treatments-xml/data/87/00/B5/8700B5F4C3D35D11A71AB708ED412EBD.xml

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<mods:title id="4C052719CB5D937E4BF4C815D704D015">Two new species of Thomasomys (Cricetidae: Sigmodontinae) from the western Andes of Ecuador and an updated phylogenetic hypothesis for the genus</mods:title>
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<mods:namePart id="71B63C08200C9A52A058088A39BC8D0F">Brito, Jorge</mods:namePart>
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<mods:namePart id="354C4D77F1E16D4CBF4FF0BB44A1464C">García, Rubí</mods:namePart>
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<mods:namePart id="360307ED66C14128C170A4BE39031DF2">Castellanos, Francisco X.</mods:namePart>
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<mods:affiliation id="3031C8384306E6ECF6754C43BC687658">Instituto Nacional de Biodiversidad (INABIO), Quito, Ecuador &amp; Department of Biological Sciences, Texas Tech University, Lubbock, Texas, United States</mods:affiliation>
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<mods:namePart id="DAB735E5392A84D24EA914302E17CFD6">Gavilanes, Gabriela</mods:namePart>
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<mods:affiliation id="2737F76E69604FCA55B50F45E9D2E665">Laboratorio de Biología Evolutiva, Colegio de Ciencias Biológicas y Ambientales COCIBA, Universidad San Francisco de Quito USFQ, Quito, Cumbayá, Ecuador</mods:affiliation>
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<mods:namePart id="0902AD17CAD485C5CD17C1D1C6BE0F52">Curay, Jenny</mods:namePart>
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<mods:affiliation id="9DB95277C312571EFEF76B54A28F8C83">Instituto Nacional de Biodiversidad (INABIO), Quito, Ecuador &amp; Instituto de Diversidad y Evolución Austral (IDEAus — CONICET), Puerto Madryn, Argentina</mods:affiliation>
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<mods:namePart id="DED348EE1E8CD2D2980CE03DF3EA958C">Carrión-Olmedo, Julio C.</mods:namePart>
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<mods:namePart id="4D17B28AEB4A120595661067AF79310F">Reyes-Barriga, Daniela</mods:namePart>
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<mods:namePart id="9A79F136A77B3CAFA03A6CDD7E83F9D5">Guayasamin, Juan M.</mods:namePart>
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<mods:namePart id="152FFE72AB7674967FDBCCD0CA1E0CB2">Salazar-Bravo, Jorge</mods:namePart>
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<mods:affiliation id="82E263B55B533A39016588E8FC26FD57">Department of Biological Sciences, Texas Tech University, Lubbock, Texas, United States</mods:affiliation>
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<mods:namePart id="7AE7D877CA33E548622969DD5D92711D">Pinto, C. Miguel</mods:namePart>
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<mods:affiliation id="DE67EC25BCB5D5B08F183C7106875318">Charles Darwin Research Station, Charles Darwin Foundation, Puerto Ayora, Galápagos, Ecuador</mods:affiliation>
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<mods:date id="485E30B3BB8D51F7960B4CA38EDAA2F0">2024</mods:date>
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<mods:number id="0D4600261061C50F54675BE5279AC54A">74</mods:number>
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<subSubSection id="CFA560C5C18F302E86C6466BCF3376AB" type="nomenclature">
<paragraph id="7E39B5549AE18A7DB793CA7981ACE598">
<taxonomicName id="808201BC5B16520E00D724AD19AB0759" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="02A479A035B30C84C9BFB7D7D80FB10F" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D51B515E63EB15F523B39B7EF916046" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZGBG" type="Holotype">
<paragraph id="A00D0765D7908AAA51E6C62BADC93431">
<heading id="D6C3AC0E257825426BB8235EC8E260C8" reason="title">
<typeStatus id="3377BF47CA9FA55FE7560F30406C08FC" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="CC15EDAD37EF82BE9300867C33805E2D">
<materialsCitation id="A470199C3F835B6C8F9FF3185B0082A3" collectingDate="2022-01-19" collectionCode="MECN" collectorName="Jorge Brito &amp; Rubi Garcia &amp; Fausto Rodriguez" country="Ecuador" elevation="3690" latitude="0.15456" location="Area de Proteccion Hidrica Otavalo Mojanda" longLatPrecision="1" longitude="-78.27536" specimenCode="MECN 6912" specimenCount="1" specimenCount-female="1" stateProvince="Provincia de Imbabura" typeStatus="holotype">
<specimenCode id="97038CD6EDD7B0929EA4BE2CC30C2EB3" collectionCode="MECN" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">
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6912
</specimenCode>
(field number JBM 2574), an 
<specimenCount id="03656B4807E963E957C266809BBB41E0" count="1" type="female">adult female</specimenCount>
collected 
<collectingDate id="93636FB6F65F698266A4AE10E6DB0F91">19 January, 2022</collectingDate>
, by 
<collectorName id="320E0A10E65859725367C48AFB1DC8AC">Jorge Brito</collectorName>
, 
<collectorName id="C839EBF8FC8E041582FF9682F860DA47">Rubí García</collectorName>
and 
<collectorName id="2E40C5896E62F47AF80C5B91DF8D00D3">Fausto Rodríguez</collectorName>
, preserved as dry skin, skull, postcranial skeleton, and muscle and liver biopsies in 95 % ethanol.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EHHBG" type="description">
<paragraph id="4E0E7E8162AB04C22F08F2BDB34F5CA3">
<heading id="BD05E81FD51EC9049408A6FFE43C9108" reason="title">
Measurements of the 
<typeStatus id="21D004FDFA28E32F2F247DCE64216D02" type="holotype">holotype</typeStatus>
(in mm, mass in g).
</heading>
</paragraph>
<paragraph id="50A882C802D3C6DD292954D9075BFBB4">
<abbrev id="ABBRID0ENHBG" xlink_title="head and body length">HB</abbrev>
= 120; 
<abbrev id="ABBRID0ERHBG" xlink_title="tail length">TL</abbrev>
= 161; 
<abbrev id="ABBRID0EVHBG" xlink_title="hind foot length">HF</abbrev>
= 29, E = 19; W = 36; 
<abbrev id="ABBRID0EZHBG" xlink_title="Condylo-incisive length">CIL</abbrev>
= 28.62; 
<abbrev id="ABBRID0E4HBG" xlink_title="length of upper diastema">LD</abbrev>
= 8.49; 
<abbrev id="ABBRID0EBIBG" xlink_title="crown length of maxillary toothrow">LM</abbrev>
= 4.7; 
<abbrev id="ABBRID0EFIBG" xlink_title="breadth of the first maxillary molar">
<collectionCode id="EA525DA75F37D37D95553D1CA647A6C6" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
1
</abbrev>
= 1.53; 
<abbrev id="ABBRID0EJIBG" xlink_title="length of incisive foramina">LIF</abbrev>
= 5.54; 
<abbrev id="ABBRID0ENIBG" xlink_title="breadth of incisive foramina">BIF</abbrev>
= 2.11; 
<abbrev id="ABBRID0ERIBG" xlink_title="breadth of the palatal bridge">BPB</abbrev>
= 3.58; 
<abbrev id="ABBRID0EVIBG" xlink_title="breadth of zygomatic plate">BZP</abbrev>
= 2.54; 
<abbrev id="ABBRID0EZIBG" xlink_title="length of incisive foramina">LIF</abbrev>
= 5.13; 
<abbrev id="ABBRID0E4IBG" xlink_title="zygomatic breadth">ZB</abbrev>
= 16.63; 
<abbrev id="ABBRID0EBJBG" xlink_title="depth of upper incisor">DI</abbrev>
= 1.54; 
<abbrev id="ABBRID0EFJBG" xlink_title="breadth across both upper incisor tips">BIT</abbrev>
= 2.03; 
<abbrev id="ABBRID0EJJBG" xlink_title="length of rostrum">LR</abbrev>
= 11.12; 
<abbrev id="ABBRID0ENJBG" xlink_title="length of nasals">LN</abbrev>
= 11.84; 
<abbrev id="ABBRID0ERJBG" xlink_title="breadth of rostrum">
<collectionCode id="415634952D148A06B38A185C88DA0468" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
</abbrev>
= 5.94; 
<abbrev id="ABBRID0EVJBG" xlink_title="orbital fossa length">OL</abbrev>
= 10.02; 
<abbrev id="ABBRID0EZJBG" xlink_title="breadth of mesopterygoid foramen">BMF</abbrev>
= 2, 29; 
<abbrev id="ABBRID0E4JBG" xlink_title="breadth between occipital condyles">BCO</abbrev>
= 7, 54; 
<abbrev id="ABBRID0EBKBG" xlink_title="braincase breadth">BCB</abbrev>
= 14.67; 
<abbrev id="ABBRID0EFKBG" xlink_title="length of mandible">LMN</abbrev>
= 19.1; 
<abbrev id="ABBRID0EJKBG" xlink_title="depth of mandibular ramus">DR</abbrev>
= 3.3. External and craniodental measurements of additional specimens are listed in Table 
<tableCitation id="82869540AE03BBFECB5CF5A26A48793F" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of morphometric measurements of all specimens in mm. Species names are accompanied by the number of analyzed individuals between parentheses. Minimum and maximum values are accompanied by mean and standard deviation between parentheses. Abbreviations of characters are detailed in the text." httpUri="http://table.plazi.org/id/B5A75659D53B773ADE5A518BE89C41B5" tableUuid="B5A75659D53B773ADE5A518BE89C41B5">4</tableCitation>
.
</paragraph>
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<paragraph id="5034F59D717E704F2C2A01E17CD54C91">
<label id="C578034D5F1BDF25E8576774B0C16B66">Table 4.</label>
</paragraph>
<paragraph id="AEBDB60641B7B0D822A098D17201FE39">Summary of morphometric measurements of all specimens in mm. Species names are accompanied by the number of analyzed individuals between parentheses. Minimum and maximum values are accompanied by mean and standard deviation between parentheses. Abbreviations of characters are detailed in the text.</paragraph>
</caption>
<paragraph id="0E907B4D4A6CE9F449BF9990D94FAB25">
<table id="TID0EHQCI" rules="all">
<tbody id="5EDBE6C253D6362956DD22CC1D9997C0">
<emphasis id="1CB054240E5588D0228920F91AEC9235" bold="true">
<tr id="BAE66E7012EF49EF8D57ACD619045B03">
<td id="A126879D7CBE7BA8732ABAA5D2A86153" colspan="1" isEmpty="true" rowspan="1">-</td>
<td id="4DDDC9FB15CBD3B1B470E71F99D6D677" colspan="1" rowspan="1">
<taxonomicName id="4844E9FD4C2933ED5188C7CE181A8376" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="C338258B3DA7EDE6FEA67F6D7A66E77E" bold="true" italics="true">T. igor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9BB8CAFAE01836EB4549592150E8458A" rank="species">sp. nov.</taxonomicNameLabel>
(4)
</td>
<td id="150383F82639617259CFC2D19740C8B7" colspan="1" rowspan="1">
<taxonomicName id="830B1109F8B9706F8619421A323EB09B" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="49392CF2DE53BB16E322D5AE358C9116" bold="true" italics="true">T. otavalo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="535509AC0814B57E898DDFC9F0D457C0" rank="species">sp. nov.</taxonomicNameLabel>
(14)
</td>
<td id="C0F2AFEAFCE7ABCC40CD3667BBF96D07" colspan="1" rowspan="1">
<taxonomicName id="FFF246B4125E6D3E17244C6C14453307" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="3DAB33AE335D4B85D91E10F613F1D21D" bold="true" italics="true">T. silvestris</emphasis>
</taxonomicName>
(30)
</td>
<td id="7429CD0FE3291934D087206E46021295" colspan="1" rowspan="1">
<taxonomicName id="102CC2210E306CA425BD50F0B03A0B29" authorityName="Voss" authorityYear="2003" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="D2CC6F8ABA93E1BC91E358B8DB622BAE" bold="true" italics="true">T. ucucha</emphasis>
</taxonomicName>
(22)
</td>
</tr>
<tr id="F664954AA530F7574791BDE46C90A105">
<td id="4B5207050ED476EC03E2E254192C1DDA" colspan="1" rowspan="1">
<abbrev id="ABBRID0EPNBG" xlink_title="head and body length">HB</abbrev>
</td>
<td id="46B36A7ECD1D221AA9D58055E8ED8391" colspan="1" rowspan="1">102–122 (113.75 ± 8.66)</td>
<td id="BA29E97B7236F08BFBF77384A965895A" colspan="1" rowspan="1">110–125 (115.79 ± 4.68)</td>
<td id="C1BDCCBFADB66B2DD4F62E3FF17FC727" colspan="1" rowspan="1">93–138 (109.17 ± 10.65)</td>
<td id="2F7089E32B318391DE8F929C85A57466" colspan="1" rowspan="1">91–117 (105.93 ± 6.64)</td>
</tr>
</emphasis>
<tr id="2276A65ADC5D130330E4AECA71834322">
<td id="B7E2B07BC504541CAB3CAB7E2C9CDF97" colspan="1" rowspan="1">
<emphasis id="CC868B5C6D35367A2C6A0E6D6F4BD521" bold="true">LT</emphasis>
</td>
<td id="263EBB9D7D733EBF4A5E8B1CF1492EC4" colspan="1" rowspan="1">133–160 (147.25 ± 14.77)</td>
<td id="71A1088924808A41C6164B0D7FCBC802" colspan="1" rowspan="1">150–180 (166.79 ± 8.66)</td>
<td id="24CA518BCFBDD5802A60FA8D122A26C1" colspan="1" rowspan="1">130–160 (146.93 ± 8.05)</td>
<td id="745509B37C9D4B516FEEEB3759431864" colspan="1" rowspan="1">127–153 (140.54 ± 5.45)</td>
</tr>
<tr id="22FBB007E6D2472D376D4FB569E1F62B">
<td id="6A765ED0FF4F6E7D2EEEE14061B6D953" colspan="1" rowspan="1">
<abbrev id="ABBRID0EYOBG" xlink_title="hind foot length">
<emphasis id="43EDE8DD53F573CF40D146A0887B594B" bold="true">HF</emphasis>
</abbrev>
</td>
<td id="AC0F3CE0839800F47D834795832AE981" colspan="1" rowspan="1">27–29 (28.25 ± 0.96)</td>
<td id="41100D60833319FC8B19912C5810E6AA" colspan="1" rowspan="1">27–31 (29.43 ± 1.02)</td>
<td id="7BD0DFD2C37F84DE0451A0A128267AF6" colspan="1" rowspan="1">24–30 (27.33 ± 1.63)</td>
<td id="495111A3F0F1E21E7FF80431EA578492" colspan="1" rowspan="1">18–30 (26.31 ± 2.39)</td>
</tr>
<tr id="D0ACC494A1B462945673CC68EF7ADBA7">
<td id="0216CB307F039E146925C432F1854F84" colspan="1" rowspan="1">
<emphasis id="56012DE29F124A04214D72A27529467E" bold="true">E</emphasis>
</td>
<td id="AFCBA6504398CF585BF19FEC6D04DAE0" colspan="1" rowspan="1">18–20 (19.25 ± 0.96)</td>
<td id="CCD2A14A1060199139A4996BDC6488F5" colspan="1" rowspan="1">18–20 (18.86 ± 0.53)</td>
<td id="06E1471151513DD21C9B01AC1D93F02B" colspan="1" rowspan="1">16–21 (18.98 ± 1.34)</td>
<td id="C1CA941EC60FE80459F5F169A75DAA17" colspan="1" rowspan="1">15–28 (19.07 ± 2.38)</td>
</tr>
<tr id="37C95915612B3CE8B8B3AB1E4AFF0329">
<td id="13FFBDE5A7724F720DDDB836083FFD5A" colspan="1" rowspan="1">
<abbrev id="ABBRID0EBQBG" xlink_title="Condylo-incisive length">
<emphasis id="9347413892AFCD8CFCB07268E5C2BAF2" bold="true">CIL</emphasis>
</abbrev>
</td>
<td id="88827F6064E86F5EC13749A886A2B3B2" colspan="1" rowspan="1">25.94–27.48 (27.06 ± 0.75)</td>
<td id="92A203C66BA9E8031A08624A6276A26A" colspan="1" rowspan="1">28.05–30.29 (28.86 ± 0.83)</td>
<td id="10606B907EB1760595649327AAD530A4" colspan="1" rowspan="1">25.21–27.66 (26.5 ± 0.55)</td>
<td id="EBED662DCCFA9FCA1E3AFC0CD31FC47F" colspan="1" rowspan="1">27.25–29.34 (28.45 ± 0.51)</td>
</tr>
<tr id="BBA995B30A1672721A8CCC1CFC198639">
<td id="7A17157EBB57F97C555941C113E31A62" colspan="1" rowspan="1">
<abbrev id="ABBRID0EXQBG" xlink_title="length of upper diastema">
<emphasis id="DCE0B37EE464611640E0AA99D8D8258E" bold="true">LD</emphasis>
</abbrev>
</td>
<td id="89885C2AA7C837EF8492636E9B6BC839" colspan="1" rowspan="1">7.53–8.54 (8.2 ± 0.47)</td>
<td id="8DDB0F42C3149A65095D152C6C260CA8" colspan="1" rowspan="1">8.13–9.98 (9.2 ± 0.7)</td>
<td id="622C6CD7AA589899458CDB1F067E510E" colspan="1" rowspan="1">6.65–8.32 (7.71 ± 0.39)</td>
<td id="2A8C9B4621A8DF2899A9317E893CBB95" colspan="1" rowspan="1">8.4–9.94 (9.1 ± 0.34)</td>
</tr>
<tr id="432E5E8440EF866C458C2BF91CD4E0B6">
<td id="4DA5CE9772FF21390151ECE45343F881" colspan="1" rowspan="1">
<abbrev id="ABBRID0ENRBG" xlink_title="crown length of maxillary toothrow">
<emphasis id="73B3DF4183C7BA04C9C4CAD2250CDC74" bold="true">LM</emphasis>
</abbrev>
</td>
<td id="D79F22748134584782CDCCCED66AB6E1" colspan="1" rowspan="1">4.74–4.83 (4.8 ± 0.04)</td>
<td id="10F6E456219B928C9AC812EEC6752E80" colspan="1" rowspan="1">4.02–5.44 (4.5 ± 0.4)</td>
<td id="91F5D142D532FC59447BCA362348FDCC" colspan="1" rowspan="1">4.41–5.05 (4.73 ± 0.14)</td>
<td id="8D440099E0909E5D9324E145BC87F1FE" colspan="1" rowspan="1">4.29–5.5 (4.57 ± 0.24)</td>
</tr>
<tr id="F1D73959FE32A9954B2659357F3DC0C1">
<td id="63B752C84715F058111AF25ADF64B891" colspan="1" rowspan="1">
<abbrev id="ABBRID0EDSBG" xlink_title="breadth of the first maxillary molar">
<emphasis id="E360AD2A91176347156247FAAA3B9646" bold="true">BM 1</emphasis>
</abbrev>
</td>
<td id="B82CB7C32F585B540F5428A909AAD940" colspan="1" rowspan="1">1.55–1.63 (1.6 ± 0.03)</td>
<td id="A0F7E1E05FA6A4EA027B0E57D46C51E2" colspan="1" rowspan="1">1.41–1.63 (1.52 ± 0.08)</td>
<td id="EE3B548C33D2601FC17B3FACF864896B" colspan="1" rowspan="1">1.47–1.68 (1.58 ± 0.06)</td>
<td id="3353000161A857321EFDB4D5A3D9BD6B" colspan="1" rowspan="1">1.43–1.7 (1.51 ± 0.06)</td>
</tr>
<tr id="2CA6F89E9F5E2C40267FF928D8BE38DE">
<td id="8E0880C089C5232B983265E01F223459" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZSBG" xlink_title="length of incisive foramina">
<emphasis id="D950B8E2BFF81A633B126FA5A7E8DD1E" bold="true">LIF</emphasis>
</abbrev>
</td>
<td id="C79DB33A796F8F389C6FCC3E81DCD0EA" colspan="1" rowspan="1">5.53–6.28 (5.86 ± 0.31)</td>
<td id="54D4CFC205D96254BFE3388E138212E2" colspan="1" rowspan="1">5.11–7.09 (5.89 ± 0.77)</td>
<td id="A69D861F6F232C3FF953333594056CC1" colspan="1" rowspan="1">4.8–5.71 (5.19 ± 0.25)</td>
<td id="66B70FD50043E28E7BAACCD6CEE3A025" colspan="1" rowspan="1">4.97–5.65 (5.25 ± 0.2)</td>
</tr>
<tr id="37D7DE5AE1EEAE175A460C7046F00F16">
<td id="E7D2F84C27D433F35A70FD8477C0DF5F" colspan="1" rowspan="1">
<abbrev id="ABBRID0EPTBG" xlink_title="breadth of incisive foramina">
<emphasis id="215E714AD445C3E7881F0C20BFD86045" bold="true">BIF</emphasis>
</abbrev>
</td>
<td id="65CAFB31520BF243EA3A5451F24999A3" colspan="1" rowspan="1">1.75–2.21 (1.98 ± 0.22)</td>
<td id="141A0C3986B7649D076B557253AB664C" colspan="1" rowspan="1">1.85–2.36 (2.17 ± 0.14)</td>
<td id="A429F24AE13EB397A36B233574E0E48B" colspan="1" rowspan="1">1.81–2.48 (2.12 ± 0.14)</td>
<td id="D3BAB4E0D1356E006FBFD8099FD4A463" colspan="1" rowspan="1">1.89–2.32 (2.15 ± 0.12)</td>
</tr>
<tr id="D1DCC612AEBF1DFA3A89E868FC3158F0">
<td id="93757884D41D1FFD9D3B0B5F7FDF7940" colspan="1" rowspan="1">
<abbrev id="ABBRID0EFUBG" xlink_title="breadth of the palatal bridge">
<emphasis id="674BD1377783343A28A112007D847C9E" bold="true">BPB</emphasis>
</abbrev>
</td>
<td id="E2F7EF4DD0DDCE180106A2A63220281A" colspan="1" rowspan="1">3.06–3.53 (3.22 ± 0.22)</td>
<td id="1F29C122D800378A2D9E009721FBED9A" colspan="1" rowspan="1">3.45–4.56 (3.96 ± 0.3)</td>
<td id="A74524CEC52B4EFB142A781EE260A831" colspan="1" rowspan="1">2.16–3.78 (3.25 ± 0.31)</td>
<td id="24F4E46FAC72C6190695C3FE488EB1F3" colspan="1" rowspan="1">3.48–4.22 (3.76 ± 0.18)</td>
</tr>
<tr id="35F7FBB46ECBC25C5715C805F67903AA">
<td id="7E9F7C3A66616B8524C5F310A4EBFFC6" colspan="1" rowspan="1">
<abbrev id="ABBRID0E2UBG" xlink_title="breadth of zygomatic plate">
<emphasis id="8B9556D872B5FA89238D6BE78F912CCB" bold="true">BZP</emphasis>
</abbrev>
</td>
<td id="7F0579104647B78F366F5F478A202E5D" colspan="1" rowspan="1">2.14–2.27 (2.17 ± 0.06)</td>
<td id="A05F221D501A7293931B1B0BFD69F4A9" colspan="1" rowspan="1">2.28–2.93 (2.53 ± 0.16)</td>
<td id="9CB3A31418E502F7061BD5A4CE44E93A" colspan="1" rowspan="1">1.69–2.28 (2.08 ± 0.13)</td>
<td id="4971AA39DEE3DE4B70102D25EB47E242" colspan="1" rowspan="1">2.36–2.73 (2.56 ± 0.11)</td>
</tr>
<tr id="86E42DE32C81F44B7EB3CC746323F73F">
<td id="5606CDE925D6CF9BB774985D8D624172" colspan="1" rowspan="1">
<abbrev id="ABBRID0ERVBG" xlink_title="least interorbital breadth">
<emphasis id="DF8E07F773B1E87CE938BDEEB1EC2DB4" bold="true">LIB</emphasis>
</abbrev>
</td>
<td id="CBB32F73FF1EAD849712A32C91AEB496" colspan="1" rowspan="1">5.15–5.21 (5.18 ± 0.03)</td>
<td id="572A6F58F8E91AB0689472DB25AB8654" colspan="1" rowspan="1">4.1–5.51 (4.75 ± 0.52)</td>
<td id="DCBBC15C2265AA547334E67053F4839A" colspan="1" rowspan="1">4.8–5.56 (5.09 ± 0.18)</td>
<td id="D54ED5C79520FD8E1171C5B130970CD2" colspan="1" rowspan="1">4.65–5.17 (4.9 ± 0.12)</td>
</tr>
<tr id="FC67847147C37A28BC285529518263C7">
<td id="6802A56579A4E058892BFA41DA52B564" colspan="1" rowspan="1">
<abbrev id="ABBRID0EHWBG" xlink_title="zygomatic breadth">
<emphasis id="76D9A62E46C1C4E4A6907F60E7C3131E" bold="true">ZB</emphasis>
</abbrev>
</td>
<td id="32BDD3C81EC9AC54117DD6806395EB67" colspan="1" rowspan="1">14.62–15.27 (15.05 ± 0.29)</td>
<td id="03703A3EB815A0634A8DADB75EA2D03F" colspan="1" rowspan="1">14.55–17.73 (16.19 ± 1.22)</td>
<td id="87EEA0FB74348FCAA79E2E8117B4F7A3" colspan="1" rowspan="1">14–15.84 (15.12 ± 0.46)</td>
<td id="BCADDC40F840E9E034E61989EACBBA81" colspan="1" rowspan="1">16–17.46 (16.69 ± 0.33)</td>
</tr>
<tr id="1AB37127C088061448EA722F5AD886BC">
<td id="261D26E5A017FB1695D0A832AA22EF4D" colspan="1" rowspan="1">
<abbrev id="ABBRID0E4WBG" xlink_title="depth of upper incisor">
<emphasis id="B98115BB102ED5EA695873E7DCACCCEC" bold="true">DI</emphasis>
</abbrev>
</td>
<td id="447641B1EB14787F787DD5679802FC95" colspan="1" rowspan="1">1.35–1.67 (1.48 ± 0.14)</td>
<td id="E76A017ACAD6C26673FA155BDEB2C8D5" colspan="1" rowspan="1">1.43–1.68 (1.53 ± 0.07)</td>
<td id="D5952854482FB0B239D33061FE523DBA" colspan="1" rowspan="1">– 3.42–2.1 (1.28 ± 0.9)</td>
<td id="AA694AA2F243BE484761E7737A516264" colspan="1" rowspan="1">1.51–159 (7.93 ± 31.47)</td>
</tr>
<tr id="CB32DFAB182C81C2DEBFCA4814790FFB">
<td id="F8F84CAC082306EAF768C6E70E91EE01" colspan="1" rowspan="1">
<abbrev id="ABBRID0ETXBG" xlink_title="breadth across both upper incisor tips">
<emphasis id="60F924E03BA7ED7AE6868269247DFEE4" bold="true">BIT</emphasis>
</abbrev>
</td>
<td id="95109F403ACC040AF19D83DDFAE65596" colspan="1" rowspan="1">1.62–1.83 (1.77 ± 0.1)</td>
<td id="34E0A3FC3128CFD575C3C0D457AF2D28" colspan="1" rowspan="1">1.75–2.14 (1.98 ± 0.11)</td>
<td id="0807389B5B34AF0BEA66278C448C807F" colspan="1" rowspan="1">1.56–1.91 (1.75 ± 0.09)</td>
<td id="5F063B8D4CE63FB876A922B8929A5107" colspan="1" rowspan="1">2.08–2.6 (2.24 ± 0.12)</td>
</tr>
<tr id="A2E5249BEDBD14E149A35DA9A6D62EBD">
<td id="D050B4507E79CA0EAFAE224A93C25C6D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EJYBG" xlink_title="length of rostrum">
<emphasis id="74E6BBDD7A2BD7E1ACA08BC6C8B4AB05" bold="true">LR</emphasis>
</abbrev>
</td>
<td id="48C7E4D5792EA520B9E16AE21A5A1FFE" colspan="1" rowspan="1">8.62–10.57 (9.58 ± 0.8)</td>
<td id="E86ED27BCA3714A2A9EC54C2A6589CC8" colspan="1" rowspan="1">9.71–11.17 (10.4 ± 0.45)</td>
<td id="2B84577259DD9234709F771A35784E24" colspan="1" rowspan="1">9.11–11.89 (10.01 ± 0.73)</td>
<td id="677A707BD2948B2AC3A5E105D4C52FF3" colspan="1" rowspan="1">9.19–10.75 (9.87 ± 0.4)</td>
</tr>
<tr id="D5BC95720CF47914C4E8E80CB85FCC35">
<td id="EADA7038E94ECF2A5A87B6286395FC18" colspan="1" rowspan="1">
<abbrev id="ABBRID0E6YBG" xlink_title="length of nasals">
<emphasis id="2BFD0118ED7D317557708762212D04DB" bold="true">LN</emphasis>
</abbrev>
</td>
<td id="90698125CE3D90D59C5185ADABBF41F9" colspan="1" rowspan="1">10.08–11.61 (11.13 ± 0.71)</td>
<td id="ED0B349BAB1545900160DED12EEB648C" colspan="1" rowspan="1">10.08–11.96 (11.14 ± 0.64)</td>
<td id="28C4E7434002EC71480DA9BBD568002F" colspan="1" rowspan="1">9.55–11.82 (10.68 ± 0.5)</td>
<td id="0C64DB2826D43F024BE0C7FD108EA0B2" colspan="1" rowspan="1">10.15–11.57 (10.89 ± 0.45)</td>
</tr>
<tr id="23A0BE4C40AE865204828D3B9F78347C">
<td id="7A3A7938F2E0DF8AF8B9DD7AE484DA03" colspan="1" rowspan="1">
<abbrev id="ABBRID0EVZBG" xlink_title="breadth of rostrum">
<emphasis id="B52E50D70950E445199BA3494FCDFF18" bold="true">BR</emphasis>
</abbrev>
</td>
<td id="06AE8363C8A3DC8CA73E88399C5DFEC9" colspan="1" rowspan="1">4.96–5.4 (5.19 ± 0.18)</td>
<td id="6E023B239BC3FB604A40A2D55F791278" colspan="1" rowspan="1">5.4–7.07 (6.12 ± 0.7)</td>
<td id="C2F6585DBAEE7E64DF8C7FB5DB35A849" colspan="1" rowspan="1">4.76–5.69 (5.19 ± 0.23)</td>
<td id="16F5E1C07C7DBD745D0D127D369DF0D6" colspan="1" rowspan="1">5.39–6.07 (5.77 ± 0.17)</td>
</tr>
<tr id="91890132457A3505AF302833527BDE87">
<td id="1273E60E3D43234E4D2BC927D91F1053" colspan="1" rowspan="1">
<abbrev id="ABBRID0EL1BG" xlink_title="orbital fossa length">
<emphasis id="1EDC0ADF3BFDC7F3CD088CE31A55C512" bold="true">OL</emphasis>
</abbrev>
</td>
<td id="68286ACEAF7FB59A8B5E240D5C338E42" colspan="1" rowspan="1">7.6–9.3 (8.1 ± 0.8)</td>
<td id="FDCED825650A8D6199EC6B83D6115F52" colspan="1" rowspan="1">9.27–10.35 (9.96 ± 0.26)</td>
<td id="9B1B9B0444C0494C735FE40BD23B2912" colspan="1" rowspan="1">7.27–9.65 (8.37 ± 0.77)</td>
<td id="8C8605D07460E56233684AEC0E135819" colspan="1" rowspan="1">9.57–10.43 (10.1 ± 0.23)</td>
</tr>
<tr id="90E0CAB1E0B865040D069577ECABC38C">
<td id="F6D06D066EE1C6D0C6A5FCE11264711D" colspan="1" rowspan="1">
<abbrev id="ABBRID0EB2BG" xlink_title="breadth between occipital condyles">
<emphasis id="C915327FD3D686BF9A5FA3DC11CAC572" bold="true">BCO</emphasis>
</abbrev>
</td>
<td id="2649128C7BEE13F9B8793FB152B197E7" colspan="1" rowspan="1">6.83–7.14 (7.04 ± 0.15)</td>
<td id="8090D34CB49AF801CCD1A1066989A1CE" colspan="1" rowspan="1">7–7.61 (7.24 ± 0.21)</td>
<td id="AF05B5415FC8C70F623C0FC08C0430D8" colspan="1" rowspan="1">6.32–7.46 (7 ± 0.25)</td>
<td id="3140761927A9E1B7AC07183EE0DE7F0B" colspan="1" rowspan="1">6.89–7.4 (7.09 ± 0.13)</td>
</tr>
<tr id="8E351439CC3616D7DD79E2BCA149ACA9">
<td id="531534FAAB76807467ABC42A33758E99" colspan="1" rowspan="1">
<abbrev id="ABBRID0EX2BG" xlink_title="breadth of mesopterygoid foramen">
<emphasis id="742215E0586E28EF1396B3CA8A33710F" bold="true">BMF</emphasis>
</abbrev>
</td>
<td id="6A92D0942ACB505D56DB9DD7E672BEAE" colspan="1" rowspan="1">1.74–2.52 (2.2 ± 0.35)</td>
<td id="AC8B08FCA85CFEBB0FC282AABA741386" colspan="1" rowspan="1">2.01–2.42 (2.24 ± 0.11)</td>
<td id="8845B004FD622FA04E100C2752669138" colspan="1" rowspan="1">1.85–2.68 (2.26 ± 0.18)</td>
<td id="23B9D3378DB716F97E4762374C45778F" colspan="1" rowspan="1">1.95–2.7 (2.29 ± 0.16)</td>
</tr>
<tr id="8C8024AE48BE7DCA941E599C745AC796">
<td id="12006ABBAACC7B1A2E3279F60427D787" colspan="1" rowspan="1">
<abbrev id="ABBRID0EN3BG" xlink_title="braincase breadth">
<emphasis id="9A4EF7243F13CC426DB3B3A8D00293F5" bold="true">BCB</emphasis>
</abbrev>
</td>
<td id="CF1C68D9759B1806C0AC55747E6EF933" colspan="1" rowspan="1">13.31–13.46 (13.39 ± 0.07)</td>
<td id="F8B909A70B2EFB9A702951A02E943648" colspan="1" rowspan="1">10–15.95 (14.23 ± 1.39)</td>
<td id="6BF48C81AE8C295388B8384DC974475A" colspan="1" rowspan="1">10.27–14.31 (13.23 ± 0.8)</td>
<td id="C2980FA179D716F242A2AC8BCFF36ADE" colspan="1" rowspan="1">13.44–14.55 (14.13 ± 0.25)</td>
</tr>
<tr id="A5EFB7A28A64377C7D2DB325EEC59550">
<td id="0E77BB5078BF7E3CC70E341235F93B5D" colspan="1" rowspan="1">
<abbrev id="ABBRID0ED4BG" xlink_title="length of mandible">
<emphasis id="B6C675B0BEE1A206B48E870365F406F3" bold="true">LMN</emphasis>
</abbrev>
</td>
<td id="B3C6CC229283068ACEEC94B28FEAC741" colspan="1" rowspan="1">14.35–15.07 (14.84 ± 0.33)</td>
<td id="AB51F7D58D8DD2F179F6073014D5FCB2" colspan="1" rowspan="1">14.95–16.64 (15.56 ± 0.42)</td>
<td id="1393EF434ECFC9749C4B38288A3817FB" colspan="1" rowspan="1">13.74–16.88 (15.08 ± 0.76)</td>
<td id="56119429AF71D418D688B31AA46815C6" colspan="1" rowspan="1">14.91–16.6 (15.98 ± 0.4)</td>
</tr>
<tr id="283C47BAF6B2073D49DF4CE1E774112E">
<td id="83E6F81056578C469B367F191EA5E843" colspan="1" rowspan="1">
<abbrev id="ABBRID0EZ4BG" xlink_title="depth of mandibular ramus">
<emphasis id="B7CD08002319F3498C62F1104C0AD3C1" bold="true">DR</emphasis>
</abbrev>
</td>
<td id="F21F1BDEC1676810639784DB508F4939" colspan="1" rowspan="1">3.05–3.25 (3.14 ± 0.09)</td>
<td id="C3387CA375C307A62B9B6AAE6FBB71C7" colspan="1" rowspan="1">2.89–4.72 (3.47 ± 0.59)</td>
<td id="BAA72917E50CD41A797A51E0D085A745" colspan="1" rowspan="1">2.92–3.6 (3.16 ± 0.13)</td>
<td id="7F2BADF848512750682AAB4D1B1A4DD0" colspan="1" rowspan="1">3.4–3.77 (3.62 ± 0.09)</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ5BG" type="type locality">
<paragraph id="5666948A56A37D6A8EB852AB468BFB58">
<heading id="A564EE1803D8AB6D4A28C668D4C8CDE1" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="531E8BFA833C9B27F9BC634056D1928D">
<collectingCountry id="036E14162AE45E5411C29E3CAF851CAB" name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion id="412B1729984722218A01C431CFBFB3CC" country="Ecuador" name="Imbabura">Provincia de Imbabura</collectingRegion>
, 
<location id="6DB3D33CB001E2867641CABEC6EA4437" LSID="urn:lsid:plazi:treatment:8700B5F4C3D35D11A71AB708ED412EBD:6DB3D33CB001E2867641CABEC6EA4437" country="Ecuador" latitude="0.15456" longLatPrecision="1" longitude="-78.27536" name="Area de Proteccion Hidrica Otavalo Mojanda" stateProvince="Provincia de Imbabura">Área de Protección Hídrica Otavalo Mojanda</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ES5BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.275360,0.154560]}">
<geoCoordinate id="5F6FC61A46036A9F2E5A15308C1DFC59" degrees="0.15456" direction="north" orientation="latitude" precision="1" value="0.15456">0.15456</geoCoordinate>
, 
<geoCoordinate id="9F451284DC8131DD603BDFAA07EDCB8D" degrees="78.27536" direction="west" orientation="longitude" precision="1" value="-78.27536">– 78.27536</geoCoordinate>
</named-content>
, WGS 84 coordinates taken by GPS at the site of collection, elevation 
<quantity id="ACE622741BF334A9FEC6199BAAE65040" metricMagnitude="3" metricUnit="m" metricValue="3.69" unit="m" value="3690.0">
<elevation id="F676A3049E7E67FACEEB367CD16D3813" metricMagnitude="3" metricUnit="m" metricValue="3.69" unit="m" value="3690.0">3,690 m</elevation>
</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EX5BG" type="Paratypes">
<paragraph id="BE128E15E53C87B238A3A9398913CE88">
<heading id="37A4398834A0697983B71BFF9746877C" reason="title">
<typeStatus id="9ABAECE8BFFDFC85B3DB30A6116906A9" type="paratype">Paratypes</typeStatus>
.
</heading>
</paragraph>
<paragraph id="5E6EEC77079EC88509A3BDD92100B992">
<named-content id="F93DE0D6BA9102D5F280F4A1EBB1022C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="1D538FB150FA9635253EFB8407B54531" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
6909, 
<named-content id="849E978D743B03B6E5A09C8D39B5853B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="52E7BD15D2CCE5225082373B60EF555E" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
6911, adult males, all preserved as dry skins and cleaned skulls, collected in 
<collectingRegion id="52DE52DF3CAB488CE38A8F1F2C87ABD6" country="Ecuador" name="Imbabura">Provincia de Imbabura</collectingRegion>
, Área de Protección Hídrica Otavalo Mojanda (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQ6BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.275360,0.154560]}">0.15456, – 78.27536</named-content>
, 
<quantity id="470E357B846F5CF15CBAB95A9DAE0E24" metricMagnitude="3" metricUnit="m" metricValue="3.6799999999999997" unit="m" value="3680.0">3,680 m</quantity>
) by J. Brito, M. Yánez and C. Paucar on 19 January, 2022. 
<named-content id="2395BA10EB81C64006E67192324BE48F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="0BB688472E760623764DAD8E269A1FB7" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
6916, adult female, preserved as dry skins and cleaned skull, collected in Área de Protección Hídrica Otavalo Mojanda (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBAAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.282590,-0.158240]}">
<geoCoordinate id="765A3D0BE83A6750894B57AAE03B48A1" degrees="0.15824" direction="south" orientation="latitude" precision="1" value="-0.15824">0.15824 ° S</geoCoordinate>
, 
<geoCoordinate id="7D0DA5FBB09C765E70DBD495519D40A3" degrees="78.28259" direction="south" orientation="latitude" precision="1" value="-78.28259">– 78.28259</geoCoordinate>
</named-content>
, 
<quantity id="8B83E7D5EA046D825B60B2556B915C93" metricMagnitude="3" metricUnit="m" metricValue="3.55" unit="m" value="3550.0">
<geoCoordinate id="5173F7726D9D21C27D58D053A2C43855" degrees="3,550" direction="east" orientation="longitude" precision="55" value="3.55">3,550</geoCoordinate>
m
</quantity>
) by J. Brito on 21 January, 2022. 
<named-content id="59CAF4E2F3A1C175289A1A3E9D209CB6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="DC413F5EB6D3D00D0C7E239A951DEDF3" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
6920, 6921, adult males, all preserved as dry skins and cleaned skulls, collected in Área de Protección Hídrica Otavalo Mojanda (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERAAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.275360,0.154560]}">0.15456, – 78.27536</named-content>
, 
<quantity id="3A8A493C655F23ABE1AE2190006D2766" metricMagnitude="3" metricUnit="m" metricValue="3.685" unit="m" value="3685.0">3,685 m</quantity>
) by J. Brito, 
<collectionCode id="725A5271CB52BA9FB6526115D4500E55" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. García and F. Rodríguez on 21 January, 2022. 
<named-content id="D9542F3A6DCD21C95AA4504B4F957B34" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="11826353C54FFA7C2EF30F0508E50B74" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
5603, juvenile female, 
<named-content id="4B0649E9AEE15ACCB487156C4FB32A21" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="E520CFACEEFFB6D349E12D02B0ED3541" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
5604, juvenile male, all preserved as dry skins and cleaned skulls, collected in Área Protegida Privada Neblina Norte (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJBAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.432619,-0.337535]}">
<geoCoordinate id="E12C21333B1ED1A2439D1590E9677F4F" degrees="0.337535" direction="south" orientation="latitude" precision="1" value="-0.337535">0.337535 ° S</geoCoordinate>
, 
<geoCoordinate id="2DBCB10F807107F0D0C48F0B559B0740" degrees="78.432619" direction="south" orientation="latitude" precision="1" value="-78.43262">– 78.432619</geoCoordinate>
</named-content>
, 
<quantity id="8B70241EB35BC5863E6FEBF6B9BE1A0F" metricMagnitude="3" metricUnit="m" metricValue="2.9899999999999998" unit="m" value="2990.0">
<geoCoordinate id="C3EBF57BEB80C762B7335D2BE9F45DA7" degrees="2,990" direction="east" orientation="longitude" precision="55" value="2.99">2,990</geoCoordinate>
m
</quantity>
) by J. Brito, J. Curay, E. Beltrán and M. Esparza on 23 March, 2017. 
<named-content id="75DAE042596A84BBAD7AA0B0482A5E7C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="05F628C31F12FF453E1B3EC5F265433C" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
5698, 
<named-content id="10617D9E05B619D08A232A2068973B45" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="B5F1A26700785C8F70BA610349D33B55" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
5700, adult males, 
<named-content id="CA1CE1C83A4C7BC20A20C0AE07E1FAB8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="A5E48360A68B4CA856845A926ABE3896" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
5701, adult female, all preserved as cleaned skulls and the rest of the body in ethanol, collected in Tabla Chupa (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJCAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.413066,0.336465]}">0.336465, – 78.413066</named-content>
, 
<quantity id="2316FD43872DCAA6F689985C50069916" metricMagnitude="3" metricUnit="m" metricValue="3.055" unit="m" value="3055.0">3,055 m</quantity>
) by J. Brito, J. Curay, E. Beltrán and M. Esparza on 24 March, 2017. 
<named-content id="8B9FB76852A09E92E34D69B24D050B52" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="78C220DA9404C0FAEFF4BD266265BC81" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7869, adult female, 
<named-content id="DDFC6ECEB18DBDF0805FDFA2EF3751D6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="F36C09A6228DADD4C81632763AB6AFE2" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7870, adult male, preserved as dry skin and cleaned skull, collected in Cayapas Chupa (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBDAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.483430,0.541170]}">0.54117, – 78.48343</named-content>
, 
<quantity id="51EC3C41A9B0A1509EFACE72EC74CFAF" metricMagnitude="3" metricUnit="m" metricValue="3.29" unit="m" value="3290.0">3,290 m</quantity>
) by J. Brito, A. Yacelga on 
<date id="45F8F5F1957FDF8ECE5A7B8962094EBF" value="2023-09-28">28 September 2023</date>
. 
<named-content id="43EE640A65EC2E09545F5108FA8576B3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="7DAB1B821018BF530FA699C329EE0D64" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7873, adult female, 
<named-content id="D190228DFBA7DB611920EAA3BFFEBEFE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="71A454D0329654E476EEDB586A667367" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7874, adult male, preserved as dry skin and cleaned skull; 
<named-content id="D5085C508B70D95E752ADF82FE3E050B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="5A8D7BAE215D3E2EA2FC9DB8A193EBE6" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7877, adult female, 
<named-content id="A81E69B83981C650FA88D15C8474D882" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="93CC3493CFEF7001937A758281E34AC1" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7881, 7882, adult males, all preserved as cleaned skulls and alcoholic bodies, all collected in Cayapas Chupa (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJEAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.475060,0.536400]}">0.53640, – 78.47506</named-content>
, 
<quantity id="657B4AF65DBF9E97D6EA71D8016B3C36" metricMagnitude="3" metricUnit="m" metricValue="3.44" unit="m" value="3440.0">3,440 m</quantity>
) by J. Brito, A. Yacelga, L. Rodríguez on 
<date id="92DBCD4317E50421D2540D987EB8947C" value="2023-10-03">03 October 2023</date>
. 
<named-content id="E493CD5390EAE1A85CC7B06D513AD46C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">
<collectionCode id="3AEBB0797BE176098139ED039F8F9083" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
</named-content>
7871, adult female, preserved as cleaned skulls and alcoholic bodies, collected in 
<collectingRegion id="42A3E366C9EE723D7C0316C3F2A6FBB4" country="Ecuador" name="Esmeraldas">Provincia de Esmeraldas</collectingRegion>
, Cordillera de Cayapas, Loma Negra (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZEAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-78.497960,0.545740]}">0.54574, – 78.49796</named-content>
, 
<quantity id="FB573162202E8CB68948F14C3A2D2A6B" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" unit="m" value="3230.0">3,230 m</quantity>
) by J. Brito, L. Rodríguez on 01–02 October, 2023.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5EAI" type="etymology">
<paragraph id="39D1763A9912E73B53DA8605399DC729">
<heading id="7F5D13283EBD7AFBB90C51BC35C270C9" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="D0309151EBC4F887E7B9B9E9BB075008">
The specific epithet “ Otavalo ” honors the Otavalo culture, here treated as a noun in apposition. The Otavalo people are recognized for their music and ability for weaving and comercializing textiles. For decades, the Otavalos have been one of the most recognizable and proud indigenous cultures of South America (
<bibRefCitation id="A79AB2284DBA5866F800D1002B122A93" author="Meisch" firstAuthor="Meisch" journalOrPublisher="University of Texas Press, Austin" refId="B40" refString="Meisch LA (2002) Andean Entrepreneurs: Otavalo Merchants and Musicians in the Global Arena. University of Texas Press, Austin, 329 pp." title="Andean Entrepreneurs: Otavalo Merchants and Musicians in the Global Arena." year="2002">Meisch 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIFAI" type="diagnosis">
<paragraph id="9E5DE34CB1FE4E2ED694AAF824F0597D">
<heading id="B5319EA5D53983B068FB7641CE5038FD" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1BD75AD78694806F372638FABA7E3521">
A species of 
<taxonomicName id="49062D2952B049A7E5BFB4972F551C1D" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="36DE085818553EE1E913E7C9FCF1D334" italics="true">Thomasomys</emphasis>
</taxonomicName>
unique due to the following combination of characters: Head-body length 
<quantity id="65B7CC9676529EA540AD4BDD33A2D1D4" metricMagnitude="-1" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" unit="mm" value="117.5" valueMax="125.0" valueMin="110.0">110–125 mm</quantity>
, very long tail (~ 144 of head-body length), with white apical portion (
<quantity id="67CA9AE1449E2E31470B46084FABB0C9" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 mm</quantity>
); narrow interorbital region with rounded supraorbital margins; narrow zygomatic arches; long incisive foramina covering approximately ~ 63 % of the diastema, but not extending posteriorly between molar series; upper first molars aligned with posterior margin of the zygomatic plate; subsquamosal fenestra larger than the postglenoid foramen; upper incisors opisthodont.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVFAI" type="description">
<paragraph id="AD2A372D1A87934B2BE1EA76919D5FAB">
<heading id="51076DAF9F77400DBA1CFC6878773796" reason="title">
Morphological description of the 
<typeStatus id="A088F14B1F58F34562075BF179F02ED2" type="holotype">holotype</typeStatus>
and variation.
</heading>
</paragraph>
<paragraph id="6799F3C05A19DEA151C45B60492C5F7A">
Pelage fine, dense, and soft, about 
<quantity id="16BEA9B37E9C533BD9F5299294ACD3BD" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10 – 12 mm</quantity>
long over the back and rump (Fig. 
<figureCitation id="7E4DD3F8009EDC22AAFFC928BDC642BE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimen of Thomasomys otavalo sp. nov. (Cordillera Cayapas, Imbabura, Ecuador), in its natural habitat (MECN 7869 paratype). Please note the color of the apical portion of the tail. Photographed by J. Brito on September 28, 2023." figureDoi="10.3897/vz.74.e128528.figure3" httpUri="https://binary.pensoft.net/fig/1168226">3</figureCitation>
); somberly colored (dark drab) with low countershading. Dorsal coloration Dark Drab (Color 45) along flanks, shading to Hair Brown (color 277) mid-dorsally. Ventral pelage Medium Neutral Gray (Color 298) basally, with superficial of Ground Cinnamon Medium Neutral Gray (Color 270); not sharply demarcated from dorsal coloration (Fig. 
<figureCitation id="16D458DEA673C46568DDBF9A7656EEB3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dry skin in dorsal (A, B), lateral (C – E), and ventral (E, F) views of Thomasomys otavalo, sp. nov. (A, C, E; MECN 6912, holotype), and T. igor sp. nov. (B, D, F; MECN 702, holotype). Scale bar: 20 mm." figureDoi="10.3897/vz.74.e128528.figure4" httpUri="https://binary.pensoft.net/fig/1168227">4 A, C, E</figureCitation>
). Mystacial vibrissae long, extending beyond pinnae when laid back alongside head. Ears sparsely covered with short, blackish hairs, not contrasting conspicuously with color of head. Hairs over metapodials and digits of manus and pes dark, but with white ungual tufts, abundant and extending well beyond the edge of the claws (Fig. 
<figureCitation id="8F38F4BE08FCD3FF0E4FCB539415F6E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selected external and soft anatomical traits of Thomasomys: A, C dorsal and (B, D) plantar surface of right pes (A, B: T. otavalo sp. nov, MECN 5604, paratype; C, D T. igor sp. nov, MECN 708, paratype); E soft palate (T. otavalo sp. nov, MECN 6921, paratype); F, G gross morphology of the stomach (T. otavalo sp. nov, MECN 5700, paratype). Abbreviations: 1–5, digits; b, bordering fold; co, cornified epithelium; d, duodenum; d 1 – d 3, diastemal rugae; ge, glandular epithelium; i, incisura angularis; i 1–15, interdental rugae. Scale bars: 5 mm." figureDoi="10.3897/vz.74.e128528.figure5" httpUri="https://binary.pensoft.net/fig/1168228">5 A, B</figureCitation>
). Tail longer than the combined length of head and body (
<collectionCode id="E88ACABF2C95BFAEBF75BEE97B629125" country="Canada" name="Université de Montréal" type="Herbarium">LT</collectionCode>
about 144 % of 
<abbrev id="ABBRID0EHGAI" xlink_title="head and body length">
<collectionCode id="9579C8F8A436E860FC90705364DCAE45" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" type="Herbarium">HB</collectionCode>
</abbrev>
), uniformly dark, with white apical portion (between 
<quantity id="2D71DE6A1AA1742E0D1331D0F06E5711" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 mm</quantity>
, Fig. 
<figureCitation id="D13D8A36B982726A0B0A77F767A0F606" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distal portion of the tail of some species of Thomasomys: A T. otavalo sp. nov. (MECN 5604, paratype); B T. ucucha (MECN 2691); C T. igor sp. nov. (MECN 708, paratype); D T. silvestris (MECN 5054). Scale bar: 10 mm." figureDoi="10.3897/vz.74.e128528.figure6" httpUri="https://binary.pensoft.net/fig/1168229">6 A</figureCitation>
); with short hairs, but end with some larger hairs; it is covered with rectangular scales (13 or 14 rows / cm near the base), with three dark brown hispid hairs emerging from the base of each scale, no longer than 1–1.5 scale rows, ventral hairs are whitish in color. Mammae six in inguinal, abdominal, and postaxial pairs.
</paragraph>
<caption id="D938B0FB68D72A053A70B779BB6A74EC" ID-DOI="10.3897/vz.74.e128528.figure3" ID-arpha="0DAE5652-CB1A-5C7F-9FAC-04FB9CCD3100" httpUri="https://binary.pensoft.net/fig/1168226" startId="F3">
<paragraph id="440ACB506EEF5EC52393089A0535AC77">
<label id="B54FFA8BB3DA9FC96E70BB0EA9A03010">Figure 3.</label>
</paragraph>
<paragraph id="4168082D6C59BD55AF45997C4EF7C48F">
Live specimen of 
<taxonomicName id="E9DB3EFC36BCD58449B2A5B6D0995D53" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="ED462A5155D008FB0A5BF27851F5570A" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="2F652F831D209606E079134BE014E523" bold="true">
<taxonomicNameLabel id="CF9E98C2D481833794A66C27C1B9B1D8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Cordillera Cayapas, Imbabura, Ecuador), in its natural habitat (
<named-content id="FC53B8FA39001C450128E0503376007C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
7869 paratype). Please note the color of the apical portion of the tail. Photographed by J. Brito on September 28, 2023.
</paragraph>
</caption>
<caption id="B9A2863A9258B5DF5BBDF091B447F90B" ID-DOI="10.3897/vz.74.e128528.figure4" ID-arpha="8809629D-18F3-55B9-B18C-911CEFDDDEF1" httpUri="https://binary.pensoft.net/fig/1168227" startId="F4">
<paragraph id="3464017C4661EB9BFD6E8C9B56CD741C">
<label id="A12DF7A3EE987A1A4029D660B3949A6B">Figure 4.</label>
</paragraph>
<paragraph id="4BBF8BBB2195DEFF4EF9BE553D90573B">
Dry skin in dorsal (
<emphasis id="2D4CEE74E06474E35EF20ED9F4A1EAA1" bold="true">A</emphasis>
, 
<emphasis id="0F536C0BC38008474CBF70315873C5EA" bold="true">B</emphasis>
), lateral (
<emphasis id="C664B17A74117FD1A392693EBBDCD92D" bold="true">C</emphasis>
– 
<emphasis id="764C55921E00FD75A6B8E593D452B23E" bold="true">E</emphasis>
), and ventral (
<emphasis id="5ACE510C38FEAE9CF288870A6ACFD0CF" bold="true">E</emphasis>
, 
<emphasis id="93901D79FF69869CFFC0BA41D71C25E6" bold="true">F</emphasis>
) views of 
<taxonomicName id="52403B7469C43FECEFE3A0C930405DE5" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="54750B7837EDD8E1F47A7DF0D2F1D3DB" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
, 
<emphasis id="6BE138A6E3C12858DEA36FE3D588C77B" bold="true">
<taxonomicNameLabel id="1916E722D7119BE4D690E79420E9D70F" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="62917F530AFF0D150B129962F4828418" bold="true">A</emphasis>
, 
<emphasis id="8BB97B113D9A42516DBDEF7E71E0CCA0" bold="true">C</emphasis>
, 
<emphasis id="670D16AF5EAC886EF54771724B9DFA4F" bold="true">E</emphasis>
; 
<named-content id="6AAC851D30EAABD0ACCB9A72F499AF1A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912, holotype), and 
<taxonomicName id="4498B6A981E3015D02575811629F9D81" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="CDE3FD465B25FECCAE055603B53177AB" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="D4C93C3B33C85DFDBB00358A9CAF749F" bold="true">
<taxonomicNameLabel id="596FA54EF64940B4DF0CF032EC759493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<emphasis id="F2C07096598411209D94FB9C73E1993A" bold="true">B</emphasis>
, 
<emphasis id="02BF6D2C71100CF454C6BFD327B0EC7B" bold="true">D</emphasis>
, 
<emphasis id="CAC3BCC59C4C59D8951588F1F5DD7501" bold="true">F</emphasis>
; 
<named-content id="8E6130DA164A799A63D7673B4048D2B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
702, holotype). Scale bar: 20 mm.
</paragraph>
</caption>
<caption id="CACA1CA922020900B240BFDCC8D963DE" ID-DOI="10.3897/vz.74.e128528.figure5" ID-arpha="38EE5543-750A-5CF2-93E6-A194C1F0716F" httpUri="https://binary.pensoft.net/fig/1168228" startId="F5">
<paragraph id="C0EDB38DD17995826A7691FF1A856648">
<label id="70D02FC1C386AB029EAF61882F6377D8">Figure 5.</label>
</paragraph>
<paragraph id="DF7E9B60AAF4B14D5E8375F8B4F05AB0">
Selected external and soft anatomical traits of 
<taxonomicName id="C209A04FE95DB988E67788C37F33269A" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="51207F10A94BDFA3502F69BDEAA7046D" italics="true">Thomasomys</emphasis>
</taxonomicName>
: 
<emphasis id="BE44E7E3B29514846722006959B34B99" bold="true">A</emphasis>
, 
<emphasis id="D15C94BF2B70EDCDF345F777402C2AB2" bold="true">C</emphasis>
dorsal and (
<emphasis id="A170AB87173D271D182C523269BF41F4" bold="true">B</emphasis>
, 
<emphasis id="BC1D35512C8CFE63B84C004F3C49C04C" bold="true">D</emphasis>
) plantar surface of right pes (
<emphasis id="D5B1A4BFF0C61739FD427C3CCE72BDE0" bold="true">A</emphasis>
, 
<emphasis id="AADE54F662265CA1B486514F4F242251" bold="true">B</emphasis>
: 
<taxonomicName id="323583FCE94CA77E23445F50BDA9D9A4" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov">
<emphasis id="489AD988A3A97F28C04B78F9A91348B2" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="B1F3DE2A9A9067AEDBC637535B0A2587" bold="true">
<taxonomicNameLabel id="DF3285EBBBCD9F925DD1969BC345EF00" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, 
<named-content id="C4D28C8ACAEEF132EFEA1EF9B99E000C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
5604, paratype; 
<emphasis id="AB7FF79F7FAA1EED774AAF4760CFEAFE" bold="true">C</emphasis>
, 
<emphasis id="3217102CFCC4723EEE49DDC57FE15481" bold="true">D</emphasis>
<taxonomicName id="7A70C3989D72F77D335BB805BF5D3E4C" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov">
<emphasis id="41617B62EBF04D71C9ACB07D87A5BF2E" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="9F92A25E5F5D5D625721563BDE280CC1" bold="true">
<taxonomicNameLabel id="D40192CF49E35AF6CF23336F70279E8B" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, 
<named-content id="75D2E052B31D94C01BDD8C0AB65C466F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
708, paratype); 
<emphasis id="066CA6D7082AE236D3C0BE2C3DB5EE9C" bold="true">E</emphasis>
soft palate (
<taxonomicName id="05F4DC913DBF61717998F666B924C471" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov">
<emphasis id="7D34DE4278A8B34B2CD1BCCA0C31ECC8" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="0F536557EE4C096232606609A5FC4B7F" bold="true">
<taxonomicNameLabel id="A179B6E50492DD7BD300BD337A688ABF" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, 
<named-content id="BD5A513A0088552D5F17882C523B64A6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6921, paratype); 
<emphasis id="23BADE78DC3028454F6A16DBDD521C88" bold="true">F</emphasis>
, 
<emphasis id="D11CF5960F5AC31894FEDE57D0676EE0" bold="true">G</emphasis>
gross morphology of the stomach (
<taxonomicName id="C45296DF21ED1F4E36DDF7ED2241EAC6" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov">
<emphasis id="CE13B535E73ED6DD0480CD80563FBB82" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="4ABF3DBFB41065BA66FA68248C3C29F3" bold="true">
<taxonomicNameLabel id="067DB3A19DA41514A779369C7CB53A07" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, 
<named-content id="54EF9C7A9FEB6D815489588BC0152C9D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
5700, paratype). Abbreviations: 1–5, digits; b, bordering fold; co, cornified epithelium; d, duodenum; d 1 – d 3, diastemal rugae; ge, glandular epithelium; i, incisura angularis; i 1–15, interdental rugae. Scale bars: 5 mm.
</paragraph>
</caption>
<caption id="22119EF5686938C380603D484B71050E" ID-DOI="10.3897/vz.74.e128528.figure6" ID-arpha="AE0C08CC-A276-5B44-A6E4-2C49853DA81D" httpUri="https://binary.pensoft.net/fig/1168229" startId="F6">
<paragraph id="0885D69C92789A1352CD2A7920043E7D">
<label id="594841E4CBDCFF368E66005176923E16">Figure 6.</label>
</paragraph>
<paragraph id="0B880F42719D1333E2B97C558FEEE992">
Distal portion of the tail of some species of 
<taxonomicName id="A9B9A6E79245289A9094CDC1CA29EF9B" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="FF046BACC00A3F41999EA2AFAA034520" italics="true">Thomasomys</emphasis>
</taxonomicName>
: 
<emphasis id="3A4BD3749968102C5103B4F575CFF9BB" bold="true">A</emphasis>
<taxonomicName id="866411FB8DFFC20A76D94B4C54319C84" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="B409499BC2D992C49FCF5632C01255A1" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="2688EE4E29980CFFE862A2C2B699D84B" bold="true">
<taxonomicNameLabel id="2BE1BA02AB7A5A0B92DFFCCE1DD91F32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="9CF3833E59F1D9C230909C2B5077160B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
5604, paratype); 
<emphasis id="AB96CD7BA4ADADD3F5E78758880C5CAA" bold="true">B</emphasis>
<taxonomicName id="D5C4ECCE0D7C20283DDF430AC00D2999" authority="(MECN 2691)" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="02CFE80088FC33C1913DD48AF80E7FAA" italics="true">T. ucucha</emphasis>
(
<named-content id="614E9139D8C25FB9466910541BADFCC6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
2691)
</taxonomicName>
; 
<emphasis id="3892A0877CA33FC9D734490D70C52690" bold="true">C</emphasis>
<taxonomicName id="94EB75AF07AC50C72C72AD30A86101AA" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="E6B8D975A3A68D79F6663D92B840B86C" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="9141A5C0D00F9E29B1D0EB934CAAD4FC" bold="true">
<taxonomicNameLabel id="592917A45472A916B4BFE93C279C00EC" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="830FA3440816D58D9AFA4A4767101551" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
708, paratype); 
<emphasis id="41A3149FAECBFD746362E23394B0B936" bold="true">D</emphasis>
<taxonomicName id="53C2B899F657B2DA04DD9AF22C2E4BA6" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="0D6F9A0B30A2508334B758DF8807B861" italics="true">T. silvestris</emphasis>
</taxonomicName>
(
<named-content id="6A22A50F5E65E223CCFF31B9C501AF2A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
5054). Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="7B68436FDFF074A1B03E33085E3D718E">
Cranium medium for the genus (
<quantity id="1CD4C12500CB971A2137FBD694D26C2C" metricMagnitude="-2" metricUnit="m" metricValue="2.915" metricValueMax="3.02" metricValueMin="2.81" unit="mm" value="29.15" valueMax="30.2" valueMin="28.1">28.1–30.2 mm</quantity>
of 
<abbrev id="ABBRID0EMRAI" xlink_title="Condylo-incisive length">
<collectionCode id="0EAA32078066BB8328BEA469CCF20BB3">CIL</collectionCode>
</abbrev>
). Long rostrum, somewhat acuminate and narrow with the nasal and premaxillary bones extending beyond the anterior face of the incisors (giving the appearance of an incipient rostral tube); poorly developed gnathic process. Posterior margin of the nasal bone not reaching the plane of the lacrimal bone. Shallow zygomatic notches. Small and rounded lacrimal bones. Narrow interorbital region with smooth outer edges, leaving the alveolar maxillary processes exposed in dorsal view (Fig. 
<figureCitation id="763AF8E1A6F4C091957C204130E3DDA5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Thomasomys otavalo sp. nov. (Área de Protección Hídrica Otavalo Mojanda, Imbabura, Ecuador): Cranium in (A) dorsal, (B) ventral, and (C) lateral views, and mandible in (D) labial view (MECN 6912 holotype). Scale bar: 10 mm." figureDoi="10.3897/vz.74.e128528.figure7" httpUri="https://binary.pensoft.net/fig/1168230">7 A</figureCitation>
). Anteriorly narrowed zygomatic arches. Supraorbital region with divergent posterior borders (sensu 
<bibRefCitation id="35DEDE6A7ABE2AB41B9848532223BFF1" author="Steppan" firstAuthor="Steppan" journalOrPublisher="Fieldiana Zoology" pagination="1-112" refId="B75" refString="Steppan S (1995) Revision of the tribe Phyllotini (Rodentia: Sigmondontinae), with a phylogenetic hypothesis for the Sigmodontinae. Revisión de la tribu Phyllotini (Rodentia: Sigmondontinae), con una hipótesis filogenética para los Sigmodontinae. Fieldiana Zoology 1464: 1–112." title="Revision of the tribe Phyllotini (Rodentia: Sigmondontinae), with a phylogenetic hypothesis for the Sigmodontinae. Revisión de la tribu Phyllotini (Rodentia: Sigmondontinae), con una hipótesis filogenética para los Sigmodontinae." volume="1464" year="1995">Steppan 1995</bibRefCitation>
). Frontoparietal suture “ 
<collectionCode id="D5508989496DE5D22CB8107C8BED5909" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” - shaped. Broad and rounded braincase, slightly flattened at the outer edges. Small and concave exoccipital. Dorsal profile (in lateral view) distinctively flattened from nasal tips to midfrontal region; anterior margin of zygomatic plate slightly tilted backward. No further development in the ethmoturbinals is distinguished. Small lambdoidal crest. First maxillary molar aligned with the posterior edge of the zygomatic plate (Fig. 
<figureCitation id="559DA6CED2F57C7EB5ACEB85A1E4CEBD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Morphological comparisons. Comparison of diastemal and palate region in four species of Thomasomys. A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Arrow in (A) indicates that the anterior border of the M 1 at the same level of the posterior end of the zygomatic plate; the lower dotted line indicates the penetration of the mesopterygoid fossa in relation to the hypoflexus of the M 3; abbreviations: hf, hypoflexid; if, incisive foramina; mpf, mesopterygoid fossa; pal, palate; ppp, posterolateral palatal pit; zp, zygomatic plate. Scale bar: 5 mm." figureDoi="10.3897/vz.74.e128528.figure8" httpUri="https://binary.pensoft.net/fig/1168231">8 A</figureCitation>
). Zygomatic arches sturdy with long jugals spanning a major segment in the zygomatic process of the maxillary bone. Alisphenoid strut wide and short. Carotid circulation primitive (Pattern 1, sensu 
<bibRefCitation id="CA6E34A13C3B1AB64922BB00B06DE627" author="Voss" firstAuthor="Voss" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="259-493" refId="B85" refString="Voss RS (1988) Systematics and ecology of ichthyomyine rodents (Muroidea): Patterns of morphological evolution in a small adaptive radiation. Bulletin of the American Museum of Natural History 188: 259–493." title="Systematics and ecology of ichthyomyine rodents (Muroidea): Patterns of morphological evolution in a small adaptive radiation." volume="188" year="1988">Voss 1988</bibRefCitation>
), as indicated by the presence of a large stapedial foramen, prominent squamosal-alisphenoid groove, and sphenofrontal foramen. Subsquamosal fenestra larger than the postglenoid foramen; hamular process of squamosal thin, long, slightly curved, and distally applied on the mastoid capsule; tegmen tympani broadly overlapping posterior suspensory process of squamosal. Upper edge of the mastoid not exceeding the edge of the subsquamosal fenestra (Fig. 
<figureCitation id="A52567F1C1106EC45389FFF6CCEFA951" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Alisphenoid-mastoid region comparisons in lateral view in four species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Arrow in (B) illustrated to upper edge of the mastoid that exceed the edge of the subsquamosal fenestra; while the oblique dotted line indicates the suture between the squamosal and parietal. Abbreviations: hp, hamular process of squamosal; exo, exoccipital; ip, interparietal; pa, parietal; lpp, lateral parietal portion; mas, mastoid; mf, mastoid fenestra; pgf, postglenoid foramen; sq, squamosal; ssf, subsquamosal fenestra; zps, zygomatic process of the squamosal bone. For ease of comparison are not included to scale" figureDoi="10.3897/vz.74.e128528.figure9" httpUri="https://binary.pensoft.net/fig/1168232">9 A</figureCitation>
). Bullae small; pars flaccida of tympanic membrane present, large; orbicular apophysis of malleus well developed. Paraoccipital process small. Hill foramen very small; long and narrow incisive foramina (averaging 64 % of diastemal length), not approaching M 1. Premaxillary process wide in the middle and narrow at the posterior end, maxillary septum of incisive foramina slender and long. Palate short and broad (sensu 
<bibRefCitation id="D07EE514659E115FC38A9C6225BA6568" author="Hershkovitz" firstAuthor="Hershkovitz" journalOrPublisher="Fieldiana Zoology" pagination="1-524" refId="B30" refString="Hershkovitz P (1962) Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group. Fieldiana Zoology 46: 1–524." title="Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group." volume="46" year="1962">Hershkovitz 1962</bibRefCitation>
), with mesopterygoid fossa reaching the hypoflexus of M 3. Posterolateral palatal pit small and inconspicuous. Mesopterygoid fossa broad, straight sided, extending anteriorly between third molars; bony roof complete or perforated by narrow, slit-like sphenopalatine openings flanking the presphenoid / basisphenoid suture. Basisphenoid wide with slightly flat edges. Foramen ovale small. Parapterygoid fossae narrow, approximately triangular, with moderately (excavated) anterior limits. Middle lacerate foramen broad. Lateral expressions of parietals present. Auditory bullae small and uninflated with large and wide Eustachian tubes. Dentary moderately long, with long and narrow coronoid process (extends beyond upper edge of condylar process); postcondylar and mental processes poorly developed; deep sigmoid notch. Semilunar recess symmetrical, with lower edge ends pointed. Capsular projection of the root of the incisor inconspicuous.
</paragraph>
<caption id="CAFEC8E709E422D535739D67789DE7D1" ID-DOI="10.3897/vz.74.e128528.figure7" ID-arpha="C2AAA69E-60EC-5603-BB81-6F511E68616B" httpUri="https://binary.pensoft.net/fig/1168230" startId="F7">
<paragraph id="06CF18364289DA06F1D0805FCD1E6637">
<label id="438C1EAB323C4C27C511D0525388B484">Figure 7.</label>
</paragraph>
<paragraph id="6440E4699BEBABEA4F2990C41C440310">
<taxonomicName id="1B4791DBC9EC11267EEE869E3F195C4C" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="8E2817B3A250BAFD05B6420C520BFF5D" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="E901266D538EB18B91F4BC64C94448A5" bold="true">
<taxonomicNameLabel id="CAF71FD0584479457CFCBFBABAEABDAE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Área de Protección Hídrica Otavalo Mojanda, Imbabura, Ecuador): Cranium in (
<emphasis id="5DE7CA45C9CAAC9A0A9DAB1F50DFA62A" bold="true">A</emphasis>
) dorsal, (
<emphasis id="BE2AAFEA3FD8D761506DB836CCF5E9A6" bold="true">B</emphasis>
) ventral, and (
<emphasis id="FB99FCC6054915B9B481BDD681CF0A4F" bold="true">C</emphasis>
) lateral views, and mandible in (
<emphasis id="B9DE3C7276FEEF5B8DC884358888825C" bold="true">D</emphasis>
) labial view (
<named-content id="201D49FB340C20A8EB3448A693EBED59" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912 holotype). Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="A5F0157B81D0D8E656AE1B6E99790CC6" ID-DOI="10.3897/vz.74.e128528.figure8" ID-arpha="302F0515-4A28-597E-9FF7-64947CE0514C" httpUri="https://binary.pensoft.net/fig/1168231" startId="F8">
<paragraph id="A7FE8F143843AD29BA02899C682C7DCA">
<label id="1997A99B160D897FF5F05F22F1A0B8DB">Figure 8.</label>
</paragraph>
<paragraph id="BF3FB91C74B0C66052CA7D97FAB3101B">
Morphological comparisons. Comparison of diastemal and palate region in four species of 
<taxonomicName id="45B4309C283BF52189ABE7E8A10041A8" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="4FBFF0F5A0D21CB015BAA7A83D647146" italics="true">Thomasomys</emphasis>
</taxonomicName>
. 
<emphasis id="1C6AD6893B410CF059CE7C6865DC331D" bold="true">A</emphasis>
<taxonomicName id="852A5EC2F9FC5E179DB61DE0DC8E5E9A" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="4F53F1B5D3FAF28C7D1EBCD1BF0DDBF3" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="242116C358F7AA79E4758404EB9BAF9F" bold="true">
<taxonomicNameLabel id="71C25A963032AB2E1CD2C1CD1A7E8230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="6C34AA037A70EB4E979EE6002E99F860" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912, holotype); 
<emphasis id="9B6F3258FF74E1E73282B371E5EB0D56" bold="true">B</emphasis>
<taxonomicName id="C104EFC915C2763C16F1C17B3BA2804F" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="0AC86B091ACE4079273C087E7D875F22" italics="true">T. ucucha</emphasis>
</taxonomicName>
(
<named-content id="7A9B39B51B2E4AA68A8BD9DF6CF8EC68" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
7937); 
<emphasis id="91199C5BCAA6004AAAA7C9C0B7009ED9" bold="true">C</emphasis>
<taxonomicName id="23E76EECB5137AC8A6D581DCF09A2EE4" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="B4954FDDC220D4C304B80F9F7E568603" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="8BE1699A48F3591C5288F3D6303DC4C2" bold="true">
<taxonomicNameLabel id="9DADE51CE221E4AEB7E47DF8B9271B66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="70FDF6DA236C01E4644D9FA693BF5D8C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
702, holotype); 
<emphasis id="ED06D59B85A5DFC8CADD83B9EDCBF19E" bold="true">D</emphasis>
<taxonomicName id="CC65786166EA186E8DBE9B3706E83677" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="1BC7AB3C47AFCF9D8E1B3AA5BEADE95A" italics="true">T. silvestris</emphasis>
</taxonomicName>
(
<named-content id="32AA8DC8822527DCE3B2853123AEE7EE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
4928). Arrow in (A) indicates that the anterior border of the M 1 at the same level of the posterior end of the zygomatic plate; the lower dotted line indicates the penetration of the mesopterygoid fossa in relation to the hypoflexus of the M 3; abbreviations: hf, hypoflexid; if, incisive foramina; mpf, mesopterygoid fossa; pal, palate; ppp, posterolateral palatal pit; zp, zygomatic plate. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="624FB0E52400D03AC66B443BB4EA3228" ID-DOI="10.3897/vz.74.e128528.figure9" ID-arpha="E321FCA8-37B8-5C34-9225-B3D740B8CCC3" httpUri="https://binary.pensoft.net/fig/1168232" startId="F9">
<paragraph id="C06185348D70E8562ADB59391F158534">
<label id="B78F74245E384CC94D91FA872F6DDF2C">Figure 9.</label>
</paragraph>
<paragraph id="4FA58FA2C3D33BF6B850D9BB47F79984">
Alisphenoid-mastoid region comparisons in lateral view in four species of 
<taxonomicName id="35F4C5923117F356840E96228CB24D66" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="C046AFF27F4D92A6FE515E5FBAB84F9E" italics="true">Thomasomys</emphasis>
</taxonomicName>
: 
<emphasis id="9245E0944BBCE81D0DDD69CD9C470493" bold="true">A</emphasis>
<taxonomicName id="6E84CE5570F9287B032B60D111A03D64" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="01F12A1E9711F90EB8EB10B4AA77F918" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="F390DE1FE7CC54C79F1C43385EB8C72C" bold="true">
<taxonomicNameLabel id="ECBADC5F9B782019479B18132C0223FA" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="3A5598C1D5C057DB0A8C54E6EEA2FE68" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912, holotype); 
<emphasis id="5C71814D3F6B2B86A1A00A6E223ED06B" bold="true">B</emphasis>
<taxonomicName id="BF9933CC5328A9B800369C9AF5B0223B" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="3380270BDB52B7494CF845117EEA39EF" italics="true">T. ucucha</emphasis>
</taxonomicName>
(
<named-content id="F1237622149E7D5C45D0599F8B1260B0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
7937); 
<emphasis id="443A8FEDB18F458326DC593C6FD70F15" bold="true">C</emphasis>
<taxonomicName id="7A3C8EDDF69986FF19F29277989CA1CB" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="EEAB95A340BBCD2163F1CC72BD7C56E6" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="E9EB4AF6BC910133314C08AEB93DC488" bold="true">
<taxonomicNameLabel id="693BD576AE5763B2D5016EEFF132318E" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="F1B949EAE0A0579FF983683310E2EAE2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
702, holotype); 
<emphasis id="0488349F6724A1FA88D668A3F3D34F2B" bold="true">D</emphasis>
<taxonomicName id="3E6CE80EE2FECFB299A6F2187A137D60" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="5A6849D293DB1A97097DCBB76ABB0537" italics="true">T. silvestris</emphasis>
</taxonomicName>
(
<named-content id="AC5FB72FB38A0DD4A43FEC5323B05367" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
4928). Arrow in (B) illustrated to upper edge of the mastoid that exceed the edge of the subsquamosal fenestra; while the oblique dotted line indicates the suture between the squamosal and parietal. Abbreviations: hp, hamular process of squamosal; exo, exoccipital; ip, interparietal; pa, parietal; lpp, lateral parietal portion; mas, mastoid; mf, mastoid fenestra; pgf, postglenoid foramen; sq, squamosal; ssf, subsquamosal fenestra; zps, zygomatic process of the squamosal bone. For ease of comparison are not included to scale
</paragraph>
</caption>
<paragraph id="773FD773E07FAC176BAA9E3F2C85C5A6">
Upper incisors large, broad, and opisthodont (Thomas’s angle of 80 °, 
<bibRefCitation id="5994882682ADABFEB413462053C10D07" DOI="10.1080/00222931908673892" author="Thomas" firstAuthor="Thomas" journalOrPublisher="Annals and Magazine of Natural History" pagination="289-290" refId="B79" refString="Thomas O (1919) The method of taking the incisive index in rodents. Annals and Magazine of Natural History 4: 289–290. https://doi.org/10.1080/00222931908673892" title="The method of taking the incisive index in rodents." volume="4" year="1919">Thomas 1919</bibRefCitation>
; see Fig. 
<figureCitation id="9932A1CF72E02746086E591AC1F384B3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Comparison of the left anterior portion of the rostrum, viewed from left side, in four species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Thomas’ angles according to incisive and molar planes are indicated as well as the extension of the molar series. Arrow in (A) point to the anterior border of the nasal bone extending beyond the anterior face of the incisors relative to (B); abbreviations: n, nasal; nc, nasolacrimal capsule; p, premaxillary; sf, supraorbital foramen; zp, zygomatic plate. Scale bar: 5 mm" figureDoi="10.3897/vz.74.e128528.figure10" httpUri="https://binary.pensoft.net/fig/1168233">10 A</figureCitation>
), with front enamel Light Chrome Orange (color 76); brachydont and pentalophodont molars (sensu 
<bibRefCitation id="49BFD8C5B78F8F99F0793E9A4B8B06B8" author="Hershkovitz" firstAuthor="Hershkovitz" journalOrPublisher="Fieldiana Zoology" pagination="1-524" refId="B30" refString="Hershkovitz P (1962) Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group. Fieldiana Zoology 46: 1–524." title="Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group." volume="46" year="1962">Hershkovitz 1962</bibRefCitation>
). Parallel upper molars small, pentalophodont, hypsodont in juveniles, and lacking cingula; coronal surfaces crested when unworn; main cusps slightly opposite and sloping backwards when viewed from side (Fig. 
<figureCitation id="0B6F3235DEFC475C07582235FDB8D3DC" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Comparison of the upper right molar series in occlusal view among some species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928); abbreviations: af, anteromedian flexus; al, anteroloph; ml, mesoloph; mfl, metaflexus; pf, paraflexus; prf, protoflexus. Scale bar: 1 mm." figureDoi="10.3897/vz.74.e128528.figure11" httpUri="https://binary.pensoft.net/fig/1168234">11 A</figureCitation>
). M 1 rectangular in outline with procingulum divided by anteromedian flexus into unequal anterolabial and anterolingual conules; thin anteroloph; wide mesoloph. M 2 squared in outline; mesoloph showing the same condition as in M 1. M 3 rounded in outline with conspicuous anteroloph; shallow paraflexus; mesolph, metaflexus and hypoflexus distinctive, when unworn. Lower molars with main cusps alternating and sloping backwards when viewed from side. First lower molar (m 1) with short anteromedian flexid that divides the procingulum into subequal anterolabial and anterolingual conules (Fig. 
<figureCitation id="428E8AE34D8E6E95A798CC179412B1D2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Comparison of the lower right molar series in occlusal view among some species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928); abbreviations: ml, mesolophid; prf, protoflexid. Scale bar: 1 mm." figureDoi="10.3897/vz.74.e128528.figure12" httpUri="https://binary.pensoft.net/fig/1168235">12 A</figureCitation>
); ectolophid thin and short mesolophid. Protoflexid of m 2 slim; thin and short mesolophid; m 3 slightly shorter than m 2.
</paragraph>
<caption id="277A11C9B6DC4A0A204BAC1A929FD8B6" ID-DOI="10.3897/vz.74.e128528.figure10" ID-arpha="FA99C00B-2A70-53A8-A716-A2E845F48177" httpUri="https://binary.pensoft.net/fig/1168233" startId="F10">
<paragraph id="2DDC3CA186DE5C63DAB53917D6CD790B">
<label id="C6E5C2C74F41038AFA739301B04EF1EF">Figure 10.</label>
</paragraph>
<paragraph id="BCC1B6EA1B37531CBC093830C194E007">
Comparison of the left anterior portion of the rostrum, viewed from left side, in four species of 
<taxonomicName id="87803AFA43F8134CA139F71DF5169A42" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="F29B0602896188A62DC9F9CD8752CF0E" italics="true">Thomasomys</emphasis>
</taxonomicName>
: 
<emphasis id="A28F850D9925B9AE4326DF9961117AEA" bold="true">A</emphasis>
<taxonomicName id="68DF671BD36EC58C6B388E511BBCEB7C" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="08271256EEA4519AB2D9FE1211535420" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="0EE66E07AAC29F0B4CCE32465845A91E" bold="true">
<taxonomicNameLabel id="D1B4BD94E8C89AD55F60C100D046FA1F" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="8DB181CBFE4121F396F06EFEBBA77C7A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912, holotype); 
<emphasis id="13DDD65F2110311267916089DDE0ABF6" bold="true">B</emphasis>
<taxonomicName id="1CBE9E1F46B9EDB78C12CD2A0546E605" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="0B76EB335F870784F4C358E7F0B9903B" italics="true">T. ucucha</emphasis>
</taxonomicName>
(
<named-content id="7C5932B0EB8D1FF961AF3AECF6B71746" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
7937); 
<emphasis id="8A798119DC1FE8190B6B6C3AB311B8B4" bold="true">C</emphasis>
<taxonomicName id="42E91CDE1F782B10858AEBC51FA62017" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="5C5B7AA74EBA2C5CFD55C284C257F56C" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="1F89A64C993AF71B0DE897CD61B8EEC7" bold="true">
<taxonomicNameLabel id="2166BB3A60F77AB9B23F2E97DCB303AE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="E685DB74D6E6B8ADABCF56824F6CA6EF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
702, holotype); 
<emphasis id="8C5CDC139874407F3B8EE928E2342537" bold="true">D</emphasis>
<taxonomicName id="7896B9DC94C75B75FB018FE8BD300A38" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="D8F4890E48B9696AAFC6297ABAF838AA" italics="true">T. silvestris</emphasis>
</taxonomicName>
(
<named-content id="E4689840D26B19DE2DD9B3D5D10619BB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
4928). Thomas’ angles according to incisive and molar planes are indicated as well as the extension of the molar series. Arrow in (A) point to the anterior border of the nasal bone extending beyond the anterior face of the incisors relative to (B); abbreviations: n, nasal; nc, nasolacrimal capsule; p, premaxillary; sf, supraorbital foramen; zp, zygomatic plate. Scale bar: 5 mm
</paragraph>
</caption>
<caption id="8AC3095D9B45A706815A83E11FC65FF4" ID-DOI="10.3897/vz.74.e128528.figure11" ID-arpha="D594EAB8-9B4B-5B63-B155-25AFB97B67E7" httpUri="https://binary.pensoft.net/fig/1168234" startId="F11">
<paragraph id="0C5AA85E88352D95DE99D8DDA0553907">
<label id="DF5D8DC2E1FE9A40C30CFB227AFA2881">Figure 11.</label>
</paragraph>
<paragraph id="A7EB3EFD8692C7164710C9E86EB3C8DF">
Comparison of the upper right molar series in occlusal view among some species of 
<taxonomicName id="64F2A858F041BBF88CA0054999C21B93" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="F870ED4DCD0DAA11018B3CE4B4C5D1A4" italics="true">Thomasomys</emphasis>
</taxonomicName>
: 
<emphasis id="F2CCCF17B3FF9B70CCA58494237D26D5" bold="true">A</emphasis>
<taxonomicName id="D04C046B5B6AB55BB7650074D2AED06A" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="3452C6A3C75EEA54666DF7AF0CE05EBF" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="AE2A8FD5EAAD685F2D9130A3B1D3DE14" bold="true">
<taxonomicNameLabel id="83466CFCE7E88CF7E4A42BBBCDADE98A" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="1CCF70B7B5ADD7ED28ADF476785B6AA9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912, holotype); 
<emphasis id="F10E96B9B8F2FD4664A6FF7BD3BD2932" bold="true">B</emphasis>
<taxonomicName id="813380EE31706AA7B3A557A59EE3A6B4" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="E9306A82308FF2947C4E6BAE17D687D1" italics="true">T. ucucha</emphasis>
</taxonomicName>
(
<named-content id="D2AFB8E71A7B493E24F0B1997CC396B9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
7937); 
<emphasis id="35AAC7693D01030D449967EB01A9E52D" bold="true">C</emphasis>
<taxonomicName id="BF104A476086B4ABCC0F44DF99C12C5D" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="006C63BB02D2FB911A663B93795D71C2" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="8A4EA26A0425916E5BEBD775B6CBD2AE" bold="true">
<taxonomicNameLabel id="5EC199D11F8578F8B5A06A3ADB57F9D3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="C27F523BCEAD9CF551FA69A1826F8E8A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
702, holotype); 
<emphasis id="A52F3E1D4755B0A8E02DF8D6042A75AE" bold="true">D</emphasis>
<taxonomicName id="6032DDFD92453F37F9C5BE5935BF641B" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="C429390884AE5C3F34B8E82F858734E6" italics="true">T. silvestris</emphasis>
</taxonomicName>
(
<named-content id="E4FA8EC1A79BE5B7B92299914405E571" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
4928); abbreviations: af, anteromedian flexus; al, anteroloph; ml, mesoloph; mfl, metaflexus; pf, paraflexus; prf, protoflexus. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="2A0AE7B397BFB3A007F539F576954CEB" ID-DOI="10.3897/vz.74.e128528.figure12" ID-arpha="ADA3F594-8668-53E1-A3F5-BFB2B7BF33A1" httpUri="https://binary.pensoft.net/fig/1168235" startId="F12">
<paragraph id="8AF8009A690707E0E212200AC65EB80F">
<label id="C11AFECFE26C36E3A6C6B3DB69999639">Figure 12.</label>
</paragraph>
<paragraph id="39245D69C2CD5DD06381E05357723424">
Comparison of the lower right molar series in occlusal view among some species of 
<taxonomicName id="98FC15640A17C845F1D531EE0D4FD1E2" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="1ABF1DD115A558A6BCB493B1984799BB" italics="true">Thomasomys</emphasis>
</taxonomicName>
: 
<emphasis id="153410904A7AAD61BDF995BCB1C67CAA" bold="true">A</emphasis>
<taxonomicName id="F5EAC6E0E4F557B29CA3F1029064962B" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="4F67246CD280CD1E27867B08F926D358" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="ECDE0B5540596C4719B356E4696CBCDB" bold="true">
<taxonomicNameLabel id="F982B67E554D20355D4EF8904C664666" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="EDA36292DFE0B348CF24B610E4661A65" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
6912, holotype); 
<emphasis id="1AA01802738B47A979C419E3617A252C" bold="true">B</emphasis>
<taxonomicName id="148D509350CC87421A1D22387895FFE3" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="6F0FC03C78F73809311C0A47367DD4BF" italics="true">T. ucucha</emphasis>
</taxonomicName>
(
<named-content id="74C61DA0C849BAF441C61EA6FF2EA4D9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
7937); 
<emphasis id="C31941B5F9289922BDBF4D7C3DEDB2A1" bold="true">C</emphasis>
<taxonomicName id="FEDA78D9C379F7A28EFDFC911C1F5238" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="61B0AD5DF625D5DB90C6A1E753D68A61" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="DDD1080F3DFC1A335DECF4705FB488FF" bold="true">
<taxonomicNameLabel id="B41A9C9375692729BE01B2BFD0E07F21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<named-content id="3C9E575D40757852CDE0BA4EA35457F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
702, holotype); 
<emphasis id="847F8A4E3A9832785738D4C8AABC08D9" bold="true">D</emphasis>
<taxonomicName id="AB0AAE1CE5B6E3E7207477F82EAE57F0" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="4CA0F584A16DA8B874E082D47C6A18CE" italics="true">T. silvestris</emphasis>
</taxonomicName>
(
<named-content id="62FEF77DA53F936910C6B310306651B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Cien­cias Naturales">MECN</named-content>
4928); abbreviations: ml, mesolophid; prf, protoflexid. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="91390E4CA22BD1648E97C52EFF2C3C33">Tuberculum of first rib articulating with transverse processes of seventh cervical and first thoracic vertebrae; second thoracic vertebra with differentially elongated neural spine; vertebral column composed of 19 thoracicolumbar, 16 th with moderately developed anapophyses and 17 th with little developed anapophyses, 4 sacrals (fused), and 42–45 caudal vertebrae; usually the second and third caudal vertebrae with small but complete hemal arches; 12 ribs.</paragraph>
<paragraph id="8BD2FD387245820D37A41E2F4FBE6474">
Soft palate with 3 slightly arched diastemal palatal rugae and 5 pairs of interdentals (Fig. 
<figureCitation id="BB112D34BA138D71AEF57E215116193E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selected external and soft anatomical traits of Thomasomys: A, C dorsal and (B, D) plantar surface of right pes (A, B: T. otavalo sp. nov, MECN 5604, paratype; C, D T. igor sp. nov, MECN 708, paratype); E soft palate (T. otavalo sp. nov, MECN 6921, paratype); F, G gross morphology of the stomach (T. otavalo sp. nov, MECN 5700, paratype). Abbreviations: 1–5, digits; b, bordering fold; co, cornified epithelium; d, duodenum; d 1 – d 3, diastemal rugae; ge, glandular epithelium; i, incisura angularis; i 1–15, interdental rugae. Scale bars: 5 mm." figureDoi="10.3897/vz.74.e128528.figure5" httpUri="https://binary.pensoft.net/fig/1168228">5 E</figureCitation>
). The second and third interdental wrinkles are well arched. The interdental wrinkles are short and leave a conspicuous midline “ channel ”. Large gall bladder present in all specimens examined (n = 11). The stomach is of the unilocular and hemiglandular 
<typeStatus id="7FCE5697593DFE99603BFFE1207D0B30">type</typeStatus>
. The cornified epithelium dominates the corpus and is characterized by its spongy surface; the glandular epithelium is widely distributed over the mostly smooth antrum (Fig. 
<figureCitation id="26240289B5EC18D060709F7F306696C7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selected external and soft anatomical traits of Thomasomys: A, C dorsal and (B, D) plantar surface of right pes (A, B: T. otavalo sp. nov, MECN 5604, paratype; C, D T. igor sp. nov, MECN 708, paratype); E soft palate (T. otavalo sp. nov, MECN 6921, paratype); F, G gross morphology of the stomach (T. otavalo sp. nov, MECN 5700, paratype). Abbreviations: 1–5, digits; b, bordering fold; co, cornified epithelium; d, duodenum; d 1 – d 3, diastemal rugae; ge, glandular epithelium; i, incisura angularis; i 1–15, interdental rugae. Scale bars: 5 mm." figureDoi="10.3897/vz.74.e128528.figure5" httpUri="https://binary.pensoft.net/fig/1168228">5 F, G</figureCitation>
). The boundary between the two epithelia (= bordering fold, sensu 
<bibRefCitation id="3B424AF48FD9A5F68FDAAB0FABD9D2CC" author="Carleton" firstAuthor="Carleton" journalOrPublisher="Miscellaneous Publications, Museum of Zoology, University of Michigan" pagination="1-43" refId="B19" refString="Carleton MD (1973) A survey of gross stomach morphology in New World Cricetinae (Rodentia, Muroidea), with comments on functional interpretations. Miscellaneous Publications, Museum of Zoology, University of Michigan 146: 1–43." title="A survey of gross stomach morphology in New World Cricetinae (Rodentia, Muroidea), with comments on functional interpretations." volume="146" year="1973">Carleton 1973</bibRefCitation>
) is manifested by a thick ridge and exceeds the level corresponding to the esophageal opening to the left. While the angular incisura is barely marked, both the esophageal canal and the angular plica are conspicuous.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETFBI" type="comparisons">
<paragraph id="171975850C1DB2B95C9282A4E7B313F2">
<heading id="19FD850000DC450EFC1941CD701C8D97" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="7A02AAEBCBC24D4244CB7B8EE9EF299C">
<taxonomicName id="0A273065F1C08F74E8BA2A912EFF4565" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="A4A22D473011277C751D514EA7059995" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="5F8C3A4BC3DE1D0A640100D22C4DF832" bold="true">
<taxonomicNameLabel id="B4A4FD61E5828F1DD1D5C13CE3915FE6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most closely related to 
<taxonomicName id="DAE0709A3EC8162F2BD4774897CE5C99" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="8B5A2AA4DCA281A632526D6CB26787E4" italics="true">T. ucucha</emphasis>
</taxonomicName>
, and a candidate species from Zuleta (Fig. 
<figureCitation id="2C1F82A827B989AAEB5320426DF8BCAE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cyt b gene tree of the genus Thomasomys and close outgroups. The tree was inferred by maximum likelihood (ML) using a nucleotide substitution model (GTR + F + G 4); it contains 222 terminals belonging to 36 named species of Thomasomys, including the two new species here described (indicated with black arrows), it also contains at least 20 putative new species of Thomasomys, and 20 outgroups of the genera Rhipidomys, Chilomys and Rhagomys. Nodal support statistics for the analyses in IQ-Tree are from: ultrafast bootstrap approximation (UF), SH-like approximate likelihood ratio test (aL), and the aBayes test (aB). White sections of the circles indicate UF and aL bootstrap frequencies &lt;75 % and aB posterior probabilities &lt;0.95, whereas black sections of the circles indicate UF and aL bootstrap frequencies ~ 75 % and aB posterior probabilities ~ 0.95. Insets represent differences in topology and branch lengths of three ML approaches. Inset A: Collapsed gene tree of the main figure. Inset B: Collapsed gene tree inferred with a codon substitution model. Inset C: Collapsed gene tree estimated with the RAxML tree with the CAT approximation; black circles represent bootstrap values ~ 75 %. Terminals are labeled with GenBank accession number, museum voucher number and scientific name — in case that the species have not been described, the sequences are indicated with ‘ sp’ and a geographical proxy — ‘ ONT ’ indicates that the sequence was obtained with MinION Oxford Nanopore Technologies." figureDoi="10.3897/vz.74.e128528.figure1" httpUri="https://binary.pensoft.net/fig/1168224">1</figureCitation>
: Part A). To construct this section, we used specimens referred to 
<taxonomicName id="49CA4F9D211B299AC41ED7D0025B5014" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="950A375158808B0E7EB6145BB3FCD738" italics="true">Thomasomys ucucha</emphasis>
</taxonomicName>
(sensu stricto) near Papallacta, 
<collectingRegion id="2E59B966E79D5320A07594608E958931" country="Ecuador" name="Napo">Provincia de Napo</collectingRegion>
, 
<typeStatus id="ADA7DBD6F69D2F90192B1C2E4DE36737">type</typeStatus>
locality of the species (
<bibRefCitation id="71E9412341359D74791F52E77027F5C5" author="Voss" firstAuthor="Voss" journalOrPublisher="American Museum Novitates" pagination="1-47" refId="B87" refString="Voss RS (2003) A new species of Thomasomys (Rodentia: Muridae) from eastern Ecuador, with remarks on mammalian diversity and biogeography in the Cordillera Oriental. American Museum Novitates 3421: 1–47." title="A new species of Thomasomys (Rodentia: Muridae) from eastern Ecuador, with remarks on mammalian diversity and biogeography in the Cordillera Oriental." volume="3421" year="2003">Voss 2003</bibRefCitation>
). 
<taxonomicName id="343A87DCB48C03FA2FAC5517BEC45B40" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="5267A5403944DE141E1B5954DEC5AAE8" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="000D699BF5CC1E17603CAF58FD3CCB05" bold="true">
<taxonomicNameLabel id="385FA1E69509281A8F8F91FDAB64AD5E" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a medium-sized species (Table 
<tableCitation id="FE80E89AE036655354E89A2CC4842465" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Selected morphological differences with species that could be confused with the new species, compiled from Voss (2003), Brito et al. (2019) and our own observations." httpUri="http://table.plazi.org/id/85B7158F42B3B32B8D6DDDB12F7DE6E7" tableUuid="85B7158F42B3B32B8D6DDDB12F7DE6E7">5</tableCitation>
), differing from 
<taxonomicName id="24AB13F701ABA000B5FDF1D27BD0AE08" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
<emphasis id="B80D2ED3ADC08985A9AD942C8A2A668D" italics="true">T. ucucha</emphasis>
</taxonomicName>
(features in parentheses) in having shorter dorsal hairs between 
<quantity id="631C3AB865651B4099DF7402C108AE1A" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
(
<quantity id="968EFF58A8BC554978A0C6D82F44DB07" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
); tail noticeably long ~ 144 % of 
<abbrev id="ABBRID0EAIBI" xlink_title="head and body length">HB</abbrev>
(~ 128 %); white tail tip 
<quantity id="FB4B8DDFD014D364DD534590CBBDF3BA" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 mm</quantity>
(&lt;
<quantity id="60961553B5D8D2A28B47FE23B3106405" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
). Qualitative craniodental differences between the two species are conspicuous: the nasal and premaxillary bones in 
<taxonomicName id="E2EA5FE9E0C5645D7E0A3FF3DE9F8F85" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="97AC2ECE14D6D61F71C7F8E59E2DA2BC" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="AE598F37D5E649C7978622E865E89C01" bold="true">
<taxonomicNameLabel id="7BA8261E7895ADA6D504B2F2B0DD00E2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are long, extending well in front of the anterior face of the incisors (short, just beyond the anterior surface of the incisors); upper incisors opisthodont (orthodont, slightly procumbent, see Fig. 
<figureCitation id="D3BC0127783BE589F08813B2B0F9F637" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Comparison of the left anterior portion of the rostrum, viewed from left side, in four species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Thomas’ angles according to incisive and molar planes are indicated as well as the extension of the molar series. Arrow in (A) point to the anterior border of the nasal bone extending beyond the anterior face of the incisors relative to (B); abbreviations: n, nasal; nc, nasolacrimal capsule; p, premaxillary; sf, supraorbital foramen; zp, zygomatic plate. Scale bar: 5 mm" figureDoi="10.3897/vz.74.e128528.figure10" httpUri="https://binary.pensoft.net/fig/1168233">10 B</figureCitation>
); Thomas angle 80 ° (86 °); M 1 goes beyond the posterior edge of the zygomatic plate (not so); M 1 with short paraflexus and indistinct mesoflexus (short and short); mesolophid of m 1 very short (large).
</paragraph>
<caption id="85B7158F42B3B32B8D6DDDB12F7DE6E7" ID-Table-UUID="85B7158F42B3B32B8D6DDDB12F7DE6E7" httpUri="http://table.plazi.org/id/85B7158F42B3B32B8D6DDDB12F7DE6E7" startId="T5" targetIsTable="true">
<paragraph id="7597EEB30C7905E82E605E207C511354">
<label id="23BA4F6C50B87F4A92EA9EDFC9BEEE73">Table 5.</label>
</paragraph>
<paragraph id="E5A45145D6D8B994815B4601C03D4E74">
Selected morphological differences with species that could be confused with the new species, compiled from 
<bibRefCitation id="F5AF7D31259FA7FF52339F283294EDD9" author="Voss RS" firstAuthor="Voss" journalOrPublisher="American Museum Novitates" pagination="1-47" refId="B87" refString="Voss RS (2003) A new species of Thomasomys (Rodentia: Muridae) from eastern Ecuador, with remarks on mammalian diversity and biogeography in the Cordillera Oriental. American Museum Novitates 3421: 1–47." title="A new species of Thomasomys (Rodentia: Muridae) from eastern Ecuador, with remarks on mammalian diversity and biogeography in the Cordillera Oriental." volume="3421" year="2003">Voss (2003)</bibRefCitation>
, 
<bibRefCitation id="1E03DE3EB0EEA5CE03654F28698E2D1A" DOI="10.31687/saremMN.19.26.2.0.04" author="Brito J &amp; Tinoco N &amp; Curay J &amp; Vargas R &amp; Reyes-Puig C &amp; Romero V &amp; Pardiñas UF" etAl="et al." firstAuthor="Brito" journalOrPublisher="Mastozoología Neotropical" pagination="308-330" refId="B15" refString="Brito J, Tinoco N, Curay J, Vargas R, Reyes-Puig C, Romero V, Pardiñas UF (2019) Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae). Mastozoología Neotropical 26: 308–330. https://doi.org/10.31687/saremMN.19.26.2.0.04" title="Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae)." volume="26" year="2019">Brito et al. (2019)</bibRefCitation>
and our own observations.
</paragraph>
</caption>
<paragraph id="D71ADA79967D16F048023C41F65A273D">
<table id="TID0ENLDI" rules="all">
<tbody id="76350FFBDD002C0A4805AA43101AA40C">
<tr id="368F413BAA8283ACBD2CA67862DA46E6">
<emphasis id="920A489CA2426AD7EC37508FE480FE60" bold="true">
<td id="31CCF91FAACC2EAA726D0229DC96DCB9" colspan="1" rowspan="1">
<taxonomicName id="7547EC0737966576F6AA21466F15D1CC" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="D792228A518BC804B2432869C770CC81" bold="true" italics="true">T. otavalo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EB25B5094ABED5C9E8C196515F1F183" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<emphasis id="A329B6C218D6BC865F2E6038D3AB7C25" bold="true" italics="true">
<td id="3A621BF2866B1E12A7A14DBDFAE297AA" colspan="1" rowspan="1">
<taxonomicName id="212C50C23B48D3FBBE3D5E0D7BC9F649" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">T. ucucha</taxonomicName>
</td>
<td id="DBD3EC84911ECCC605B6CB225EAEBB0F" colspan="1" rowspan="1">
<taxonomicName id="38CE5AA73696FAE97B8C9E576D7D2317" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">T. igor</taxonomicName>
<taxonomicNameLabel id="3F1D983209E774098A4EFE9F3C8ADEBA" rank="species">sp. nov.</taxonomicNameLabel>
</td>
</emphasis>
<td id="CBD687A0FF7A1D8560565008531F9160" colspan="1" rowspan="1">
<taxonomicName id="B2053411B70B773815BC70F671B19CDD" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="597CD96FED2C2706B653F88EBED9DBE3" bold="true" italics="true">T. silvestris</emphasis>
</taxonomicName>
</td>
</emphasis>
</tr>
<tr id="88AD8C29C7553F1080FB4BBD7C24D349">
<td id="411A0A7BC8DCC7B4424D3F090E1EBCB9" colspan="1" rowspan="1">
Tail very long (LT about 144 % of 
<abbrev id="ABBRID0EYLBI" xlink_title="head and body length">HB</abbrev>
)
</td>
<td id="4CA2C9EDFAD203EA174CBFC147BEF731" colspan="1" rowspan="1">
Tail long (LT about 128 % of 
<abbrev id="ABBRID0EAMBI" xlink_title="head and body length">HB</abbrev>
)
</td>
<td id="E28A7A53E53233C82F8D39555195160E" colspan="1" rowspan="1">
Tail long (LT about 130 % of 
<abbrev id="ABBRID0EIMBI" xlink_title="head and body length">HB</abbrev>
)
</td>
<td id="894CD9C49AC9BC1A67270C13764E1D5A" colspan="1" rowspan="1">
Tail long (LT about 140 % of 
<abbrev id="ABBRID0EQMBI" xlink_title="head and body length">HB</abbrev>
)
</td>
</tr>
<tr id="010288800FFC546F1401415104B93A55">
<td id="6FB5C20C15329B68D678DF1F4F6B0B7D" colspan="1" rowspan="1">White tail tip (15–35 mm)</td>
<td id="E0E15E2D983E1EDD4AF4A81E0FC9806B" colspan="1" rowspan="1">Tail tip usually not white (when present &lt;5 mm)</td>
<td id="F346A4012CD79E9A92CDE33CC3E93D13" colspan="1" rowspan="1">Tail tip usually not white (when present &lt;5 mm)</td>
<td id="711E2EE8A7105A0164838EA7B2C5DD1A" colspan="1" rowspan="1">Tail tip usually not white (when present &lt;5 mm)</td>
</tr>
<tr id="9451E3CBF53FB2AC21433B227F5FB082">
<td id="7C9D528271D4427D5FC2E69A53251C73" colspan="1" rowspan="1">Dorsal pelage uniformly grayish brown</td>
<td id="3F4C77410FF16F39030E0C71A3415133" colspan="1" rowspan="1">Dorsal pelage uniformly dark grayish brown</td>
<td id="907468FAFF817CEA1E3BDC68731DE1B8" colspan="1" rowspan="1">Dorsal pelage verona Brown, darker in the midline</td>
<td id="D3FE1F992B2B6FA4CE49D624524135F6" colspan="1" rowspan="1">Dorsal pelage uniformly dark grayish brown</td>
</tr>
<tr id="7D5145566EB1DBB89CA7854A0F87A653">
<td id="96A7DB175D24AD2BE755008FDBB8CD44" colspan="1" rowspan="1">Incipient rostral tube present</td>
<td id="010579866E84599F09EBB735B4BD8529" colspan="1" rowspan="1">Rostral tube absent</td>
<td id="D4C891D47C7A01B89491B3C66829A3CC" colspan="1" rowspan="1">Rostral tube absent</td>
<td id="835973FCDA864645512E41C677DFDAA5" colspan="1" rowspan="1">Rostral tube absent</td>
</tr>
<tr id="AE553CEAA22582FD2FBFA21C1BF7F0AF">
<td id="25A17B2E17E3BA94B14E0F144D3B90E0" colspan="1" rowspan="1">Thomas angle 80 °</td>
<td id="A7ACB507D7762F0D20B753020285A0F9" colspan="1" rowspan="1">Thomas angle 86 °</td>
<td id="61B3C44A086968BB21113E88A49A64BD" colspan="1" rowspan="1">Thomas angle 80 °</td>
<td id="4017DC7537695EAED0DE7972F941A39B" colspan="1" rowspan="1">Thomas angle 80 °</td>
</tr>
<tr id="9DC785997E3261F92086B3DCD0B6F2EC">
<td id="B91C3973A42169FE3409BF147B3CC7E0" colspan="1" rowspan="1">Zygomatic arches converge anteriorly</td>
<td id="25D6052B9FCFE4A7E3EF3A492E53CC5B" colspan="1" rowspan="1">Widely flaring zygomatic arches</td>
<td id="AD66EC2D65C7FF889551EF5644C79CB9" colspan="1" rowspan="1">Zygomatic arches converge anteriorly</td>
<td id="2B845010ADE05E6EB6600469A3CC6864" colspan="1" rowspan="1">Zygomatic arches converge anteriorly</td>
</tr>
<tr id="529FD89FA3DC52D03D4EB4ABEB6591DD">
<td id="27625F69147650E24E07DE31ABE86649" colspan="1" rowspan="1">Upper first molar aligned with posterior edge of the zygomatic plate</td>
<td id="B752133F35CC808CE61263A760AC7710" colspan="1" rowspan="1">First upper molar does not reach the posterior edge of the zygomatic plate</td>
<td id="417976C2AB2C9DEC4A686C4BDFD04BB1" colspan="1" rowspan="1">First upper molar does not reach the posterior edge of the zygomatic plate</td>
<td id="65F172FC14242CC894CA8C615D916074" colspan="1" rowspan="1">Upper first molar reaches beyond the posterior edge of the zygomatic plate</td>
</tr>
<tr id="27F8A784F47550DAF632EC966DDC1AB0">
<td id="C97A0405A0205C37451A22291AA0B524" colspan="1" rowspan="1">
Incisive foramina long (
<abbrev id="ABBRID0EQPBI" xlink_title="length of incisive foramina">LIF</abbrev>
about 63 % of 
<abbrev id="ABBRID0EUPBI" xlink_title="length of upper diastema">LD</abbrev>
), usually not extending posteriorly between molar alveoli
</td>
<td id="CECFBEAB6D8E55B05515E8B4B51D7415" colspan="1" rowspan="1">
Incisive foramina vey short (
<abbrev id="ABBRID0E3PBI" xlink_title="length of incisive foramina">LIF</abbrev>
about 57 % of 
<abbrev id="ABBRID0EAQBI" xlink_title="length of upper diastema">LD</abbrev>
), not approaching level of molar alveoli
</td>
<td id="4B691D03C40FAB81AF2D07BA9BE003E3" colspan="1" rowspan="1">
Incisive foramina long (
<abbrev id="ABBRID0EIQBI" xlink_title="length of incisive foramina">LIF</abbrev>
about 72 % of 
<abbrev id="ABBRID0EMQBI" xlink_title="length of upper diastema">LD</abbrev>
), usually not extending posteriorly between molar alveoli
</td>
<td id="6FF2B449971995E9E094ADEC413549D4" colspan="1" rowspan="1">
Incisive foramina long (
<abbrev id="ABBRID0EUQBI" xlink_title="length of incisive foramina">LIF</abbrev>
about 65 % of 
<abbrev id="ABBRID0EYQBI" xlink_title="length of upper diastema">LD</abbrev>
), usually not extending posteriorly between molar alveoli
</td>
</tr>
<tr id="1C8215EC98544CF41DA6FD19978E1C44">
<td id="9208C2D87FFAC737448724E4CA91DEC3" colspan="1" rowspan="1">Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra</td>
<td id="A2AC4ACF09E8630E68D43B3CAD22361B" colspan="1" rowspan="1">The superior border of the mastoid goes above the subsquamosal fenestra.</td>
<td id="4E866AAF6C5CF6D0224DDF4CB392E7C6" colspan="1" rowspan="1">Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra</td>
<td id="BE4079CD94A9B54204BC668D2932748C" colspan="1" rowspan="1">Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra</td>
</tr>
<tr id="9EB815A3824F084BF89D2D4E7F7B05D5">
<td id="C7F14CC1B1386D3EFD997D9589712D86" colspan="1" rowspan="1">Capsular process of lower incisor alveolus usually indistinct</td>
<td id="37F2B1C4405208E3B5F213D93D778D0A" colspan="1" rowspan="1">Capsular process of lower incisor alveolus usually distinct</td>
<td id="EA52A5EAC27F6E2DB4DC2E72E8B5F91E" colspan="1" rowspan="1">Capsular process of lower incisor alveolus indistinct or absent</td>
<td id="BB72D665E4F1D46B06BFBCCABB1AF816" colspan="1" rowspan="1">Capsular process of lower incisor alveolus indistinct or absent</td>
</tr>
<tr id="EDA918B63768F09D4844BD28DDD4B305">
<td id="07F03DF6813E6A120F85F8F486F3087B" colspan="1" rowspan="1">Upper incisors opisthodont, not procumbent</td>
<td id="4BC2F42EA36B95F935883B3E56A29CD0" colspan="1" rowspan="1">Upper incisors orthodont, conspicuously procumbent</td>
<td id="52BB997563C2E94C2BEE7A4856097BFA" colspan="1" rowspan="1">Upper incisors opisthodont, not procumbent</td>
<td id="47290CF4CBDE776BBCCBB0B8A38D7DD6" colspan="1" rowspan="1">Upper incisors opisthodont, not procumbent</td>
</tr>
<tr id="D65DB0D0F0E57D5C1CD9C2DBEBD258E6">
<td id="84B6EB1EEB41B13D12585426B2312077" colspan="1" rowspan="1">Subsquamosal fenestra larger than the postglenoid foramen</td>
<td id="78693E7FB699D13C7494A46AB730D26A" colspan="1" rowspan="1">Subsquamosal fenestra usually smaller than postglenoid foramen</td>
<td id="151A5DAB87BA502B2FB533C3556D3A68" colspan="1" rowspan="1">Subsquamosal fenestra larger than the postglenoid foramen</td>
<td id="3D154C2C21A1918A7CA6A1AA4A8F9F86" colspan="1" rowspan="1">Subsquamosal fenestra larger than the postglenoid foramen</td>
</tr>
<tr id="B2391FC8247F7F50620259C64023E952">
<td id="61C74228BC1269D678B6342794AB5052" colspan="1" rowspan="1">M 3 Paraflexo short and metaflexo indistinct</td>
<td id="CB47B51717769190E3603C9CB4E3632E" colspan="1" rowspan="1">Paraflexus and metaflexus short</td>
<td id="317477249E8900359DCFF1535BE2C76F" colspan="1" rowspan="1">Paraflexus and metaflexus length and fused</td>
<td id="55CCB559F4FD759F62DEE9DFA43B10EB" colspan="1" rowspan="1">Paraflexus and mesoflexus short</td>
</tr>
<tr id="608DF740262248A6FC7850B7CDA2BECE">
<td id="39F5C875B5F555459B872A017D04CC42" colspan="1" rowspan="1">M 3 comparatively large</td>
<td id="16AE5265F3EAD65174AD4AB46619773E" colspan="1" rowspan="1">M 3 comparatively small</td>
<td id="58B1038A89376721CC72BD9EC13D1A10" colspan="1" rowspan="1">M 3 comparatively large</td>
<td id="C5C30FB2D28AB602FF524E50FECF2F56" colspan="1" rowspan="1">M 3 comparatively small</td>
</tr>
<tr id="247F82A2AF82ED4E2278AABF811231EB">
<td id="9513135EFD93F2C9FB2DB28D49B511EB" colspan="1" rowspan="1">Procingulum of m 1 divided</td>
<td id="4920D3DE03240C7BAD872AFF7D7D737B" colspan="1" rowspan="1">Procingulum of m 1 undivided</td>
<td id="7B29D2A52B3CF569763EFFFB4D92E679" colspan="1" rowspan="1">Procingulum of m 1 divided</td>
<td id="E57264AF2889D55285FAB2A5042B749F" colspan="1" rowspan="1">Procingulum of m 1 undivided or indistinct</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="2383018ED892C849E1932820DBCF02D3">
Other 
<taxonomicName id="8E21263BA995D812358EC8A3921104D8">
<emphasis id="77B6A0E8267C539AB2EDC0914A827574" italics="true">Thomasomys</emphasis>
species
</taxonomicName>
that could be confused with 
<taxonomicName id="1DB71117C925A62C0D6D6799A74DA026" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="DE965AE056489205BADD446968CBF991" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="4AC3BB80DEE87452D43336FA47427F84" bold="true">
<taxonomicNameLabel id="771569AAF23EB70EAFA2FC5070819401" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are 
<taxonomicName id="B3FAF09BC219EF11012B73123AE2137E" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="A8EE0364F24879FCE9C8EDA953AEB765" italics="true">T. silvestris</emphasis>
</taxonomicName>
and 
<taxonomicName id="ED8E6B11E9BAEDD6B750B8F1143019DC" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
<emphasis id="4F475CC7815A627E948DD8C75C88DAD1" italics="true">T. igor</emphasis>
</taxonomicName>
<emphasis id="8E5A64A7787C4CF9FE43003995C6B312" bold="true">
<taxonomicNameLabel id="E2F7A220C3D12CB51AA5CA50ABC09988" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, 
<taxonomicName id="1A298A8DF4AD537DB5B557094E4FFAA6" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="C661F1984E10C92AC4837E7A2E407B5E" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="A02F2C667AE046DEA536380F33E29562" bold="true">
<taxonomicNameLabel id="5C1B76E29F8A31E9C0384F3450664093" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from these species by the white tail tip between 
<quantity id="8387DF6BE02E18085BBAD00D940876D9" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 mm</quantity>
(&lt;
<quantity id="5696909A89F15F32F566D7F9A02966F0" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
in 
<taxonomicName id="6D8EBFA4E47F7EA289CCCB66F2FCF378" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="0354491C665C4FE8509F7F4B24132471" italics="true">T. silvestris</emphasis>
</taxonomicName>
and 
<taxonomicName id="432DFDFF9AC212C1618AB31A45631338" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor">
<emphasis id="25947AA3E41159B8866624D3A71E1D11" italics="true">T. igor</emphasis>
</taxonomicName>
). Regarding the cranium, 
<taxonomicName id="CF2003E55D098131527699F5F7DAF9F7" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="D7DDC4748D7387F4A346C1927D8555CC" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="7611B53631E24918B31A5AD9A1215942" bold="true">
<taxonomicNameLabel id="EBB7D216F917E5E8AF771E6EE99B1D5B" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by the incisive foramina (64 % of 
<abbrev id="ABBRID0EBXBI" xlink_title="length of upper diastema">
<collectionCode id="49E708B1E21DB9B2FDDC132519E13B98" country="Sweden" name="Lund University" type="Herbarium">LD</collectionCode>
</abbrev>
), while 
<taxonomicName id="34EEB076A770F1841CA457F69ADE786A" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="BA1B82D9C19CF727281A0BA7A601CD32" italics="true">T. silvestris</emphasis>
</taxonomicName>
and 
<taxonomicName id="96EF3C6E28F9B00BB75DD84B6EC3E197" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor">
<emphasis id="48BDC4C1AF1B4579A7F558194C309033" italics="true">T. igor</emphasis>
</taxonomicName>
are longer (65 and 71 % of 
<abbrev id="ABBRID0E2XBI" xlink_title="length of upper diastema">LD</abbrev>
, respectively); in 
<taxonomicName id="7AA0BB79BA81D93EB8512CF047B23260" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo">
<emphasis id="C61BE85CC831808663CA0E354F5002B3" italics="true">T. otavalo</emphasis>
</taxonomicName>
the alisphenoid strut slender while in 
<taxonomicName id="262B3661F5B6C419D4EF81A6F51C145C" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="9D06D46321A979B6F55792CBE1E5C6EA" italics="true">T. silvestris</emphasis>
</taxonomicName>
and 
<taxonomicName id="4934D45E576BD2F7EC9C6CA8B16B9535" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor">
<emphasis id="768B94A9455934D3AF61A50D07288DFE" italics="true">T. igor</emphasis>
</taxonomicName>
they are wide. As for the molars, in 
<taxonomicName id="C44D0B7D11EF7B36F70EBC5D1D2897DF" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="B150A7105BC646255597F426738BD9F4" italics="true">T. otavalo</emphasis>
</taxonomicName>
<emphasis id="182B1F448060B21CB2AF2F8A879DEC1B" bold="true">
<taxonomicNameLabel id="629387600849AD0372DD10F628BD0658" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and in 
<taxonomicName id="73A0F7809B29A8EB8B791B9F538F7605" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
<emphasis id="4488DDE6A3841F0FED721C3AD5AE5F75" italics="true">T. silvestris</emphasis>
</taxonomicName>
the M 3 paraflexus short and mesoflexus indistinct, while in 
<taxonomicName id="A04D3934BD92E71733A76A6835F7099C" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor">
<emphasis id="0DA07AF4A435DF27C5F8AD608BFE2D14" italics="true">T. igor</emphasis>
</taxonomicName>
they are long and fused giving the appearance of a horseshoe. Another species with which 
<taxonomicName id="1A73BA573D15ABBE52B5699EF807EBB1" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="F9B630FF7215674ECC8A094B2C5CBAF9" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="7AC67B2C299FF333F778752A5939A776" bold="true">
<taxonomicNameLabel id="7F943C932F5B8E0226151902B802A243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could be confused is 
<taxonomicName id="7B37F6467A26A04C6602DB721E28D970" baseAuthorityName="Thomas" baseAuthorityYear="1898" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="vulcani">
<emphasis id="09BDFCCE2FABF3714066B6448157D2AB" italics="true">Thomasomys vulcani</emphasis>
</taxonomicName>
(sympatric species), however they are easily distinguished because the new species has noticeably long tail ~ 144 % of 
<abbrev id="ABBRID0E21BI" xlink_title="head and body length">HB</abbrev>
(~ 90 %).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E61BI" type="distribution">
<paragraph id="52B10216F61C63542C081DC5507324B1">
<heading id="424B3E2009442010FA4C8141FD21C59E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="4851F54DFC0AE8FAEDFD9803522A33FF">
Known so far from less than 10 localities, all between the Otavalo-Mojanda Hydric Protected Area (Área de Protección Hídrica Otavalo Mojanda), from the highlands of Intag in 
<collectingRegion id="0D2C9163A2D45C04C096E6967972210F" country="Ecuador" name="Imbabura">Provincia de Imbabura</collectingRegion>
, up to the Cordillera de Cayapas at the provincial boundary between 
<collectingRegion id="C7653FD2023D14583990DE4AE28FFBC8" country="Ecuador" name="Esmeraldas">Esmeraldas</collectingRegion>
and 
<collectingRegion id="775FAF6D3F86DDD1DE73FB13AC062C58" country="Ecuador" name="Imbabura">Imbabura</collectingRegion>
, at elevations of 
<quantity id="B1A9BD6C12C3F00689C90F6D175E7CDD" metricMagnitude="3" metricUnit="m" metricValue="2.9875" metricValueMax="3.685" metricValueMin="2.29" unit="m" value="2987.5" valueMax="3685.0" valueMin="2290.0">
<geoCoordinate id="95AE73B0DB02320C0712857014BF217C" degrees="2,290" direction="north" orientation="latitude" precision="55" value="2.29">2,290</geoCoordinate>
<geoCoordinate id="5A30BD318E22735252C64EE99B5741C5" degrees="3,685" direction="west" orientation="longitude" precision="55" value="-3.685">–3,685</geoCoordinate>
m
</quantity>
. 
<taxonomicName id="8404A3008FF299AAE1697DF5723E9090" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="365E08E055E5F5538D02783120B1E09B" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="0EA70EF5FFF88AA591950D45E048C983" bold="true">
<taxonomicNameLabel id="92E76420D95EE4700D048A3AECCC18DF" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is geographically delimited by the basins of the rivers Mira (north), Guayllabamba (south), to the west by the inter-Andean valley and to the east by the tropical rainforest.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES2BI" type="natural history">
<paragraph id="BE8651EBA2B993437E8B04831EC7BFC8">
<heading id="AAD31C8508F6D19120F9810BACB7C015" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="832B4FC55237FCD4228001F000A43041">
<taxonomicName id="EFCADD2DD180F9420EF77AEFB8524AEC" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="C603FD86C6A80792D76FEE14574A63B6" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="0A0254F3E151D8068DD919A7C9BB6E0A" bold="true">
<taxonomicNameLabel id="98B934DA8A931DEA4321F5A953072B0D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is, thus far, endemic to the temperate and high Andean zoogeographic areas (
<bibRefCitation id="3747B07434AFABB0463D6BBB4FD32427" author="Albuja" etAl="et al." firstAuthor="Albuja" journalOrPublisher="Escuela Politécnica Nacional, Quito" refId="B1" refString="Albuja L, Almendáriz A, Barriga R, Montalvo LD, Cáceres F, Román JL (2012) Fauna de vertebrados del Ecuador. Escuela Politécnica Nacional, Quito, 490 pp. http://bibdigital.epn.edu.ec/handle/15000/6686" title="Fauna de vertebrados del Ecuador." url="http://bibdigital.epn.edu.ec/handle/15000/6686" year="2012">Albuja et al. 2012</bibRefCitation>
) of the montane forest (
<bibRefCitation id="0C2FDD818F801E5A8D3ECC7F7009E452" journalOrPublisher="Subsecretaría de Patrimonio Natural, Quito" refId="B42" refString="Ministerio del Ambiente del Ecuador (2013) Sistema de clasificación de los ecosistemas del Ecuador continental. Subsecretaría de Patrimonio Natural, Quito, 143 pp." title="Sistema de clasificación de los ecosistemas del Ecuador continental." year="2013">
Ministerio del Ambiente del 
<collectingCountry id="EA8A35426F8F8007874F351EF049757E" name="Ecuador">Ecuador</collectingCountry>
2013
</bibRefCitation>
), an environment characterized by trees with abundant orchids, ferns, and bromeliads. 
<taxonomicName id="D819BC5F86D3A1B75C1485DA62DE1EF5" authorityName="Brito &amp; García &amp; Castellanos &amp; Gavilanes &amp; Curay &amp; Carrión-Olmedo &amp; Reyes-Barriga &amp; Guayasamin &amp; Salazar-Bravo &amp; Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo" status="sp. nov.">
<emphasis id="9538E89CF009B8131C763B02AB0D06E0" italics="true">Thomasomys otavalo</emphasis>
</taxonomicName>
<emphasis id="AA033D5E58E33BDC565680FB60DFC645" bold="true">
<taxonomicNameLabel id="C82E1A75694248CFA5A5DACB8FA40C03" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected in mature and disturbed forest where the understory is visually dominated by herbaceous families such as 
<taxonomicName id="5AF55D382677AA9BC972F6043A70EE5C" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
(
<taxonomicName id="9414432F63CC5A5DA98EECDD23483472" class="Liliopsida" family="Poaceae" genus="Chusquea" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="8D91560FB42F4384FD7948A6C0DEAA47" italics="true">Chusquea</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="37AE99C3CCCA5619CCEF7AB3DAF22C10" class="Liliopsida" family="Poaceae" genus="Neurolepis" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="DD263F7525E715BAD6D852EADB9912A6" italics="true">Neurolepis</emphasis>
sp.
</taxonomicName>
), 
<taxonomicName id="6E495C30C282165F4ABC6F8C245303BA" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
and 
<taxonomicName id="738D1F857BC1354298D26B9DA54B1F7F" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
. Visually the canopy is dominated by encino (
<taxonomicName id="0E0C3627190D6B18266525D5B2CABE6B" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" kingdom="Plantae" order="Oxalidales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="E848855D3B6D8753894C0A2F97F88C9E" italics="true">Weinmannia</emphasis>
spp.
</taxonomicName>
), árbol de papel (
<taxonomicName id="C590003E5F38F241D371A0F8EA414365" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="F8D552592ACFEF5CB85DF89E84A92B09" italics="true">Polylepis</emphasis>
spp.
</taxonomicName>
) and guandera (
<taxonomicName id="5C09A4C420A65BA7DAF64B62C143C340" class="Insecta" family="Clusiidae" genus="Clusia" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E2DC9216E6128F2915EDFCCF29CBF93E" italics="true">Clusia</emphasis>
spp.
</taxonomicName>
). The species was collected in sympatry with the shrew-opossum, 
<taxonomicName id="CCA7D70E27206D15282E054A03998A92" authority="(Tomes, 1863)" baseAuthorityName="Tomes" baseAuthorityYear="1863" class="Mammalia" family="Caenolestidae" genus="Caenolestes" kingdom="Animalia" order="Paucituberculata" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="F72E5A250F1EC07EA4626575063A97A3" italics="true">Caenolestes fuliginosus</emphasis>
(Tomes, 1863)
</taxonomicName>
, the shrew 
<taxonomicName id="6673DEC25B53E07301078F092CADBAF4" authority="(Thomas, 1912)" baseAuthorityName="Thomas" baseAuthorityYear="1912" class="Mammalia" family="Soricidae" genus="Cryptotis" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="equatoris">
<emphasis id="9F2A0E76C30E7C7F682061BCAF8C1D34" italics="true">Cryptotis equatoris</emphasis>
(
<bibRefCitation id="76651CE3BFD7572FA4ED9B122CF3BACE" DOI="10.1080/00222931208693196" author="Thomas O" journalOrPublisher="Annals and Magazine of Natural History" pagination="44-48" refId="B78" refString="Thomas O (1912) Small mammals from South America. Annals and Magazine of Natural History 10: 44 - 48. https: // doi. org / 10.1080 / 00222931208693196" title="Small mammals from South America." volume="10" year="1912">Thomas, 1912</bibRefCitation>
)
</taxonomicName>
, and the rodents 
<taxonomicName id="F9205A3BB503D5D362294DD6AD7F8489" authority="Thomas, 1894" authorityName="Thomas" authorityYear="1894" class="Mammalia" family="Cricetidae" genus="Akodon" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="mollis">
<emphasis id="6073DEF104FBE4B53157AD1F36E2DD6F" italics="true">Akodon mollis</emphasis>
Thomas, 1894
</taxonomicName>
, 
<taxonomicName id="9059902B8AB671B145C1AA230770D80B" authority="(Thomas, 1914)" baseAuthorityName="Thomas" baseAuthorityYear="1914" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="moerex">
<emphasis id="A18AF4D708DD3E9F0EF282F992DD5636" italics="true">Nephelomys moerex</emphasis>
(Thomas, 1914)
</taxonomicName>
, 
<emphasis id="6AF64258561D61181DF8E5F857864B1F" italics="true">Neomicroxus latebricola</emphasis>
(Anthony, 1924), 
<taxonomicName id="29AB2EB76FF7C8CF43312BA5FA41166C" authority="(Tomes, 1860)" baseAuthorityName="Tomes" baseAuthorityYear="1860" class="Mammalia" family="Cricetidae" genus="Microryzomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="minutus">
<emphasis id="A722C0BA4D4DD780F95947B06BBEA8CD" italics="true">Microryzomys minutus</emphasis>
(Tomes, 1860)
</taxonomicName>
, 
<taxonomicName id="20CEC8CD80EB4937BF7CBDFC73B29B44" authority="(Thomas, 1898)" baseAuthorityName="Thomas" baseAuthorityYear="1898" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="vulcani">
<emphasis id="F1534CCBAEB1795A35F6E69CA101E964" italics="true">T. vulcani</emphasis>
(
<bibRefCitation id="6D2FE1342FB81715A7A9902EC88CB597" DOI="10.1080/00222939808678001" author="Thomas O" journalOrPublisher="Annals and Magazine of Natural History" pagination="451-457" refId="B77" refString="Thomas O (1898) On seven new small mammals from Ecuador and Venezuela. Annals and Magazine of Natural History 7: 451 - 457. https: // doi. org / 10.1080 / 00222939808678001" title="On seven new small mammals from Ecuador and Venezuela." volume="7" year="1898">Thomas, 1898</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="8A8C704BB90733B8504E349C3B5CCC91" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="CBAD9EB3B9BC53793EA07C6D10754E7A" italics="true">Thomasomys</emphasis>
sp.
</taxonomicName>
Mojanda (
<taxonomicName id="A2F7D2197DCD90F0BCCF54FA8F66D136" baseAuthorityName="Thomas" baseAuthorityYear="1882" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="F4DEE878CA042ED24C6885C7EE82BFA9" italics="true">cinereus</emphasis>
</taxonomicName>
group), and 
<taxonomicName id="6E50DD8F75D0A6B32CF8D56F3DC30585" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="DC710FC26AED4192ACBDA5BAA0EE79DE" italics="true">Thomasomys</emphasis>
sp.
</taxonomicName>
Imbabura-Mojanda (
<taxonomicName id="D942F94C098A18B0403AA9C8EAB2644B" baseAuthorityName="Tomes" baseAuthorityYear="1860" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="aureus">
<emphasis id="FC203CC7E02B2A8737572FFD9E03BBA2" italics="true">aureus</emphasis>
</taxonomicName>
group).
</paragraph>
</subSubSection>
</treatment>
</document>