]" italics="true" pageId="6" pageNumber="483">Pictonautilus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="CC284B2CFFDBBE68FBBFF8A6FB79F894" author="GRULKE W." box="[1046,1241,1874,1901]" pageId="6" pageNumber="483" refId="ref5854" refString="GRULKE W. 2016. - Nautilus, Beautiful Survivor: 500 Million Years of Evolutionary. @ One Edition, Sherborne, 224 p." type="book" year="2016">Grulke 2016: 141</bibRefCitation>
|
||
),
|
||
<taxonomicName id="6FB94D5EFFDBBE68FB45F8A6FCFFF874" authorityName="Branger" authorityYear="2023" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFDBBE68FB45F8A6FCFFF874" italics="true" pageId="6" pageNumber="483">Micronautilus evolutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDBBE68FCC1F886FBA2F874" box="[872,1026,1906,1932]" pageId="6" pageNumber="483" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
has not previously been illustrated. At the same stratigraphic level, the closest small sized nautilid with a compressed shell belongs to the earliest representatives of
|
||
<taxonomicName id="6FB94D5EFFDBBE68FC28F825FBB3F813" authorityName="Branger" authorityYear="2023" box="[897,1043,2001,2027]" class="Cephalopoda" family="Pseudonautilidae" genus="Pseudaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="483" phylum="Mollusca" rank="genus">
|
||
<emphasis id="9ACDEACFFFDBBE68FC28F825FBB3F813" box="[897,1043,2001,2027]" italics="true" pageId="6" pageNumber="483">Pseudaganides</emphasis>
|
||
</taxonomicName>
|
||
(ongoing studies from personal data). This genus is not considered here to be closely related due to its closed umbilicus and suture lines that are much more sinuous than
|
||
<taxonomicName id="6FB94D5EFFDABE69FEBFFC41FE08FC37" authorityName="Branger" authorityYear="2023" box="[278,424,949,975]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="484" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="9ACDEACFFFDABE69FEBFFC41FE08FC37" box="[278,424,949,975]" italics="true" pageId="7" pageNumber="484">Micronautilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDABE69FE06FC42FE59FC28" box="[431,505,950,976]" pageId="7" pageNumber="484" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="FCC66655FFDABE69FF7AFCDBFAC3FCB8" box="[211,1379,815,832]" pageId="7" pageNumber="484" startId="7.[211,221,815,832]" targetBox="[155,1436,218,778]" targetPageId="7" targetType="figure">
|
||
<paragraph id="A80636DDFFDABE69FF7AFCDBFAC3FCB8" blockId="7.[211,1379,815,832]" box="[211,1379,815,832]" pageId="7" pageNumber="484">
|
||
<smallCapsWord id="AEE0A001FFDABE69FF7AFCDBFF4BFCB8" baselines="828,829" box="[211,235,815,832]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="7" pageNumber="484">FIG</smallCapsWord>
|
||
. 5. — Septum distances of
|
||
<taxonomicName id="6FB94D5EFFDABE69FE60FCDBFDDEFCB8" authorityName="Branger" authorityYear="2023" box="[457,638,815,832]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="484" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFDABE69FE60FCDBFDDEFCB8" box="[457,638,815,832]" italics="true" pageId="7" pageNumber="484">Micronautilus evolutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDABE69FD29FCDBFD50FCB8" box="[640,752,815,832]" pageId="7" pageNumber="484" rank="species">n.gen.,n. sp.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="9ACDEACFFFDABE69FD5FFCDBFCA3FCB8" bold="true" box="[758,771,815,832]" pageId="7" pageNumber="484">A</emphasis>
|
||
, 2021.BR.N.066;
|
||
<emphasis id="9ACDEACFFFDABE69FC38FCDBFC3EFCB8" bold="true" box="[913,926,815,832]" pageId="7" pageNumber="484">B</emphasis>
|
||
, 2021.BR.N.091;
|
||
<emphasis id="9ACDEACFFFDABE69FB85FCDBFB9AFCB8" bold="true" box="[1068,1082,815,832]" pageId="7" pageNumber="484">C</emphasis>
|
||
, 2021.BR.N.136;
|
||
<emphasis id="9ACDEACFFFDABE69FB60FCDBFB77FCB8" bold="true" box="[1225,1239,815,832]" pageId="7" pageNumber="484">D</emphasis>
|
||
, 2021.BR.N.138.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A80636DDFFDABE69FF32FC21FE5AFA36" blockId="7.[130,777,885,1486]" pageId="7" pageNumber="484">
|
||
Despite of their small size, the specimens of
|
||
<taxonomicName id="6FB94D5EFFDABE69FD24FC22FCA8FC17" authorityName="Branger" authorityYear="2023" box="[653,776,981,1007]" class="Cephalopoda" family="Paracenoceratidae" genus="Micronautilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="484" phylum="Mollusca" rank="species" species="evolutus" status="n. gen., n. sp.">
|
||
<emphasis id="9ACDEACFFFDABE69FD24FC22FCA8FC17" box="[653,776,981,1007]" italics="true" pageId="7" pageNumber="484">M. evolutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81FE57B4FFDABE69FF2DFC01FEBCFBF7" box="[132,284,1013,1039]" pageId="7" pageNumber="484" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
are regarded as mature shells because of two reasons. The first one is related to the crowding of the last septum at the end of the phragmocone. The septal spacing has been measured on
|
||
<specimenCount id="BEBFFD54FFDABE69FEFBFBA1FE59FB97" box="[338,505,1109,1135]" count="4" pageId="7" pageNumber="484" type="generic">four specimens</specimenCount>
|
||
, 2021.BR.N.066, 2021. BR.N.091, 2021.BR.N.136 and 2021.BR.N.138 (
|
||
<figureCitation id="30822A58FFDABE69FD0DFB80FD43FB77" box="[676,739,1140,1167]" captionStart="FIG" captionStartId="6.[813,824,1682,1699]" captionTargetBox="[867,1426,1113,1641]" captionTargetId="figure-471@6.[811,1456,1090,1587]" captionTargetPageId="6" captionText="FIG. 4. — Measurement of the septal spacing in the studied Micronautilus n. gen. Scale bar: 1 cm." pageId="7" pageNumber="484">Figs 4</figureCitation>
|
||
;
|
||
<figureCitation id="30822A58FFDABE69FD59FB80FD5EFB77" box="[752,766,1140,1167]" captionStart="FIG" captionStartId="7.[211,221,815,832]" captionTargetBox="[155,1436,218,778]" captionTargetId="graphics-781@7.[213,1413,225,697]" captionTargetPageId="7" captionText="FIG. 5. — Septum distances of Micronautilus evolutusn.gen.,n. sp.: A, 2021.BR.N.066;B, 2021.BR.N.091;C, 2021.BR.N.136;D, 2021.BR.N.138." pageId="7" pageNumber="484">5</figureCitation>
|
||
) according the method used by
|
||
<bibRefCitation id="CC284B2CFFDABE69FE7EFB60FD3EFB57" author="KRAFT S. & KORN D. & KLUG C." box="[471,670,1172,1199]" pageId="7" pageNumber="484" pagination="31 - 44" refId="ref5995" refString="KRAFT S., KORN D. & KLUG C. 2008. - Patterns of ontogenic septal spacing in Carboniferous ammonoids. Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 250 (1): 31 - 44. https: // doi. org / 10.1127 / 0077 - 7749 / 2008 / 0250 - 0031" type="journal article" year="2008">
|
||
Kraft
|
||
<emphasis id="9ACDEACFFFDABE69FDBFFB61FDEBFB56" box="[534,587,1172,1198]" italics="true" pageId="7" pageNumber="484">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
. As inner whorls are small and hidden by overlapping whorls, they cannot be observed clearly and the measurements have been only done from 180° to 20° before the last septum. All of them show an angular distance that varies from 12 to 20° (
|
||
<figureCitation id="30822A58FFDABE69FD21FAE0FD66FAD6" box="[648,710,1300,1326]" captionStart="FIG" captionStartId="6.[813,824,1682,1699]" captionTargetBox="[867,1426,1113,1641]" captionTargetId="figure-471@6.[811,1456,1090,1587]" captionTargetPageId="6" captionText="FIG. 4. — Measurement of the septal spacing in the studied Micronautilus n. gen. Scale bar: 1 cm." pageId="7" pageNumber="484">Fig. 4</figureCitation>
|
||
). The two last one are approximated like on the modern nautiluses, the angular distance falls under 10°. The second indication of maturity is shown by the increase in the relative umbilical width close to adulthood size when U/D reaches 20% on the largest specimens (Table 1;
|
||
<figureCitation id="30822A58FFDABE69FE05FA47FE49FA35" box="[428,489,1459,1485]" captionStart="FIG" captionStartId="8.[223,234,711,728]" captionTargetBox="[355,1236,218,670]" captionTargetId="graphics-609@8.[445,1223,226,598]" captionTargetPageId="8" captionText="FIG. 6. — Ratio of the width of the umbilicus compared with the size of the shell of Micronautilus evolutus n. gen., n. sp. (2021.BR.N.066)." pageId="7" pageNumber="484">Fig. 6</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |