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<document id="DE44BADA5BDF8B50AC7107C9964E4FB5" ID-CLB-Dataset="27714" ID-DOI="10.11646/zootaxa.4543.1.1" ID-GBIF-Dataset="22ef92b8-1d9c-4e9f-b01c-ce1f5d3faa7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617607" ID-ZooBank="31ABFA6E-6126-4603-B84F-4BEC7632D1E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927968043" checkinUser="plazi" docAuthor="Fresneda, Javier, Faille, Arnaud, Fery, Hans &amp; Ribera, Ignacio" docDate="2019" docId="03FB8C32F809FFA5FF60FD64FC764AEC" docLanguage="en" docName="zootaxa.4543.1.1.pdf" docOrigin="Zootaxa 4543 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speonemadus orchesioides" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="FFC2F44AF819FFB4FFF7FF98FFAD4B19" masterDocTitle="A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="17" updateTime="1698715325246" updateUser="ExternalLinkService">
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<mods:title id="A7CDBD3AE2F3A08A017EB0F1E074E802">A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)</mods:title>
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<mods:namePart id="ADFC6B7B2A3FF521BE253D0FE3AF1236">Fresneda, Javier</mods:namePart>
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<mods:namePart id="6045F8D5A480203025461E25F2D84A23">Fery, Hans</mods:namePart>
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<treatment id="03FB8C32F809FFA5FF60FD64FC764AEC" ID-DOI="http://doi.org/10.5281/zenodo.5927138" ID-GBIF-Taxon="154529613" ID-Zenodo-Dep="5927138" LSID="urn:lsid:plazi:treatment:03FB8C32F809FFA5FF60FD64FC764AEC" httpUri="http://treatment.plazi.org/id/03FB8C32F809FFA5FF60FD64FC764AEC" lastPageId="17" lastPageNumber="17" pageId="16" pageNumber="17">
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<paragraph id="8BED3D24F809FFA4FF60FD64FD13480F" blockId="16.[151,702,764,823]" box="[151,702,764,790]" pageId="16" pageNumber="17">
<heading id="D0A58A48F809FFA4FF60FD64FD13480F" bold="true" box="[151,702,764,790]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C5246A7F809FFA4FF60FD64FD13480F" ID-CoL="4YHZ3" authority="(Fairmaire, 1879)" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[151,702,764,790]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FF60FD64FD13480F" bold="true" box="[151,702,764,790]" pageId="16" pageNumber="17">
<emphasis id="B926E136F809FFA4FF60FD64FE76480F" bold="true" box="[151,475,764,790]" italics="true" pageId="16" pageNumber="17">Speonemadus orchesioides</emphasis>
(
<bibRefCitation id="EFC340D5F809FFA4FE1CFD64FD1B480F" author="Fairmaire, L." box="[491,694,764,790]" pageId="16" pageNumber="17" pagination="155 - 172" refId="ref12762" refString="Fairmaire, L. (1879) Descriptions de coleopteres nouveaux du nord de l'Afrique. Annales de la Societe Entomologique de France, 5 eme Serie, 9, 155 - 172." type="journal article" year="1879">Fairmaire, 1879</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3486EAFF809FFA4FF60FC86FE9D482E" box="[151,304,798,823]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BED3D24F809FFA4FF60FC86FE9D482E" blockId="16.[151,702,764,823]" box="[151,304,798,823]" pageId="16" pageNumber="17">
<figureCitation id="136921A1F809FFA4FF60FC86FF73482E" box="[151,222,798,823]" captionStart="FIGURES 5–6" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 5–6. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="16" pageNumber="17">Figs 5</figureCitation>
,
<figureCitation id="136921A1F809FFA4FF1DFC87FEA5482E" box="[234,264,799,823]" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="16" pageNumber="17">13</figureCitation>
,
<figureCitation id="136921A1F809FFA4FEE3FC87FE9D482E" box="[276,304,799,823]" captionStart="FIGURE 30" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[196,1405,228,1095]" captionTargetId="figure@14.[151,1436,193,1124]" captionTargetPageId="14" captionText="FIGURE 30. Lectotype of Choleva orchesioides (scale bar: 2 mm)." httpUri="https://zenodo.org/record/2617623/files/figure.png" pageId="16" pageNumber="17">30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF809FFA4FF60FCFEFD154882" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BED3D24F809FFA4FF60FCFEFDFF4865" blockId="16.[151,954,870,1015]" box="[151,594,870,892]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB421A0AF809FFA4FF60FCFEFDFF4865" box="[151,594,870,892]" pageId="16" pageNumber="17">
<taxonomicName id="4C5246A7F809FFA4FF60FCFEFDE34865" ID-CoL="5XYGK" authority="Fairmaire, 1879: 165" authorityName="Fairmaire" authorityPageNumber="165" authorityYear="1879" box="[151,590,870,892]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FF60FCFEFEDC4865" box="[151,369,870,892]" italics="true" pageId="16" pageNumber="17">Choleva orchesioides</emphasis>
<treatmentCitation id="0AF31B35F809FFA4FE81FCFEFDE34865" author="Fairmaire, L." box="[374,590,870,892]" page="165" pageId="16" pageNumber="17" year="1879">
<bibRefCitation id="EFC340D5F809FFA4FE81FCFEFDB14865" author="Fairmaire, L." box="[374,540,870,892]" pageId="16" pageNumber="17" pagination="155 - 172" refId="ref12762" refString="Fairmaire, L. (1879) Descriptions de coleopteres nouveaux du nord de l'Afrique. Annales de la Societe Entomologique de France, 5 eme Serie, 9, 155 - 172." type="journal article" year="1879">Fairmaire, 1879</bibRefCitation>
: 165
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BED3D24F809FFA4FF60FC1DFD154882" blockId="16.[151,954,870,1015]" box="[151,696,901,923]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB421A0AF809FFA4FF60FC1DFD154882" box="[151,696,901,923]" pageId="16" pageNumber="17">
<taxonomicName id="4C5246A7F809FFA4FF60FC1DFDAF4882" authority="(Fairmaire)" baseAuthorityName="Fairmaire" box="[151,514,901,923]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FF60FC1DFE284882" box="[151,389,901,923]" italics="true" pageId="16" pageNumber="17">Anemadus orchesioides</emphasis>
(Fairmaire)
</taxonomicName>
;
<treatmentCitation id="0AF31B35F809FFA4FDFBFC1DFD194882" author="Reitter, E." box="[524,692,901,923]" page="61" pageId="16" pageNumber="17" year="1885">
<bibRefCitation id="EFC340D5F809FFA4FDFBFC1DFD234882" author="Reitter, E." box="[524,654,901,923]" pageId="16" pageNumber="17" pagination="3 - 122" refId="ref14647" refString="Reitter, E. (1885) Bestimmungs-Tabellen der europaischen Coleopteren, 12 Necrophaga. Ferhandlungen des Naturforschenden Fereines in Brunn, 23 (1884), 3 - 122." type="journal article" year="1885">Reitter 1885</bibRefCitation>
: 61
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF809FFA4FF60FC3DFC1748EE" pageId="16" pageNumber="17" type="description">
<paragraph id="8BED3D24F809FFA4FF60FC3DFD4F48A3" blockId="16.[151,954,870,1015]" box="[151,738,932,954]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB421A0AF809FFA4FF60FC3DFD4F48A3" box="[151,738,932,954]" pageId="16" pageNumber="17">
<taxonomicName id="4C5246A7F809FFA4FF60FC3DFDB448A3" authority="(Fairmaire)" baseAuthorityName="Fairmaire" box="[151,537,932,954]" class="Insecta" family="Leiodidae" genus="Hormosacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FF60FC3DFE3148A3" box="[151,412,932,954]" italics="true" pageId="16" pageNumber="17">Hormosacus orchesioides</emphasis>
(Fairmaire)
</taxonomicName>
;
<treatmentCitation id="0AF31B35F809FFA4FDD4FC3CFD7348A3" author="Jeannel, R." box="[547,734,932,954]" page="213" pageId="16" pageNumber="17" year="1936">
<bibRefCitation id="EFC340D5F809FFA4FDD4FC3CFD0148A3" author="Jeannel, R." box="[547,684,932,954]" pageId="16" pageNumber="17" pagination="1 - 433" refId="ref13829" refString="Jeannel, R. (1936) Monographie des Catopides. Memoires du Museum National d'Histoire Naturelle, 1, 1 - 433." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
: 213
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3D24F809FFA4FF60FC5BFD4548C1" blockId="16.[151,954,870,1015]" box="[151,744,962,984]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB421A0AF809FFA4FF60FC5BFD4548C1" box="[151,744,962,984]" pageId="16" pageNumber="17">
<taxonomicName id="4C5246A7F809FFA4FF60FC5BFE4948C1" authority="Jeannel" authorityName="Jeannel" box="[151,484,962,984]" class="Insecta" family="Leiodidae" genus="Hormosacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="vandalitiae">
<emphasis id="B926E136F809FFA4FF60FC5BFE2248C1" box="[151,399,962,984]" italics="true" pageId="16" pageNumber="17">Hormosacus vandalitiae</emphasis>
Jeannel
</taxonomicName>
;
<treatmentCitation id="0AF31B35F809FFA4FE19FC5AFD2548C1" author="Blas, M." box="[494,648,962,984]" page="252" pageId="16" pageNumber="17" year="1984">
<bibRefCitation id="EFC340D5F809FFA4FE19FC5AFDF548C1" author="Blas, M." box="[494,600,962,984]" pageId="16" pageNumber="17" pagination="251 - 255" refId="ref12162" refString="Blas, M. (1984) El genero Hormosacus Jeannel, 1936 y su distribucion geografica. (Col. Catopidae). Memoires de Biospeologie, 11, 251 - 255." type="journal article" year="1984">Blas 1984</bibRefCitation>
: 252
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3D24F809FFA4FF60FC79FC1748EE" blockId="16.[151,954,870,1015]" box="[151,954,993,1015]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB421A0AF809FFA4FF60FC79FC1748EE" box="[151,954,993,1015]" pageId="16" pageNumber="17">
<taxonomicName id="4C5246A7F809FFA4FF60FC79FD8948EE" ID-CoL="4YHZ3" authority="(Fairmaire)" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[151,548,993,1015]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FF60FC79FE0A48EE" box="[151,423,993,1015]" italics="true" pageId="16" pageNumber="17">Speonemadus orchesioides</emphasis>
(Fairmaire)
</taxonomicName>
;
<treatmentCitation id="0AF31B35F809FFA4FDD9FC79FCF748EE" author="Giachino, P. M. &amp; Vailati, D." box="[558,858,993,1015]" page="174" pageId="16" pageNumber="17" year="1993">
<bibRefCitation id="EFC340D5F809FFA4FDD9FC79FC8748EE" author="Giachino, P. M. &amp; Vailati, D." box="[558,810,993,1015]" pageId="16" pageNumber="17" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
: 174
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<materialsCitation id="3B3A3779F809FFA4FF60FBBCFCD34F25" ID-GBIF-Occurrence="2147524157" box="[151,894,1060,1085]" country="Algeria" location="Algeria" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<emphasis id="B926E136F809FFA4FF60FBBCFE944F24" bold="true" box="[151,313,1060,1085]" pageId="16" pageNumber="17">
<typeStatus id="54E98386F809FFA4FF60FBBCFF7F4F24" box="[151,210,1060,1085]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3457DB4F809FFA4FEB7FBBCFE3A4F24" box="[320,407,1060,1085]" name="Algeria" pageId="16" pageNumber="17">Algeria</collectingCountry>
, &quot;Daya, province d'Oran&quot; (
<bibRefCitation id="EFC340D5F809FFA4FD33FBBCFCDB4F25" author="Fairmaire, L." box="[708,886,1060,1085]" pageId="16" pageNumber="17" pagination="155 - 172" refId="ref12762" refString="Fairmaire, L. (1879) Descriptions de coleopteres nouveaux du nord de l'Afrique. Annales de la Societe Entomologique de France, 5 eme Serie, 9, 155 - 172." type="journal article" year="1879">Fairmaire 1879</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BED3D24F809FFA4FF30FBDFFDDD4EF5" blockId="16.[151,1436,1060,2021]" pageId="16" pageNumber="17">
<emphasis id="B926E136F809FFA4FF30FBDFFEE14F79" bold="true" box="[199,332,1095,1120]" pageId="16" pageNumber="17">Type series</emphasis>
.
<materialsCitation id="3B3A3779F809FFA4FEADFBDFFDDD4EF5" ID-GBIF-Occurrence="2147524634" collectionCode="DEI, SDEI, MNHN, MUSEUM, PARIS, COLL, TYPE" collectorName="L. v. Heyden &amp; D. Vailati" country="Algeria" location="Daya" municipality="Bedel" pageId="16" pageNumber="17" specimenCount="6" specimenCount-female="6" typeStatus="lectotype">
<emphasis id="B926E136F809FFA4FEADFBDFFE7D4F79" bold="true" box="[346,464,1095,1120]" pageId="16" pageNumber="17">
<typeStatus id="54E98386F809FFA4FEADFBDFFE7D4F79" box="[346,464,1095,1120]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
</emphasis>
(
<figureCitation id="136921A1F809FFA4FE28FBDFFD994F79" box="[479,564,1095,1120]" captionStart="FIGURE 30" captionStartId="14.[151,250,1146,1168]" captionTargetBox="[196,1405,228,1095]" captionTargetId="figure@14.[151,1436,193,1124]" captionTargetPageId="14" captionText="FIGURE 30. Lectotype of Choleva orchesioides (scale bar: 2 mm)." httpUri="https://zenodo.org/record/2617623/files/figure.png" pageId="16" pageNumber="17">Fig. 30</figureCitation>
):
<specimenCount id="9D54F6ADF809FFA4FDBFFBD0FDF04F46" box="[584,605,1096,1119]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
, &quot;orchesioides / Fairm. /
<location id="8E8D6BFFF809FFA4FC8EFBD0FC1A4F79" LSID="urn:lsid:plazi:treatment:03FB8C32F809FFA5FF60FD64FC764AEC:8E8D6BFFF809FFA4FC8EFBD0FC1A4F79" box="[889,951,1096,1120]" country="Algeria" municipality="Bedel" name="Daya" pageId="16" pageNumber="17">Daya</location>
,
<collectingCountry id="F3457DB4F809FFA4FC35FBD0FC574F79" box="[962,1018,1096,1120]" name="Algeria" pageId="16" pageNumber="17">Oran</collectingCountry>
/
<collectingMunicipality id="6B89A75EF809FFA4FBE6FBDFFBFF4F79" box="[1041,1106,1095,1120]" pageId="16" pageNumber="17">Bedel</collectingMunicipality>
/ typ.&quot; [hw], &quot;
<typeStatus id="54E98386F809FFA4FB05FBD0FAF24F79" box="[1266,1375,1096,1120]" pageId="16" pageNumber="17" type="syntype">Syntypus</typeStatus>
&quot; [red label, printed], &quot;coll.
<collectorName id="26A758F2F809FFA4FE73FBF5FDB94F9C" box="[388,532,1132,1157]" pageId="16" pageNumber="17">L. v. Heyden</collectorName>
/
<collectionCode id="ED43A5E1F809FFA4FDDEFBF5FDFA4F9D" box="[553,599,1133,1156]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="16" pageNumber="17" type="Other type of research institution/biorepository">DEI</collectionCode>
Eberswalde&quot; (printed), &quot;
<typeStatus id="54E98386F809FFA4FC9CFBF4FC474F9D" box="[875,1002,1132,1156]" pageId="16" pageNumber="17" type="lectotype">Lectotypus</typeStatus>
<specimenCount id="9D54F6ADF809FFA4FC06FBF5FBAB4F9D" box="[1009,1030,1133,1156]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
/
<taxonomicName id="4C5246A7F809FFA4FBEBFBF4FBD44F9C" box="[1052,1145,1132,1157]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="31" phylum="Arthropoda" rank="genus">Choleva</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FB79FBF4FA204F9C" authority="Fairmaire" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[1166,1421,1132,1157]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">orchesioides Fairmaire</taxonomicName>
/, P. M. GIACHINO &amp; /
<collectorName id="26A758F2F809FFA4FE45FB08FD8D4FB1" box="[434,544,1167,1192]" pageId="16" pageNumber="17">D. Vailati</collectorName>
des. 1992&quot; [red label, printed] (
<collectionCode id="ED43A5E1F809FFA4FC63FB08FC754FB1" box="[916,984,1168,1192]" pageId="16" pageNumber="17">SDEI</collectionCode>
). We have found four specimens in the general collection of the
<collectionCode id="ED43A5E1F809FFA4FE3EFB2DFDB34FD5" box="[457,542,1205,1228]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
that should be considered
<typeStatus id="54E98386F809FFA4FC99FB2CFBBE4FD4" box="[878,1043,1204,1229]" pageId="16" pageNumber="17" type="paralectotype">paralectotypes</typeStatus>
, with data: (1)
<specimenCount id="9D54F6ADF809FFA4FB2FFB2CFAA54FD5" box="[1240,1288,1204,1228]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, &quot;
<taxonomicName id="4C5246A7F809FFA4FADEFB2CFA2B4FD4" box="[1321,1414,1204,1229]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F809FFA4FADEFB2CFA2B4FD4" box="[1321,1414,1204,1229]" italics="true" pageId="16" pageNumber="17">Choleva</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FF60FB4FFE894FE9" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[151,292,1239,1264]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FF60FB4FFE894FE9" box="[151,292,1239,1264]" italics="true" pageId="16" pageNumber="17">orchesioides</emphasis>
</taxonomicName>
/ Frm. Daya&quot; [hw Fairmaire], &quot;
<collectionCode id="ED43A5E1F809FFA4FD56FB40FCD54FE9" box="[673,888,1240,1264]" pageId="16" pageNumber="17">MUSEUM PARIS</collectionCode>
/
<collectionCode id="ED43A5E1F809FFA4FC60FB40FC494FE9" box="[919,996,1240,1264]" pageId="16" pageNumber="17">COLL</collectionCode>
. L. FAIRMAIRE 1906&quot; [green label, printed], &quot;
<collectionCode id="ED43A5E1F809FFA4FEF0FB64FEFD4E0D" box="[263,336,1276,1300]" pageId="16" pageNumber="17">TYPE</collectionCode>
&quot; [red label, printed], &quot;
<typeStatus id="54E98386F809FFA4FDB9FB64FCE14E0D" box="[590,844,1276,1300]" pageId="16" pageNumber="17" type="paralectotype">PARALECTOTYPUS</typeStatus>
[printed] /
<taxonomicName id="4C5246A7F809FFA4FC39FB64FB864E0C" box="[974,1067,1276,1301]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F809FFA4FC39FB64FB864E0C" box="[974,1067,1276,1301]" italics="true" pageId="16" pageNumber="17">Choleva</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FBB4FB64FB624E0C" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[1091,1231,1276,1301]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FBB4FB64FB624E0C" box="[1091,1231,1276,1301]" italics="true" pageId="16" pageNumber="17">orchesioides</emphasis>
</taxonomicName>
[hw]&quot; [red label], (coll. générale in
<collectionCode id="ED43A5E1F809FFA4FE91FAB8FE104E21" box="[358,445,1312,1336]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
) (
<figureCitation id="136921A1F809FFA4FE2FFAB8FD9D4E21" box="[472,560,1312,1336]" captionStart="FIGURES 31–33" captionStartId="14.[151,264,1996,2018]" captionTargetBox="[178,1414,1241,1950]" captionTargetId="figure@14.[151,1436,1217,1974]" captionTargetPageId="14" captionText="FIGURES 31–33. Labels of the paralectotypes of Choleva orchesioides." httpUri="https://zenodo.org/record/2617625/files/figure.png" pageId="16" pageNumber="17">Fig. 31</figureCitation>
); (2)
<specimenCount id="9D54F6ADF809FFA4FD83FAB8FD0F4E2E" box="[628,674,1312,1336]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
, &quot;
<taxonomicName id="4C5246A7F809FFA4FD48FA87FCB14E21" box="[703,796,1311,1336]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F809FFA4FD48FA87FCB14E21" box="[703,796,1311,1336]" italics="true" pageId="16" pageNumber="17">Choleva</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FCCDFA87FC6A4E21" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[826,967,1311,1336]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="34" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FCCDFA87FC6A4E21" box="[826,967,1311,1336]" italics="true" pageId="16" pageNumber="17">orchesioÏdes</emphasis>
</taxonomicName>
/ Fairm. type / Daya&quot; [hw, Fairmaire], &quot;
<collectionCode id="ED43A5E1F809FFA4FF68FADCFEDB4E45" box="[159,374,1348,1372]" pageId="16" pageNumber="17">MUSEUM PARIS</collectionCode>
/
<collectionCode id="ED43A5E1F809FFA4FE62FADCFE4F4E45" box="[405,482,1348,1372]" pageId="16" pageNumber="17">COLL</collectionCode>
. ABEILLE DE PERRIN / 1919&quot; [green label, printed], &quot;
<collectionCode id="ED43A5E1F809FFA4FB74FADCFB634E45" box="[1155,1230,1348,1372]" pageId="16" pageNumber="17">TYPE</collectionCode>
&quot; [printed in red], &quot;Daya&quot; [hw Jeannel], &quot;
<typeStatus id="54E98386F809FFA4FE6BFAF0FD374E99" box="[412,666,1384,1408]" pageId="16" pageNumber="17" type="paralectotype">PARALECTOTYPUS</typeStatus>
[printed] /
<taxonomicName id="4C5246A7F809FFA4FCE9FAFFFCD64E99" box="[798,891,1383,1408]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F809FFA4FCE9FAFFFCD64E99" box="[798,891,1383,1408]" italics="true" pageId="16" pageNumber="17">Choleva</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FC62FAFFFB8F4E99" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[917,1058,1383,1408]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FC62FAFFFB8F4E99" box="[917,1058,1383,1408]" italics="true" pageId="16" pageNumber="17">orchesioides</emphasis>
</taxonomicName>
[hw]&quot; [red label], (coll. générale
<collectionCode id="ED43A5E1F809FFA4FF60FA14FF424EBD" box="[151,239,1420,1444]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
) (
<figureCitation id="136921A1F809FFA4FEFFFA14FEF24EBD" box="[264,351,1420,1444]" captionStart="FIGURES 31–33" captionStartId="14.[151,264,1996,2018]" captionTargetBox="[178,1414,1241,1950]" captionTargetId="figure@14.[151,1436,1217,1974]" captionTargetPageId="14" captionText="FIGURES 31–33. Labels of the paralectotypes of Choleva orchesioides." httpUri="https://zenodo.org/record/2617625/files/figure.png" pageId="16" pageNumber="17">Fig. 32</figureCitation>
); (3&amp;4)
<specimenCount id="9D54F6ADF809FFA4FE33FA14FDAB4EBA" box="[452,518,1420,1444]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
on the same pin, &quot;
<collectionCode id="ED43A5E1F809FFA4FD15FA14FC1A4EBD" box="[738,951,1420,1444]" pageId="16" pageNumber="17">MUSEUM PARIS</collectionCode>
/
<collectionCode id="ED43A5E1F809FFA4FC25FA14FBB24EBD" box="[978,1055,1420,1444]" pageId="16" pageNumber="17">COLL</collectionCode>
. ABEILLE DE PERRIN / 1919&quot; [green label, printed], &quot;Daya&quot; [hw Jeannel], &quot;
<typeStatus id="54E98386F809FFA4FD6EFA28FC3A4ED1" box="[665,919,1456,1480]" pageId="16" pageNumber="17" type="paralectotype">PARALECTOTYPUS</typeStatus>
[printed] /
<taxonomicName id="4C5246A7F809FFA4FBEDFA37FBDA4ED1" box="[1050,1143,1455,1480]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F809FFA4FBEDFA37FBDA4ED1" box="[1050,1143,1455,1480]" italics="true" pageId="16" pageNumber="17">Choleva</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FB66FA37FAB34ED1" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[1169,1310,1455,1480]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FB66FA37FAB34ED1" box="[1169,1310,1455,1480]" italics="true" pageId="16" pageNumber="17">orchesioides</emphasis>
</taxonomicName>
[hw]&quot; [red label] (coll. générale in
<collectionCode id="ED43A5E1F809FFA4FE55FA4CFE544EF5" box="[418,505,1492,1516]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
) (
<figureCitation id="136921A1F809FFA4FDE7FA4CFDCE4EF5" box="[528,611,1492,1517]" captionStart="FIGURES 31–33" captionStartId="14.[151,264,1996,2018]" captionTargetBox="[178,1414,1241,1950]" captionTargetId="figure@14.[151,1436,1217,1974]" captionTargetPageId="14" captionText="FIGURES 31–33. Labels of the paralectotypes of Choleva orchesioides." httpUri="https://zenodo.org/record/2617625/files/figure.png" pageId="16" pageNumber="17">Fig. 33</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF809FFA4FF30FA6FFD904D41" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BED3D24F809FFA4FF30FA6FFD904D41" blockId="16.[151,1436,1060,2021]" pageId="16" pageNumber="17">
<emphasis id="B926E136F809FFA4FF30FA6FFEEC4D09" bold="true" box="[199,321,1527,1552]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
The species is characterised by: parameres simple, apex acute, only slightly turned outwards (
<figureCitation id="136921A1F809FFA4FA99FA6FFF184D2D" captionStart="FIGURES 13–28" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
); apex of median lobe roughly pentagonal (
<figureCitation id="136921A1F809FFA4FD52F984FD474D2C" box="[677,746,1564,1589]" captionStart="FIGURES 5–6" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 5–6. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
). The most similar species seems to be
<taxonomicName id="4C5246A7F809FFA4FB5BF985FAB24D2C" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[1196,1311,1564,1589]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F809FFA4FB5BF985FAB24D2C" box="[1196,1311,1564,1589]" italics="true" pageId="16" pageNumber="17">S. brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F809FFA4FAD0F984FACA4D2C" bold="true" box="[1319,1383,1564,1589]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A2155C4DF809FFA4FAD0F984FACA4D2C" box="[1319,1383,1564,1589]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(see the description of this species above).
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF809FFA4FF30F9FCFAE84C84" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BED3D24F809FFA4FF30F9FCFAE84C84" blockId="16.[151,1436,1060,2021]" pageId="16" pageNumber="17">
<emphasis id="B926E136F809FFA4FF30F9FCFE994D64" bold="true" box="[199,308,1636,1661]" pageId="16" pageNumber="17">Remarks</emphasis>
. The type series of
<taxonomicName id="4C5246A7F809FFA4FDEBF9FDFD654D64" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[540,712,1636,1661]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FDEBF9FDFD654D64" box="[540,712,1636,1661]" italics="true" pageId="16" pageNumber="17">S. orchesioides</emphasis>
</taxonomicName>
is divided between the SDEI (
<typeStatus id="54E98386F809FFA4FBDBF9FCFB3B4D64" box="[1068,1174,1636,1661]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
) and the MNHN (four
<typeStatus id="54E98386F809FFA4FF60F91FFE934DB9" box="[151,318,1671,1696]" pageId="16" pageNumber="17" type="paralectotype">paralectotypes</typeStatus>
). All five specimens are females, so males of the species had to be identified from non-typical material. We consider that the male specimen figured by
<bibRefCitation id="EFC340D5F809FFA4FCEDF934FC6D4DDC" author="Jeannel, R." box="[794,960,1708,1733]" pageId="16" pageNumber="17" pagination="1 - 433" refId="ref13829" refString="Jeannel, R. (1936) Monographie des Catopides. Memoires du Museum National d'Histoire Naturelle, 1, 1 - 433." type="journal article" year="1936">Jeannel (1936)</bibRefCitation>
from Ammi-Moussa (a locality ca.
<quantity id="4CAA90C1F809FFA4FAA2F934FA314DDC" box="[1365,1436,1708,1733]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="17" unit="km" value="80.0">80 km</quantity>
east of Daya, the type locality) belongs to this species. This specimen and a microscopic slide with its dissected aedeagus are deposited in the general collection of the MNHN; the slide is labelled with Jeannel's manuscript handwritting &quot;
<taxonomicName id="4C5246A7F809FFA4FECFF881FE684C29" authorityName="Jeannel" authorityYear="1936" box="[312,453,1817,1840]" class="Insecta" family="Leiodidae" genus="Hormosacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F809FFA4FECFF881FE684C29" box="[312,453,1817,1840]" italics="true" pageId="16" pageNumber="17">Hormosacus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FE2CF881FD284C29" authority="Peyer." authorityName="Peyerimhoff" authorityYear="1917" box="[475,645,1816,1840]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B926E136F809FFA4FE2CF881FD954C29" box="[475,568,1817,1840]" italics="true" pageId="16" pageNumber="17">tenuipes</emphasis>
Peyer.
</taxonomicName>
/
<taxonomicName id="4C5246A7F809FFA4FD6DF88FFCCE4C36" authority="Frm." authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[666,867,1815,1840]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FD6DF88FFC8A4C29" box="[666,807,1815,1840]" italics="true" pageId="16" pageNumber="17">orchesioides</emphasis>
Frm.
</taxonomicName>
/ Ammi-moussa&quot;, &quot;org. cop. [= organe copulateur]
<specimenCount id="9D54F6ADF809FFA4FF60F8A4FF024C4D" box="[151,175,1852,1876]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
&quot;. The same preparation has the aedeagus of a specimen of
<taxonomicName id="4C5246A7F809FFA4FCB1F8A5FC124C4D" authorityName="Peyerimhoff" authorityYear="1917" box="[838,959,1853,1876]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B926E136F809FFA4FCB1F8A5FC124C4D" box="[838,959,1853,1876]" italics="true" pageId="16" pageNumber="17">S. tenuipes</emphasis>
</taxonomicName>
from the same locality.
<emphasis id="B926E136F809FFA4FB38F8A4FAAA4C4C" bold="true" box="[1231,1287,1852,1877]" pageId="16" pageNumber="17">Note</emphasis>
: A specimen figured by
<bibRefCitation id="EFC340D5F809FFA4FEEDF8C7FDE14C61" author="Giachino, P. M. &amp; Vailati, D." box="[282,588,1887,1912]" pageId="16" pageNumber="17" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati (1993)</bibRefCitation>
as
<taxonomicName id="4C5246A7F809FFA4FD8DF8F8FC854C61" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[634,808,1887,1912]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F809FFA4FD8DF8F8FC854C61" box="[634,808,1887,1912]" italics="true" pageId="16" pageNumber="17">S. orchesioides</emphasis>
</taxonomicName>
is from Djebel Babor, some
<quantity id="4CAA90C1F809FFA4FB73F8C7FB734C61" box="[1156,1246,1887,1912]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="17" unit="km" value="600.0">600 km</quantity>
east of Daya, a considerable distance from the type locality, and representing a different species (
<taxonomicName id="4C5246A7F809FFA4FBDFF81DFB1A4C84" baseAuthorityName="Reitter" baseAuthorityYear="1885" box="[1064,1207,1924,1949]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B926E136F809FFA4FBDFF81DFB1A4C84" box="[1064,1207,1924,1949]" italics="true" pageId="16" pageNumber="17">S. pulchellus</emphasis>
</taxonomicName>
, see below).
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF809FFA5FF30F830FC764AEC" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BED3D24F809FFA5FF30F830FC764AEC" blockId="16.[151,1436,1060,2021]" lastBlockId="17.[151,1437,151,501]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B926E136F809FFA4FF30F830FEF44CD8" bold="true" box="[199,345,1960,1985]" pageId="16" pageNumber="17">Distribution</emphasis>
(
<figureCitation id="136921A1F809FFA4FE9EF830FE104CD9" box="[361,445,1960,1985]" captionStart="FIGURE 38" captionStartId="21.[151,250,1234,1256]" captionTargetBox="[259,1364,471,1140]" captionTargetId="figure@21.[205,1395,421,1189]" captionTargetPageId="21" captionText="FIGURE 38. Distribution maps of: (38) Speonemadus brusteli sp. n., S. comasi sp. n., S. orchesioides, S. pulchellus, S. tenuipes (empty circles are dubious records of which no material could be studied; the records from continental Italy, given originally as S. orchesoides, are considered S. pulchellus, P.M. Giachino, personal communication 2018; see main text). See Appendix for detailed locality data." httpUri="https://zenodo.org/record/2617639/files/figure.png" pageId="16" pageNumber="17">Fig. 38</figureCitation>
; Appendix).
<collectingCountry id="F3457DB4F809FFA4FDB8F830FD0E4CD8" box="[591,675,1960,1985]" name="Algeria" pageId="16" pageNumber="17">Algeria</collectingCountry>
(Tell Atlas) (
<bibRefCitation id="EFC340D5F809FFA4FCCDF830FC464CD9" author="Fairmaire, L." box="[826,1003,1960,1985]" pageId="16" pageNumber="17" pagination="155 - 172" refId="ref12762" refString="Fairmaire, L. (1879) Descriptions de coleopteres nouveaux du nord de l'Afrique. Annales de la Societe Entomologique de France, 5 eme Serie, 9, 155 - 172." type="journal article" year="1879">Fairmaire 1879</bibRefCitation>
;
<bibRefCitation id="EFC340D5F809FFA4FC00F830FB2B4CD8" author="Reitter, E." box="[1015,1158,1960,1985]" pageId="16" pageNumber="17" pagination="3 - 122" refId="ref14647" refString="Reitter, E. (1885) Bestimmungs-Tabellen der europaischen Coleopteren, 12 Necrophaga. Ferhandlungen des Naturforschenden Fereines in Brunn, 23 (1884), 3 - 122." type="journal article" year="1885">Reitter 1885</bibRefCitation>
;
<bibRefCitation id="EFC340D5F809FFA4FB64F830FA844CD8" author="Jeannel, R." box="[1171,1321,1960,1985]" pageId="16" pageNumber="17" pagination="1 - 433" refId="ref13829" refString="Jeannel, R. (1936) Monographie des Catopides. Memoires du Museum National d'Histoire Naturelle, 1, 1 - 433." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
;
<bibRefCitation id="EFC340D5F809FFA4FAC2F830FEEC4CFD" author="Giachino, P. M. &amp; Vailati, D." pageId="16" pageNumber="17" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
;
<bibRefCitation id="EFC340D5F809FFA4FEB9F855FE454CFD" author="Perreau, M." box="[334,488,1996,2020]" pageId="16" pageNumber="17" pagination="1 - 460" refId="ref14397" refString="Perreau, M. (2000) Catalogue des coleopteres Leiodidae Cholevinae et Platypsyllinae. Memoires de la Societe Entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau 2000</bibRefCitation>
,
<bibRefCitation id="EFC340D5F809FFA4FE03F854FD824CFD" author="Perreau, M." box="[500,559,1996,2020]" pageId="16" pageNumber="17" pagination="133 - 203" refId="ref14426" refString="Perreau, M. (2004) Leiodidae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Apollo Books, Stenstrup, pp. 133 - 203." type="book chapter" year="2004">2004</bibRefCitation>
;
<bibRefCitation id="EFC340D5F809FFA4FDCBF854FCB04CFD" author="Salgado, J. M. &amp; Blas, M. &amp; Fresneda, J." box="[572,797,1996,2021]" pageId="16" pageNumber="17" refId="ref15024" refString="Salgado, J. M., Blas, M. &amp; Fresneda, J. (2008) Coleoptera, Cholevidae. Fauna Iberica. Fol. 31. Museo Nacional de Ciencias Naturales, CSIC, Madrid, 799 pp." type="book" year="2008">
Salgado
<emphasis id="B926E136F809FFA4FD69F855FD7C4CFC" box="[670,721,1996,2021]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. 2008
</bibRefCitation>
). The records from the Iberian Peninsula,
<bibRefCitation id="EFC340D5F809FFA4FB0AF855FA314CFC" author="Fuente, J. M. de la" box="[1277,1436,1996,2021]" pageId="16" pageNumber="17" pagination="93 - 117" refId="ref13138" refString="Fuente, J. M. de la (1925) Catalogo sistematico-geografico de los coleopteros observados en la peninsula Iberica, Pirineos propiamente dichos y Baleares. Boletin de la Real Sociedad Entomologica de Espana, 7 (2 - 3), 93 - 117. [1924]" type="journal article" year="1925">Fuente (1925)</bibRefCitation>
and
<bibRefCitation id="EFC340D5F808FFA5FF3FFF0FFECA4BA9" author="Seabra, A. F. de" box="[200,359,151,176]" pageId="17" pageNumber="18" pagination="1 - 151" refId="ref15176" refString="Seabra, A. F. de (1943) Contribucoes para o inventario da fauna lusitanica, Insecta, Coleoptera. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra, 142, 1 - 151." type="journal article" year="1943">Seabra (1943)</bibRefCitation>
for
<collectingCountry id="F3457DB4F808FFA5FE6FFF0FFE574BA9" box="[408,506,151,176]" name="Portugal" pageId="17" pageNumber="18">Portugal</collectingCountry>
, and
<bibRefCitation id="EFC340D5F808FFA5FDC0FF0FFCCC4BA9" author="Giachino, P. M. &amp; Vailati, D." box="[567,865,151,176]" pageId="17" pageNumber="18" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati (1993)</bibRefCitation>
for
<collectingCountry id="F3457DB4F808FFA5FC65FF0FFC784BA9" box="[914,981,151,176]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
, are unreliable; it seems likely that they should be referred to
<taxonomicName id="4C5246A7F808FFA5FE7CFF25FE504BCC" authorityName="Kraatz" authorityYear="1870" box="[395,509,188,213]" class="Insecta" family="Leiodidae" genus="Choleva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B926E136F808FFA5FE7CFF25FE504BCC" box="[395,509,188,213]" italics="true" pageId="17" pageNumber="18">S. gracilis</emphasis>
</taxonomicName>
(see Discussion below). According to
<bibRefCitation id="EFC340D5F808FFA5FC41FF24FB4C4BCD" author="Giachino, P. M. &amp; Vailati, D." box="[950,1249,188,213]" pageId="17" pageNumber="18" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati (1993)</bibRefCitation>
, the record of
<taxonomicName id="4C5246A7F808FFA5FA70FF25FE894BE1" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F808FFA5FA70FF25FE894BE1" italics="true" pageId="17" pageNumber="18">S. orchesioides</emphasis>
</taxonomicName>
from continental
<collectingCountry id="F3457DB4F808FFA5FE00FF47FD864BE1" box="[503,555,223,248]" name="Italy" pageId="17" pageNumber="18">Italy</collectingCountry>
by
<bibRefCitation id="EFC340D5F808FFA5FDAAFF47FCAB4BE1" author="Jeannel, R." box="[605,774,223,248]" pageId="17" pageNumber="18" pagination="1 - 433" refId="ref13829" refString="Jeannel, R. (1936) Monographie des Catopides. Memoires du Museum National d'Histoire Naturelle, 1, 1 - 433." type="journal article" year="1936">Jeannel (1936)</bibRefCitation>
(&quot;
<collectingRegion id="4996F3C6F808FFA5FCD5FF47FC394BE1" box="[802,916,223,248]" country="Italy" name="Abruzzo" pageId="17" pageNumber="18">Abruzzes</collectingRegion>
: Gran Sasso, une femelle (Schatzmayr!)&quot;) is doubtful. However, this record was confirmed by newly obtained material from different provinces in continental
<collectingCountry id="F3457DB4F808FFA5FF60FEBFFF664A59" box="[151,203,295,320]" name="Italy" pageId="17" pageNumber="18">Italy</collectingCountry>
by
<bibRefCitation id="EFC340D5F808FFA5FF0CFEBFFE554A59" author="Fabbri, R. A. &amp; Zoia, S." box="[251,504,295,320]" pageId="17" pageNumber="18" pagination="78" refId="ref12512" refString="Fabbri, R. A. &amp; Zoia, S. (1996) Segnalazioni faunistiche n ° 10 - Speonemadus orchesioides (Fairmaire, 1879) (Insecta, Coleoptera, Cholevidae). Quaderni di Studi e Notizie di Storia Naturale della Romagna, 6, 78." type="journal article" year="1996">Fabbri &amp; Zoia (1996)</bibRefCitation>
and
<bibRefCitation id="EFC340D5F808FFA5FDC1FEBFFCF24A59" author="Giachino, P. M. &amp; Vailati, D." box="[566,863,295,320]" pageId="17" pageNumber="18" pagination="119 - 124" refId="ref13350" refString="Giachino, P. M. &amp; Vailati, D. (2000) Nuovi dati sugli Anemadinae con descrizione di Anemadus whiteheadi sp. n. (Coleotera Cholevidae). Elytron, 14, 119 - 124." type="journal article" year="2000">Giachino &amp; Vailati (2000</bibRefCitation>
,
<bibRefCitation id="EFC340D5F808FFA5FC99FEB0FC074A59" author="Giachino, P. M. &amp; Vailati, D." box="[878,938,296,320]" pageId="17" pageNumber="18" pagination="365 - 420" refId="ref13390" refString="Giachino, P. M. &amp; Vailati, D. (2007) I Coleotteri Cholevidi dell'Appennino settentrionale e centrale: inventario, analisi faunistica e origine del popolamento (Coleoptera Cholevidae). Biogeographia, 28, 365 - 420." type="journal article" year="2007">2007</bibRefCitation>
), as well as specimens in the SMNS (see Appdenxi), still identified as
<taxonomicName id="4C5246A7F808FFA5FE1AFED5FD344A7C" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[493,665,332,357]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F808FFA5FE1AFED5FD344A7C" box="[493,665,332,357]" italics="true" pageId="17" pageNumber="18">S. orchesioides</emphasis>
</taxonomicName>
. These records likely correspond to
<taxonomicName id="4C5246A7F808FFA5FBBEFED5FB714A7C" baseAuthorityName="Reitter" baseAuthorityYear="1885" box="[1097,1244,332,357]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B926E136F808FFA5FBBEFED5FB714A7C" box="[1097,1244,332,357]" italics="true" pageId="17" pageNumber="18">S. pulchellus</emphasis>
</taxonomicName>
(P.M. Giachino, personal communication 2018; see below under
<taxonomicName id="4C5246A7F808FFA5FD31FEE8FCF54A91" baseAuthorityName="Reitter" baseAuthorityYear="1885" box="[710,856,367,392]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="B926E136F808FFA5FD31FEE8FCF54A91" box="[710,856,367,392]" italics="true" pageId="17" pageNumber="18">S. pulchellus</emphasis>
</taxonomicName>
). The same happens with all the records from the Aurès mountains in
<collectingCountry id="F3457DB4F808FFA5FE72FE0CFE744AB4" box="[389,473,404,429]" name="Algeria" pageId="17" pageNumber="18">Algeria</collectingCountry>
and
<collectingCountry id="F3457DB4F808FFA5FDEFFE0CFDC64AB4" box="[536,619,404,429]" name="Tunisia" pageId="17" pageNumber="18">Tunisia</collectingCountry>
(
<bibRefCitation id="EFC340D5F808FFA5FD8AFE0CFCBA4AB4" author="Jeannel, R." box="[637,791,404,429]" pageId="17" pageNumber="18" pagination="1 - 433" refId="ref13829" refString="Jeannel, R. (1936) Monographie des Catopides. Memoires du Museum National d'Histoire Naturelle, 1, 1 - 433." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
) and the Eastern half of
<collectingCountry id="F3457DB4F808FFA5FBB4FE0CFB3A4AB4" box="[1091,1175,404,429]" name="Algeria" pageId="17" pageNumber="18">Algeria</collectingCountry>
(
<bibRefCitation id="EFC340D5F808FFA5FB5DFE0CFAF44AB4" author="Giachino, P. M." box="[1194,1369,404,429]" pageId="17" pageNumber="18" pagination="337 - 348" refId="ref13257" refString="Giachino, P. M. (1985) Choleva (s. str.) gabriellae e Sciodrepoides casalei nuove specie della Kabylia e note corologiche sui Catopidi d'Algeria (Coleoptera, Catopidae). Bollettino del Museo Regionale di Scienze Naturali, Torino, 3, 337 - 348." type="journal article" year="1985">Giachino 1985</bibRefCitation>
). The records from the Moroccan Rif (
<bibRefCitation id="EFC340D5F808FFA5FDF1FE2FFD304AC9" author="Jeannel, R." box="[518,669,439,464]" pageId="17" pageNumber="18" pagination="1 - 433" refId="ref13829" refString="Jeannel, R. (1936) Monographie des Catopides. Memoires du Museum National d'Histoire Naturelle, 1, 1 - 433." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
) may correspond to
<taxonomicName id="4C5246A7F808FFA5FC70FE20FC544AC9" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[903,1017,439,464]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F808FFA5FC70FE20FC544AC9" box="[903,1017,439,464]" italics="true" pageId="17" pageNumber="18">S. brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F808FFA5FBF6FE2FFBEC4AC9" bold="true" box="[1025,1089,439,464]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A2155C4DF808FFA5FBF6FE2FFBEC4AC9" box="[1025,1089,439,464]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but the identity of those from the Middle Atlas, also in
<collectingCountry id="F3457DB4F808FFA5FE45FE45FDBA4AED" box="[434,535,477,500]" name="Morocco" pageId="17" pageNumber="18">Morocco</collectingCountry>
(
<bibRefCitation id="EFC340D5F808FFA5FDD2FE44FC904AED" author="Giachino, P. M. &amp; Vailati, D." box="[549,829,476,501]" pageId="17" pageNumber="18" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
), is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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