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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne &amp; Haring, Elisabeth" docDate="2021" docId="BA38914980815E608676622E40244051" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Charpentieria stenzii subsp. letochana" docType="treatment" docVersion="5" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1732485878053" updateUser="ExternalLinkService">
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<mods:title id="CEE9026BE46F7196DC1E281FBD10A764">Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:namePart id="73AC691C59A4F628FE298C2223E84470">De Mattia, Willy</mods:namePart>
<mods:nameIdentifier id="E85313BFA51336794AA38DF6BA74DC59" type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation id="B3A365D149735A524C73189FEFD2DC8E">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier id="E78F2391462B81073B3DA0796923B5B6" type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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<mods:namePart id="09D6955D1551907FCA8AB0A82C48687A">Reier, Susanne</mods:namePart>
<mods:affiliation id="4FCDF8186F4ED1AEFE1DBA6B9CB23DAE">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:namePart id="20E14E3D8CFE0DEABBFB3648D9E2DA25">Haring, Elisabeth</mods:namePart>
<mods:affiliation id="A4EE694D21EB2A900C669B2D6F473A84">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:date id="B78EC9464E98051A9F3FCB86DCB1DF36">2021</mods:date>
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<mods:number id="A557DD64B35AA5A48C34BC55FE20716B">1077</mods:number>
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<mods:identifier id="254FC48235AD846AAD50525C7C3B693A" type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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<treatment id="BA38914980815E608676622E40244051" ID-GBIF-Taxon="191645944" LSID="urn:lsid:plazi:treatment:BA38914980815E608676622E40244051" httpUri="http://treatment.plazi.org/id/BA38914980815E608676622E40244051" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Clausiliidae" scope_infraOrder="Clausilioidei" scope_order="Stylommatophora" scope_phylum="Mollusca" scope_superFamily="Clausilioidea">
<subSubSection id="34C631CE219CC23E6661CB739BC60456" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="2E9E030A0B029C9C088A59EF61C34F04" pageId="0" pageNumber="1">
<taxonomicName id="A5432EE0C4740CAA114455A52551C06F" ID-CoL="7GNW3" LSID="BA389149-8081-5E60-8676-622E40244051" authority="(Gredler, 1874)" baseAuthorityName="Gredler" baseAuthorityYear="1874" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. letochana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="letochana">Charpentieria stenzii letochana (Gredler, 1874)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3C9A9391280B14B727A9BA8590FE828D" pageId="0" pageNumber="1" type="description">
<paragraph id="0601833BDE5C310948BF6DFC5ED06245" pageId="0" pageNumber="1">
<figureCitation id="79EAE65F5B2D0DC252796DE18E24F2CF" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Charpentieria stenzii letochana (Gredler, 1874), Val Fonda, Belluno 54.1 whole distal genital organs 54.2 internal distal part of genital organs 54.3 detail of the hemipapilla: uncut 54.4 detail of the hemipapilla: cut funnel 54.5 cross section of the pseudopapilla 54.6 cross section of the epiphallus." figureDoi="10.3897/zookeys.1077.67081.figure54" httpUri="https://binary.pensoft.net/fig/622846" pageId="0" pageNumber="1">Figs 54.1-54.6, 56.5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="390E28076D306D7B8C0038086D4AED1D" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="FDC76E3134F783C741878B3BF57C8FDD" pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph id="CF4C1BD1A1F0D19FDEC675AB83D537AE" pageId="0" pageNumber="1">
<materialsCitation id="A48E17E3A13A6B55EB9305AFC0004BFB" accessionNumber="MW758957" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Lab ID, W. De Mattia, J. Macor" country="Italy" county="Belluno" elevation="1500" latitude="46.607616" location="Val Fonda." longLatPrecision="1" longitude="12.205558" municipality="Misurina" specimenCount="1" stateProvince="Veneto">
<collectingCountry id="9FEDDFAE50CA62427B2FEDD50FDD1EF6" name="Italy">Italy</collectingCountry>
,
<collectingRegion id="2162B97B10A855E6C70ECA799CEC3CB9" country="Italy" name="Veneto">Veneto</collectingRegion>
,
<collectingCounty id="89343C5E037BE6DDEB826FD45880CB48">Belluno</collectingCounty>
,
<collectingMunicipality id="C691198731090E86B2FCEB5431E2984B">Misurina</collectingMunicipality>
,
<location id="33E4D158BEA366E347AA702AE5C53073" LSID="urn:lsid:plazi:treatment:BA38914980815E608676622E40244051:33E4D158BEA366E347AA702AE5C53073" country="Italy" county="Belluno" latitude="46.607616" longLatPrecision="1" longitude="12.205558" municipality="Misurina" name="Val Fonda." stateProvince="Veneto">Val Fonda.</location>
<elevation id="C03C5BCA4F9DDDC49BEC9D29A92D3567" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
<quantity id="0D155453BB960C91D5A21423E61781E3" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</quantity>
asl
</elevation>
,
<geoCoordinate id="DDCC2597C48D398525C102D84218A6B0" degrees="46" direction="north" minutes="36" orientation="latitude" precision="1" seconds="27.42" value="46.607616">46°36'27.42&quot;N</geoCoordinate>
,
<geoCoordinate id="08C02486EBD64923064EBD15417A249D" degrees="12" direction="east" minutes="12" orientation="longitude" precision="1" seconds="20.01" value="12.205558">12°12'20.01&quot;E</geoCoordinate>
, [
<collectorName id="77A76B5FF9617CBBDD669542DA996D5F">Lab ID</collectorName>
C19_1, COI:
<accessionNumber id="9A4962BA1189A6643F5E1AE2C15D1022" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758957">MW758957</accessionNumber>
],
<collectorName id="97D261E4D543E6894B17479E05C87AE1">W. De Mattia</collectorName>
and
<collectorName id="B1A8775141D6DCD452CDF70567F340B5">J. Macor</collectorName>
leg. and det., 2 dissected spm
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D61334AFA3FD0203658D782653499008" pageId="0" pageNumber="1" type="distribution">
<paragraph id="AB86D6BC8DC2493859D6714C5BE85934" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="5E1EAC18C6E455C1CFB4D71C20272427" pageId="0" pageNumber="1">
<taxonomicName id="2D99AC7009AD44309A228403F714A62A" baseAuthorityName="Gredler" baseAuthorityYear="1874" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. letochana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="letochana">
<emphasis id="0438BC7C2A21B7059501AE917F2D7ED8" italics="true" pageId="0" pageNumber="1">Charpentieria stenzii letochana</emphasis>
</taxonomicName>
is exclusively known from Val Fonda (Belluno, Veneto) and Val Pusteria (
<normalizedToken id="F3CABCF36775D2D52E2039A563753C99" originalValue="Schönleitental">Schoenleitental</normalizedToken>
) (Bolzano, Trentino-Alto Adige), Italy (
<bibRefCitation id="0039118A1F6B04B681AFA8C37B0C64BB" author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">Nordsieck 1963b</bibRefCitation>
: 198).
</paragraph>
</subSubSection>
<subSubSection id="F39B331ED229B9F80DEAA141EF226C8A" pageId="0" pageNumber="1" type="external morphology of the genital organs">
<paragraph id="DF7F52D2C2D886AEEA89D84600800975" pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
<paragraph id="46BBFB6E379FF9E700E7E148DC847F23" pageId="0" pageNumber="1">
<emphasis id="D21BF48DB37D580FBF032BA5DC4A580C" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="9DE37BD9158BF4E12BC74FDE0BDFE811" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Charpentieria stenzii letochana (Gredler, 1874), Val Fonda, Belluno 54.1 whole distal genital organs 54.2 internal distal part of genital organs 54.3 detail of the hemipapilla: uncut 54.4 detail of the hemipapilla: cut funnel 54.5 cross section of the pseudopapilla 54.6 cross section of the epiphallus." figureDoi="10.3897/zookeys.1077.67081.figure54" httpUri="https://binary.pensoft.net/fig/622846" pageId="0" pageNumber="1">54</figureCitation>
.1).
</emphasis>
The V is shorter than the FO. The FDBC is as long as the SDBC+BC. The BC+SDBC is slim, club-like in shape, with a clear distinction between the SDBC and the BC. The D is much longer than the SDBC+BC and thinner. The V is short and thin in diameter. The PC is ~ 2
<normalizedToken id="06BCF30BFAEA3680113DD73332CDBF3D" originalValue="×">x</normalizedToken>
longer than the V. The P is cylindrical and slightly swollen along its proximal part. The transition between P and EP is clearly visible. The PR is long and robust. The E is as long as the P but much thinner in diameter.
</paragraph>
</subSubSection>
<subSubSection id="AAF65E4EB9F49A3FCBAA96D31E8C5366" pageId="0" pageNumber="1" type="internal morphology of the genital organs">
<paragraph id="4E832FBF3756C5B7BC9062C723EB7655" pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
<paragraph id="355AAC09377C35C38E49835A39EF6D6E" pageId="0" pageNumber="1">
<emphasis id="524982C416A297080318C6C82823091A" bold="true" pageId="0" pageNumber="1">
(Figs
<figureCitation id="D9266B5768BC212A12164C0548C32AB9" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Charpentieria stenzii letochana (Gredler, 1874), Val Fonda, Belluno 54.1 whole distal genital organs 54.2 internal distal part of genital organs 54.3 detail of the hemipapilla: uncut 54.4 detail of the hemipapilla: cut funnel 54.5 cross section of the pseudopapilla 54.6 cross section of the epiphallus." figureDoi="10.3897/zookeys.1077.67081.figure54" httpUri="https://binary.pensoft.net/fig/622846" pageId="0" pageNumber="1">54</figureCitation>
.2-54.6).
</emphasis>
The V presents irregular smooth pleats. The A has a fleshy, mushroom-like fold. The P shows many irregular smooth to fringed pleats. The hemipapilla is short, smooth to moderately folded. The outlet of the hemipapilla is slit-like. The E shows two moderately fringed ELP. They proximally fade before the VD.
</paragraph>
</subSubSection>
</treatment>
</document>