treatments-xml/data/37/29/87/3729879BFF0FFF1BFA93FC71D1741C30.xml
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<document id="B8FAF3441BBF04097773DED3F1F72D54" ID-DOI="10.11646/zootaxa.4978.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4820443" ID-ZooBank="9C7C1723-73EB-4FBE-A47A-54627DEB8F93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622139238996" checkinUser="plazi" docAuthor="Kim, Il-Hoi &amp; Boxshall, Geoff A." docDate="2021" docId="3729879BFF0FFF1BFA93FC71D1741C30" docLanguage="en" docName="zootaxa.4978.1.1.pdf" docOrigin="Zootaxa 1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Enterocola nodulosus Kim &amp; Boxshall 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="236" masterDocId="CB10FFE3FFE5FFF7FA04FF87D4011D72" masterDocTitle="Copepods (Cyclopoida) associated with ascidian hosts: Ascidicolidae, Buproridae, Botryllophilidae, and Enteropsidae, with descriptions of 84 new species" masterLastPageNumber="286" masterPageNumber="1" pageNumber="235" updateTime="1698948597350" updateUser="plazi">
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<mods:title id="FC91FD31188C65FF058A50E7F061F1E2">Copepods (Cyclopoida) associated with ascidian hosts: Ascidicolidae, Buproridae, Botryllophilidae, and Enteropsidae, with descriptions of 84 new species</mods:title>
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<mods:name id="0CBBF7B5A7ED01CE2A309EEE088FF504" type="personal">
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<mods:namePart id="A5644079243ECB3280AFC80127557C12">Kim, Il-Hoi</mods:namePart>
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<mods:namePart id="238FF87AB6F290E604B466832B5A6249">Boxshall, Geoff A.</mods:namePart>
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<mods:title id="5A4F7D13ECE1839562CD4EAE066728E8">Zootaxa</mods:title>
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<mods:part id="25F259529CFEF8EAA22A44DD06D31A9D">
<mods:date id="73A0FB2C5AB98D184F122398AB3C976F">2021</mods:date>
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<mods:classification id="73A77F90CAE8721B3775BDB5584A7FD9">journal article</mods:classification>
<mods:identifier id="7A7B8C4DFA6C3ACCC67F8711E43798B7" type="DOI">10.11646/zootaxa.4978.1.1</mods:identifier>
<mods:identifier id="0519864BC631F3CFB3DC2A0A11B69575" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3729879BFF0FFF1BFA93FC71D1741C30" ID-DOI="http://doi.org/10.5281/zenodo.5047155" ID-Zenodo-Dep="5047155" LSID="urn:lsid:plazi:treatment:3729879BFF0FFF1BFA93FC71D1741C30" httpUri="http://treatment.plazi.org/id/3729879BFF0FFF1BFA93FC71D1741C30" lastPageId="236" lastPageNumber="236" pageId="234" pageNumber="235">
<subSubSection id="F79A6506FF0FFF1DFA93FC71D6041963" box="[151,517,1014,1041]" pageId="234" pageNumber="235" type="nomenclature">
<paragraph id="BF3F368DFF0FFF1DFA93FC71D6041963" blockId="234.[151,517,1014,1077]" box="[151,517,1014,1041]" pageId="234" pageNumber="235">
<heading id="E47781E1FF0FFF1DFA93FC71D6041963" bold="true" box="[151,517,1014,1041]" fontSize="11" level="2" pageId="234" pageNumber="235" reason="4">
<emphasis id="8DF4EA9FFF0FFF1DFA93FC71D6041963" bold="true" box="[151,517,1014,1041]" pageId="234" pageNumber="235">
<taxonomicName id="78804D0EFF0FFF1DFA93FC71D5A11963" authority="Kim &amp; Boxshall, 2021" authorityName="Kim &amp; Boxshall" authorityYear="2021" box="[151,416,1014,1041]" class="Maxillopoda" family="Ascidicolidae" genus="Enterocola" kingdom="Animalia" order="Cyclopoida" pageId="234" pageNumber="235" phylum="Arthropoda" rank="species" species="nodulosus" status="sp. nov.">
<emphasis id="8DF4EA9FFF0FFF1DFA93FC71D5A11963" bold="true" box="[151,416,1014,1041]" italics="true" pageId="234" pageNumber="235">Enterocola nodulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96C757E4FF0FFF1DFBA3FC70D6041963" box="[423,517,1015,1041]" pageId="234" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FF0FFF1DFA93FB9CD53E1947" box="[151,319,1050,1077]" pageId="234" pageNumber="235" type="description">
<paragraph id="BF3F368DFF0FFF1DFA93FB9CD53E1947" blockId="234.[151,517,1014,1077]" box="[151,319,1050,1077]" pageId="234" pageNumber="235">
(
<figureCitation id="27BB2A08FF0FFF1DFAA4FB9CD5011947" box="[160,256,1050,1077]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153</figureCitation>
,
<figureCitation id="27BB2A08FF0FFF1DFB08FB9DD5361947" box="[268,311,1050,1077]" captionStart="FIG" captionStartId="235.[151,198,1125,1149]" captionTargetBox="[155,1430,385,1094]" captionTargetId="figure-122@235.[151,1436,381,1101]" captionTargetPageId="235" captionText="FIG. 154. bnterçcçla nçdulçsus sp. nov., female.A, leg 1; B, leg 3; C, leg 4. Scale bars: 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821867" httpUri="https://zenodo.org/record/4821867/files/figure.png" pageId="234" pageNumber="235">154</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FF0FFF1DFA93FBE4D5EF19D3" pageId="234" pageNumber="235" type="materials_examined">
<paragraph id="BF3F368DFF0FFF1DFA93FBE4D5EF19D3" blockId="234.[151,1437,1122,2013]" pageId="234" pageNumber="235">
<emphasis id="8DF4EA9FFF0FFF1DFA93FBE4D545190E" bold="true" box="[151,324,1122,1149]" pageId="234" pageNumber="235">Type material.</emphasis>
<materialsCitation id="0FE83CD0FF0FFF1DFB4FFBE4D5EF19D3" country="Indonesia" location="Indonesia" pageId="234" pageNumber="235" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="603B882FFF0FFF1DFB4FFBE4D5B2190F" box="[331,435,1123,1149]" pageId="234" pageNumber="235" type="holotype">Holotype</typeStatus>
<specimenCount id="A986FD04FF0FFF1DFBBDFBE4D5CF190F" box="[441,462,1123,1149]" pageId="234" pageNumber="235" type="female"></specimenCount>
(MNHN-IU-2014-21563, dissected and mounted on a slide) from
<emphasis id="8DF4EA9FFF0FFF1DFEB7FBE3D12A190D" box="[1203,1323,1124,1151]" italics="true" pageId="234" pageNumber="235">budẚstçma</emphasis>
sp.;
<collectingCountry id="C797761DFF0FFF1DFF64FBE4D4D219D3" name="Indonesia" pageId="234" pageNumber="235">Indonesia</collectingCountry>
, no other collection data.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FF0FFF1DFAC3FB2CD548199B" pageId="234" pageNumber="235" type="etymology">
<paragraph id="BF3F368DFF0FFF1DFAC3FB2CD548199B" blockId="234.[151,1437,1122,2013]" pageId="234" pageNumber="235">
<emphasis id="8DF4EA9FFF0FFF1DFAC3FB2CD54F19B7" bold="true" box="[199,334,1195,1221]" pageId="234" pageNumber="235">Etymology.</emphasis>
The name of the new species alludes to the presence of a distal nodule on the endopods of the swimming legs.
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FF0FFF1CFAC3FB75D55919B7" lastPageId="235" lastPageNumber="236" pageId="234" pageNumber="235" type="description">
<paragraph id="BF3F368DFF0FFF1DFAC3FB75D5321B5F" blockId="234.[151,1437,1122,2013]" pageId="234" pageNumber="235">
<emphasis id="8DF4EA9FFF0FFF1DFAC3FB75D5CC187E" bold="true" box="[199,461,1266,1292]" pageId="234" pageNumber="235">Description of female.</emphasis>
Body (
<figureCitation id="27BB2A08FF0FFF1DF81BFB74D67A187F" box="[543,635,1266,1293]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153</figureCitation>
A-C) eruciform, curved dorsally; body length 909 μm; maximum width 350 μm (across second pedigerous somite). Cephalosome with convex posterodorsal margin. Trunk unsegmented, but pedigerous somites defined from one another by dorsal and lateral constrictions; second to fourth pedigerous somites each bearing pair of dorsal tergal folds (
<figureCitation id="27BB2A08FF0FFF1DF8A9FAD8D741180B" box="[685,832,1374,1401]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153A, C</figureCitation>
) and single, large, plate-like ventral interpodal protrusion between left and right legs (
<figureCitation id="27BB2A08FF0FFF1DF80AFA04D68518EF" box="[526,644,1410,1437]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153B</figureCitation>
). Genitoabdomen (
<figureCitation id="27BB2A08FF0FFF1DF958FA04D7D018EF" box="[860,977,1410,1437]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153D</figureCitation>
) unsegmented dorsally but 4-segmented ventrally; segmentation indistinct between genital and first free abdominal somites. Free abdominal somites covered with ornamentation of minute spinules on all surfaces; anal prominence distinct. Caudal rami (
<figureCitation id="27BB2A08FF0FFF1DFEBAFA4CD1301897" box="[1214,1329,1482,1509]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153D</figureCitation>
) straight, incompletely articulated from anal somite, gradually narrowing distally, unarmed, about 2.6 times longer than wide (100×38 μm).
</paragraph>
<paragraph id="BF3F368DFF0FFF1DFAC3F9B0D05E1B93" blockId="234.[151,1437,1122,2013]" pageId="234" pageNumber="235">
Rostrum absent. Antennule (
<figureCitation id="27BB2A08FF0FFF1DF800F9B0D6751B23" box="[516,628,1590,1617]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153E</figureCitation>
) leaf-like, 39×27 μm, inflated posteriorly, with convex posterior margin; armed with 5 small, subequal setae at apex. Antenna (
<figureCitation id="27BB2A08FF0FFF1DF8F5F9DCD7611B07" box="[753,864,1626,1653]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153F</figureCitation>
) about 2.6 times longer than wide, incompletely 2- segmented, slightly curved medially; proximal segment unarmed; distal segment ornamented with minute spinules on convex surface; armed with 6 thin setae, lengths of setae I-VI 20, 18, 23, 19, 32, and 64 μm, respectively; seta I naked, but other 5 setae weakly pinnate; setae I-III widely separated from setae IV-VI.
</paragraph>
<paragraph id="BF3F368DFF0FFF1DFAC3F96CD11B1ACB" blockId="234.[151,1437,1122,2013]" pageId="234" pageNumber="235">
Labrum (
<figureCitation id="27BB2A08FF0FFF1DFB37F96CD5A81A77" box="[307,425,1770,1797]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153G</figureCitation>
) with spinulose, distally swollen palps. Mandible (
<figureCitation id="27BB2A08FF0FFF1DF9F9F96CD0751A77" box="[1021,1140,1770,1797]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153G</figureCitation>
) spinulose, tapering, narrower than labral palp. Maxillule consisting of precoxa (
<figureCitation id="27BB2A08FF0FFF1DF90DF888D77D1A5B" box="[777,892,1806,1833]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153H</figureCitation>
) and palm-shaped palp (
<figureCitation id="27BB2A08FF0FFF1DFE89F888D0F91A5B" box="[1165,1272,1806,1833]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153I</figureCitation>
); precoxa with bifurcate distal part, 1 small tubercle subdistally, and endite in middle bearing 1 spinulose seta and more than 20 thin spinules; palp with 6 spinulose spines. Maxilla (
<figureCitation id="27BB2A08FF0FFF1DF8F3F8D0D7651A03" box="[759,868,1878,1905]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="234" pageNumber="235">Fig. 153J</figureCitation>
) with bifurcate, spinulose mediodistal process on proximal segment; distal segment with 1 robust, basally articulated, spinulose spine subdistally, 1 transverse row of minute spinules on middle of anterior surface, and 1 small seta on posterior surface. Maxilliped absent.
</paragraph>
<paragraph id="BF3F368DFF0FFF1CFAC3F844D06D1C33" blockId="234.[151,1437,1122,2013]" lastBlockId="235.[151,1437,150,357]" lastPageId="235" lastPageNumber="236" pageId="234" pageNumber="235">
Legs 1-4 each consisting of unarmed, 2-segmented protopod and unsegmented rami. Exopods bearing patch of spinules proximally on lateral margin ((
<figureCitation id="27BB2A08FF0EFF1CF858FF10D6BA1DC3" box="[604,699,150,177]" captionStart="FIG" captionStartId="235.[151,198,1125,1149]" captionTargetBox="[155,1430,385,1094]" captionTargetId="figure-122@235.[151,1436,381,1101]" captionTargetPageId="235" captionText="FIG. 154. bnterçcçla nçdulçsus sp. nov., female.A, leg 1; B, leg 3; C, leg 4. Scale bars: 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821867" httpUri="https://zenodo.org/record/4821867/files/figure.png" pageId="235" pageNumber="236">Fig. 154</figureCitation>
A-C). Endopods armed distally with 2 naked setae and 1 nodule. Sizes of endopods 55×25, 55×24, 55×32, and 48×27 μm, respectively, in legs 1-4; endopods of legs 3 and 4 with markedly protruded lateral margin. Lengths of laterodistal and mediodistal setae 56 and 50, 55 and 43, 55 and 40, and 58 and 39 μm, respectively, in legs 1-4. Laterodistal setae longer than or as long as endopodal segments and about 1.1, 1.3, 1.4, and 1.5 times longer than mediodistal setae in legs 1-4, respectively.
</paragraph>
<paragraph id="BF3F368DFF0EFF1CFAC3FECCD0251C17" blockId="235.[151,1437,150,357]" box="[199,1060,330,357]" pageId="235" pageNumber="236">
Leg 5 (
<figureCitation id="27BB2A08FF0EFF1CFB13FECCD58A1C17" box="[279,395,330,357]" captionStart="FIG" captionStartId="233.[151,198,1943,1967]" captionTargetBox="[210,1372,191,1907]" captionTargetId="figure-19@233.[201,1387,181,1919]" captionTargetPageId="233" captionText="FIG. 153. bnterçcçla nçdulçsus sp. nov., female. A, habitus, dorsal; B, habitus, ventral; C, habitus, right; D, genitoabdomen, ventral; E, antennule; F, antenna; G, labrum and mandibles; H, precoxa of maxillule; I, palp of maxillule; J, maxilla; K, leg 5. Scale bars: A-C, 0.1 mm; D, K, 0.05 mm; E-J, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4821855" httpUri="https://zenodo.org/record/4821855/files/figure.png" pageId="235" pageNumber="236">Fig. 153K</figureCitation>
) lamellate, unarmed, much wider than long (129×255 μm).
</paragraph>
<caption id="EBFF6605FF0EFF1CFA93FBE2D083190F" ID-DOI="http://doi.org/10.5281/zenodo.4821867" ID-Zenodo-Dep="4821867" box="[151,1154,1125,1149]" httpUri="https://zenodo.org/record/4821867/files/figure.png" pageId="235" pageNumber="236" startId="235.[151,198,1125,1149]" targetBox="[155,1430,385,1094]" targetPageId="235">
<paragraph id="BF3F368DFF0EFF1CFA93FBE2D083190F" blockId="235.[151,1154,1125,1149]" box="[151,1154,1125,1149]" pageId="235" pageNumber="236">
<emphasis id="8DF4EA9FFF0EFF1CFA93FBE2D4FC190F" bold="true" box="[151,253,1125,1149]" pageId="235" pageNumber="236">FIG. 154.</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFB00FBE2D5E0190F" box="[260,481,1125,1149]" italics="true" pageId="235" pageNumber="236">bnterçcçla nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFBE3FBE2D636190F" bold="true" box="[487,567,1125,1149]" pageId="235" pageNumber="236">sp. nov.</emphasis>
, female. A, leg 1; B, leg 3; C, leg 4. Scale bars: 0.02 mm.
</paragraph>
</caption>
<paragraph id="BF3F368DFF0EFF1CFA93FB2CD55919B7" blockId="235.[151,344,1195,1221]" box="[151,344,1195,1221]" pageId="235" pageNumber="236">
<emphasis id="8DF4EA9FFF0EFF1CFA93FB2CD4DA19B7" bold="true" box="[151,219,1195,1221]" pageId="235" pageNumber="236">
<collectingRegion id="7D44F86FFF0EFF1CFA93FB2CD4D719B7" box="[151,214,1195,1221]" country="Maldives" name="Maale" pageId="235" pageNumber="236">Male</collectingRegion>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F79A6506FF0EFF1BFA93FB74D1741C30" lastPageId="236" lastPageNumber="237" pageId="235" pageNumber="236" type="discussion">
<paragraph id="BF3F368DFF0EFF1CFA93FB74D0AE1B5C" blockId="235.[151,1437,1267,2014]" pageId="235" pageNumber="236">
<emphasis id="8DF4EA9FFF0EFF1CFA93FB74D50A187F" bold="true" box="[151,267,1267,1293]" pageId="235" pageNumber="236">Remarks.</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFB17FB74D608187F" box="[275,521,1267,1293]" italics="true" pageId="235" pageNumber="236">bnterçcçla nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CF814FB74D66A187F" bold="true" box="[528,619,1267,1293]" pageId="235" pageNumber="236">sp. nov.</emphasis>
is characterized by the presence of a nodule (small tubercle) distally on the endopods of swimming legs 1 to 4. In addition, the unsegmented antennule of the new species is inflated and leaf-like. This form of the antennule (bulbous or D-shaped) is shared with ten known species of
<emphasis id="8DF4EA9FFF0EFF1CFED6FABCD14C1827" box="[1234,1357,1339,1365]" italics="true" pageId="235" pageNumber="236">bnterçcçla</emphasis>
(
<tableCitation id="F2020336FF0EFF1CFF59FABCD4A61808" captionStart="TABLE 6" captionStartId="191.[151,239,1139,1163]" captionTargetPageId="191" captionText="TABLE 6. Summary of key differences between species of bnterçcçla. (= indicates approximate ratio or number)" httpUri="http://table.plazi.org/id/EBFF6605FF5AFF48FA93FBF4D15919F9" pageId="235" pageNumber="236" tableUuid="EBFF6605FF5AFF48FA93FBF4D15919F9">Table 6</tableCitation>
). Interestingly, in species with this form of antennule, the body is small (mostly less than 1.0 mm long), and the laterodistal setae on the endopods of swimming legs are shorter than endopodal segments but as long as or longer than the mediodistal setae. Six of those species (
<emphasis id="8DF4EA9FFF0EFF1CF8C8FA20D6DC18B3" box="[716,733,1447,1473]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF8E9FA20D7AB18B3" box="[749,938,1447,1473]" italics="true" pageId="235" pageNumber="236">parapterçphçrus</emphasis>
,
<emphasis id="8DF4EA9FFF0EFF1CF9BFFA20D7CD18B3" box="[955,972,1447,1473]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF9D9FA2ED05918B6" box="[989,1112,1449,1476]" italics="true" pageId="235" pageNumber="236">dẚcaudatus</emphasis>
,
<emphasis id="8DF4EA9FFF0EFF1CFE6DFA20D07B18B3" box="[1129,1146,1447,1473]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFE8FFA2ED10C18B6" box="[1163,1293,1449,1476]" italics="true" pageId="235" pageNumber="236">quadrẚsetus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFF13FA20D17218B3" bold="true" box="[1303,1395,1447,1473]" pageId="235" pageNumber="236">sp. nov.</emphasis>
,
<emphasis id="8DF4EA9FFF0EFF1CFF80FA20D19418B3" box="[1412,1429,1447,1473]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFA93FA4AD514189A" box="[151,277,1485,1512]" italics="true" pageId="235" pageNumber="236">mabulensẚs</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFB1FFA4CD5751897" bold="true" box="[283,372,1483,1509]" pageId="235" pageNumber="236">sp. nov.</emphasis>
,
<emphasis id="8DF4EA9FFF0EFF1CFB85FA4CD5931897" box="[385,402,1483,1509]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFB9BFA4BD5EA1897" box="[415,491,1484,1509]" italics="true" pageId="235" pageNumber="236">parvus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFBF6FA4CD64B1897" bold="true" box="[498,586,1483,1509]" pageId="235" pageNumber="236">
<taxonomicNameLabel id="96C757E4FF0EFF1CFBF6FA4CD64B1897" box="[498,586,1483,1509]" pageId="235" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<emphasis id="8DF4EA9FFF0EFF1CF882FA4CD6961897" box="[646,663,1483,1509]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF8A0FA4AD753189A" box="[676,850,1485,1512]" italics="true" pageId="235" pageNumber="236">neçcaledçnẚcus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CF95DFA4CD7B01897" bold="true" box="[857,945,1483,1509]" pageId="235" pageNumber="236">sp. nov.</emphasis>
) can be eliminated from further comparison because they have fewer than 6 setae on the antenna (cf. 6 setae in
<emphasis id="8DF4EA9FFF0EFF1CF97BFA68D7911B7B" box="[895,912,1519,1545]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF99AFA68D0111B7B" box="[926,1040,1519,1545]" italics="true" pageId="235" pageNumber="236">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFE12FA68D0711B7B" bold="true" box="[1046,1136,1519,1545]" pageId="235" pageNumber="236">sp. nov.</emphasis>
), or 1 or 2 dorsal tubercles or processes on the cephalosome or pedigerous somites (cf. lacking in
<emphasis id="8DF4EA9FFF0EFF1CF9A9F994D7BF1B5F" box="[941,958,1555,1581]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF9C8F994D03F1B5F" box="[972,1086,1555,1581]" italics="true" pageId="235" pageNumber="236">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFE41F994D09F1B5F" bold="true" box="[1093,1182,1555,1581]" pageId="235" pageNumber="236">sp. nov.</emphasis>
).
</paragraph>
<paragraph id="BF3F368DFF0EFF1BFAC3F9B0D1741C30" blockId="235.[151,1437,1267,2014]" lastBlockId="236.[151,1437,151,322]" lastPageId="236" lastPageNumber="237" pageId="235" pageNumber="236">
The remaining four species (
<emphasis id="8DF4EA9FFF0EFF1CF801F9B0D6171B23" box="[517,534,1591,1617]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF827F9BED6AF1B26" box="[547,686,1593,1620]" italics="true" pageId="235" pageNumber="236">mammẚferus</emphasis>
,
<emphasis id="8DF4EA9FFF0EFF1CF8BFF9B0D6CD1B23" box="[699,716,1591,1617]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF8DDF9B0D7601B23" box="[729,865,1591,1617]" italics="true" pageId="235" pageNumber="236">pterçphçrus</emphasis>
,
<emphasis id="8DF4EA9FFF0EFF1CF96AF9B0D77E1B23" box="[878,895,1591,1617]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF988F9B0D7EA1B23" box="[908,1003,1591,1617]" italics="true" pageId="235" pageNumber="236">rçbustus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CF9F6F9B0D04B1B23" bold="true" box="[1010,1098,1591,1617]" pageId="235" pageNumber="236">sp. nov.</emphasis>
, and
<taxonomicName id="78804D0EFF0EFF1CFE82F9B0D1091B23" authority="Kim &amp; Boxshall, 2021" authorityName="Kim &amp; Boxshall" authorityYear="2021" box="[1158,1288,1591,1618]" class="Maxillopoda" family="Ascidicolidae" genus="Enterocola" kingdom="Animalia" order="Cyclopoida" pageId="235" pageNumber="92" phylum="Arthropoda" rank="species" species="angustus" status="sp. nov.">
<emphasis id="8DF4EA9FFF0EFF1CFE82F9B0D0961B23" box="[1158,1175,1591,1617]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFEA0F9BFD1091B23" box="[1188,1288,1592,1617]" italics="true" pageId="235" pageNumber="236">angustus</emphasis>
</taxonomicName>
<emphasis id="8DF4EA9FFF0EFF1CFF09F9B0D1671B23" bold="true" box="[1293,1382,1591,1617]" pageId="235" pageNumber="236">
<taxonomicNameLabel id="96C757E4FF0EFF1CFF09F9B0D1671B23" box="[1293,1382,1591,1617]" pageId="235" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) can be distinguished from the new species as follows: in
<emphasis id="8DF4EA9FFF0EFF1CF8EEF9DCD6FA1B07" box="[746,763,1627,1653]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF90DF9DAD7951B0A" box="[777,916,1629,1656]" italics="true" pageId="235" pageNumber="236">mammẚferus</emphasis>
the caudal rami are wider than long (cf. about 2.6 times longer than wide in
<emphasis id="8DF4EA9FFF0EFF1CFBE6F9F8D5F21BEB" box="[482,499,1663,1689]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF804F9F8D6731BEB" box="[512,626,1663,1689]" italics="true" pageId="235" pageNumber="236">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CF87CF9F8D6D01BEB" bold="true" box="[632,721,1663,1689]" pageId="235" pageNumber="236">sp. nov.</emphasis>
), the antennule is armed with 4 setae (cf. 5 setae), the 6 setae on the antenna are subequal in length (cf. setae unequal), and the 2 distal setae on the endopods of legs 1-4 are almost equal in length, as described or illustrated by
<bibRefCitation id="DB114B7CFF0EFF1CF892F940D7421B90" author="Ooishi, S." box="[662,835,1735,1762]" pageId="235" pageNumber="236" pagination="312 - 323" refId="ref144518" refString="Ooishi, S. (2010 b) bntercccla brement ẚ Illg &amp; Dudley (Copepoda: Cyclopoida: Ascidicolidae) living in the compound ascidian Apl ẚdẚum pall ẚdum (Verrill), compared with b. mamm ẚferus Chatton &amp; Harant living in A. sperum Drasche. mrcceed ẚngs cf the B ẚclcgẚcal Scc ẚety cf tash ẚngtcn, 123 (4), 312 - 323. https: // doi. org / 10.2988 / 09 - 27.1" type="journal article" year="2010">Ooishi (2010b)</bibRefCitation>
. In
<emphasis id="8DF4EA9FFF0EFF1CF969F940D77F1B93" box="[877,894,1735,1761]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF988F940D0151B93" box="[908,1044,1735,1761]" italics="true" pageId="235" pageNumber="236">pterçphçrus</emphasis>
(as redescribed above), the ventral interpodal protrusion is absent on the pedigerous somites (cf. present, large in second to fourth pedigerous somites of
<emphasis id="8DF4EA9FFF0EFF1CFAB3F888D4C91A5B" box="[183,200,1807,1833]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFAD3F888D5481A5B" box="[215,329,1807,1833]" italics="true" pageId="235" pageNumber="236">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFB55F888D5AD1A5B" bold="true" box="[337,428,1807,1833]" pageId="235" pageNumber="236">sp. nov.</emphasis>
), the lateral seta (seta VI) of the antenna is 34 μm long, distinctly shorter than that of
<emphasis id="8DF4EA9FFF0EFF1CFF80F888D1941A5B" box="[1412,1429,1807,1833]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFA93F8B4D5081A3F" box="[151,265,1843,1869]" italics="true" pageId="235" pageNumber="236">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFB14F8B4D56A1A3F" bold="true" box="[272,363,1843,1869]" pageId="235" pageNumber="236">sp. nov.</emphasis>
(cf. 64 μm long), the labral palp is less expanded, and the mediodistal element of the proximal maxillary segment is articulated at base and simple, not bifurcate. In
<emphasis id="8DF4EA9FFF0EFF1CF988F8D0D79C1A03" box="[908,925,1879,1905]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CF9AEF8D0D0081A03" box="[938,1033,1879,1905]" italics="true" pageId="235" pageNumber="236">rçbustus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CFE0BF8D0D0661A03" bold="true" box="[1039,1127,1879,1905]" pageId="235" pageNumber="236">sp. nov.</emphasis>
the antennule is armed with 3 setae ((cf. 5 setae in
<emphasis id="8DF4EA9FFF0EFF1CFB91F8FCD5A71AE7" box="[405,422,1915,1941]" italics="true" pageId="235" pageNumber="236">b</emphasis>
.
<emphasis id="8DF4EA9FFF0EFF1CFBB7F8FCD6241AE7" box="[435,549,1915,1941]" italics="true" pageId="235" pageNumber="236">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF0EFF1CF82FF8FCD6851AE7" bold="true" box="[555,644,1915,1941]" pageId="235" pageNumber="236">sp. nov.</emphasis>
), the mediodistal element of the proximal maxillary segment is simple, not bifurcate, the setae on the antenna are shorter, the longest lateral seta (seta VI) is only 22 μm long (cf. 64 μm long), and the distal setae on the endopods of legs 1-4 are distinctly shorter than endopodal segments (cf. the setae are equal or subequal in length to endopodal segments in
<emphasis id="8DF4EA9FFF09FF1BF92EFF10D73A1DC3" box="[810,827,151,177]" italics="true" pageId="236" pageNumber="237">b</emphasis>
.
<emphasis id="8DF4EA9FFF09FF1BF94FFF10D7BC1DC3" box="[843,957,151,177]" italics="true" pageId="236" pageNumber="237">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF09FF1BF9C1FF10D0211DC3" bold="true" box="[965,1056,151,177]" pageId="236" pageNumber="237">sp. nov.</emphasis>
). Finally, in
<taxonomicName id="78804D0EFF09FF1BFEB0FF10D1381DC2" authority="Kim &amp; Boxshall, 2021" authorityName="Kim &amp; Boxshall" authorityYear="2021" box="[1204,1337,151,177]" class="Maxillopoda" family="Ascidicolidae" genus="Enterocola" kingdom="Animalia" order="Cyclopoida" pageId="236" pageNumber="92" phylum="Arthropoda" rank="species" species="angustus" status="sp. nov.">
<emphasis id="8DF4EA9FFF09FF1BFEB0FF10D0C41DC3" box="[1204,1221,151,177]" italics="true" pageId="236" pageNumber="237">b</emphasis>
.
<emphasis id="8DF4EA9FFF09FF1BFED1FF10D1381DC2" box="[1237,1337,151,176]" italics="true" pageId="236" pageNumber="237">angustus</emphasis>
</taxonomicName>
<emphasis id="8DF4EA9FFF09FF1BFF45FF10D19D1DC3" bold="true" box="[1345,1436,151,177]" pageId="236" pageNumber="237">
<taxonomicNameLabel id="96C757E4FF09FF1BFF45FF10D19D1DC3" box="[1345,1436,151,177]" pageId="236" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the caudal rami are strongly tapering and about 1.9 times longer than wide (cf. about 2.6 times longer than wide in
<emphasis id="8DF4EA9FFF09FF1BFA93FF58D4A91D8B" box="[151,168,223,249]" italics="true" pageId="236" pageNumber="237">b</emphasis>
.
<emphasis id="8DF4EA9FFF09FF1BFAB3FF58D5281D8B" box="[183,297,223,249]" italics="true" pageId="236" pageNumber="237">nçdulçsus</emphasis>
<emphasis id="8DF4EA9FFF09FF1BFB34FF58D58A1D8B" bold="true" box="[304,395,223,249]" pageId="236" pageNumber="237">sp. nov.</emphasis>
), the longest lateral seta (seta VI) of the antenna is 29 μm long (cf. 64 μm long), the ventral interpodal protrusions on pedigerous somites are obscure, weakly developed (cf. large and well developed), and the distal setae on the endopods of legs 1-4 are distinctly shorter than endopodal segment (cf. as long as or longer).
</paragraph>
</subSubSection>
</treatment>
</document>