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<document id="BEC45476F7E99FC90FE6D2D584A67AD5" ID-DOI="10.11646/zootaxa.4978.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4820443" ID-ZooBank="9C7C1723-73EB-4FBE-A47A-54627DEB8F93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622139238996" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2021" docId="3729879BFFBBFF95FA93FB24D66618BA" docLanguage="en" docName="zootaxa.4978.1.1.pdf" docOrigin="Zootaxa 1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Botryllophilus coniorhynchus Kim &amp; Boxshall 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="99" masterDocId="CB10FFE3FFE5FFF7FA04FF87D4011D72" masterDocTitle="Copepods (Cyclopoida) associated with ascidian hosts: Ascidicolidae, Buproridae, Botryllophilidae, and Enteropsidae, with descriptions of 84 new species" masterLastPageNumber="286" masterPageNumber="1" pageNumber="95" updateTime="1698948597350" updateUser="plazi">
<mods:mods id="070DB7CD22DE5E5CE7D3C1FCEA5C7D0F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14587D54DB00A664238A5995A0ACFDFD">
<mods:title id="A2EF693816A664473FCD4CB667F2753A">Copepods (Cyclopoida) associated with ascidian hosts: Ascidicolidae, Buproridae, Botryllophilidae, and Enteropsidae, with descriptions of 84 new species</mods:title>
</mods:titleInfo>
<mods:name id="F84F6CC8FB3939563515D22A1A682801" type="personal">
<mods:role id="45F5D096A8F46178649C2F9445B7CDC3">
<mods:roleTerm id="BCE33488FF3E5603E7ABC850B875B9EC">Author</mods:roleTerm>
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<mods:namePart id="11C7260581A99CE0B7D8F27B5351990F">Kim, Il-Hoi</mods:namePart>
<mods:nameIdentifier id="692412DAAEE24337921BFEBBEAE6A3E8" type="email">m@gwnu</mods:nameIdentifier>
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<mods:name id="81FEED7F2C515B5009E311A9EC6546E4" type="personal">
<mods:role id="7CD5EDD322147564FC9AA809C49DA123">
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<mods:namePart id="B18555B4D22648B03AECF510282AC1A5">Boxshall, Geoff A.</mods:namePart>
<mods:nameIdentifier id="2E7829787BB42C4908BE8E481E88A91C" type="email">m@gwnu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="6DDA1F70CA6989B8A803199435476B3A">text</mods:typeOfResource>
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<mods:titleInfo id="E00DE851751C06254814285F43E19A55">
<mods:title id="B9FAF23AABE011B4F7EA960BE57EC72A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F49F13C8C1AA9DF9161C67C1F9B96C00">
<mods:date id="DB2345E50201913AC0F90C439A9A0D2D">2021</mods:date>
<mods:detail id="9E3D145CA14A8EAB01DEFF6C479739CA" type="issue">
<mods:number id="0227D14E79762B9D7A36FB3B795956D6">1</mods:number>
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<mods:start id="49965052916B5C288BCEBF448D20C529">1</mods:start>
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<mods:classification id="7480DE089FD053A061F1C1EFEB5722E7">journal article</mods:classification>
<mods:identifier id="6C5747E4B6CA147A42EB887EBC065E3B" type="DOI">10.11646/zootaxa.4978.1.1</mods:identifier>
<mods:identifier id="17F20E93B06D9A5C0FC568A7F57EFA6A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C484087613AB7151945D1DE38D80CFF" type="Zenodo-Dep">4820443</mods:identifier>
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<treatment id="3729879BFFBBFF95FA93FB24D66618BA" ID-DOI="http://doi.org/10.5281/zenodo.4822494" ID-Zenodo-Dep="4822494" LSID="urn:lsid:plazi:treatment:3729879BFFBBFF95FA93FB24D66618BA" httpUri="http://treatment.plazi.org/id/3729879BFFBBFF95FA93FB24D66618BA" lastPageId="98" lastPageNumber="99" pageId="94" pageNumber="95">
<subSubSection id="F79A6506FFBBFFA9FA93FB24D66419CC" box="[151,613,1187,1214]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="BF3F368DFFBBFFA9FA93FB24D66419CC" blockId="94.[151,613,1187,1251]" box="[151,613,1187,1214]" pageId="94" pageNumber="95">
<heading id="E47781E1FFBBFFA9FA93FB24D66419CC" bold="true" box="[151,613,1187,1214]" fontSize="11" level="2" pageId="94" pageNumber="95" reason="4">
<emphasis id="8DF4EA9FFFBBFFA9FA93FB24D66419CC" bold="true" box="[151,613,1187,1214]" pageId="94" pageNumber="95">
<taxonomicName id="78804D0EFFBBFFA9FA93FB24D60119CC" authority="Kim &amp; Boxshall, 2021" authorityName="Kim &amp; Boxshall" authorityYear="2021" box="[151,512,1187,1214]" class="Maxillopoda" family="Ascidicolidae" genus="Botryllophilus" kingdom="Animalia" order="Cyclopoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="coniorhynchus" status="sp. nov.">
<emphasis id="8DF4EA9FFFBBFFA9FA93FB24D60119CC" bold="true" box="[151,512,1187,1214]" italics="true" pageId="94" pageNumber="95">Botryllophilus coniorhynchus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96C757E4FFBBFFA9F80CFB23D66419CC" box="[520,613,1188,1214]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FFBBFFA9FA93FB4FD5281991" box="[151,297,1224,1251]" pageId="94" pageNumber="95" type="description">
<paragraph id="BF3F368DFFBBFFA9FA93FB4FD5281991" blockId="94.[151,613,1187,1251]" box="[151,297,1224,1251]" pageId="94" pageNumber="95">
(
<figureCitation id="27BB2A08FFBBFFA9FA9BFB4FD5201991" box="[159,289,1224,1251]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="95.[151,198,1943,1967]" captionStartId-1="96.[151,198,1943,1967]" captionStartId-2="97.[151,198,1943,1967]" captionTargetBox-0="[209,1381,189,1901]" captionTargetBox-1="[201,1378,191,1903]" captionTargetBox-2="[204,1386,196,1906]" captionTargetId-0="figure-19@95.[192,1395,181,1919]" captionTargetId-1="figure-23@96.[189,1398,181,1919]" captionTargetId-2="figure-19@97.[191,1397,181,1919]" captionTargetPageId-0="95" captionTargetPageId-1="96" captionTargetPageId-2="97" captionText-0="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." captionText-1="FIG. 61. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female.A, rostrum; B, maxillule; C, maxilla; D, right leg 1; E, left leg 1; F, right leg 2; G, left leg 2. Scale bars: 0.02 mm." captionText-2="FIG. 62. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, maxilliped; B, right leg 3; C, left leg 3; D, right leg 4; E, left leg 4; F, leg 5; G, genital aperture. Scale bars: A-E, G, 0.02 mm; F, 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4820979" figureDoi-1="http://doi.org/10.5281/zenodo.4820994" figureDoi-2="http://doi.org/10.5281/zenodo.4821004" httpUri-0="https://zenodo.org/record/4820979/files/figure.png" httpUri-1="https://zenodo.org/record/4820994/files/figure.png" httpUri-2="https://zenodo.org/record/4821004/files/figure.png" pageId="94" pageNumber="95">Figs. 60-62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FFBBFFA9FA93FA97D03C18E4" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="BF3F368DFFBBFFA9FA93FA97D03C18E4" blockId="94.[151,1437,1296,2043]" pageId="94" pageNumber="95">
<emphasis id="8DF4EA9FFFBBFFA9FA93FA97D5461858" bold="true" box="[151,327,1296,1322]" pageId="94" pageNumber="95">Type material.</emphasis>
<materialsCitation id="0FE83CD0FFBBFFA9FB4BFA97D03C18E4" collectingDate="2008-01-15" collectionCode="MNHN, APL, RV" elevation="420" latitude="-66.166664" longLatPrecision="998" longitude="139.68333" pageId="94" pageNumber="95" specimenCode="A1" specimenCount="6" specimenCount-female="6" typeStatus="holotype">
<typeStatus id="603B882FFFBBFFA9FB4BFA97D5B61858" box="[335,439,1296,1322]" pageId="94" pageNumber="95" type="holotype">Holotype</typeStatus>
<specimenCount id="A986FD04FFBBFFA9FBC4FA97D5D41858" box="[448,469,1296,1322]" pageId="94" pageNumber="95" type="female">♀</specimenCount>
(MNHN-IU-2018-1973),
<specimenCount id="A986FD04FFBBFFA9F8FBFA97D73E1858" box="[767,831,1296,1322]" pageId="94" pageNumber="95" type="female">3 ♀♀</specimenCount>
<typeStatus id="603B882FFFBBFFA9F94CFA97D7B21858" box="[840,947,1296,1322]" pageId="94" pageNumber="95" type="paratype">paratypes</typeStatus>
(intact, MNHN-IU-2018-1974), and
<specimenCount id="A986FD04FFBBFFA9FF58FA97D19D1858" box="[1372,1436,1296,1322]" pageId="94" pageNumber="95" type="female">2 ♀♀</specimenCount>
<typeStatus id="603B882FFFBBFFA9FA93FAB3D503183C" box="[151,258,1332,1358]" pageId="94" pageNumber="95" type="paratype">paratypes</typeStatus>
(dissected, MNHN-IU-2014-17376) from
<emphasis id="8DF4EA9FFFBBFFA9F8FCFAB2D7FF1822" box="[760,1022,1333,1360]" italics="true" pageId="94" pageNumber="95">Aplẚdẚum falklandẚcum</emphasis>
Millar, 1960 (MNHN-IT-2008-386 =
<collectionCode id="D991AE48FFBBFFA9FAB7FADFD5081800" box="[179,265,1368,1394]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="94" pageNumber="95">MNHN</collectionCode>
<specimenCode id="EF269EF6FFBBFFA9FB17FADFD5331801" box="[275,306,1368,1395]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="94" pageNumber="95">A1</specimenCode>
/
<collectionCode id="D991AE48FFBBFFA9FB3FFADFD5771801" box="[315,374,1368,1395]" pageId="94" pageNumber="95">APL</collectionCode>
.B/532); Antarctic, Terre Adélie, CEAMARC cruise,
<collectionCode id="D991AE48FFBBFFA9F9E0FADFD0091800" box="[996,1032,1368,1394]" country="Italy" name="Collection of Leptospira Strains" pageId="94" pageNumber="95">RV</collectionCode>
“Aurore Australis”, Stn 18EV479 (
<geoCoordinate id="DAB4504AFFBBFFA9FAA4FAFBD4FD18E4" box="[160,252,1404,1430]" degrees="66" direction="south" minutes="10" orientation="latitude" pageId="94" pageNumber="95" precision="925" value="-66.166664">66º10’S</geoCoordinate>
,
<geoCoordinate id="DAB4504AFFBBFFA9FB0CFAFBD57518E4" box="[264,372,1404,1430]" degrees="139" direction="east" minutes="41" orientation="longitude" pageId="94" pageNumber="95" precision="925" value="139.68333">139º41’E</geoCoordinate>
), depth
<quantity id="78789B68FFBBFFA9FBCAFAFBD64B18E4" box="[462,586,1404,1430]" metricMagnitude="2" metricUnit="m" metricValue="4.195" metricValueMax="4.37" metricValueMin="4.02" pageId="94" pageNumber="95" unit="m" value="419.5" valueMax="437.0" valueMin="402.0">
<elevation id="34ADD1BEFFBBFFA9FBCAFAFBD64B18E4" box="[462,586,1404,1430]" metricMagnitude="2" metricUnit="m" metricValue="4.195" metricValueMax="4.37" metricValueMin="4.02" pageId="94" pageNumber="95" unit="m" value="419.5" valueMax="437.0" valueMin="402.0">402-437 m</elevation>
</quantity>
, IPEV-AAD-MNHN coll.,
<date id="CB3E104DFFBBFFA9F978FAFBD03818E4" box="[892,1081,1404,1431]" pageId="94" pageNumber="95" value="2008-01-15">
<collectingDate id="DB7AE9A5FFBBFFA9F978FAFBD03818E4" box="[892,1081,1404,1431]" pageId="94" pageNumber="95" value="2008-01-15">15 January 2008</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FFBBFFA9FAC3FA27D5DA18AC" pageId="94" pageNumber="95" type="etymology">
<paragraph id="BF3F368DFFBBFFA9FAC3FA27D5DA18AC" blockId="94.[151,1437,1296,2043]" pageId="94" pageNumber="95">
<emphasis id="8DF4EA9FFFBBFFA9FAC3FA27D54F18C8" bold="true" box="[199,334,1440,1466]" pageId="94" pageNumber="95">Etymology.</emphasis>
The name of the new species is derived from the Greek
<emphasis id="8DF4EA9FFFBBFFA9F9BAFA26D7EE18CE" box="[958,1007,1441,1468]" italics="true" pageId="94" pageNumber="95">cçnẚ</emphasis>
(= a cone) and
<emphasis id="8DF4EA9FFFBBFFA9FE9FFA27D0E918C8" box="[1179,1256,1440,1466]" italics="true" pageId="94" pageNumber="95">rhynch</emphasis>
(= a snout), and refers to the conical rostrum.
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FFBBFF95FAC3FA6FD55919B9" lastPageId="98" lastPageNumber="99" pageId="94" pageNumber="95" type="description">
<paragraph id="BF3F368DFFBBFFA9FAC3FA6FD6EA1BC4" blockId="94.[151,1437,1296,2043]" pageId="94" pageNumber="95">
<emphasis id="8DF4EA9FFFBBFFA9FAC3FA6FD5D31B70" bold="true" box="[199,466,1512,1538]" pageId="94" pageNumber="95">Description of female.</emphasis>
Body (
<figureCitation id="27BB2A08FFBBFFA9F82EFA6FD6B41B70" box="[554,693,1512,1538]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="94" pageNumber="95">Fig. 60A, B</figureCitation>
) asymmetrical,
<quantity id="78789B68FFBBFFA9F96AFA6FD7D41B70" box="[878,981,1512,1539]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="94" pageNumber="95" unit="mm" value="1.15">1.15 mm</quantity>
long. Anterior part of body comprising cephalosome to fifth pedigerous somite,
<quantity id="78789B68FFBBFFA9F86CF98BD6CC1B54" box="[616,717,1548,1574]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="94" pageNumber="95" unit="mm" value="0.73">0.73 mm</quantity>
long, invariably slightly curved to right side, unsegmented but with 4 weak constrictions dorsally and laterally; lateral margins parallel. Body width
<quantity id="78789B68FFBBFFA9FE6BF9B7D0D41B38" box="[1135,1237,1584,1610]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="94" pageNumber="95" unit="mm" value="0.33">0.33 mm</quantity>
across middle of anterior part. Posterior part of body (
<figureCitation id="27BB2A08FFBBFFA9F846F9D3D6A81B1D" box="[578,681,1620,1647]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="94" pageNumber="95">Fig. 60C</figureCitation>
) consisting of genital and 4 abdominal somites. Genital apertures positioned dorsally on genital somite. Caudal ramus (
<figureCitation id="27BB2A08FFBBFFA9F8F5F9FFD7591BE1" box="[753,856,1656,1683]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="94" pageNumber="95">Fig. 60D</figureCitation>
) armed with 4 claws and 1 seta; 1 outer distal claw bluntly tipped, other claw not articulated with ramus.
</paragraph>
<paragraph id="BF3F368DFFBBFFA9FAC3F947D79F1A88" blockId="94.[151,1437,1296,2043]" pageId="94" pageNumber="95">
Rostrum (
<figureCitation id="27BB2A08FFBBFFA9FB3EF947D5A21BA9" box="[314,419,1728,1755]" captionStart="FIG" captionStartId="96.[151,198,1943,1967]" captionTargetBox="[201,1378,191,1903]" captionTargetId="figure-23@96.[189,1398,181,1919]" captionTargetPageId="96" captionText="FIG. 61. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female.A, rostrum; B, maxillule; C, maxilla; D, right leg 1; E, left leg 1; F, right leg 2; G, left leg 2. Scale bars: 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820994" httpUri="https://zenodo.org/record/4820994/files/figure.png" pageId="94" pageNumber="95">Fig. 61A</figureCitation>
) distinct, conical, longer than wide (57×42 μm). Antennule (
<figureCitation id="27BB2A08FFBBFFA9FE6DF947D0CE1BA9" box="[1129,1231,1728,1755]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="94" pageNumber="95">Fig. 60E</figureCitation>
) short, about 115 μm long, strongly tapering, with markedly broadened first and second segments; terminal segment subdivided on posterior side; armature formula 9 (3 large and 6 small), 5 (2 large and 3 small), 2 (1 large and 1 small), and 12. Left antenna (
<figureCitation id="27BB2A08FFBBFFA9FB31F8ABD5991A35" box="[309,408,1836,1863]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="94" pageNumber="95">Fig. 60F</figureCitation>
) apparently 5-segmented, consisting of coxa, basis, and 3-segmented endopod; proximal 3 segments unarmed; second endopodal segment (fourth segment) 46×18 μm, with 2 bluntly tipped setae on inner margin; third endopodal segment 17×16 μm, with 4 (2 shorter inner and 2 longer outer) bluntly tipped setae on distal margin. Right antenna similar in size and segmentation to left antenna; 2 setae on second endopodal segment and 2 inner distal setae on third endopodal segment slightly longer than those of left antenna and distally spinulose (
<figureCitation id="27BB2A08FFBBFFA9FF6BF83BD4CD1A89" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="94" pageNumber="95">Fig. 60G</figureCitation>
); 2 outer distal setae distinctly shorter than those of left antenna.
</paragraph>
<caption id="EBFF6605FFBAFFA8FA93F810D5291A85" ID-DOI="http://doi.org/10.5281/zenodo.4820979" ID-Zenodo-Dep="4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,198,1943,1967]" targetBox="[209,1381,189,1901]" targetPageId="95">
<paragraph id="BF3F368DFFBAFFA8FA93F810D5291A85" blockId="95.[151,1436,1943,2040]" pageId="95" pageNumber="96">
<emphasis id="8DF4EA9FFFBAFFA8FA93F810D4F01ADD" bold="true" box="[151,241,1943,1967]" pageId="95" pageNumber="96">FIG. 60.</emphasis>
<emphasis id="8DF4EA9FFFBAFFA8FAF3F810D6221AC2" box="[247,547,1943,1968]" italics="true" pageId="95" pageNumber="96">Bçtryllçphẚlus cçnẚçrhynchus</emphasis>
<emphasis id="8DF4EA9FFFBAFFA8F82DF81FD67B1AC2" bold="true" box="[553,634,1944,1968]" pageId="95" pageNumber="96">sp. nov.</emphasis>
, female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm.
</paragraph>
</caption>
<caption id="EBFF6605FF85FF97FA93F810D6331AA1" ID-DOI="http://doi.org/10.5281/zenodo.4820994" ID-Zenodo-Dep="4820994" httpUri="https://zenodo.org/record/4820994/files/figure.png" pageId="96" pageNumber="97" startId="96.[151,198,1943,1967]" targetBox="[201,1378,191,1903]" targetPageId="96">
<paragraph id="BF3F368DFF85FF97FA93F810D6331AA1" blockId="96.[151,1436,1943,2004]" pageId="96" pageNumber="97">
<emphasis id="8DF4EA9FFF85FF97FA93F810D4F11ADD" bold="true" box="[151,240,1943,1967]" pageId="96" pageNumber="97">FIG. 61.</emphasis>
<emphasis id="8DF4EA9FFF85FF97FAF1F810D6211AC2" box="[245,544,1943,1968]" italics="true" pageId="96" pageNumber="97">Bçtryllçphẚlus cçnẚçrhynchus</emphasis>
<emphasis id="8DF4EA9FFF85FF97F821F81FD6751AC2" bold="true" box="[549,628,1944,1968]" pageId="96" pageNumber="97">sp. nov.</emphasis>
, female.A, rostrum; B, maxillule; C, maxilla; D, right leg 1; E, left leg 1; F, right leg 2; G, left leg 2. Scale bars: 0.02 mm.
</paragraph>
</caption>
<caption id="EBFF6605FF84FF96FA93F810D7651AA1" ID-DOI="http://doi.org/10.5281/zenodo.4821004" ID-Zenodo-Dep="4821004" httpUri="https://zenodo.org/record/4821004/files/figure.png" pageId="97" pageNumber="98" startId="97.[151,198,1943,1967]" targetBox="[204,1386,196,1906]" targetPageId="97">
<paragraph id="BF3F368DFF84FF96FA93F810D7651AA1" blockId="97.[151,1437,1943,2004]" pageId="97" pageNumber="98">
<emphasis id="8DF4EA9FFF84FF96FA93F810D4F01ADD" bold="true" box="[151,241,1943,1967]" pageId="97" pageNumber="98">FIG. 62.</emphasis>
<emphasis id="8DF4EA9FFF84FF96FAF3F810D6251AC2" box="[247,548,1943,1968]" italics="true" pageId="97" pageNumber="98">Bçtryllçphẚlus cçnẚçrhynchus</emphasis>
<emphasis id="8DF4EA9FFF84FF96F82EF81FD67B1AC2" bold="true" box="[554,634,1944,1968]" pageId="97" pageNumber="98">sp. nov.</emphasis>
, female. A, maxilliped; B, right leg 3; C, left leg 3; D, right leg 4; E, left leg 4; F, leg 5; G, genital aperture. Scale bars: A-E, G, 0.02 mm; F, 0.05 mm.
</paragraph>
</caption>
<paragraph id="BF3F368DFF87FF95FAC3FF10D7051C87" blockId="98.[151,1437,151,718]" pageId="98" pageNumber="99">
Labrum (
<figureCitation id="27BB2A08FF87FF95FB2BFF10D5951DC0" box="[303,404,151,178]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="98" pageNumber="99">Fig. 60H</figureCitation>
) with large, semicircular posteromedian lobe. Mandible (
<figureCitation id="27BB2A08FF87FF95FE10FF10D06F1DC0" box="[1044,1134,151,178]" captionStart="FIG" captionStartId="95.[151,198,1943,1967]" captionTargetBox="[209,1381,189,1901]" captionTargetId="figure-19@95.[192,1395,181,1919]" captionTargetPageId="95" captionText="FIG. 60. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, habitus, dorsal; B, habitus, left; C, posterior region of body, dorsal; D, caudal ramus; E, antennule; F, left antenna; G, distal part of right antenna; H, labrum; I, mandible. Scale bars: A-C, 0.1 mm; D-I, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820979" httpUri="https://zenodo.org/record/4820979/files/figure.png" pageId="98" pageNumber="99">Fig. 60I</figureCitation>
) with 1 bifurcate tooth and 2 shorter teeth on coxal gnathobase; palp with 2 lobes in middle of inner margin, armed with 9 setae arranged as 3, 2, 2, and 2. Maxillule (
<figureCitation id="27BB2A08FF87FF95FBBEFF58D61E1D88" box="[442,543,223,250]" captionStart="FIG" captionStartId="96.[151,198,1943,1967]" captionTargetBox="[201,1378,191,1903]" captionTargetId="figure-23@96.[189,1398,181,1919]" captionTargetPageId="96" captionText="FIG. 61. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female.A, rostrum; B, maxillule; C, maxilla; D, right leg 1; E, left leg 1; F, right leg 2; G, left leg 2. Scale bars: 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820994" httpUri="https://zenodo.org/record/4820994/files/figure.png" pageId="98" pageNumber="99">Fig. 61B</figureCitation>
) consisting of precoxa and palp; precoxa with 5 distinct setae on arthrite; palp 3-segmented, consisting of coxa, basis, and endopod; coxa unarmed (lacking seta of epipodite); basis with 2 setae on medial margin, 3 setae on outer margin, and setulose distal lobe; endopod articulated from basis, armed with 3 setae on distal margin. Maxilla (
<figureCitation id="27BB2A08FF87FF95F819FECCD6821C14" box="[541,643,331,358]" captionStart="FIG" captionStartId="96.[151,198,1943,1967]" captionTargetBox="[201,1378,191,1903]" captionTargetId="figure-23@96.[189,1398,181,1919]" captionTargetPageId="96" captionText="FIG. 61. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female.A, rostrum; B, maxillule; C, maxilla; D, right leg 1; E, left leg 1; F, right leg 2; G, left leg 2. Scale bars: 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820994" httpUri="https://zenodo.org/record/4820994/files/figure.png" pageId="98" pageNumber="99">Fig. 61C</figureCitation>
) indistinctly 3-segmented and armed with 2, 4, and 3 setae on first to third segments, respectively. Maxilliped (
<figureCitation id="27BB2A08FF87FF95F86AFEE8D6D71CF8" box="[622,726,367,394]" captionStart="FIG" captionStartId="97.[151,198,1943,1967]" captionTargetBox="[204,1386,196,1906]" captionTargetId="figure-19@97.[191,1397,181,1919]" captionTargetPageId="97" captionText="FIG. 62. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, maxilliped; B, right leg 3; C, left leg 3; D, right leg 4; E, left leg 4; F, leg 5; G, genital aperture. Scale bars: A-E, G, 0.02 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4821004" httpUri="https://zenodo.org/record/4821004/files/figure.png" pageId="98" pageNumber="99">Fig. 62A</figureCitation>
) 4-segmented; syncoxa large, with 1 small seta proximally on inner margin and patch of spinules subdistally on inner side; basis with 2 small isolated setae; short first endopodal segment unarmed; second endopodal segment with 2 small setules and 1 pointed inner distal process; terminal claw small, about half as long as second endopodal segment.
</paragraph>
<paragraph id="BF3F368DFF87FF95FAC3FE78D6A51FBF" blockId="98.[151,1437,151,718]" pageId="98" pageNumber="99">
Legs 1-4 (
<figureCitation id="27BB2A08FF87FF95FB3BFE78D59D1F6B" box="[319,412,511,537]" captionStart="FIG" captionStartId="96.[151,198,1943,1967]" captionTargetBox="[201,1378,191,1903]" captionTargetId="figure-23@96.[189,1398,181,1919]" captionTargetPageId="96" captionText="FIG. 61. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female.A, rostrum; B, maxillule; C, maxilla; D, right leg 1; E, left leg 1; F, right leg 2; G, left leg 2. Scale bars: 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4820994" httpUri="https://zenodo.org/record/4820994/files/figure.png" pageId="98" pageNumber="99">Figs. 61</figureCitation>
D-G, 62B-E) biramous, with 1-segmented exopods and 2-segmented endopods; coxa unarmed; basis with 1 seta on outer margin. Basis of legs 1-3 with 1 prominent, spiniform process at inner distal corner and 2 to 4 spinules on distal margin near base of endopod; basis of leg 4 with 2 patches of several spinules at each inner distal corner and on distal margin. Spines on exopods of right legs 1-3 elongate. Exopods of right legs 1 and 2 armed with 1 seta (outer proximal element) and 5 spines. Numbers of spines (Roman numerals) and setae (Arabic numerals) on right and left legs 1-4 as follows:
</paragraph>
<paragraph id="BF3F368DFF87FF95FB99FC84D0C11EB9" pageId="98" pageNumber="99">
<table id="CD80C42DFF870008FAA7FC84D1211EBE" box="[163,1312,771,972]" colsContinueFrom="91.[163,1326,1669,1884]" colsContinueIn="101.[163,1305,914,1112]" gridcols="5" gridrows="5" pageId="98" pageNumber="99">
<tr id="01B034CFFF870008FAA7FC84D1211E69" box="[163,1312,771,795]" gridrow="0" pageId="98" pageNumber="99" rowspan-0="1">
<th id="42615DB3FF870008FB99FC84D6261E69" box="[413,551,771,795]" gridcol="1" gridrow="0" pageId="98" pageNumber="99">Right exopod</th>
<th id="42615DB3FF870008F89CFC84D72C1E69" box="[664,813,771,795]" gridcol="2" gridrow="0" pageId="98" pageNumber="99">Right endopod</th>
<th id="42615DB3FF870008F9ADFC84D0241E69" box="[937,1061,771,795]" gridcol="3" gridrow="0" pageId="98" pageNumber="99">Left exopod</th>
<th id="42615DB3FF870008FE9DFC84D1211E69" box="[1177,1312,771,795]" gridcol="4" gridrow="0" pageId="98" pageNumber="99">Left endopod</th>
</tr>
<tr id="01B034CFFF870008FAA7FCA8D1211E35" box="[163,1312,815,839]" gridrow="1" pageId="98" pageNumber="99">
<th id="42615DB3FF870008FAA7FCA8D4DC1E35" box="[163,221,815,839]" gridcol="0" gridrow="1" pageId="98" pageNumber="99">Leg 1</th>
<td id="42615DB3FF870008FB99FCA8D6261E35" box="[413,551,815,839]" gridcol="1" gridrow="1" pageId="98" pageNumber="99">V+1</td>
<td id="42615DB3FF870008F89CFCA8D72C1E35" box="[664,813,815,839]" gridcol="2" gridrow="1" pageId="98" pageNumber="99">1; 7</td>
<td id="42615DB3FF870008F9ADFCA8D0241E35" box="[937,1061,815,839]" gridcol="3" gridrow="1" pageId="98" pageNumber="99">5</td>
<td id="42615DB3FF870008FE9DFCA8D1211E35" box="[1177,1312,815,839]" gridcol="4" gridrow="1" pageId="98" pageNumber="99">1; 7</td>
</tr>
<tr id="01B034CFFF870008FAA7FCDCD1211E06" box="[163,1312,859,884]" gridrow="2" pageId="98" pageNumber="99">
<th id="42615DB3FF870008FAA7FCDCD4DC1E06" box="[163,221,859,884]" gridcol="0" gridrow="2" pageId="98" pageNumber="99">Leg 2</th>
<td id="42615DB3FF870008FB99FCDCD6261E06" box="[413,551,859,884]" gridcol="1" gridrow="2" pageId="98" pageNumber="99">V+1</td>
<td id="42615DB3FF870008F89CFCDCD72C1E06" box="[664,813,859,884]" gridcol="2" gridrow="2" pageId="98" pageNumber="99">1; 7</td>
<td id="42615DB3FF870008F9ADFCDCD0241E06" box="[937,1061,859,884]" gridcol="3" gridrow="2" pageId="98" pageNumber="99">5</td>
<td id="42615DB3FF870008FE9DFCDCD1211E06" box="[1177,1312,859,884]" gridcol="4" gridrow="2" pageId="98" pageNumber="99">1; 7</td>
</tr>
<tr id="01B034CFFF870008FAA7FC00D1211ED2" box="[163,1312,903,928]" gridrow="3" pageId="98" pageNumber="99">
<th id="42615DB3FF870008FAA7FC00D4DC1ED2" box="[163,221,903,928]" gridcol="0" gridrow="3" pageId="98" pageNumber="99">Leg 3</th>
<td id="42615DB3FF870008FB99FC00D6261ED2" box="[413,551,903,928]" gridcol="1" gridrow="3" pageId="98" pageNumber="99">VI</td>
<td id="42615DB3FF870008F89CFC00D72C1ED2" box="[664,813,903,928]" gridcol="2" gridrow="3" pageId="98" pageNumber="99">1; 5</td>
<td id="42615DB3FF870008F9ADFC00D0241ED2" box="[937,1061,903,928]" gridcol="3" gridrow="3" pageId="98" pageNumber="99">5</td>
<td id="42615DB3FF870008FE9DFC00D1211ED2" box="[1177,1312,903,928]" gridcol="4" gridrow="3" pageId="98" pageNumber="99">1; 5</td>
</tr>
<tr id="01B034CFFF870008FAA7FC34D1211EBE" box="[163,1312,947,972]" gridrow="4" pageId="98" pageNumber="99">
<th id="42615DB3FF870008FAA7FC34D4DC1EBE" box="[163,221,947,972]" gridcol="0" gridrow="4" pageId="98" pageNumber="99">Leg 4</th>
<td id="42615DB3FF870008FB99FC34D6261EBE" box="[413,551,947,972]" gridcol="1" gridrow="4" pageId="98" pageNumber="99">IV</td>
<td id="42615DB3FF870008F89CFC34D72C1EBE" box="[664,813,947,972]" gridcol="2" gridrow="4" pageId="98" pageNumber="99">1; 5</td>
<td id="42615DB3FF870008F9ADFC34D0241EBE" box="[937,1061,947,972]" gridcol="3" gridrow="4" pageId="98" pageNumber="99">4</td>
<td id="42615DB3FF870008FE9DFC34D1211EBE" box="[1177,1312,947,972]" gridcol="4" gridrow="4" pageId="98" pageNumber="99">1; 5</td>
</tr>
</table>
</paragraph>
<paragraph id="BF3F368DFF87FF95FAC3FC79D76319F1" blockId="98.[151,1437,1021,1156]" pageId="98" pageNumber="99">
Leg 5 (
<figureCitation id="27BB2A08FF87FF95FB1FFC7AD57F196A" box="[283,382,1021,1048]" captionStart="FIG" captionStartId="97.[151,198,1943,1967]" captionTargetBox="[204,1386,196,1906]" captionTargetId="figure-19@97.[191,1397,181,1919]" captionTargetPageId="97" captionText="FIG. 62. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, maxilliped; B, right leg 3; C, left leg 3; D, right leg 4; E, left leg 4; F, leg 5; G, genital aperture. Scale bars: A-E, G, 0.02 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4821004" httpUri="https://zenodo.org/record/4821004/files/figure.png" pageId="98" pageNumber="99">Fig. 62F</figureCitation>
) positioned dorsolaterally on fifth pedigerous somite, evenly tapering, 2.4 times longer than wide (140×58 μm), with 1 small seta proximally and 2 small plus 1 larger seta distally; larger distal seta longer than leg (exopodal) segment. Leg 6 (
<figureCitation id="27BB2A08FF87FF95FBFAFBC2D6651912" box="[510,612,1093,1120]" captionStart="FIG" captionStartId="97.[151,198,1943,1967]" captionTargetBox="[204,1386,196,1906]" captionTargetId="figure-19@97.[191,1397,181,1919]" captionTargetPageId="97" captionText="FIG. 62. Bçtryllçphẚlus cçnẚçrhynchus sp. nov., female. A, maxilliped; B, right leg 3; C, left leg 3; D, right leg 4; E, left leg 4; F, leg 5; G, genital aperture. Scale bars: A-E, G, 0.02 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4821004" httpUri="https://zenodo.org/record/4821004/files/figure.png" pageId="98" pageNumber="99">Fig. 62G</figureCitation>
) represented by 1 spinule and 2 spinule-like processes on genital operculum; 9 small denticle-like elements on surface adjacent to leg 6.
</paragraph>
<paragraph id="BF3F368DFF87FF95FA93FB36D55919B9" blockId="98.[151,344,1201,1227]" box="[151,344,1201,1227]" pageId="98" pageNumber="99">
<emphasis id="8DF4EA9FFF87FF95FA93FB36D4DA19B9" bold="true" box="[151,219,1201,1227]" pageId="98" pageNumber="99">
<collectingRegion id="7D44F86FFF87FF95FA93FB36D4D719B9" box="[151,214,1201,1227]" country="Maldives" name="Maale" pageId="98" pageNumber="99">Male</collectingRegion>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FF87FF95FA93FB7ED66618BA" pageId="98" pageNumber="99" type="discussion">
<paragraph id="BF3F368DFF87FF95FA93FB7ED66618BA" blockId="98.[151,1436,1273,1481]" pageId="98" pageNumber="99">
<emphasis id="8DF4EA9FFF87FF95FA93FB7ED50A1861" bold="true" box="[151,267,1273,1299]" pageId="98" pageNumber="99">Remarks.</emphasis>
Three outstanding features serve to characterize
<emphasis id="8DF4EA9FFF87FF95F92FFB7DD7EF1867" box="[811,1006,1274,1301]" italics="true" pageId="98" pageNumber="99">BK cçnẚçrhynchus</emphasis>
<emphasis id="8DF4EA9FFF87FF95F9F0FB7ED04D1861" bold="true" box="[1012,1100,1273,1299]" pageId="98" pageNumber="99">sp. nov.</emphasis>
as follows: (1) the exopods of right legs 1 and 2 each carry 1 seta in addition to spines; (2) the combination of armature elements on right legs 1-4 is 6-6-6-4, and (3) the endopod of the antenna is apparently 3-segmented, the segmentation has not been recorded previously in
<emphasis id="8DF4EA9FFF87FF95FB36FAE1D5D318F3" box="[306,466,1382,1409]" italics="true" pageId="98" pageNumber="99">Bçtryllçphẚlus</emphasis>
and, as it appears incomplete, it may represent a distinct fold in the cuticle rather than a true articulation. The combination of armature elements on the exopods of left legs 1-4 (5-5-5-4) also is unusual, known previously only in
<emphasis id="8DF4EA9FFF87FF95FBB9FA29D5CF18B5" box="[445,462,1454,1479]" italics="true" pageId="98" pageNumber="99">B</emphasis>
.
<emphasis id="8DF4EA9FFF87FF95FBD8FA29D66118BB" box="[476,608,1454,1481]" italics="true" pageId="98" pageNumber="99">banyulensẚs</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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