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<document id="B225C2D3A4B27E736C0120806A2F765E" ID-CLB-Dataset="32047" ID-DOI="10.11646/zootaxa.4323.1.1" ID-GBIF-Dataset="27a65bbd-651d-438b-9f2b-93cfd23d2a1f" ID-ISSN="1175-5326" ID-Zenodo-Dep="897285" ID-ZooBank="94D28Ee0-6D90-401F-B060-A2Deff2Ada21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505881797759" checkinUser="plazi" docAuthor="Vidal, Nicolas" docDate="2017" docId="29190528FFC6C273FF1FFA9DFD1EF9A4" docLanguage="en" docName="zootaxa.4323.1.1.pdf" docOrigin="Zootaxa 4323 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anilios leptosoma" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="D5207D50FFCEC278FF88FF8EFF94FFDA" masterDocTitle="Worms in the sand: Systematic revision of the Australian blindsnake Anilios leptosoma (Robb, 1972) species complex (Squamata: Scolecophidia: Typhlopidae) from the Geraldton Sandplain, with description of two new species" masterLastPageNumber="24" masterPageNumber="1" pageNumber="9" updateTime="1698468714414" updateUser="ExternalLinkService">
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<mods:title id="FC6929DC59CFAC6EF7EECE345F7B97B3">Worms in the sand: Systematic revision of the Australian blindsnake Anilios leptosoma (Robb, 1972) species complex (Squamata: Scolecophidia: Typhlopidae) from the Geraldton Sandplain, with description of two new species</mods:title>
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<mods:namePart id="581D11C6703FE44FFBE91C33F591E56F">Vidal, Nicolas</mods:namePart>
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<mods:title id="AA48DCA54FD61C6FF37BB4D8DB4E8B19">Zootaxa</mods:title>
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<mods:part id="6E7DD27525682E6EA00C16056A5B2A55">
<mods:date id="7A7F37B5A759D8D5F4D5A8B96646BDA9">2017</mods:date>
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<mods:number id="DD13D55BD403E7D56EBEED0D0B6E88B8">2017-09-20</mods:number>
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<mods:number id="4A16333669FD70FD9E7CE81253B1E01B">4323</mods:number>
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<mods:identifier id="66C1811285F3AF34C272CB8BC419E240" type="DOI">10.11646/zootaxa.4323.1.1</mods:identifier>
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<treatment id="29190528FFC6C273FF1FFA9DFD1EF9A4" ID-DOI="http://doi.org/10.5281/zenodo.6002024" ID-GBIF-Taxon="134313800" ID-Zenodo-Dep="6002024" LSID="urn:lsid:plazi:treatment:29190528FFC6C273FF1FFA9DFD1EF9A4" httpUri="http://treatment.plazi.org/id/29190528FFC6C273FF1FFA9DFD1EF9A4" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="E9AAE7B5FFC6C270FF1FFA9DFE34FA95" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="A10FB43EFFC6C270FF1FFA9DFD83FAF4" blockId="8.[151,535,1299,1395]" box="[151,535,1299,1326]" pageId="8" pageNumber="9">
<heading id="FA470352FFC6C270FF1FFA9DFD83FAF4" bold="true" box="[151,535,1299,1326]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="66B0CFBDFFC6C270FF1FFA9DFD83FAF4" ID-CoL="66Q6N" authority="(Robb, 1972)" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[151,535,1299,1326]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC6C270FF1FFA9DFD83FAF4" bold="true" box="[151,535,1299,1326]" pageId="8" pageNumber="9">
<emphasis id="93C4682CFFC6C270FF1FFA9DFEE4FAF7" bold="true" box="[151,368,1299,1325]" italics="true" pageId="8" pageNumber="9">Anilios leptosoma</emphasis>
(
<bibRefCitation id="C521C9CFFFC6C270FEF6FA9AFD9BFAF4" author="Robb" box="[382,527,1300,1326]" pageId="8" pageNumber="9" refString="Robb, J. (1972) A new species of the genus Ramphotyphlops (Serpentes: Typhlopidae) from Western Australia. Journal of the Royal Society of Western Australia, 55, 39 - 40." type="journal article" year="1972">Robb, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A10FB43EFFC6C270FF1FFAB8FE34FA95" blockId="8.[151,535,1299,1395]" box="[151,416,1334,1359]" pageId="8" pageNumber="9">Murchison Blindsnakes</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC6C270FF1FFAD4FEC5FAA9" box="[151,337,1370,1395]" pageId="8" pageNumber="9" type="description">
<paragraph id="A10FB43EFFC6C270FF1FFAD4FEC5FAA9" blockId="8.[151,535,1299,1395]" box="[151,337,1370,1395]" pageId="8" pageNumber="9">
<figureCitation id="398BA8BBFFC6C270FF1FFAD4FEA3FAA9" box="[151,311,1370,1395]" captionStart="FIGURE 3" captionStartId="7.[151,250,1907,1929]" captionTargetBox="[183,1436,262,1802]" captionTargetId="figure@7.[151,1436,233,1830]" captionTargetPageId="7" captionText="FIGURE 3. Line drawings showing snout morphology and head scalation variation within the Anilios leptosoma complex in dorsal and lateral view. A – B) A. leptosoma holotype (WAM R 29623); C – D) A. systenos sp. nov. holotype (WAM R 114892); E – F) A. obtusifrons sp. nov. holotype (WAM R 146400). Irregularity of some head scale sutures and edges illustrated represent scale damage and scratches of holotype specimens." httpUri="https://zenodo.org/record/897291/files/figure.png" pageId="8" pageNumber="9">Figures 3A–B</figureCitation>
,
<figureCitation id="398BA8BBFFC6C270FECBFAD5FEC5FAA9" box="[323,337,1371,1395]" captionStart="FIGURE 4" captionStartId="10.[151,250,1654,1676]" captionTargetBox="[151,1436,805,1633]" captionTargetId="figure@10.[151,1436,805,1633]" captionTargetPageId="10" captionText="FIGURE 4. Anilios leptosoma (Robb, 1972) holotype (WAM R 29623). Scale bar = 10 mm." httpUri="https://zenodo.org/record/897293/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC6C270FF1FFA2CFD36F9FD" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="A10FB43EFFC6C270FF1FFA2CFBEFFA05" blockId="8.[151,1436,1442,2007]" pageId="8" pageNumber="9">
<materialsCitation id="11D8BE63FFC6C270FF1FFA2CFBEFFA05" ID-GBIF-Occurrence="1639433962" collectingDate="1967-09" collectionCode="WAM" collectorName="R. B. Humphries." county="Kalbarri" latitude="-27.55" location="The Loop" longLatPrecision="1236" longitude="114.46667" municipality="lower Murchison River" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="93C4682CFFC6C270FF1FFA2CFE9FFA61" bold="true" box="[151,267,1442,1467]" pageId="8" pageNumber="9">
<typeStatus id="7E0B0A9CFFC6C270FF1FFA2CFE92FA61" box="[151,262,1442,1467]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="C7A12CFBFFC6C270FE9BFA2DFECCFA61" box="[275,344,1443,1467]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="9">WAM</collectionCode>
R29623, an adult male collected from “
<location id="A46FE2E5FFC6C270FC95FA2CFC06FA61" LSID="urn:lsid:plazi:treatment:29190528FFC6C273FF1FFA9DFD1EF9A4:A46FE2E5FFC6C270FC95FA2CFC06FA61" box="[797,914,1442,1467]" county="Kalbarri" latitude="-27.55" longLatPrecision="1236" longitude="114.46667" municipality="lower Murchison River" name="The Loop" pageId="8" pageNumber="9" stateProvince="Western Australia">The Loop</location>
”,
<collectingMunicipality id="416B2E44FFC6C270FC20FA2CFB25FA61" box="[936,1201,1442,1467]" pageId="8" pageNumber="9">lower Murchison River</collectingMunicipality>
,
<quantity id="664819DBFFC6C270FB35FA2CFA91FA61" box="[1213,1285,1442,1467]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="9" unit="km" value="35.0">35 km</quantity>
north-east of
<collectingCounty id="486ECCB2FFC6C270FF1FFA48FF60FA05" box="[151,244,1478,1503]" pageId="8" pageNumber="9">Kalbarri</collectingCounty>
(
<geoCoordinate id="C484D2F9FFC6C270FE8CFA49FECBFA05" box="[260,351,1479,1503]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-27.55">27°33'S</geoCoordinate>
;
<geoCoordinate id="C484D2F9FFC6C270FEE3FA49FE41FA05" box="[363,469,1479,1503]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="114.46667">114°28'E</geoCoordinate>
),
<collectingRegion id="63747ADCFFC6C270FE61FA49FD21FA05" box="[489,693,1478,1503]" country="Australia" name="Western Australia" pageId="8" pageNumber="9">
<collectingRegion id="63747ADCFFC6C270FE61FA49FDD0FA04" box="[489,580,1479,1502]" country="American Samoa" name="Western" pageId="8" pageNumber="9">Western</collectingRegion>
Australia
</collectingRegion>
, in
<date id="D50E92FEFFC6C270FD54FA48FC07FA05" box="[732,915,1478,1503]" pageId="8" pageNumber="9" value="1967-09">
<collectingDate id="C54A6B16FFC6C270FD54FA48FC07FA05" box="[732,915,1478,1503]" pageId="8" pageNumber="9" value="1967-09">September 1967</collectingDate>
</date>
by
<collectorName id="0C45D1E8FFC6C270FC35FA49FBEFFA05" box="[957,1147,1478,1503]" pageId="8" pageNumber="9">R.B. Humphries.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="A10FB43EFFC6C270FF4FFA64FD36F9FD" blockId="8.[151,1436,1442,2007]" pageId="8" pageNumber="9">
<materialsCitation id="11D8BE63FFC6C270FF4FFA64FD36F9FD" ID-GBIF-Occurrence="1639433963" collectingDate="1967-09" collectionCode="WAM" county="Kalbarri" latitude="-27.55" location="The Loop'" longLatPrecision="1236" longitude="114.46667" municipality="lower Murchison River" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<emphasis id="93C4682CFFC6C270FF4FFA64FEAFF9D9" bold="true" box="[199,315,1514,1539]" pageId="8" pageNumber="9">
<typeStatus id="7E0B0A9CFFC6C270FF4FFA64FEA2F9D9" box="[199,310,1514,1539]" pageId="8" pageNumber="9" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectionCode id="C7A12CFBFFC6C270FECCFA65FE1DF9D9" box="[324,393,1515,1539]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="9">WAM</collectionCode>
R29624, from ‘
<location id="A46FE2E5FFC6C270FDB5FA64FD29F9D9" LSID="urn:lsid:plazi:treatment:29190528FFC6C273FF1FFA9DFD1EF9A4:A46FE2E5FFC6C270FDB5FA64FD29F9D9" box="[573,701,1514,1539]" county="Kalbarri" latitude="-27.55" longLatPrecision="1236" longitude="114.46667" municipality="lower Murchison River" name="The Loop'" pageId="8" pageNumber="9" stateProvince="Western Australia">The Loop’</location>
,
<collectingMunicipality id="416B2E44FFC6C270FD43FA64FC42F9D9" box="[715,982,1514,1539]" pageId="8" pageNumber="9">lower Murchison River</collectingMunicipality>
,
<quantity id="664819DBFFC6C270FC6AFA64FBBFF9D9" box="[994,1067,1514,1539]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="9" unit="km" value="35.0">35 km</quantity>
north-east of
<collectingCounty id="486ECCB2FFC6C270FB45FA64FABEF9D9" box="[1229,1322,1514,1539]" pageId="8" pageNumber="9">Kalbarri</collectingCounty>
(
<geoCoordinate id="C484D2F9FFC6C270FAB4FA65FA0CF9D9" box="[1340,1432,1515,1539]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-27.55">27°33'S</geoCoordinate>
;
<geoCoordinate id="C484D2F9FFC6C270FF1FF981FF6AF9FD" box="[151,254,1551,1575]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="114.46667">114°28'E</geoCoordinate>
)
<collectingRegion id="63747ADCFFC6C270FE85F981FE4DF9FD" box="[269,473,1550,1575]" country="Australia" name="Western Australia" pageId="8" pageNumber="9">
<collectingRegion id="63747ADCFFC6C270FE85F981FEFCF9FC" box="[269,360,1551,1574]" country="American Samoa" name="Western" pageId="8" pageNumber="9">Western</collectingRegion>
Australia
</collectingRegion>
,
<date id="D50E92FEFFC6C270FE6CF980FD0AF9FD" box="[484,670,1550,1575]" pageId="8" pageNumber="9" value="1967-09">
<collectingDate id="C54A6B16FFC6C270FE6CF980FD0AF9FD" box="[484,670,1550,1575]" pageId="8" pageNumber="9" value="1967-09">September 1967</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC6C270FF4FF9BCFCD0F901" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="A10FB43EFFC6C270FF4FF9BCFCD0F901" blockId="8.[151,1436,1442,2007]" pageId="8" pageNumber="9">
<emphasis id="93C4682CFFC6C270FF4FF9BCFED5F991" bold="true" box="[199,321,1586,1611]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
A moderately long, slender
<taxonomicName id="66B0CFBDFFC6C270FD1AF9BDFD75F991" authorityName="Gray" authorityYear="1845" box="[658,737,1587,1611]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="93C4682CFFC6C270FD1AF9BDFD75F991" box="[658,737,1587,1611]" italics="true" pageId="8" pageNumber="9">Anilios</emphasis>
</taxonomicName>
to about
<quantity id="664819DBFFC6C270FCDDF9BDFC22F990" box="[853,950,1587,1611]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="400.0">400 mm</quantity>
total length. Distinguished from all other
<taxonomicName id="66B0CFBDFFC6C270FF1FF9D8FF72F9B4" authorityName="Gray" authorityYear="1845" box="[151,230,1622,1646]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="93C4682CFFC6C270FF1FF9D8FF72F9B4" box="[151,230,1622,1646]" italics="true" pageId="8" pageNumber="9">Anilios</emphasis>
</taxonomicName>
by a combination of midbody scales in 16 rows, dorsal body scales 583–781, snout in profile prominent with obtusely angular horizontal edge, snout rounded and weakly trilobed in dorsal view, nasal cleft originating from second supralabial, extending anteriorly to nostril and terminating at rostral scale, presence of a terminal tail spine and lack of any black colouration on head, body or tail.
</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC6C273FF4FF968FE86FEC6" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="description">
<paragraph id="A10FB43EFFC6C270FF4FF968FB7CF8F9" blockId="8.[151,1436,1442,2007]" pageId="8" pageNumber="9">
<emphasis id="93C4682CFFC6C270FF4FF968FD32F925" bold="true" box="[199,678,1766,1791]" pageId="8" pageNumber="9">
Description of
<typeStatus id="7E0B0A9CFFC6C270FEFEF968FE49F925" box="[374,477,1766,1791]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(WAM R29623).
</emphasis>
A slender male
<taxonomicName id="66B0CFBDFFC6C270FCD4F968FC3FF924" authorityName="Gray" authorityYear="1845" box="[860,939,1766,1790]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="93C4682CFFC6C270FCD4F968FC3FF924" box="[860,939,1766,1790]" italics="true" pageId="8" pageNumber="9">Anilios</emphasis>
</taxonomicName>
, measurements and counts: SVL
<quantity id="664819DBFFC6C270FAABF968FA0CF924" box="[1315,1432,1766,1791]" metricMagnitude="-1" metricUnit="m" metricValue="2.496" pageId="8" pageNumber="9" unit="mm" value="249.6">249.6 mm</quantity>
; TailL
<quantity id="664819DBFFC6C270FF52F884FEA4F8F8" box="[218,304,1802,1827]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="9" unit="mm" value="8.5">8.5 mm</quantity>
(3.3% of SVL); BW
<quantity id="664819DBFFC6C270FDA9F884FDEEF8F8" box="[545,634,1802,1827]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="9" unit="mm" value="2.5">2.5 mm</quantity>
; MBSR 16; TDS 605; DBS 648; SCS 26; anal scales 4.
</paragraph>
<paragraph id="A10FB43EFFC6C271FF4FF8A0FB79FB1F" blockId="8.[151,1436,1442,2007]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Head in dorsal view rounded and elongate, gradually widening posteriorly, not distinct from body; snout tapering anteriorly from posterior to eye to a distinctly narrowed but broadly rounded rostral protrusion anterior to rostral-nasal suture, faintly trilobed anterior to rostral-nasal suture; rostral extends anteriorly beyond anterior edge of nasal scales for ~one-quarter of rostral length; head bluntly angular in profile, evenly rounded dorsally to anteriormost point of rostral protrusion, hardened rostral edge extends anteriorly and ventrally slightly forming blunt protrusion, not hooked, ventral surface of snout from anterior rostral edge to lower jaw flat; rostral scale in dorsal view oval in shape, much longer than wide (~1.8 x
<quantity id="664819DBFFC7C271FCECFF32FC51FF0E" box="[868,965,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="10" unit="mm" value="1.2">1.2 mm</quantity>
), widest at rostral-nasal suture, anterior protrusion from rostral-nasal suture shallowly concave giving faint trilobed appearance, posterior edge of rostral terminating anterior to level of eye, overlapping frontal and nasals; frontal wider than long, ~2 times the area of postfrontal, overlapping postfrontal and supraoculars; postfrontal wider than long, overlapping parietals; postfrontal and interparietal separated by parietals coming into contact between them; interparietal wider than long, similar in area to frontal, overlapping four unevenly sized body scales (including first of vertebral body scale row), one of which represents an aberrant asymmetrical scale not part of the regular body scale series presented by other specimens of the species; supraoculars larger than frontal, ~1.5–1.75 the size of frontal, overlapping postfrontal, parietal and ocular, narrowly separated by postfrontal scale, separation ~one-third of postfrontal width at widest point; parietals much wider than long, ~half the size of supraoculars, slightly wider than supraoculars, ~half the length of supraoculars, overlapping interparietal, one body scale and upper postocular; nasal scales ~2.0–2.5 times higher than wide, widest just dorsal to nostril, overlapping frontal, supraocular, preocular, first and second supralabials, narrowly separated by frontal, separation ~one-quarter frontal width at widest point, ~equal to width of supraocular separation; preoculars ~4 times higher than wide, ~one-quarter the size of nasal scale, overlapping supraocular, ocular, second and third supralabials; ocular scales higher than wide, ~3 times the size of preocular, shallowly wedged between third and fourth supralabials, widest near level of eye, overlapping parietal, two postoculars and fourth supralabial; postoculars two, slightly larger in size to body scales; eyes dark, small and inconspicuous, ~3–4 times the size of nostril, positioned under ocular and preocular scales, mostly in ocular and below sutures of preocular, supraocular and ocular scales; nostrils oblong in shape, anteriorly offset to middle of nasal scale, positioned ventrolaterally below dorsolateral narial prominences, much closer to rostral than preocular, distance from rostral-nasal suture ~one-half of nostril width, not visible from above; nasal cleft originates from anterodorsal edge of second supralabial, extending dorsally then anteriorly in a gradual curve to posteroventral edge of nostril, along ventral edge of nostril, from anteroventral edge of nostril dorsally and anteriorly to reach rostral scale, not visible from above; supralabials four, first smallest and overlapped by nasal, second ~4 times the size of the first, overlapped by nasal and preocular, third ~3 times the size of second, overlapped by preocular and overlapping ocular, fourth largest, ~3.0–3.5 times the size of third, overlapped by ocular, posterior edge terminates just posterior to posteriormost edge of ocular scale; supralabial imbrication pattern
<typeStatus id="7E0B0A9CFFC7C271FBF1FBEBFB3DFBA7" box="[1145,1193,1125,1149]" pageId="9" pageNumber="10">type</typeStatus>
T-III
<emphasis id="93C4682CFFC7C271FB7EFBEBFAA0FBA6" box="[1270,1332,1125,1148]" italics="true" pageId="9" pageNumber="10">sensu</emphasis>
<bibRefCitation id="C521C9CFFFC7C271FAC9FBEAFF71FB7A" author="Wallach" pageId="9" pageNumber="10" refString="Wallach, V. (1993) The supralabial imbrication pattern of the Typhlopoidea (Reptilia: Serpentes). Journal of Herpetology, 27, 214 - 218. https: // doi. org / 10.2307 / 1564940" type="journal article" year="1993">Wallach (1993)</bibRefCitation>
; mental scale about 1.5 times wider than long; postmental wider than long, slightly larger than mental; scale organs sparsely over rostral and nasals, becoming more sparsely distributed over scales posteriorly.
</paragraph>
<paragraph id="A10FB43EFFC7C271FF4FFB41FCF1FA8F" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
Body width constant between head and tail; body scales smooth, lacking texture, imbricate, ~2.0–2.5 times wider than long, posterior edge overlapping anterior edge of proceeding scale; tail width tapering gradually from cloaca for length of tail, tapering more abruptly posteriorly to blunt rounded tip, terminating in short (&lt;
<quantity id="664819DBFFC7C271FAD8FA96FA00FAEA" box="[1360,1428,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1.0">1 mm</quantity>
) blunt apical spine; apical spine surrounded at base by six scales.
</paragraph>
<paragraph id="A10FB43EFFC7C271FF5EFAD1FC9AFA46" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
<emphasis id="93C4682CFFC7C271FF5EFAD1FEFAFAA2" bold="true" box="[214,366,1375,1400]" pageId="9" pageNumber="10">Colouration.</emphasis>
Colour in preservative pale cream-tan, anterior half slightly darker than posterior half, colouration of tail consistent with posterior half of SVL.
</paragraph>
<paragraph id="A10FB43EFFC7C271FF4FFA29FC99F9D2" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
<emphasis id="93C4682CFFC7C271FF4FFA29FEA8FA1A" bold="true" box="[199,316,1447,1472]" pageId="9" pageNumber="10">Variation.</emphasis>
Table 1 presents ranges for meristic characters of all specimens examined. SVL up to
<quantity id="664819DBFFC7C271FABDFA26FA0CFA1A" box="[1333,1432,1448,1472]" metricMagnitude="-1" metricUnit="m" metricValue="3.91" pageId="9" pageNumber="10" unit="mm" value="391.0">391 mm</quantity>
; TailL 1.2–4.0% of SVL, mean 2.0% (N = 24); TDS 598–798, mean 670 (N = 23); BDS 583–78, mean 652 (N = 23); SCS 13–29, mean 19 (N = 29); MBSR 16 (N = 29).
</paragraph>
<paragraph id="A10FB43EFFC7C271FF4FF99AFCBEF942" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
Head in dorsal view rounded, faintly trilobed in most specimens (N = 23), occasionally more prominently trilobed (N = 5, R26012, R66343, R136311, R142023 and R146455). Head shape in lateral view bluntly angular with hardened anterior rostral edge. Hardened rostral edge protruding downwards slightly (N = 24), not as well developed or absent on small individuals (&lt;
<quantity id="664819DBFFC7C271FD08F9F1FD4AF942" box="[640,734,1663,1688]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="mm" value="150.0">150 mm</quantity>
SVL).
</paragraph>
<paragraph id="A10FB43EFFC7C272FF4FF92AFC03FEBE" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,752]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">Rostral oval in shape, much longer than wide, ~1.5–2.0 times longer than wide, posterior edge of scale terminates level with eyes (N = 19) or anterior to them (N = 9). Frontal scale usually larger than postfrontal (N = 24), rarely equal (N = 1). Frontal often 2.5 times larger than postfrontal (N = 13), sometimes 2 times (N = 5) or 3 times (N = 4), rarely 5 times (N = 1) or 1.5 times (N = 1) the size of postfrontal. Frontal often equal to interparietal in area (N =16), sometimes 1.5 times larger (N = 4) or smaller than, ~0.75 times the size of interparietal (N = 5). Scale size and position could not accurately be determined for some specimens due to damage or missing scales (N = 3). Nasal scales narrowly to widely separated by frontal, distance between nasals approximately one-third (N = 9), one-quarter (N = 6) or two-fifths (N = 6) the width of frontal scale, occasionally one-fifth (N = 1), half (N = 1) or entire width of frontal (N = 1), rarely in contact (N = 1). Supraoculars narrow to widely separated (N = 22), sometimes in contact (N = 3); separation between supraoculars approximately one-third (N = 8), half (N = 6), onefifth (N = 3), two-fifths (N = 3) or one-quarter (N = 2) of postfrontal width. Parietal scales narrowly to widely separated (N = 23), sometimes in contact (N = 2); separation between parietals approximately one-fifth (N = 8), one-sixth (N = 6), two-fifths (N = 5), one-third (N = 3), one-quarter (N =2) or half (N = 1) the width of interparietal scale. Eye positioned below ocular, preocular and supraocular (N = 15) or preocular and ocular (N = 11) scale sutures, occasionally below preocular and nasal scale suture (N = 2).</paragraph>
<paragraph id="A10FB43EFFC4C272FF4FFEE1FE45FD5F" blockId="10.[151,1437,151,752]" pageId="10" pageNumber="11">
There are no known images of the species in life; the description below is based on preserved specimens. Colouration pale cream to tan with dorsum slightly darker than ventrum (N = 12), uniform colouration (N = 10) or with anterior portion of body slightly darker than posterior portion and tail (N = 7). Variation of colouration does not correspond with geographic distribution. The four smallest specimens (R13462, R55038, R55039 and R
<date id="D50E92FEFFC4C272FF22FD8EFF6BFDC2" box="[170,255,512,536]" pageId="10" pageNumber="11" value="09-11-30">113009</date>
, 125–
<quantity id="664819DBFFC4C272FEC2FE71FE3BFDCD" box="[330,431,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.44" pageId="10" pageNumber="11" unit="mm" value="144.0">144 mm</quantity>
SVL) have uniform colouration; however, the remaining larger specimens vary in colouration in the following ways. A) uniform pale colouration (N = 12), anterior portion darker, gradually becoming lighter posteriorly, lightest on tail (N = 8), C) dorsum darker, gradually becoming lighter ventrally, lightest on ventrum (N = 6).
</paragraph>
<paragraph id="A10FB43EFFC4C272FF4FFD01FDD4FD2A" blockId="10.[151,1437,151,752]" pageId="10" pageNumber="11">Sexual dimorphism is not obvious, although male SCS count and TailL in relation to SVL (%) are higher than in females: male SCS 22–29, mean 26, TailL 2.6–4.2% of SVL, mean 3.5% (N = 7); female SCS 13–18, mean 16, TailL 1.3–2.4%, mean 1.8% (N = 21).</paragraph>
<caption id="F5CFE4B6FFC4C272FF1FF9F8FBAEF956" box="[151,1082,1654,1676]" httpUri="https://zenodo.org/record/897293/files/figure.png" pageId="10" pageNumber="11" targetBox="[151,1436,805,1633]" targetPageId="10">
<paragraph id="A10FB43EFFC4C272FF1FF9F8FBAEF956" blockId="10.[151,1082,1654,1676]" box="[151,1082,1654,1676]" pageId="10" pageNumber="11">
<emphasis id="93C4682CFFC4C272FF1FF9F8FE86F951" bold="true" box="[151,274,1654,1676]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
<taxonomicName id="66B0CFBDFFC4C272FE91F9F8FDC8F956" authority="(Robb, 1972)" authorityName="Robb" authorityYear="1972" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[281,604,1654,1676]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC4C272FE91F9F8FE5BF956" box="[281,463,1654,1676]" italics="true" pageId="10" pageNumber="11">Anilios leptosoma</emphasis>
(Robb, 1972)
</taxonomicName>
holotype (WAM R29623). Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="A10FB43EFFC4C272FF4FF93DFB25F879" blockId="10.[151,1436,1715,2028]" pageId="10" pageNumber="11">
<emphasis id="93C4682CFFC4C272FF4FF93DFDAAF916" bold="true" box="[199,574,1715,1740]" pageId="10" pageNumber="11">Comparison with other species.</emphasis>
<taxonomicName id="66B0CFBDFFC4C272FDCDF93DFC9AF911" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[581,782,1715,1739]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC4C272FDCDF93DFC9AF911" box="[581,782,1715,1739]" italics="true" pageId="10" pageNumber="11">Anilios leptosoma</emphasis>
</taxonomicName>
can be distinguished from all but three
<taxonomicName id="66B0CFBDFFC4C272FB44F93DFA8FF911" authorityName="Gray" authorityYear="1845" box="[1228,1307,1715,1739]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="93C4682CFFC4C272FB44F93DFA8FF911" box="[1228,1307,1715,1739]" italics="true" pageId="10" pageNumber="11">Anilios</emphasis>
</taxonomicName>
species (
<emphasis id="93C4682CFFC4C272FA0CF93AFE3CF935" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="66B0CFBDFFC4C272FA0CF93AFE8AF935" baseAuthorityName="Aplin" baseAuthorityYear="1998" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="longissimus">A. longissimus</taxonomicName>
,
<taxonomicName id="66B0CFBDFFC4C272FEA1F956FE3CF935" baseAuthorityName="Kinghorn" baseAuthorityYear="1929" box="[297,424,1752,1775]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="minimus">A. minimus</taxonomicName>
</emphasis>
and
<taxonomicName id="66B0CFBDFFC4C272FE57F956FDAEF935" baseAuthorityName="Shea &amp; Horner" baseAuthorityYear="1997" box="[479,570,1752,1775]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nema">
<emphasis id="93C4682CFFC4C272FE57F956FDAEF935" box="[479,570,1752,1775]" italics="true" pageId="10" pageNumber="11">A. nema</emphasis>
</taxonomicName>
) by its slender elongate body and low MBSR counts (16
<emphasis id="93C4682CFFC4C272FB49F956FB4CF935" box="[1217,1240,1752,1775]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
18, 20, 22 or 24) which does not differ within any species of the genus. Of the remaining three
<taxonomicName id="66B0CFBDFFC4C272FB9CF975FBF7F8C9" authorityName="Gray" authorityYear="1845" box="[1044,1123,1787,1811]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="93C4682CFFC4C272FB9CF975FBF7F8C9" box="[1044,1123,1787,1811]" italics="true" pageId="10" pageNumber="11">Anilios</emphasis>
</taxonomicName>
species with 16 MBSR (
<emphasis id="93C4682CFFC4C272FA0CF972FE3CF8ED" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="66B0CFBDFFC4C272FA0CF972FE8AF8ED" baseAuthorityName="Aplin" baseAuthorityYear="1998" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="longissimus">A. longissimus</taxonomicName>
,
<taxonomicName id="66B0CFBDFFC4C272FEA1F8AEFE3CF8ED" baseAuthorityName="Kinghorn" baseAuthorityYear="1929" box="[297,424,1824,1847]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="minimus">A. minimus</taxonomicName>
</emphasis>
and
<taxonomicName id="66B0CFBDFFC4C272FE57F8AEFDAFF8ED" baseAuthorityName="Shea &amp; Horner" baseAuthorityYear="1997" box="[479,571,1824,1847]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nema">
<emphasis id="93C4682CFFC4C272FE57F8AEFDAFF8ED" box="[479,571,1824,1847]" italics="true" pageId="10" pageNumber="11">A. nema</emphasis>
</taxonomicName>
),
<taxonomicName id="66B0CFBDFFC4C272FDDBF8AEFD71F8ED" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[595,741,1823,1847]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC4C272FDDBF8AEFD71F8ED" box="[595,741,1823,1847]" italics="true" pageId="10" pageNumber="11">A. leptosoma</emphasis>
</taxonomicName>
can be distinguished by the termination point of nasal cleft at rostral completely dividing the nasal scale (
<emphasis id="93C4682CFFC4C272FD0DF8CAFD08F881" box="[645,668,1860,1883]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
terminating at nostril in
<taxonomicName id="66B0CFBDFFC4C272FC3FF8CAFBC8F881" baseAuthorityName="Aplin" baseAuthorityYear="1998" box="[951,1116,1859,1883]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="93C4682CFFC4C272FC3FF8CAFBC8F881" box="[951,1116,1859,1883]" italics="true" pageId="10" pageNumber="11">A. longissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="66B0CFBDFFC4C272FB1CF8CAFA87F881" baseAuthorityName="Kinghorn" baseAuthorityYear="1929" box="[1172,1299,1860,1883]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="minimus">
<emphasis id="93C4682CFFC4C272FB1CF8CAFA87F881" box="[1172,1299,1860,1883]" italics="true" pageId="10" pageNumber="11">A. minimus</emphasis>
</taxonomicName>
, or midway between nostril and rostral in
<taxonomicName id="66B0CFBDFFC4C272FE70F8E6FDC3F8A5" baseAuthorityName="Shea &amp; Horner" baseAuthorityYear="1997" box="[504,599,1896,1919]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nema">
<emphasis id="93C4682CFFC4C272FE70F8E6FDC3F8A5" box="[504,599,1896,1919]" italics="true" pageId="10" pageNumber="11">A. nema</emphasis>
</taxonomicName>
), higher number of dorsal body scale rows (583–781
<emphasis id="93C4682CFFC4C272FB5CF8E6FB7FF8A5" box="[1236,1259,1896,1919]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
<quantity id="664819DBFFC4C272FB7EF8E9FAEDF85A" box="[1270,1401,1895,1920]" metricMagnitude="1" metricUnit="m" metricValue="1.06426" metricValueMax="1.16078" metricValueMin="0.96774" pageId="10" pageNumber="11" unit="in" value="419.0" valueMax="457.0" valueMin="381.0">381–457 in</quantity>
<taxonomicName id="66B0CFBDFFC4C272FA0CF8E6FF62F879" baseAuthorityName="Kinghorn" baseAuthorityYear="1929" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="minimus">
<emphasis id="93C4682CFFC4C272FA0CF8E6FF62F879" italics="true" pageId="10" pageNumber="11">A. minimus</emphasis>
</taxonomicName>
and
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<taxonomicName id="66B0CFBDFFC4C272FE3AF802FD99F879" baseAuthorityName="Shea &amp; Horner" baseAuthorityYear="1997" box="[434,525,1932,1955]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="nema">
<emphasis id="93C4682CFFC4C272FE3AF802FD99F879" box="[434,525,1932,1955]" italics="true" pageId="10" pageNumber="11">A. nema</emphasis>
</taxonomicName>
) and a less depressed head in comparison to
<taxonomicName id="66B0CFBDFFC4C272FB8EF802FB3DF879" baseAuthorityName="Aplin" baseAuthorityYear="1998" box="[1030,1193,1931,1955]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="longissimus">
<emphasis id="93C4682CFFC4C272FB8EF802FB3DF879" box="[1030,1193,1931,1955]" italics="true" pageId="10" pageNumber="11">A. longissimus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A10FB43EFFC4C272FF4FF821FA0FF836" blockId="10.[151,1436,1715,2028]" pageId="10" pageNumber="11">
Within its distribution,
<taxonomicName id="66B0CFBDFFC4C272FE68F83EFDEDF81D" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[480,633,1967,1991]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC4C272FE68F83EFDEDF81D" box="[480,633,1967,1991]" italics="true" pageId="10" pageNumber="11">A. leptosoma</emphasis>
</taxonomicName>
is most similar in general appearance to
<taxonomicName id="66B0CFBDFFC4C272FBF4F83EFA94F81D" authority="Vidal, 2017" authorityName="Vidal" authorityYear="2017" box="[1148,1280,1968,1991]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="systenos" status="sp. nov.">
<emphasis id="93C4682CFFC4C272FBF4F83EFA94F81D" box="[1148,1280,1968,1991]" italics="true" pageId="10" pageNumber="11">A. systenos</emphasis>
</taxonomicName>
<emphasis id="93C4682CFFC4C272FA86F821FAFBF812" bold="true" box="[1294,1391,1967,1992]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="88F7D557FFC4C272FA86F821FAFBF812" box="[1294,1391,1967,1992]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="66B0CFBDFFC4C272FA0CF83EFE80F836" authority="Vidal, 2017" authorityName="Vidal" authorityYear="2017" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="obtusifrons" status="sp. nov.">
<emphasis id="93C4682CFFC4C272FA0CF83EFE80F836" italics="true" pageId="10" pageNumber="11">A. obtusifrons</emphasis>
</taxonomicName>
<emphasis id="93C4682CFFC4C272FE94F85DFEE2F836" bold="true" box="[284,374,2003,2028]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="88F7D557FFC4C272FE94F85DFEE2F836" box="[284,374,2003,2028]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="66B0CFBDFFC4C272FE27F85AFD8FF831" baseAuthorityName="Waite" baseAuthorityYear="1918" box="[431,539,2004,2027]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="grypus">
<emphasis id="93C4682CFFC4C272FE27F85AFD8FF831" box="[431,539,2004,2027]" italics="true" pageId="10" pageNumber="11">A. grypus</emphasis>
</taxonomicName>
; however, it is easily diagnosed by the low MBSR (16
<emphasis id="93C4682CFFC4C272FB1BF85AFB3EF831" box="[1171,1194,2004,2027]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
<quantity id="664819DBFFC4C272FB3BF85AFB79F836" box="[1203,1261,2003,2028]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="10" pageNumber="11" unit="in" value="18.0">18 in</quantity>
<taxonomicName id="66B0CFBDFFC4C272FB7DF85AFAE6F831" authority="Vidal, 2017" authorityName="Vidal" authorityYear="2017" box="[1269,1394,2004,2027]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="systenos" status="sp.">
<emphasis id="93C4682CFFC4C272FB7DF85AFAE6F831" box="[1269,1394,2004,2027]" italics="true" pageId="10" pageNumber="11">A. systenos</emphasis>
</taxonomicName>
<emphasis id="93C4682CFFC4C272FAF2F85DFA0FF836" bold="true" box="[1402,1435,2003,2028]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="88F7D557FFC4C272FAF2F85DFA0FF836" box="[1402,1435,2003,2028]" pageId="10" pageNumber="11">sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="A10FB43EFFC5C273FF1FFF19FE86FEC6" blockId="11.[151,1437,151,1553]" pageId="11" pageNumber="12">
<emphasis id="93C4682CFFC5C273FF1FFF19FF5CFF6A" bold="true" box="[151,200,151,176]" pageId="11" pageNumber="12">nov.</emphasis>
,
<taxonomicName id="66B0CFBDFFC5C273FF5EFF16FEE7FF6A" authority="Vidal, 2017" authorityName="Vidal" authorityYear="2017" box="[214,371,152,176]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="obtusifrons" status="sp. nov.">
<emphasis id="93C4682CFFC5C273FF5EFF16FEE7FF6A" box="[214,371,152,176]" italics="true" pageId="11" pageNumber="12">A. obtusifrons</emphasis>
</taxonomicName>
<emphasis id="93C4682CFFC5C273FEF2FF19FE40FF6A" bold="true" box="[378,468,151,176]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="88F7D557FFC5C273FEF2FF19FE40FF6A" box="[378,468,151,176]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="66B0CFBDFFC5C273FD83FF16FDE3FF6A" baseAuthorityName="Waite" baseAuthorityYear="1918" box="[523,631,152,176]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="grypus">
<emphasis id="93C4682CFFC5C273FD83FF16FDE3FF6A" box="[523,631,152,176]" italics="true" pageId="11" pageNumber="12">A. grypus</emphasis>
</taxonomicName>
) and a combination of absence of a hooked beak (
<emphasis id="93C4682CFFC5C273FB27FF17FB52FF6A" box="[1199,1222,153,176]" italics="true" pageId="11" pageNumber="12">vs</emphasis>
distinct hook in
<taxonomicName id="66B0CFBDFFC5C273FA0CFF16FF77FF0E" baseAuthorityName="Waite" baseAuthorityYear="1918" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="grypus">
<emphasis id="93C4682CFFC5C273FA0CFF16FF77FF0E" italics="true" pageId="11" pageNumber="12">A. grypus</emphasis>
</taxonomicName>
), termination point of the nasal cleft at the rostral scale (midway between nostril and rostral in
<taxonomicName id="66B0CFBDFFC5C273FA0CFF32FE80FF22" authority="Vidal, 2017" authorityName="Vidal" authorityYear="2017" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="obtusifrons" status="sp. nov.">
<emphasis id="93C4682CFFC5C273FA0CFF32FE80FF22" italics="true" pageId="11" pageNumber="12">A. obtusifrons</emphasis>
</taxonomicName>
<emphasis id="93C4682CFFC5C273FE93FF51FEE1FF22" bold="true" box="[283,373,223,248]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="88F7D557FFC5C273FE93FF51FEE1FF22" box="[283,373,223,248]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and absence of any black pigment on the head or tail (
<emphasis id="93C4682CFFC5C273FC6FFF6FFC6AFF22" box="[999,1022,225,248]" italics="true" pageId="11" pageNumber="12">vs</emphasis>
black pigment on head and/or tail in
<taxonomicName id="66B0CFBDFFC5C273FF1FFE8AFE96FEC6" baseAuthorityName="Waite" baseAuthorityYear="1918" box="[151,258,260,284]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="grypus">
<emphasis id="93C4682CFFC5C273FF1FFE8AFE96FEC6" box="[151,258,260,284]" italics="true" pageId="11" pageNumber="12">A. grypus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC5C273FF4FFEA9FADEFC85" pageId="11" pageNumber="12" type="distribution">
<paragraph id="A10FB43EFFC5C273FF4FFEA9FE16FDE5" blockId="11.[151,1437,151,1553]" pageId="11" pageNumber="12">
<emphasis id="93C4682CFFC5C273FF4FFEA9FEF4FE9A" bold="true" box="[199,352,295,320]" pageId="11" pageNumber="12">Distribution.</emphasis>
<taxonomicName id="66B0CFBDFFC5C273FEE4FEA6FDADFE9A" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[364,569,296,320]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC5C273FEE4FEA6FDADFE9A" box="[364,569,296,320]" italics="true" pageId="11" pageNumber="12">Anilios leptosoma</emphasis>
</taxonomicName>
is known from the mid-west of
<collectingRegion id="63747ADCFFC5C273FC4BFEA6FB05FE9A" box="[963,1169,295,320]" country="Australia" name="Western Australia" pageId="11" pageNumber="12">
<collectingRegion id="63747ADCFFC5C273FC4BFEA6FB8AFEE5" box="[963,1054,296,319]" country="American Samoa" name="Western" pageId="11" pageNumber="12">Western</collectingRegion>
<collectingCountry id="D9A7F4AEFFC5C273FBA2FEA9FB05FE9A" box="[1066,1169,295,320]" name="Australia" pageId="11" pageNumber="12">Australia</collectingCountry>
</collectingRegion>
from
<quantity id="664819DBFFC5C273FB56FEA9FAA5FE9A" box="[1246,1329,295,320]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="12" unit="km" value="7.5">7.5 km</quantity>
south of Northampton (WAM R13642 &amp; R
<date id="D50E92FEFFC5C273FDADFEC2FDEFFEBE" box="[549,635,332,356]" pageId="11" pageNumber="12" value="07-11-30">113007</date>
), north to Wooramel, approximately
<quantity id="664819DBFFC5C273FBA0FEC2FBEBFEBF" box="[1064,1151,332,357]" metricMagnitude="5" metricUnit="m" metricValue="1.14" pageId="11" pageNumber="12" unit="km" value="114.0">114 km</quantity>
south-east of Carnarvon (WAM R55038 &amp; R55039), and from the west coast to Meeberrie Station, approximately
<quantity id="664819DBFFC5C273FB09FEFEFB41FE52" box="[1153,1237,367,392]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="km" value="200.0">200 km</quantity>
east north-east of Kalbarri (WAM R51097) in the east (
<figureCitation id="398BA8BBFFC5C273FDB5FE1AFDEAFE76" box="[573,638,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1961,1983]" captionTargetBox="[180,1408,210,1933]" captionTargetId="figure@2.[173,1415,195,1951]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of the Anilios leptosoma species complex in Western Australia. Localities marked with an asterik (*) denote localities for specimens used in molecular analysis by Marin et al. (2013 a)." httpUri="https://zenodo.org/record/897287/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
). Based on the extent of specimen records and using The International Union for Conservation of Nature (IUCN) Red List Guidelines (
<bibRefCitation id="C521C9CFFFC5C273FC06FE36FF47FE2D" author="IUCN" pageId="11" pageNumber="12" refString="IUCN Standards and Petitions Subcommittee (2017) Guidelines for Using the IUCN Red List Categories and Criteria. Version 12. Gland, Switzerland: International Commission for Conservation of Nature. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 20 November 2016)" type="journal article" year="2017">IUCN Standards and Petitions Subcommittee 2017</bibRefCitation>
), a rough calculation of the species’ extent of occurrence (EOO) is approximately
<quantity id="664819DBFFC5C273FB04FE51FA92FE2D" box="[1164,1286,478,503]" metricMagnitude="7" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="12" unit="km" value="32000.0">32,000 km</quantity>
2 of which its area of occupancy (AOO) is likely to be much smaller. No limiting factors that may restrict the species distribution have been identified.
</paragraph>
<paragraph id="A10FB43EFFC5C273FF4FFDC4FADEFC85" blockId="11.[151,1437,151,1553]" pageId="11" pageNumber="12">
<emphasis id="93C4682CFFC5C273FF4FFDC4FE55FDB9" bold="true" box="[199,449,586,611]" pageId="11" pageNumber="12">Habitat and ecology.</emphasis>
Collector’s notes associated with specimens indicated specimens were collected from habitats supporting loose sands or loam substrates of various colour. One specimen was collected in open mallee woodland with
<taxonomicName id="66B0CFBDFFC5C273FEC0FD1DFE60FD71" box="[328,500,659,683]" class="Magnoliopsida" family="Proteaceae" genus="Banksia" kingdom="Plantae" order="Proteales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="ashbyi">
<emphasis id="93C4682CFFC5C273FEC0FD1DFE60FD71" box="[328,500,659,683]" italics="true" pageId="11" pageNumber="12">Banksia ashbyi</emphasis>
</taxonomicName>
to
<quantity id="664819DBFFC5C273FD93FD1CFDDCFD70" box="[539,584,658,683]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="5.0">5 m</quantity>
with a canopy cover of less than 10% over
<emphasis id="93C4682CFFC5C273FBCAFD1DFA89FD71" box="[1090,1309,659,683]" italics="true" pageId="11" pageNumber="12">Spinifex longifolius</emphasis>
and mixed small to medium shrubs to
<quantity id="664819DBFFC5C273FE4FFD38FD93FD15" box="[455,519,694,719]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="m" value="1.5">1.5 m</quantity>
with cover ranging from 30–70%, from soil below a clump of spinifex (R66343). One specimen was collected from amongst the roots of spinifex in a burnt eucalypt woodland on red soil (R57545) and another was found in a mallee woodland with low
<taxonomicName id="66B0CFBDFFC5C273FC98FD71FCCAFCCC" box="[784,862,767,790]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="93C4682CFFC5C273FC98FD71FCCAFCCC" box="[784,862,767,790]" italics="true" pageId="11" pageNumber="12">Acacia</emphasis>
</taxonomicName>
on yellowish-brown sand. Specimens from Binnu were raked from red sandy loam spoil heaps in
<taxonomicName id="66B0CFBDFFC5C273FD37FCADFC99FCE0" box="[703,781,803,826]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="93C4682CFFC5C273FD37FCADFC99FCE0" box="[703,781,803,826]" italics="true" pageId="11" pageNumber="12">Acacia</emphasis>
</taxonomicName>
and
<taxonomicName id="66B0CFBDFFC5C273FCC0FCADFC2AFCE0" box="[840,958,803,826]" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="93C4682CFFC5C273FCC0FCADFC2AFCE0" box="[840,958,803,826]" italics="true" pageId="11" pageNumber="12">Casuarina</emphasis>
</taxonomicName>
shrubland (R146454–56, R146459). Two specimens (R55038 and R55039) were collected from under a cement slab at Wooramel homestead garden.
</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC5C273FF4FFCE4FD33FBED" pageId="11" pageNumber="12" type="etymology">
<paragraph id="A10FB43EFFC5C273FF4FFCE4FD33FBED" blockId="11.[151,1437,151,1553]" pageId="11" pageNumber="12">
<emphasis id="93C4682CFFC5C273FF4FFCE4FEDAFC59" bold="true" box="[199,334,874,899]" pageId="11" pageNumber="12">Etymology.</emphasis>
Derived from the Greek words
<emphasis id="93C4682CFFC5C273FD39FCE5FD60FC59" box="[689,756,875,899]" italics="true" pageId="11" pageNumber="12">leptos</emphasis>
meaning fine or thin and
<emphasis id="93C4682CFFC5C273FB9BFCE5FBDAFC58" box="[1043,1102,875,898]" italics="true" pageId="11" pageNumber="12">soma</emphasis>
meaning body in reference to the thin thread-like appearance of the species. The amendment to the specific epithet to
<emphasis id="93C4682CFFC5C273FB36FC01FAE7FC7D" box="[1214,1395,911,935]" italics="true" pageId="11" pageNumber="12">A. ‘leptosomus’</emphasis>
by
<bibRefCitation id="C521C9CFFFC5C273FF1FFC3CFE38FC11" box="[151,428,946,971]" pageId="11" pageNumber="12" refString="McDiarmid, R. W., Campbell, J. A. &amp; Toure, T. S. A. (1999) Snake Species of the World: a Taxonomic and Geographic Reference. Vol. 1. Herpetologists' League, Washington, D. C., 511 pp." type="book">
McDiarmid
<emphasis id="93C4682CFFC5C273FEA9FC3DFEC7FC11" box="[289,339,947,971]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (1999)
</bibRefCitation>
and subsequently accepted by other authors (
<bibRefCitation id="C521C9CFFFC5C273FC25FC3CFB12FC11" author="Hedges" box="[941,1158,946,971]" pageId="11" pageNumber="12" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. &amp; Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article" year="2014">
Hedges
<emphasis id="93C4682CFFC5C273FB82FC3DFBD7FC11" box="[1034,1091,947,971]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="C521C9CFFFC5C273FB19FC3DFA0CFC11" author="Pyron" box="[1169,1432,946,971]" pageId="11" pageNumber="12" refString="Pyron, R. A. &amp; Wallach, V. (2014) Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa, 3829 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 3829.1.1" type="journal article" year="2014">Pyron &amp; Wallach 2014</bibRefCitation>
; Wallach
<emphasis id="93C4682CFFC5C273FF74FC56FEA3FC35" box="[252,311,983,1007]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014) is not warranted (
<bibRefCitation id="C521C9CFFFC5C273FDDFFC58FD42FC35" author="Shea" box="[599,726,982,1007]" pageId="11" pageNumber="12" refString="Shea, G. M. (2015) A new species of Anilios (Scolecophidia: Typhlopidae) from central Australia. Zootaxa, 4033 (1), 103 - 116. https: // doi. org / 10.11646 / zootaxa. 4033.1.5" type="journal article" year="2015">Shea 2015</bibRefCitation>
). As
<bibRefCitation id="C521C9CFFFC5C273FC9CFC58FC30FC35" author="Robb" box="[788,932,982,1007]" pageId="11" pageNumber="12" refString="Robb, J. (1972) A new species of the genus Ramphotyphlops (Serpentes: Typhlopidae) from Western Australia. Journal of the Royal Society of Western Australia, 55, 39 - 40." type="journal article" year="1972">Robb (1972)</bibRefCitation>
did not state explicitly the use of the word ‘
<emphasis id="93C4682CFFC5C273FF28FC75FF4FFBC8" box="[160,219,1019,1042]" italics="true" pageId="11" pageNumber="12">soma</emphasis>
’ as a noun or adjective, it is to be treated as a noun and does not change from
<taxonomicName id="66B0CFBDFFC5C273FB13FC75FAA5FBC9" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[1179,1329,1019,1043]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC5C273FB13FC75FAA5FBC9" box="[1179,1329,1019,1043]" italics="true" pageId="11" pageNumber="12">A. leptosoma</emphasis>
</taxonomicName>
with the resurrection of
<taxonomicName id="66B0CFBDFFC5C273FECAFB91FE05FBED" authorityName="Gray" authorityYear="1845" box="[322,401,1055,1079]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="93C4682CFFC5C273FECAFB91FE05FBED" box="[322,401,1055,1079]" italics="true" pageId="11" pageNumber="12">Anilios</emphasis>
</taxonomicName>
by
<bibRefCitation id="C521C9CFFFC5C273FE33FB90FD37FBED" box="[443,675,1054,1079]" pageId="11" pageNumber="12" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. &amp; Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article">
Hedges
<emphasis id="93C4682CFFC5C273FD9EFBAEFDDAFBED" box="[534,590,1055,1079]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AAE7B5FFC5C273FF4FFBCCFD1EF9A4" pageId="11" pageNumber="12" type="discussion">
<paragraph id="A10FB43EFFC5C273FF4FFBCCFEE5FA7F" blockId="11.[151,1437,151,1553]" pageId="11" pageNumber="12">
<emphasis id="93C4682CFFC5C273FF4FFBCCFEAFFB81" bold="true" box="[199,315,1090,1115]" pageId="11" pageNumber="12">Remarks.</emphasis>
There are insufficient data to assess
<taxonomicName id="66B0CFBDFFC5C273FD68FBCDFCE0FB81" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[736,884,1091,1115]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC5C273FD68FBCDFCE0FB81" box="[736,884,1091,1115]" italics="true" pageId="11" pageNumber="12">A. leptosoma</emphasis>
</taxonomicName>
under IUCN Red List assessment criteria A and C–E; however, confirmed records of the species allow partial assessment under criterion B. The EOO for
<taxonomicName id="66B0CFBDFFC5C273FA0CFBE9FE9EFB7F" baseAuthorityName="Robb" baseAuthorityYear="1972" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC5C273FA0CFBE9FE9EFB7F" italics="true" pageId="11" pageNumber="12">A. leptosoma</emphasis>
</taxonomicName>
is approximately
<quantity id="664819DBFFC5C273FE6EFB00FDF1FB7C" box="[486,613,1165,1190]" metricMagnitude="7" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="12" unit="km" value="32000.0">32,000 km</quantity>
2 which exceeds values for any threatened categories and its area of occupancy (based on 29 confirmed records) equals
<quantity id="664819DBFFC5C273FD7AFB3AFCA8FB17" box="[754,828,1204,1229]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="km" value="60.0">60 km</quantity>
2 which partially meets criterion B2 for endangered; however, due to the lack of any population data, threats or potential decline for the species it cannot be assessed any further under criterion B. Despite the limited number of confirmed specimens, the species has been recorded from habitats that are relatively widespread across most of its EOO and the AOO is therefore considered to be underestimated. Due to the limited knowledge of the species, particularly in regards to its population/s and any major threats or evidence of decline,
<taxonomicName id="66B0CFBDFFC5C273FDCEFAE7FD4FFA5A" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[582,731,1384,1408]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC5C273FDCEFAE7FD4FFA5A" box="[582,731,1384,1408]" italics="true" pageId="11" pageNumber="12">A. leptosoma</emphasis>
</taxonomicName>
is considered to be Least Concern under the IUCN Red List assessment criteria.
</paragraph>
<paragraph id="A10FB43EFFC5C273FF4FFA3EFE4CF9CB" blockId="11.[151,1437,151,1553]" pageId="11" pageNumber="12">
The line drawings of the head scalation for
<taxonomicName id="66B0CFBDFFC5C273FD5CFA3FFCFFFA12" baseAuthorityName="Robb" baseAuthorityYear="1972" box="[724,875,1456,1480]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leptosoma">
<emphasis id="93C4682CFFC5C273FD5CFA3FFCFFFA12" box="[724,875,1456,1480]" italics="true" pageId="11" pageNumber="12">A. leptosoma</emphasis>
</taxonomicName>
in the original description by
<bibRefCitation id="C521C9CFFFC5C273FB53FA3EFAFAFA13" author="Robb" box="[1243,1390,1456,1481]" pageId="11" pageNumber="12" refString="Robb, J. (1972) A new species of the genus Ramphotyphlops (Serpentes: Typhlopidae) from Western Australia. Journal of the Royal Society of Western Australia, 55, 39 - 40." type="journal article" year="1972">Robb (1972)</bibRefCitation>
are presumably based on the
<typeStatus id="7E0B0A9CFFC5C273FE33FA5BFD8FFA37" box="[443,539,1493,1517]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
specimen (R29624) which shows closer affinities with the scale configuration illustrated than the
<typeStatus id="7E0B0A9CFFC5C273FEE6FA76FE40F9CB" box="[366,468,1528,1553]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="A10FB43EFFC5C273FF1FF9EDFD1EF9A4" blockId="11.[151,650,1635,1662]" box="[151,650,1635,1662]" pageId="11" pageNumber="12">
<taxonomicName id="66B0CFBDFFC5C273FF1FF9EDFECCF9A7" authority="Vidal, 2017" authorityName="Vidal" authorityYear="2017" box="[151,344,1635,1661]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="systenos" status="sp. nov.">
<emphasis id="93C4682CFFC5C273FF1FF9EDFECCF9A7" bold="true" box="[151,344,1635,1661]" italics="true" pageId="11" pageNumber="12">Anilios systenos</emphasis>
</taxonomicName>
<emphasis id="93C4682CFFC5C273FED7F9EBFD1EF9A4" bold="true" box="[351,650,1636,1662]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="88F7D557FFC5C273FED7F9EBFE2DF9A4" box="[351,441,1637,1662]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
Ellis &amp; Doughty
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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