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<document id="9ECDF080E229F9F1721C8C75627DBA17" ID-DOI="10.11646/zootaxa.2319.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5317868" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629657556386" checkinUser="felipe" docAuthor="Müller, Hans-Georg &amp; Krapp, Franz" docDate="2009" docId="03E687F82977FF807ADC17C89CCBFEC1" docLanguage="en" docName="zt02319p138.pdf" docOrigin="Zootaxa 2319 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2319.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Tanystylum isthmiacum Stock 1955" docType="treatment" docVersion="5" lastPageNumber="51" masterDocId="FFDFFF802947FFB27A4B175F9C1FFF8D" masterDocTitle="The pycnogonid fauna (Pycnogonida, Arthropoda) of the Tayrona National Park and adjoining areas on the Caribbean coast of Colombia 2319" masterLastPageNumber="138" masterPageNumber="1" pageNumber="49" updateTime="1699090509478" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D0EE383932567A649158360F8963726F">The pycnogonid fauna (Pycnogonida, Arthropoda) of the Tayrona National Park and adjoining areas on the Caribbean coast of Colombia 2319</mods:title>
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<mods:namePart id="205D2270A6F1D97171AF6AFD066F9315">Müller, Hans-Georg</mods:namePart>
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<mods:namePart id="AAF881511F8A14C9FE461CD21F45FF5F">Krapp, Franz</mods:namePart>
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<paragraph id="8BF036EE2977FF827ADC17C89E54FF3C" blockId="48.[151,587,151,212]" box="[151,587,151,177]" pageId="48" pageNumber="49">
<heading id="D0B881822977FF827ADC17C89E54FF3C" bold="true" box="[151,587,151,177]" fontSize="11" level="1" pageId="48" pageNumber="49" reason="1">
<taxonomicName id="4C4F4D6D2977FF827ADC17C89E54FF3C" ID-CoL="54QGF" authority="Stock, 1955" authorityName="Stock" authorityYear="1955" box="[151,587,151,177]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="isthmiacum">
<emphasis id="B93BEAFC2977FF827ADC17C89E54FF3C" bold="true" box="[151,587,151,177]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827ADC17C89DA8FF3C" bold="true" box="[151,439,151,177]" italics="true" pageId="48" pageNumber="49">Tanystylum isthmiacum</emphasis>
Stock, 1955
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C35565652977FF827ADC17E59CF6FF59" box="[151,233,186,212]" pageId="48" pageNumber="49" type="description">
<paragraph id="8BF036EE2977FF827ADC17E59CF6FF59" blockId="48.[151,587,151,212]" box="[151,233,186,212]" pageId="48" pageNumber="49">
<figureCitation id="13742A6B2977FF827ADC17E59CF6FF59" box="[151,233,186,212]" captionStart="FIGURE 23" captionStartId="49.[151,256,1760,1784]" captionTargetBox="[184,1407,193,1732]" captionTargetId="figure-107@49.[176,1408,176,1754]" captionTargetPageId="49" captionText="FIGURE 23. Tanystylum isthmiacum Stock, 1955: A male dorsal, B male lateral, C male proboscis ventral, D female proboscis dorsal, E another female proboscis dorsal, F male palp, G male oviger, H male leg 3, I cement gland duct, male leg 3, J female oviger." figureDoi="http://doi.org/10.5281/zenodo.5362480" httpUri="https://zenodo.org/record/5362480/files/figure.png" pageId="48" pageNumber="49">Fig. 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35565652977FF827ADC16579ECBFEF2" pageId="48" pageNumber="49" type="reference_group">
<paragraph id="8BF036EE2977FF827ADC16579F2FFE92" blockId="48.[151,1436,264,383]" box="[151,816,264,287]" pageId="48" pageNumber="49">
<taxonomicName id="4C4F4D6D2977FF827ADC16579E53FE92" ID-CoL="54QGF" authority="Stock, 1955: 247" authorityName="Stock" authorityPageNumber="247" authorityYear="1955" box="[151,588,264,287]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="isthmiacum">
<emphasis id="B93BEAFC2977FF827ADC16579D8EFE92" box="[151,401,264,287]" italics="true" pageId="48" pageNumber="49">Tanystylum isthmiacum</emphasis>
Stock, 1955: 247
</taxonomicName>
.—Müller 1990: 280.
</paragraph>
<paragraph id="8BF036EE2977FF827ADC16779D63FED2" blockId="48.[151,1436,264,383]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="AB5F11C02977FF827ADC16779D63FED2" pageId="48" pageNumber="49">
<taxonomicName id="4C4F4D6D2977FF827ADC16779DF7FEB2" ID-CoL="5F6Z7" authorityName="Stock" authorityYear="1966" box="[151,488,296,319]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="isthmiacum" subSpecies="difficile">
<emphasis id="B93BEAFC2977FF827ADC16779DF7FEB2" box="[151,488,296,319]" italics="true" pageId="48" pageNumber="49">Tanystylum isthmiacum difficile</emphasis>
</taxonomicName>
—Fage &amp; Stock, 1966: 318. Stock: 389. Stock 1975: 74.
<treatmentCitation id="0AEE10FF2977FF827E03167798F4FEB2" author="Child, C. A." box="[1096,1259,296,319]" page="29" pageId="48" pageNumber="49" year="1979">
<bibRefCitation id="EFDE4B1F2977FF827E03167798F4FEB2" author="Child, C. A." box="[1096,1259,296,319]" pageId="48" pageNumber="49" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child 1979: 29</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AEE10FF2977FF827EBE16779988FEB2" author="Stock, J. H." box="[1269,1431,296,319]" page="11" pageId="48" pageNumber="49" year="1979">
<bibRefCitation id="EFDE4B1F2977FF827EBE16779988FEB2" author="Stock, J. H." box="[1269,1431,296,319]" pageId="48" pageNumber="49" pagination="1 - 32" refId="ref77628" refString="Stock, J. H. (1979) Pycnogonida from the mediolittoral and infralittoral zones in the tropical Western Atlantic. Studies in the Fauna of Curacao and other Caribbean Islands, 59 (184), 1 - 32." type="journal article" year="1979">Stock 1979: 11</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AEE10FF2977FF827A8C16179D67FED2" author="Stock, J. H." box="[199,376,328,351]" page="402" pageId="48" pageNumber="49" year="1986">
<bibRefCitation id="EFDE4B1F2977FF827A8C16179D67FED2" author="Stock, J. H." box="[199,376,328,351]" pageId="48" pageNumber="49" pagination="399 - 441" refId="ref77709" refString="Stock, J. H. (1986) Pycnogonida from the Caribbean and the straits of Florida. Bulletin of marine Science, 38 (3), 399 - 441." type="journal article" year="1986">Stock 1986: 402</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF036EE2977FF827ADC16379ECBFEF2" blockId="48.[151,1436,264,383]" box="[151,724,360,383]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="AB5F11C02977FF827ADC16379ECBFEF2" box="[151,724,360,383]" pageId="48" pageNumber="49">
<taxonomicName id="4C4F4D6D2977FF827ADC16379E0DFEF2" ID-CoL="54QGF" authority="isthmiacum" box="[151,530,360,383]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="isthmiacum" subSpecies="isthmiacum">
<emphasis id="B93BEAFC2977FF827ADC16379E0DFEF2" box="[151,530,360,383]" italics="true" pageId="48" pageNumber="49">Tanystylum isthmiacum isthmiacum</emphasis>
</taxonomicName>
<treatmentCitation id="0AEE10FF2977FF82786616379ECFFEF2" author="Child, C. A." box="[557,720,360,383]" page="29" pageId="48" pageNumber="49" year="1979">
<bibRefCitation id="EFDE4B1F2977FF82786616379ECFFEF2" author="Child, C. A." box="[557,720,360,383]" pageId="48" pageNumber="49" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child 1979: 29</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BF036EE2977FF827ADC16F198ADFE45" blockId="48.[151,1437,430,1971]" box="[151,1202,430,456]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827ADC16F19D1BFE45" bold="true" box="[151,260,430,456]" pageId="48" pageNumber="49">Material</emphasis>
:
<emphasis id="B93BEAFC2977FF827B5816F19D3DFE45" bold="true" box="[275,290,430,456]" pageId="48" pageNumber="49">4</emphasis>
.—
<materialsCitation id="3B273CB32977FF827B0016F198B1FE45" box="[331,1198,430,456]" collectingDate="1985-05-24" collectionCode="SMF" pageId="48" pageNumber="49" specimenCode="SMF 1399" specimenCount="2" specimenCount-male="1">
<specimenCount id="9D49FD672977FF827B0016F19D86FE45" box="[331,409,430,456]" pageId="48" pageNumber="49" type="male">1 male</specimenCount>
,
<specimenCount id="9D49FD672977FF827BEF16F19DF9FE45" box="[420,486,430,456]" pageId="48" pageNumber="49" type="generic">1 juv.</specimenCount>
(
<specimenCode id="DBE99E952977FF827BBF16F19E69FE45" box="[500,630,430,456]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="48" pageNumber="49">SMF 1399</specimenCode>
), algal growth on rocks, mesolittoral,
<date id="FFF1102E2977FF827E7816F198B1FE45" box="[1075,1198,430,456]" pageId="48" pageNumber="49" value="1985-05-24">
<collectingDate id="EFB5E9C62977FF827E7816F198B1FE45" box="[1075,1198,430,456]" pageId="48" pageNumber="49" value="1985-05-24">24.V.1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D168A98BDFE62" blockId="48.[151,1437,430,1971]" box="[198,1186,469,495]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D168A9CFCFE62" bold="true" box="[198,227,469,495]" pageId="48" pageNumber="49">16</emphasis>
.—
<materialsCitation id="3B273CB32977FF827B42168A9881FE62" box="[265,1182,469,495]" collectingDate="1985-07-15" collectionCode="SMF" pageId="48" pageNumber="49" specimenCode="SMF 1400" specimenCount="3" specimenCount-male="3">
<specimenCount id="9D49FD672977FF827B42168A9D7DFE62" box="[265,354,469,495]" pageId="48" pageNumber="49" type="male">3 males</specimenCount>
(
<specimenCode id="DBE99E952977FF827B39168A9DEBFE62" box="[370,500,469,495]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="48" pageNumber="49">SMF 1400</specimenCode>
), brown and red algae on rocks, mesolittoral,
<date id="FFF1102E2977FF827E40168A9881FE62" box="[1035,1182,469,495]" pageId="48" pageNumber="49" value="1985-07-15">
<collectingDate id="EFB5E9C62977FF827E40168A9881FE62" box="[1035,1182,469,495]" pageId="48" pageNumber="49" value="1985-07-15">15.VII.1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D16A49898FD9B" blockId="48.[151,1437,430,1971]" box="[198,1159,507,534]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D16A49CFCFD98" bold="true" box="[198,227,507,533]" pageId="48" pageNumber="49">17</emphasis>
.—
<materialsCitation id="3B273CB32977FF827B4716A3989CFD9B" box="[268,1155,508,534]" collectingDate="1985-11-08" collectionCode="ZFMK" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D49FD672977FF827B4716A39D48FD9B" box="[268,343,508,534]" pageId="48" pageNumber="49" type="male">1 male</specimenCount>
(ov.) (
<collectionCode id="ED5EAE2B2977FF827BEF16A39DE3FD9B" box="[420,508,508,534]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="48" pageNumber="49" type="Museum">ZFMK</collectionCode>
), from
<taxonomicName id="4C4F4D6D2977FF82781B16A39EEBFD98" authorityName="Splettstoesser" authorityYear="1929" box="[592,756,508,533]" class="Hydrozoa" family="Thyroscyphidae" genus="Cnidoscyphus" kingdom="Animalia" order="Leptothecata" pageId="48" pageNumber="49" phylum="Cnidaria" rank="genus">
<emphasis id="B93BEAFC2977FF82781B16A39EEBFD98" box="[592,756,508,533]" italics="true" pageId="48" pageNumber="49">Cnidoscyphus</emphasis>
</taxonomicName>
on rocks, mesolittoral,
<date id="FFF1102E2977FF827E4216A3989CFD9B" box="[1033,1155,508,534]" pageId="48" pageNumber="49" value="1985-11-08">
<collectingDate id="EFB5E9C62977FF827E4216A3989CFD9B" box="[1033,1155,508,534]" pageId="48" pageNumber="49" value="1985-11-08">8.XI.1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D157D9E5BFD07" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D157D9CFCFDB1" bold="true" box="[198,227,546,572]" pageId="48" pageNumber="49">20</emphasis>
.—
<materialsCitation id="3B273CB32977FF827B47157D98C5FDB1" box="[268,1242,546,572]" collectingDate="1985-10-02" collectionCode="SMF" elevation="3" pageId="48" pageNumber="49" specimenCode="SMF 1401" specimenCount="1">
<specimenCount id="9D49FD672977FF827B47157D9D54FDB1" box="[268,331,546,572]" pageId="48" pageNumber="49" type="generic">1 juv.</specimenCount>
(
<specimenCode id="DBE99E952977FF827B17157D9DC0FDB1" box="[348,479,546,572]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="48" pageNumber="49">SMF 1401</specimenCode>
), from
<taxonomicName id="4C4F4D6D2977FF827873157D9F31FDB6" box="[568,814,546,571]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" kingdom="Chromista" order="Fucales" pageId="48" pageNumber="49" phylum="Ochrophyta" rank="species" species="cymosum">
<emphasis id="B93BEAFC2977FF827873157D9F31FDB6" box="[568,814,546,571]" italics="true" pageId="48" pageNumber="49">Sargassum cymosum</emphasis>
</taxonomicName>
, 5 l substratum,
<quantity id="4CB79B0B2977FF8279B3157D9843FDB1" box="[1016,1116,546,572]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="48" pageNumber="49" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">
<elevation id="0062D1DD2977FF8279B3157D9843FDB1" box="[1016,1116,546,572]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="48" pageNumber="49" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">00.5 m</elevation>
</quantity>
,
<date id="FFF1102E2977FF827E21157D98C5FDB1" box="[1130,1242,546,572]" pageId="48" pageNumber="49" value="1985-10-02">
<collectingDate id="EFB5E9C62977FF827E21157D98C5FDB1" box="[1130,1242,546,572]" pageId="48" pageNumber="49" value="1985-10-02">2.X.1985</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B273CB32977FF827EA3157D9FDFFDEE" collectingDate="1985-11-08" collectionCode="SMF" pageId="48" pageNumber="49" specimenCode="SMF 1402" specimenCount="1">
1 fem. (gravid) (
<specimenCode id="DBE99E952977FF827AD415169D3AFDEE" box="[159,293,585,611]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="48" pageNumber="49">SMF 1402</specimenCode>
), from
<taxonomicName id="4C4F4D6D2977FF827B3615169E68FDEF" box="[381,631,585,610]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" kingdom="Chromista" order="Fucales" pageId="48" pageNumber="49" phylum="Ochrophyta" rank="species" species="cymosum">
<emphasis id="B93BEAFC2977FF827B3615169E68FDEF" box="[381,631,585,610]" italics="true" pageId="48" pageNumber="49">Sargassum cymosum</emphasis>
</taxonomicName>
, 5 l substratum,
<date id="FFF1102E2977FF82790815169FDFFDEE" box="[835,960,585,611]" pageId="48" pageNumber="49" value="1985-11-08">
<collectingDate id="EFB5E9C62977FF82790815169FDFFDEE" box="[835,960,585,611]" pageId="48" pageNumber="49" value="1985-11-08">8.XI.1985</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B273CB32977FF82798615169E20FD07" collectingDate="1986-03-03" collectionCode="SMF" pageId="48" pageNumber="49" specimenCode="SMF 1403" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D49FD672977FF82798615169836FDEE" box="[973,1065,585,611]" pageId="48" pageNumber="49" type="male">2 males</specimenCount>
(
<specimenCode id="DBE99E952977FF827E70151698DFFDEE" box="[1083,1216,585,611]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="48" pageNumber="49">SMF 1403</specimenCode>
), from
<taxonomicName id="4C4F4D6D2977FF827F5115169D1BFD04" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" kingdom="Chromista" order="Fucales" pageId="48" pageNumber="49" phylum="Ochrophyta" rank="species" species="cymosum">
<emphasis id="B93BEAFC2977FF827F5115169D1BFD04" italics="true" pageId="48" pageNumber="49">Sargassum cymosum</emphasis>
</taxonomicName>
, 5 l substratum,
<date id="FFF1102E2977FF827B8C152F9E20FD07" box="[455,575,624,650]" pageId="48" pageNumber="49" value="1986-03-03">
<collectingDate id="EFB5E9C62977FF827B8C152F9E20FD07" box="[455,575,624,650]" pageId="48" pageNumber="49" value="1986-03-03">3.III.1986</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D15C998C0FD3D" blockId="48.[151,1437,430,1971]" box="[198,1247,662,688]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D15C99CFCFD3D" bold="true" box="[198,227,662,688]" pageId="48" pageNumber="49">21</emphasis>
.—
<materialsCitation id="3B273CB32977FF827B4715C998C4FD3D" box="[268,1243,662,688]" collectingDate="1985-11-08" collectionCode="ZMA" pageId="48" pageNumber="49" specimenCode="ZMA 3381" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D49FD672977FF827B4715C99D48FD3D" box="[268,343,662,688]" pageId="48" pageNumber="49" type="male">1 male</specimenCount>
(
<specimenCode id="DBE99E952977FF827B2D15C99DF0FD3D" box="[358,495,662,688]" collectionCode="ZMA" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="48" pageNumber="49">ZMA 3381</specimenCode>
), from
<taxonomicName id="4C4F4D6D2977FF82780F15C89F1CFD3D" box="[580,771,663,688]" class="Aves" genus="Digenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="simplex">
<emphasis id="B93BEAFC2977FF82780F15C89F1CFD3D" box="[580,771,663,688]" italics="true" pageId="48" pageNumber="49">Digenia simplex</emphasis>
</taxonomicName>
, 5 l substratum, mesolittoral,
<date id="FFF1102E2977FF827E2A15C998C4FD3D" box="[1121,1243,662,688]" pageId="48" pageNumber="49" value="1985-11-08">
<collectingDate id="EFB5E9C62977FF827E2A15C998C4FD3D" box="[1121,1243,662,688]" pageId="48" pageNumber="49" value="1985-11-08">8.XI.1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35565652977FF827A8D15E29F1DF90B" pageId="48" pageNumber="49" type="description">
<paragraph id="8BF036EE2977FF827A8D15E29F0AFC16" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D15E29DA3FD5A" bold="true" box="[198,444,701,727]" pageId="48" pageNumber="49">Description of male</emphasis>
: Dorsal trunk outline circular, crurigers touching along their entire length; crurigers 13 with short, dorso-distal spines, mostly on flat-rounded antero-lateral tubercle; all trunk segments fused. Anterior rim of cephalon with two short lateral spines. Ocular process higher than broad, with a distal pointed tip. Eyes relatively large, distinctly pigmented. Abdomen rather short and directed obliquely dorsally, with 5 short spines in distal half, abdomen surpasses the 4
<superScript id="7C3A9BA62977FF8278A614079EE6FCEB" attach="left" box="[749,761,856,870]" fontSize="6" pageId="48" pageNumber="49">th</superScript>
crurigers by ca. 1/4 of its length. Proboscis rather robust, bottle-shaped, somewhat narrowed in distal third only.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D14F49F3AFBE8" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
Cheliphores 1/43/5 length of proboscis, distally rounded, with 34 short setae. Palp 6-articled, 4
<superScript id="7C3A9BA62977FF827F7514F79955FC3B" attach="left" box="[1342,1354,936,950]" fontSize="6" pageId="48" pageNumber="49">th</superScript>
article distinctly elongate, 3
<superScript id="7C3A9BA62977FF827BC0148E9D87FC52" attach="left" box="[395,408,977,991]" fontSize="6" pageId="48" pageNumber="49">rd</superScript>
article very short; terminal article twice as long as penultimate, with about 16 spines on lateral, ventral and distal aspect. Oviger 10-articled, robust; articles 4 and 5 elongate, the fifth slightly longer than the fourth; 7
<superScript id="7C3A9BA62977FF827B29137D9D71FBBD" attach="left" box="[354,366,1058,1072]" fontSize="6" pageId="48" pageNumber="49">th</superScript>
article with a pronounced roundish tubercle which bears ca. 5 short spines; terminal article the shortest and bearing a pair of denticulate flat spines.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D132D9CF7FA16" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">Legs robust, articles with relatively few short spines and setae; coxae 1 of all legs on distal rim with an antero-lateral, a dorsal and frequently a postero-lateral round tubercle; these tubercles armed with 1 to 3 short setae; coxae 1 and coxae 3 of equal length, coxae 2 slightly longer; cement gland tube of femur dorso-distal, with a short and nearly straight terminal region; tibia 1 and tibia 2 of equal length, barely longer than femur, with two dorsal flat tubercles; tarsus as long as broad, with 5 short ventral setae; propodus moderately curved and without pronounced heel; proximal third of sole armed with 3 robust spines; distally with ca. 5 short setae; main claw moderately curved of 1/2 length of propodus; auxiliary claws feebly curved, 1/4 of propodus length.</paragraph>
<paragraph id="8BF036EE2977FF827A8D12F59E1AF99F" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D12F59D62FA49" bold="true" box="[198,381,1450,1476]" pageId="48" pageNumber="49">Measurements</emphasis>
: Length 0.61; width (across 2
<superScript id="7C3A9BA62977FF8278BB12F89F1FFA38" attach="left" box="[752,768,1447,1461]" fontSize="6" pageId="48" pageNumber="49">nd</superScript>
crurigers) 0.63; abdomen 0.17; proboscis 0.36. Leg 3: coxa 1, 0.18; coxa 2, 0.19; coxa 3, 0.18; femur 0.41; tibia 1, 0.39; tibia 2, 0.39; tarsus 0.04; propodus 0.35; main claw 0.17; auxiliary 0.09.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D11419F1DF90B" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D11419D3EF9B5" bold="true" box="[198,289,1566,1592]" pageId="48" pageNumber="49">Female</emphasis>
: In contrast to to the previously decribed species without any apparent salient sexual dimorphism. Oviger conspicuously shorter and slenderer than in male; terminal article tiny, with one pair of denticulated foliose spines which are longer than the article itself.
</paragraph>
</subSubSection>
<subSubSection id="C35565652977FF837A8D11CD9F65F840" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="discussion">
<paragraph id="8BF036EE2977FF827A8D11CD9831F8AD" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
<emphasis id="B93BEAFC2977FF827A8D11CD9D27F921" bold="true" box="[198,312,1682,1708]" pageId="48" pageNumber="49">Remarks</emphasis>
:
<taxonomicName id="4C4F4D6D2977FF827B0211CC9DF0F921" authorityName="Stock" authorityYear="1955" box="[329,495,1683,1708]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="isthmiacum">
<emphasis id="B93BEAFC2977FF827B0211CC9DF0F921" box="[329,495,1683,1708]" italics="true" pageId="48" pageNumber="49">T. isthmiacum</emphasis>
</taxonomicName>
is morphologically very variable, which induced Fage &amp; Stock (1966: 318) to split it into 2 subspecies,
<taxonomicName id="4C4F4D6D2977FF827B8211E69F1DF95F" authority="isthmiacum" box="[457,770,1721,1746]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="isthmiacum" subSpecies="isthmiacum">
<emphasis id="B93BEAFC2977FF827B8211E69F1DF95F" box="[457,770,1721,1746]" italics="true" pageId="48" pageNumber="49">T. isthmiacum isthmiacum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D6D2977FF82797511E69856F95F" authorityName="Stock" authorityYear="1966" box="[830,1097,1721,1746]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="isthmiacum" subSpecies="difficile">
<emphasis id="B93BEAFC2977FF82797511E69856F95F" box="[830,1097,1721,1746]" italics="true" pageId="48" pageNumber="49">T. isthmiacum difficile</emphasis>
</taxonomicName>
. Here we do not follow this procedure as a clear separation is too difficult. The first author is convinced that an erection of subspecies is not helpful while even the distinction of species in this genus is problematical.
</paragraph>
<paragraph id="8BF036EE2977FF827A8D10729E49F83E" blockId="48.[151,1437,430,1971]" pageId="48" pageNumber="49">
Closer relations may exist with
<taxonomicName id="4C4F4D6D2977FF8278061072982FF8CA" authority="Hilton, 1942" authorityName="Hilton" authorityYear="1942" box="[589,1072,1837,1863]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="oculospinosum">
<emphasis id="B93BEAFC2977FF82780610729F92F8CB" box="[589,909,1837,1862]" italics="true" pageId="48" pageNumber="49">Tanystylum oculospinosum</emphasis>
Hilton, 1942
</taxonomicName>
from
<collectingRegion id="498BF80C2977FF827ECB10729928F8CA" box="[1152,1335,1837,1863]" country="Mexico" name="Baja California" pageId="48" pageNumber="49">Baja California</collectingRegion>
and the
<collectingRegion id="498BF80C2977FF827ADC100E9D0CF8E6" box="[151,275,1873,1899]" country="Ecuador" name="Galapagos" pageId="48" pageNumber="49">Galápagos</collectingRegion>
Archipelago (see Child &amp; Hedgpeth 1971: 619, fig.5). Based on the figures of Child &amp; Hedgpeth,
<taxonomicName id="4C4F4D6D2977FF827ADC102A9D24F803" authorityName="Stock" authorityYear="1955" box="[151,315,1909,1934]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="isthmiacum">
<emphasis id="B93BEAFC2977FF827ADC102A9D24F803" box="[151,315,1909,1934]" italics="true" pageId="48" pageNumber="49">T. isthmiacum</emphasis>
</taxonomicName>
is differentiated from
<taxonomicName id="4C4F4D6D2977FF82780F102A9F0EF803" authorityName="Hilton" authorityYear="1942" box="[580,785,1909,1934]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="oculospinosum">
<emphasis id="B93BEAFC2977FF82780F102A9F0EF803" box="[580,785,1909,1934]" italics="true" pageId="48" pageNumber="49">T. oculospinosum</emphasis>
</taxonomicName>
by more feebly developed tubercles on coxae 1 of legs and by more robust femora and tibiae.
</paragraph>
<caption id="DF3066662976FF837ADC11BF9D9CF8B1" ID-DOI="http://doi.org/10.5281/zenodo.5362480" ID-Zenodo-Dep="5362480" httpUri="https://zenodo.org/record/5362480/files/figure.png" pageId="49" pageNumber="50" startId="49.[151,256,1760,1784]" targetBox="[184,1407,193,1732]" targetPageId="49">
<paragraph id="8BF036EE2976FF837ADC11BF9D9CF8B1" blockId="49.[151,1436,1760,1852]" pageId="49" pageNumber="50">
<emphasis id="B93BEAFC2976FF837ADC11BF9D36F975" bold="true" box="[151,297,1760,1784]" pageId="49" pageNumber="50">FIGURE 23.</emphasis>
<taxonomicName id="4C4F4D6D2976FF837B7A11BF9EA2F97A" authority="Stock, 1955" authorityName="Stock" authorityYear="1955" box="[305,701,1760,1783]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="isthmiacum">
<emphasis id="B93BEAFC2976FF837B7A11BF9E2FF97A" box="[305,560,1760,1783]" italics="true" pageId="49" pageNumber="50">Tanystylum isthmiacum</emphasis>
Stock, 1955
</taxonomicName>
: A male dorsal, B male lateral, C male proboscis ventral, D female proboscis dorsal, E another female proboscis dorsal, F male palp, G male oviger, H male leg 3, I cement gland duct, male leg 3, J female oviger.
</paragraph>
</caption>
<paragraph id="8BF036EE2976FF837A8D10349F65F840" blockId="49.[151,1436,1899,1997]" pageId="49" pageNumber="50">
In the research area
<taxonomicName id="4C4F4D6D2976FF837BF510349E7AF809" authorityName="Stock" authorityYear="1955" box="[446,613,1899,1924]" class="Pycnogonida" family="Ammotheidae" genus="Tanystylum" kingdom="Animalia" order="Pantopoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="isthmiacum">
<emphasis id="B93BEAFC2976FF837BF510349E7AF809" box="[446,613,1899,1924]" italics="true" pageId="49" pageNumber="50">T. isthmiacum</emphasis>
</taxonomicName>
belongs among the more rarely found species. Mostly diverse algal stands from the mesolittoral to ca.
<quantity id="4CB79B0B2976FF83787010D09E67F824" box="[571,632,1935,1961]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="49" pageNumber="50" unit="m" value="0.5">0.5m</quantity>
depth were colonized. This comes as a surprise, as almost all data from the literature report this species from deeper water layers.
</paragraph>
</subSubSection>
<subSubSection id="C35565652975FF807A8D17C89CCBFEC1" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BF036EE2975FF807A8D17C89CCBFEC1" blockId="50.[151,1436,151,332]" pageId="50" pageNumber="51">
<emphasis id="B93BEAFC2975FF807A8D17C89D40FF3C" bold="true" box="[198,351,151,177]" pageId="50" pageNumber="51">Distribution</emphasis>
: From the Pacific there are records from
<collectingCountry id="F358767E2975FF80791417C79FFDFF3F" box="[863,994,152,178]" name="Costa Rica" pageId="50" pageNumber="51">Costa Rica</collectingCountry>
and
<collectingCountry id="F358767E2975FF807E6917C79864FF3F" box="[1058,1147,152,178]" name="Mexico" pageId="50" pageNumber="51">Mexico</collectingCountry>
(
<bibRefCitation id="EFDE4B1F2975FF807EC517C79955FF3F" author="Child, C. A." box="[1166,1354,152,178]" pageId="50" pageNumber="51" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child 1979: 29</bibRefCitation>
), from
<collectingCountry id="F358767E2975FF807ADC17E19CEBFF55" box="[151,244,190,216]" name="Panama" pageId="50" pageNumber="51">Panamá</collectingCountry>
(Stock 1955b: 247) and
<collectingCountry id="F358767E2975FF80785317E19E66FF55" box="[536,633,190,216]" name="Ecuador" pageId="50" pageNumber="51">Ecuador</collectingCountry>
(Stock 1975c: 74). In the western Atlantic the species was recorded from
<collectingCountry id="F358767E2975FF807A9317BA9D2AFF72" box="[216,309,229,255]" name="Panama" pageId="50" pageNumber="51">Panamá</collectingCountry>
(
<bibRefCitation id="EFDE4B1F2975FF807B0E17BA9DE5FF72" author="Child, C. A." box="[325,506,229,255]" pageId="50" pageNumber="51" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child 1979: 29</bibRefCitation>
),
<collectingCountry id="F358767E2975FF80785B17BA9E48FF72" box="[528,599,229,255]" name="Brazil" pageId="50" pageNumber="51">Brazil</collectingCountry>
(Stock 1975a: 984),
<collectingCountry id="F358767E2975FF80791B17BA9FAEFF72" box="[848,945,229,255]" name="Curaçao" pageId="50" pageNumber="51">Curaçao</collectingCountry>
(
<bibRefCitation id="EFDE4B1F2975FF80798A17BA986AFF72" author="Stock, J. H." box="[961,1141,229,255]" pageId="50" pageNumber="51" pagination="1 - 32" refId="ref77628" refString="Stock, J. H. (1979) Pycnogonida from the mediolittoral and infralittoral zones in the tropical Western Atlantic. Studies in the Fauna of Curacao and other Caribbean Islands, 59 (184), 1 - 32." type="journal article" year="1979">Stock 1979: 11</bibRefCitation>
) and Martinique (
<bibRefCitation id="EFDE4B1F2975FF807F0117BA9D3CFEAB" author="Muller, H. - G." pageId="50" pageNumber="51" pagination="277 - 285" refId="ref77186" refString="Muller, H. - G. (1990 d) Shallow-water Pycnogonida from Martinique, French Antilles, with description of Nymphon macabou n. sp. Bonner zoologische Beitrage, 41 (3 - 4), 277 - 285." type="journal article" year="1990">Müller 1990d: 280</bibRefCitation>
). From the eastern Atlantic there is one find from the Islands of
<collectingCountry id="F358767E2975FF807E7A165398A4FEAB" box="[1073,1211,268,294]" name="Cape Verde" pageId="50" pageNumber="51">Cape Verde</collectingCountry>
only (Stock 1966: 389).
</paragraph>
</subSubSection>
</treatment>
</document>