339 lines
49 KiB
XML
339 lines
49 KiB
XML
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<mods:title id="6203360F80BED90928425CDC06A846D4">An appraisal of Typilobus Stoliczka, 1871 (Crustacea, Brachyura, Leucosioidea), with description of a new family and genus</mods:title>
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<mods:namePart id="605E60DA94A3B3876EE1F421C405B8AA">Hyžný, Matúš</mods:namePart>
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<mods:title id="CBDAE0A19CE169AB584BC5403902A0C0">Zootaxa</mods:title>
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<treatment id="03DB1737FF8BFFB4FF6442B9633B6CE6" ID-DOI="http://doi.org/10.5281/zenodo.6082117" ID-GBIF-Taxon="120735461" ID-Zenodo-Dep="6082117" LSID="urn:lsid:plazi:treatment:03DB1737FF8BFFB4FF6442B9633B6CE6" httpUri="http://treatment.plazi.org/id/03DB1737FF8BFFB4FF6442B9633B6CE6" lastPageId="5" lastPageNumber="392" pageId="3" pageNumber="390">
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<subSubSection id="C368F5AAFF8BFFB2FF6442B967CB6DF8" box="[151,462,152,178]" pageId="3" pageNumber="390" type="nomenclature">
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<paragraph id="8BCDA621FF8BFFB2FF6442B967CB6DF8" blockId="3.[151,462,152,178]" box="[151,462,152,178]" pageId="3" pageNumber="390">
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<heading id="D085114DFF8BFFB2FF6442B967CB6DF8" bold="true" box="[151,462,152,178]" fontSize="11" level="1" pageId="3" pageNumber="390" reason="1">
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<taxonomicName id="4C72DDA2FF8BFFB2FF6442B967CB6DF8" ID-CoL="9F4M9" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[151,462,152,178]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="390" phylum="Arthropoda" rank="genus">
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<emphasis id="B9067A33FF8BFFB2FF6442B967CB6DF8" bold="true" box="[151,462,152,178]" pageId="3" pageNumber="390">
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<emphasis id="B9067A33FF8BFFB2FF6442B967086DF8" bold="true" box="[151,269,152,178]" italics="true" pageId="3" pageNumber="390">Typilobus</emphasis>
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<bibRefCitation id="EFE3DBD0FF8BFFB2FEE742B967CB6DF8" author="Stoliczka" box="[276,462,152,178]" pageId="3" pageNumber="390" refString="Stoliczka, F. (1871) Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica, series 7, 14, volume 1, part 1, 16 p., 5 pls." type="book" year="1871">Stoliczka, 1871</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C368F5AAFF8BFFB2FF6442FE64A76DBE" box="[151,674,222,244]" pageId="3" pageNumber="390" type="materials_examined">
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<paragraph id="8BCDA621FF8BFFB2FF6442FE64A76DBE" blockId="3.[151,674,222,244]" box="[151,674,222,244]" pageId="3" pageNumber="390">
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<typeStatus id="54C91883FF8BFFB2FF6442FE66CC6DBE" box="[151,201,223,244]" pageId="3" pageNumber="390">Type</typeStatus>
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species:
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<taxonomicName id="4C72DDA2FF8BFFB2FED542FF64A76DBE" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[294,674,222,244]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="granulosus">
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<emphasis id="B9067A33FF8BFFB2FED542FF67FA6DBE" box="[294,511,222,244]" italics="true" pageId="3" pageNumber="390">Typilobus granulosus</emphasis>
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<bibRefCitation id="EFE3DBD0FF8BFFB2FDF642FF64A76DBE" author="Stoliczka" box="[517,674,222,244]" pageId="3" pageNumber="390" refString="Stoliczka, F. (1871) Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica, series 7, 14, volume 1, part 1, 16 p., 5 pls." type="book" year="1871">Stoliczka, 1871</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C368F5AAFF8BFFB2FF64430066EB6FA0" pageId="3" pageNumber="390" type="diagnosis">
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<paragraph id="8BCDA621FF8BFFB2FF64430066EB6FA0" blockId="3.[151,1437,289,746]" pageId="3" pageNumber="390">
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<emphasis id="B9067A33FF8BFFB2FF644300678B6C70" bold="true" box="[151,398,289,314]" pageId="3" pageNumber="390">Emended diagnosis.</emphasis>
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Carapace transversely oval, moderately convex; front very narrow, bilobed, scarcely projected; orbits small, subcircular, directed forward; anterolateral margins arched, with small indentation at level of the cervical groove; posterolateral margins nearly straight, converging posteriorly, bearing small protuberance at level of cardiac region; posterior margin narrow, straight; main dorsal regions not subdivided; gastric region large, swollen, bounded by conspicuous grooves; hepatic region small, flat, subtriangular; branchial region large, swollen, bearing small protuberance in posterior portion; cardiac region transversely ovate, fairly swollen; intestinal region small, depressed; gastrohepatic, cervical, cardiac grooves well marked; sternum fairly broad, sternites 1–4 fused, sternites 5–6 subparallel, short, broad, sternite 7 somewhat inclined, sternite 8 inclined, reduced, small portion of it not totally covered by male pleon; male pleon narrow, long, reaching sternite 3, pleonal segments 3–5 fused, telson long, narrow, with rounded tip; female abdomen narrow, long, nearly reaching sternite 2, pleonal segments 3–6 fused, bearing 3 conspicuous longitudinal rows of subtle protuberances. Exopodite, endopodite of Mxp3 long, equally broad, both completely covering buccal cavity; epistome as wide as frontal margin.
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</paragraph>
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</subSubSection>
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<caption id="DF0DF6A9FF8BFFB2FF644518671A6AA2" httpUri="https://zenodo.org/record/266706/files/figure.png" pageId="3" pageNumber="390" targetBox="[176,1408,794,1819]" targetPageId="3">
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<paragraph id="8BCDA621FF8BFFB2FF644518671A6AA2" blockId="3.[151,1436,1849,2024]" pageId="3" pageNumber="390">
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<emphasis id="B9067A33FF8BFFB2FF64451867176A05" bold="true" box="[151,274,1849,1871]" pageId="3" pageNumber="390">FIGURE 1</emphasis>
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.
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<taxonomicName id="4C72DDA2FF8BFFB2FED7451864B76A05" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[292,690,1849,1871]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="granulosus">
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<emphasis id="B9067A33FF8BFFB2FED7451864076A05" box="[292,514,1849,1871]" italics="true" pageId="3" pageNumber="390">Typilobus granulosus</emphasis>
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Stoliczka, 1871
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</taxonomicName>
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, the type species of
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<taxonomicName id="4C72DDA2FF8BFFB2FC6A451862AE6A05" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[921,1195,1849,1871]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="390" phylum="Arthropoda" rank="genus">
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<emphasis id="B9067A33FF8BFFB2FC6A451865FE6A05" box="[921,1019,1849,1871]" italics="true" pageId="3" pageNumber="390">Typilobus</emphasis>
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Stoliczka, 1871
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</taxonomicName>
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, from the Miocene of Pakistan. A, NHMUK In28638 in dorsal (A1), ventral (A2), and posterior (A3) views; B, NHMUK In22990 (figured by Withers 1925: pl. 3, figs. 1–2) in lateral (B1), dorsal (B2), ventral (B3), posterior (B4), and frontal (B5) views; C–D, digital images of Stoliczka (1871: pl. 3, figs. 3, 5) showing male (C) and female (D) specimens. Note that schematic drawings of pleon (C2, D2) do not fully correspond to the figures of entire specimens (C1, D1). Specimens in A and B are to scale. Scale bar equals 5 mm.
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</paragraph>
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</caption>
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<caption id="DF0DF6A9FF8CFFB5FF644128640B6EF3" httpUri="https://zenodo.org/record/266707/files/figure.png" pageId="4" pageNumber="391" targetBox="[154,1431,195,750]" targetPageId="4">
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<paragraph id="8BCDA621FF8CFFB5FF644128640B6EF3" blockId="4.[151,1437,777,953]" pageId="4" pageNumber="391">
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<emphasis id="B9067A33FF8CFFB5FF64412867146E55" bold="true" box="[151,273,777,799]" pageId="4" pageNumber="391">FIGURE 2</emphasis>
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. Morphological variation shown in several species assigned to
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<taxonomicName id="4C72DDA2FF8CFFB5FC4A4128621E6E55" box="[953,1051,777,799]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
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<emphasis id="B9067A33FF8CFFB5FC4A4128621E6E55" box="[953,1051,777,799]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
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</taxonomicName>
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. A,
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<taxonomicName id="4C72DDA2FF8CFFB5FBBC412867226E74" authority="Morris & Collins, 1991" authorityName="Morris & Collins" authorityYear="1991" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="marginatus">
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<emphasis id="B9067A33FF8CFFB5FBBC4128632A6E55" box="[1103,1327,777,799]" italics="true" pageId="4" pageNumber="391">Typilobus marginatus</emphasis>
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Morris & Collins, 1991
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</taxonomicName>
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, holotype NHMUK In62163 in dorsal (A1) and ventral view (A2); B–C,
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<taxonomicName id="4C72DDA2FF8CFFB5FBBA4109639D6E74" authority="Withers, 1925" authorityName="Withers" authorityYear="1925" box="[1097,1432,808,830]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="sadeki">
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<emphasis id="B9067A33FF8CFFB5FBBA410962F26E74" box="[1097,1271,808,830]" italics="true" pageId="4" pageNumber="391">Typilobus sadeki</emphasis>
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Withers, 1925
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</taxonomicName>
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, holotype NHMUK In22991 (B) and In22992 (C) in dorsal (B1, C1) and ventral views (B2, C2); D,
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<taxonomicName id="4C72DDA2FF8CFFB5FB69416767206E31" authority="Quayle & Collins, 1981" authorityName="Quayle & Collins" authorityYear="1981" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="belli">
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<emphasis id="B9067A33FF8CFFB5FB69416763356E16" box="[1178,1328,838,860]" italics="true" pageId="4" pageNumber="391">Typilobus belli</emphasis>
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Quayle & Collins, 1981
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</taxonomicName>
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, holotype NHMUK In61707; E,
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<taxonomicName id="4C72DDA2FF8CFFB5FD76414465E26E31" authority="Muller, 1993" authorityName="Muller" authorityYear="1993" box="[645,999,869,891]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="moralejai">
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<emphasis id="B9067A33FF8CFFB5FD76414465566E31" box="[645,851,869,891]" italics="true" pageId="4" pageNumber="391">Typilobus moralejai</emphasis>
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Müller, 1993
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</taxonomicName>
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, holotype MGSB 51918; F,
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<taxonomicName id="4C72DDA2FF8CFFB5FAE5414467B46ED0" authority="Noetling, 1885" authorityName="Noetling" authorityYear="1885" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="corrodatus">
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<emphasis id="B9067A33FF8CFFB5FAE54144637D6E31" box="[1302,1400,869,891]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
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cf.
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<emphasis id="B9067A33FF8CFFB5FF6441A567006ED0" box="[151,261,900,922]" italics="true" pageId="4" pageNumber="391">corrodatus</emphasis>
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Noetling, 1885
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</taxonomicName>
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, BSPG 1981 XI 15. Specimens in E and F were coated with ammonium chloride prior the photography. Scale bar equals 5 mm.
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</paragraph>
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</caption>
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<subSubSection id="C368F5AAFF8CFFB4FF3441C1633B6CE6" lastPageId="5" lastPageNumber="392" pageId="4" pageNumber="391" type="discussion">
|
||
<paragraph id="8BCDA621FF8CFFB5FF3441C162BD6853" blockId="4.[151,1437,992,2025]" pageId="4" pageNumber="391">
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<emphasis id="B9067A33FF8CFFB5FF3441C1673E6EB3" bold="true" box="[199,315,992,1017]" pageId="4" pageNumber="391">Remarks.</emphasis>
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||
<bibRefCitation id="EFE3DBD0FF8CFFB5FEB141C1644E6EB2" author="Schweitzer" box="[322,587,992,1017]" pageId="4" pageNumber="391" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. & Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222. http: // dx. doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
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Schweitzer
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<emphasis id="B9067A33FF8CFFB5FE3541C067FD6EB3" box="[454,504,992,1017]" italics="true" pageId="4" pageNumber="391">et al</emphasis>
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||
. (2010
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||
</bibRefCitation>
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||
: 91) listed sixteen species assigned to
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<taxonomicName id="4C72DDA2FF8CFFB5FC0F41C1626D6EB3" box="[1020,1128,992,1017]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
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<emphasis id="B9067A33FF8CFFB5FC0F41C1626D6EB3" box="[1020,1128,992,1017]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
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</taxonomicName>
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, nine of them questionably indicating a problematic assignment. The rather broadly defined genus has been considered as embracing a wide range of morphotypes, as already noted by
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<bibRefCitation id="EFE3DBD0FF8CFFB5FD7346096540690B" author="Via" box="[640,837,1064,1089]" pageId="4" pageNumber="391" refString="Via Boada, L. (1969) Crustaceos decapodos del Eoceno espanol. Pirineos, 91 - 94, 1 - 469." type="book chapter" year="1969">
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||
Vía
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<bibRefCitation id="EFE3DBD0FF8CFFB5FD4346096540690B" author="Boada" box="[688,837,1064,1089]" pageId="4" pageNumber="391" refString="Via Boada, L. (1969) Crustaceos decapodos del Eoceno espanol. Pirineos, 91 - 94, 1 - 469." type="book chapter" year="1969">Boada (1969</bibRefCitation>
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||
</bibRefCitation>
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||
: 158),
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||
<bibRefCitation id="EFE3DBD0FF8CFFB5FC614609625A690A" author="Muller" box="[914,1119,1064,1089]" pageId="4" pageNumber="391" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993: 10)</bibRefCitation>
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, and
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||
<bibRefCitation id="EFE3DBD0FF8CFFB5FB6F4609639D690A" author="Feldmann" box="[1180,1432,1064,1089]" pageId="4" pageNumber="391" refString="Feldmann, R. M., Schweitzer, C. E., Bennett, O., Frantescu, O., Resar, N. & Trudeau, A. (2011) New Eocene Brachyura (Crustacea: Decapoda) from Egypt. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 262, 323 - 353. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B9067A33FF8CFFB5FAE046086341690B" box="[1299,1348,1064,1089]" italics="true" pageId="4" pageNumber="391">et al</emphasis>
|
||
. (2011
|
||
</bibRefCitation>
|
||
: 327). Therefore, the emended generic diagnosis provided herein is based solely on the
|
||
<typeStatus id="54C91883FF8CFFB5FB61466C62C7692F" box="[1170,1218,1101,1125]" pageId="4" pageNumber="391">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FADC466D671169C3" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9067A33FF8CFFB5FADC466D671169C3" italics="true" pageId="4" pageNumber="391">Typilobus granulosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1349BAA4FF8CFFB5FED74651676269C2" box="[292,359,1136,1161]" captionStart="FIGURE 1" captionStartId="3.[151,250,1849,1871]" captionTargetBox="[176,1408,794,1819]" captionTargetId="figure@3.[170,1418,784,1830]" captionTargetPageId="3" captionText="FIGURE 1. Typilobus granulosus Stoliczka, 1871, the type species of Typilobus Stoliczka, 1871, from the Miocene of Pakistan. A, NHMUK In 28638 in dorsal (A 1), ventral (A 2), and posterior (A 3) views; B, NHMUK In 22990 (figured by Withers 1925: pl. 3, figs. 1 – 2) in lateral (B 1), dorsal (B 2), ventral (B 3), posterior (B 4), and frontal (B 5) views; C – D, digital images of Stoliczka (1871: pl. 3, figs. 3, 5) showing male (C) and female (D) specimens. Note that schematic drawings of pleon (C 2, D 2) do not fully correspond to the figures of entire specimens (C 1, D 1). Specimens in A and B are to scale. Scale bar equals 5 mm." httpUri="https://zenodo.org/record/266706/files/figure.png" pageId="4" pageNumber="391">Fig. 1</figureCitation>
|
||
). The diagnosis of
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FDB44651654069C2" author="Feldmann" box="[583,837,1136,1161]" pageId="4" pageNumber="391" refString="Feldmann, R. M., Schweitzer, C. E., Bennett, O., Frantescu, O., Resar, N. & Trudeau, A. (2011) New Eocene Brachyura (Crustacea: Decapoda) from Egypt. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 262, 323 - 353. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B9067A33FF8CFFB5FD4C465064F769C3" box="[703,754,1136,1161]" italics="true" pageId="4" pageNumber="391">et al</emphasis>
|
||
. (2011
|
||
</bibRefCitation>
|
||
: 327) stated that the male pleonal segments are free. This observation was apparently based on the inaccurate drawings in
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FC5746B566B2699A" author="Stoliczka" pageId="4" pageNumber="391" refString="Stoliczka, F. (1871) Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica, series 7, 14, volume 1, part 1, 16 p., 5 pls." type="book" year="1871">
|
||
Stoliczka (1871: pl. 3, figs. 3, 5, see
|
||
<figureCitation id="1349BAA4FF8CFFB5FAB246B5637A69E6" box="[1345,1407,1172,1197]" captionStart="FIGURE 1" captionStartId="3.[151,250,1849,1871]" captionTargetBox="[176,1408,794,1819]" captionTargetId="figure@3.[170,1418,784,1830]" captionTargetPageId="3" captionText="FIGURE 1. Typilobus granulosus Stoliczka, 1871, the type species of Typilobus Stoliczka, 1871, from the Miocene of Pakistan. A, NHMUK In 28638 in dorsal (A 1), ventral (A 2), and posterior (A 3) views; B, NHMUK In 22990 (figured by Withers 1925: pl. 3, figs. 1 – 2) in lateral (B 1), dorsal (B 2), ventral (B 3), posterior (B 4), and frontal (B 5) views; C – D, digital images of Stoliczka (1871: pl. 3, figs. 3, 5) showing male (C) and female (D) specimens. Note that schematic drawings of pleon (C 2, D 2) do not fully correspond to the figures of entire specimens (C 1, D 1). Specimens in A and B are to scale. Scale bar equals 5 mm." httpUri="https://zenodo.org/record/266706/files/figure.png" pageId="4" pageNumber="391">Fig. 1</figureCitation>
|
||
C, D)
|
||
</bibRefCitation>
|
||
, which indicates all male and female abdominal segments free. According to the original description of
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FF6446FD677B69BF" author="Stoliczka" box="[151,382,1244,1269]" pageId="4" pageNumber="391" refString="Stoliczka, F. (1871) Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica, series 7, 14, volume 1, part 1, 16 p., 5 pls." type="book" year="1871">Stoliczka (1871: 16)</bibRefCitation>
|
||
, however, the male pleonal segments 3–5 are fused, whereas female pleonal segments appear “to consist of seven separate joints, but they are not very well defined in the single specimen”.
|
||
</paragraph>
|
||
<paragraph id="8BCDA621FF8CFFB5FF344705676D6A7E" blockId="4.[151,1437,992,2025]" pageId="4" pageNumber="391">
|
||
The outline of the carapace varies among the species of
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FCC4470565A66877" box="[823,931,1316,1341]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FCC4470565A66877" box="[823,931,1316,1341]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FC424705630A6877" authority="Withers, 1925" authorityName="Withers" authorityYear="1925" box="[945,1295,1316,1341]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="sadeki">
|
||
<emphasis id="B9067A33FF8CFFB5FC42470562696877" box="[945,1132,1316,1341]" italics="true" pageId="4" pageNumber="391">Typilobus sadeki</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FB814705630A6877" author="Withers" box="[1138,1295,1316,1341]" pageId="4" pageNumber="391" refString="Withers, T. H. (1925) Typibolus Sadeki, a new crab from the Miocene of Sinai. Appendix to Moon, F. W. and Sadek, H., Preliminary Geologic Report on Gebel Khosera Area. Ministry of Finances Egypt, Petroleum Research Bulletin, 9, 37 - 40, pl. 3." type="journal article" year="1925">Withers, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1349BAA4FF8CFFB5FAED470563646876" box="[1310,1377,1316,1341]" captionStart="FIGURE 2" captionStartId="4.[151,250,777,799]" captionTargetBox="[154,1431,195,750]" captionTargetId="figure@4.[150,1437,193,756]" captionTargetPageId="4" captionText="FIGURE 2. Morphological variation shown in several species assigned to Typilobus. A, Typilobus marginatus Morris & Collins, 1991, holotype NHMUK In 62163 in dorsal (A 1) and ventral view (A 2); B – C, Typilobus sadeki Withers, 1925, holotype NHMUK In 22991 (B) and In 22992 (C) in dorsal (B 1, C 1) and ventral views (B 2, C 2); D, Typilobus belli Quayle & Collins, 1981, holotype NHMUK In 61707; E, Typilobus moralejai Müller, 1993, holotype MGSB 51918; F, Typilobus cf. corrodatus Noetling, 1885, BSPG 1981 XI 15. Specimens in E and F were coated with ammonium chloride prior the photography. Scale bar equals 5 mm." httpUri="https://zenodo.org/record/266707/files/figure.png" pageId="4" pageNumber="391">Fig. 2</figureCitation>
|
||
B, C) and
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FF3447686413682A" authority="Karasawa, 1998" authorityName="Karasawa" authorityYear="1998" box="[199,534,1352,1377]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="kishimotoi">
|
||
<emphasis id="B9067A33FF8CFFB5FF344768675D682B" box="[199,344,1352,1377]" italics="true" pageId="4" pageNumber="391">T. kishimotoi</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FEAC47686413682A" author="Karasawa" box="[351,534,1352,1376]" pageId="4" pageNumber="391" refString="Karasawa, H. (1998) Typilobus kishimotoi, a new leucosiid crab (Crustacea: Decapoda: Brachyura) from the Miocene Katsuta Group, Japan. Proceedings of the Biological Society of Washington, 111, 97 - 101." type="journal article" year="1998">Karasawa, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, have a carapace with a transversely oval outline, similar to the
|
||
<typeStatus id="54C91883FF8CFFB5FB1B4768631D682A" box="[1256,1304,1353,1376]" pageId="4" pageNumber="391">type</typeStatus>
|
||
species; all three species are from Miocene strata and exhibit a similar distribution of dorsal regions and grooves. Other species assigned to the genus present a subcircular or longitudinally subelliptical carapace outline, similar grooves, and similar distribution of dorsal regions, e.g.,
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FD85479465DC6886" authority="Lorenthey, 1898" authorityName="Lorenthey" authorityYear="1898" box="[630,985,1460,1485]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="semseyanus">
|
||
<emphasis id="B9067A33FF8CFFB5FD85479465136886" box="[630,790,1461,1484]" italics="true" pageId="4" pageNumber="391">T. semseyanus</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FCED479565DC6886" author="Lorenthey" box="[798,985,1460,1485]" pageId="4" pageNumber="391" refString="Lorenthey, E. (1898) Beitrage zur Decapodenfauna der ungarischen Tertiars. Termeszetrajzi Fuzetek, 21, 1 - 133." type="journal article" year="1898">Lőrenthey, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FC174794632C6887" authority="Noetling, 1885" authorityName="Noetling" authorityYear="1885" box="[996,1321,1460,1485]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="corrodatus">
|
||
<emphasis id="B9067A33FF8CFFB5FC174794627F6887" box="[996,1146,1460,1485]" italics="true" pageId="4" pageNumber="391">T. corrodatus</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FB724795632C6887" author="Noetling" box="[1153,1321,1460,1485]" pageId="4" pageNumber="391" refString="Noetling, F. (1885) Die Fauna des samlandischen Tertiars. Abhandlungen zue Geologischen Spezialkarte von Preussen vol. 6, heft II, I. Teil, II: 112 - 172." type="journal article" year="1885">Noetling, 1885</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1349BAA4FF8CFFB5FACA479563786886" box="[1337,1405,1460,1485]" captionStart="FIGURE 2" captionStartId="4.[151,250,777,799]" captionTargetBox="[154,1431,195,750]" captionTargetId="figure@4.[150,1437,193,756]" captionTargetPageId="4" captionText="FIGURE 2. Morphological variation shown in several species assigned to Typilobus. A, Typilobus marginatus Morris & Collins, 1991, holotype NHMUK In 62163 in dorsal (A 1) and ventral view (A 2); B – C, Typilobus sadeki Withers, 1925, holotype NHMUK In 22991 (B) and In 22992 (C) in dorsal (B 1, C 1) and ventral views (B 2, C 2); D, Typilobus belli Quayle & Collins, 1981, holotype NHMUK In 61707; E, Typilobus moralejai Müller, 1993, holotype MGSB 51918; F, Typilobus cf. corrodatus Noetling, 1885, BSPG 1981 XI 15. Specimens in E and F were coated with ammonium chloride prior the photography. Scale bar equals 5 mm." httpUri="https://zenodo.org/record/266707/files/figure.png" pageId="4" pageNumber="391">Fig. 2</figureCitation>
|
||
F), or
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FF4447F8641068BA" authority="Lorenthey, 1907" authorityName="Lorenthey" authorityYear="1907" box="[183,533,1496,1521]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="B9067A33FF8CFFB5FF4447F8674A68BA" box="[183,335,1497,1520]" italics="true" pageId="4" pageNumber="391">T. trispinosus</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FEAA47F9641068BA" author="Lorenthey" box="[345,533,1496,1521]" pageId="4" pageNumber="391" refString="Lorenthey, I. [E.] (1907) Palaeontologiai tanulmanyok a harmadkor rakok korebol. V. Adatok Egyptom eocenkori rakfaunajahoz. Mathematikai es Termeszettudomanyi Kozlemenyek, 29 (2), 195 - 242, pls. 1, 2." type="journal article" year="1907">Lőrenthey, 1907</bibRefCitation>
|
||
</taxonomicName>
|
||
, all reported from Paleogene strata. Observation that the general outline of the Paleogene species assigned to
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FDF647DD64776B5F" box="[517,626,1532,1557]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FDF647DD64776B5F" box="[517,626,1532,1557]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of species reported from the Miocene strata were presented by
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FEDE440164276B72" author="Muller" box="[301,546,1568,1593]" pageId="4" pageNumber="391" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993: 10, 11)</bibRefCitation>
|
||
. Moreover, in his view, the
|
||
<typeStatus id="54C91883FF8CFFB5FCAA4400658C6B73" box="[857,905,1569,1593]" pageId="4" pageNumber="391">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FC03440163926B72" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[1008,1431,1568,1593]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9067A33FF8CFFB5FC03440162E56B73" box="[1008,1248,1568,1593]" italics="true" pageId="4" pageNumber="391">Typilobus granulosus</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FB14440163926B72" author="Stoliczka" box="[1255,1431,1568,1593]" pageId="4" pageNumber="391" refString="Stoliczka, F. (1871) Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica, series 7, 14, volume 1, part 1, 16 p., 5 pls." type="book" year="1871">Stoliczka, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
, is probably identical with
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FE31446465F26B16" authority="Bachmayer & Mohanti, 1973" authorityName="Bachmayer & Mohanti" authorityYear="1973" box="[450,1015,1604,1629]" class="Malacostraca" family="Leucosiidae" genus="Nucia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="baripadensis">
|
||
<emphasis id="B9067A33FF8CFFB5FE314464649B6B17" box="[450,670,1604,1629]" italics="true" pageId="4" pageNumber="391">Nucia baripadensis</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FD55446565F26B16" author="Bachmayer" box="[678,1015,1604,1629]" pageId="4" pageNumber="391" refString="Bachmayer, F. & Mohanti, M. (1973) Neue fossile Krebse aus dem Tertiar von Ost-Indien. Annalen der Naturhistorischen Museums in Wien, 77, 63 - 67, pls. 1 - 3." type="journal article" year="1973">Bachmayer & Mohanti, 1973</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Miocene of
|
||
<collectingCountry id="F365E6B1FF8CFFB5FB0B446563316B17" box="[1272,1332,1604,1629]" name="India" pageId="4" pageNumber="391">India</collectingCountry>
|
||
, and that the relatively narrow female pleon of
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FDA7444864F46BCB" box="[596,753,1640,1665]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9067A33FF8CFFB5FDA7444864F46BCB" box="[596,753,1640,1665]" italics="true" pageId="4" pageNumber="391">T. granulosus</emphasis>
|
||
</taxonomicName>
|
||
reflects the immature stage of the
|
||
<typeStatus id="54C91883FF8CFFB5FB7D444862BB6BCB" box="[1166,1214,1641,1665]" pageId="4" pageNumber="391">type</typeStatus>
|
||
specimens.
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FAA3444966E46BEF" author="Muller" pageId="4" pageNumber="391" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993)</bibRefCitation>
|
||
suggested that
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FE6744AC64606BEF" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[404,613,1676,1701]" class="Malacostraca" family="Leucosiidae" genus="Nucia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FE6744AC67D26BEE" box="[404,471,1677,1700]" italics="true" pageId="4" pageNumber="391">Nucia</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FE2C44AC64606BEF" author="Dana" box="[479,613,1676,1701]" pageId="4" pageNumber="391" refString="Dana, J. D. (1852) Crustacea. Part I. United States Exploring Expedition. During the years 1838, 1839, 1840, 1841, 1842. Under the command of Charles Wilkes, U. S. N. Vol. 13. Philadelphia, C. Sherman, 685 pp." type="book" year="1852">Dana, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
, seemed to be very close to the Miocene
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FBB344AD62A96BEF" box="[1088,1196,1676,1701]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FBB344AD62A96BEF" box="[1088,1196,1676,1701]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
species, whereas the Paleogene species of
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FE60449164056B83" box="[403,512,1712,1737]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FE60449164056B83" box="[403,512,1712,1737]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
might belong to an undescribed new genus. One of them,
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FB44449163726B83" box="[1207,1399,1712,1737]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="boscoi">
|
||
<emphasis id="B9067A33FF8CFFB5FB44449163726B83" box="[1207,1399,1712,1737]" italics="true" pageId="4" pageNumber="391">Typilobus boscoi</emphasis>
|
||
</taxonomicName>
|
||
, is recognized herein as representing a separate genus (see below). We agree with
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FBB944F562E86BA7" author="Muller" box="[1098,1261,1748,1773]" pageId="4" pageNumber="391" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993)</bibRefCitation>
|
||
that
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FADC44F567166A5B" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="granulosus">
|
||
<emphasis id="B9067A33FF8CFFB5FADC44F567166A5B" italics="true" pageId="4" pageNumber="391">Typilobus granulosus</emphasis>
|
||
</taxonomicName>
|
||
, is close to
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FE6944D867D86A5A" box="[410,477,1785,1808]" class="Malacostraca" family="Leucosiidae" genus="Nucia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FE6944D867D86A5A" box="[410,477,1785,1808]" italics="true" pageId="4" pageNumber="391">Nucia</emphasis>
|
||
</taxonomicName>
|
||
spp., but we refrain from synonymising
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FC5444D962116A5B" box="[935,1044,1784,1809]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FC5444D962116A5B" box="[935,1044,1784,1809]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FBA744D862926A5A" box="[1108,1175,1785,1808]" class="Malacostraca" family="Leucosiidae" genus="Nucia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FBA744D862926A5A" box="[1108,1175,1785,1808]" italics="true" pageId="4" pageNumber="391">Nucia</emphasis>
|
||
</taxonomicName>
|
||
, considering
|
||
<taxonomicName id="4C72DDA2FF8CFFB5FADC44D9639F6A5B" box="[1327,1434,1784,1809]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8CFFB5FADC44D9639F6A5B" box="[1327,1434,1784,1809]" italics="true" pageId="4" pageNumber="391">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
as a distinct taxon.
|
||
</paragraph>
|
||
<paragraph id="8BCDA621FF8CFFB4FF344561654F6DB2" blockId="4.[151,1437,992,2025]" lastBlockId="5.[151,1436,151,429]" lastPageId="5" lastPageNumber="392" pageId="4" pageNumber="391">
|
||
There is a high degree of morphological distinction in the distribution and shape of dorsal regions and grooves, and identifying gastric, cardiac, and intestinal regions is often difficult. Except for
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FBBD454463106A37" author="Via" box="[1102,1301,1892,1917]" pageId="4" pageNumber="391" refString="Via Boada, L. (1969) Crustaceos decapodos del Eoceno espanol. Pirineos, 91 - 94, 1 - 469." type="book chapter" year="1969">
|
||
Vía
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB5FB8C454563106A37" author="Boada" box="[1151,1301,1892,1917]" pageId="4" pageNumber="391" refString="Via Boada, L. (1969) Crustaceos decapodos del Eoceno espanol. Pirineos, 91 - 94, 1 - 469." type="book chapter" year="1969">Boada (1969</bibRefCitation>
|
||
</bibRefCitation>
|
||
: 156), most authors consider the cardiac region to be the swollen region situated at the rear of the carapace; the gastric region is described as equidimensionally hexagonal in some species, whereas the region is very much elongated in others, which is confusing. On the basis of the posterior placement, this region is recognized as intestinal (e.g.,
|
||
<bibRefCitation id="EFE3DBD0FF8CFFB4FA9745F166E56DFA" author="Galil" lastPageId="5" lastPageNumber="392" pageId="4" pageNumber="391" refString="Galil B. S. (2003 a) Four new genera of leucosiid crabs (Crustacea: Brachyura: Leucosiidae) for three new species and nine species previously in the genus Randallia Stimpson, 1857, with a redescription of the type species, R. ornata (Randall, 1939). Proceedings of the Biological Society of Washington, 116, 395 - 422." type="journal article" year="2003" yearSuffix="a">Galil 2003a</bibRefCitation>
|
||
). The gastric region is described as nearly equidimensionally hexagonal in
|
||
<taxonomicName id="4C72DDA2FF8DFFB4FBC542B863996DFA" authority="Quayle & Collins, 1981" authorityName="Quayle & Collins" authorityYear="1981" box="[1078,1436,151,176]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="392" phylum="Arthropoda" rank="species" species="belli">
|
||
<emphasis id="B9067A33FF8DFFB4FBC542B862866DFA" box="[1078,1155,151,176]" italics="true" pageId="5" pageNumber="392">T. belli</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FB7E42B663996DFA" author="Quayle" box="[1165,1436,151,176]" pageId="5" pageNumber="392" refString="Quayle, W. J. & Collins, J. S. H. (1981) New Eocene crabs from the Hampshire Basin. Palaeontology, 24, 733 - 758." type="journal article" year="1981">Quayle & Collins, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1349BAA4FF8DFFB4FF6C429D66E66D9E" box="[159,227,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,777,799]" captionTargetBox="[154,1431,195,750]" captionTargetId="figure@4.[150,1437,193,756]" captionTargetPageId="4" captionText="FIGURE 2. Morphological variation shown in several species assigned to Typilobus. A, Typilobus marginatus Morris & Collins, 1991, holotype NHMUK In 62163 in dorsal (A 1) and ventral view (A 2); B – C, Typilobus sadeki Withers, 1925, holotype NHMUK In 22991 (B) and In 22992 (C) in dorsal (B 1, C 1) and ventral views (B 2, C 2); D, Typilobus belli Quayle & Collins, 1981, holotype NHMUK In 61707; E, Typilobus moralejai Müller, 1993, holotype MGSB 51918; F, Typilobus cf. corrodatus Noetling, 1885, BSPG 1981 XI 15. Specimens in E and F were coated with ammonium chloride prior the photography. Scale bar equals 5 mm." httpUri="https://zenodo.org/record/266707/files/figure.png" pageId="5" pageNumber="392">Fig. 2</figureCitation>
|
||
D),
|
||
<taxonomicName id="4C72DDA2FF8DFFB4FEFD429C65456D9E" authority="Beschin, De Angeli & Zorzin, 2009" authorityName="Beschin, De Angeli & Zorzin" authorityYear="2009" box="[270,832,188,213]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="392" phylum="Arthropoda" rank="species" species="alponensis">
|
||
<emphasis id="B9067A33FF8DFFB4FEFD429C67A46D9F" box="[270,417,188,213]" italics="true" pageId="5" pageNumber="392">T. alponensis</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FE5A429D65456D9E" author="Beschin" box="[425,832,188,213]" pageId="5" pageNumber="392" refString="Beschin, C., De Angeli, A. & Zorzin, G. (2009) I decapodi dell'Eocene inferiore di Contrada Gecchelina (Vicenza, Italia settentrionale) (Anomura e Brachyura). Museo di Archeologia e Scienze Naturale " G. Zannato ", Montecchio Maggiore (Vicenza), 1 - 76." type="book chapter" year="2009">Beschin, De Angeli & Zorzin, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C72DDA2FF8DFFB4FC88429C62CF6D9F" authority="Via Boada, 1959" authorityName="Via Boada" authorityYear="1959" box="[891,1226,188,213]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="392" phylum="Arthropoda" rank="species" species="modregoi">
|
||
<emphasis id="B9067A33FF8DFFB4FC88429C62046D9F" box="[891,1025,188,213]" italics="true" pageId="5" pageNumber="392">T. modregoi</emphasis>
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FBFA429D62CF6D9F" author="Via" box="[1033,1226,188,213]" pageId="5" pageNumber="392" refString="Via Boada, L. (1959) Decapodos fosiles del Eoceno espanol. Boletin del Instituto Geologico y Minero de Espana, 70, 331 - 402." type="journal article" year="1959">Vía Boada, 1959</bibRefCitation>
|
||
</taxonomicName>
|
||
, whereas it is very much elongated in
|
||
<taxonomicName id="4C72DDA2FF8DFFB4FE9F42C065406DB2" authority="Feldmann et al. 2011: 328" authorityName="Feldmann et al." authorityPageNumber="328" authorityYear="2011" box="[364,837,223,248]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="392" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="B9067A33FF8DFFB4FE9F42C064066DB2" box="[364,515,225,248]" italics="true" pageId="5" pageNumber="392">T. trispinosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FDE242FE65016DB2" author="Feldmann" box="[529,772,223,248]" pageId="5" pageNumber="392" refString="Feldmann, R. M., Schweitzer, C. E., Bennett, O., Frantescu, O., Resar, N. & Trudeau, A. (2011) New Eocene Brachyura (Crustacea: Decapoda) from Egypt. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 262, 323 - 353. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B9067A33FF8DFFB4FD7942C064BE6DB2" box="[650,699,223,248]" italics="true" pageId="5" pageNumber="392">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
: 328)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCDA621FF8DFFB4FF344324633B6CE6" blockId="5.[151,1436,151,429]" pageId="5" pageNumber="392">
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FF34432467D06C56" author="Karasawa" box="[199,469,260,285]" pageId="5" pageNumber="392" refString="Karasawa, H. (1998) Typilobus kishimotoi, a new leucosiid crab (Crustacea: Decapoda: Brachyura) from the Miocene Katsuta Group, Japan. Proceedings of the Biological Society of Washington, 111, 97 - 101." type="journal article" year="1998">Karasawa (1998: fig. 3)</bibRefCitation>
|
||
presented a table with a summary of stratigraphic and geographic distribution of the genus. Based on his conclusion, the genus originated in the Western Tethys during the Eocene and subsequently migrated eastward to the Indo-West Pacific region in the Miocene. Although this scenario partly applies to other decapod taxa (e.g.,
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FE9C435167FF6CC2" author="Hyzny" box="[367,506,368,392]" pageId="5" pageNumber="392" refString="Hyzny, M. (2011) Revision of Jaxea kuemeli Bachmayer, 1954 (Decapoda: Gebiidea: Laomediidae) from the Miocene of Europe, with remarks on the palaeobiogeography of the genus Jaxea NARDO, 1847. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 260, 173 - 184. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2011 / 0165" type="journal article" year="2011">Hyžný 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FDF6435164FA6CC2" author="Hyzny" box="[517,767,367,392]" pageId="5" pageNumber="392" refString="Hyzny, M. & Muller, P. M. (2012) The fossil record of Glypturus Stimpson, 1866 (Crustacea, Decapoda, Axiidea, Callianassidae) revisited, with notes on palaeoecology and palaeobiogeography. Palaeontology, 55, 967 - 993. http: // dx. doi. org / 10.1111 / j. 1475 - 4983.2012.01169. x" type="journal article" year="2012">Hyžný & Müller 2012</bibRefCitation>
|
||
) and parallels the “Hopping Biodiversity Hotspots” concept of
|
||
<bibRefCitation id="EFE3DBD0FF8DFFB4FF4643B467A66CE6" box="[181,419,404,429]" pageId="5" pageNumber="392" refString="Renema, W., Bellwood, D. R., Braga, J. C., Bromfield, K., Hall, R., Johnson, K. G., Lunt, P., Meyer, C. P., McMonagle, L. B., Morley, R. J., O'Rea, A., Todd, J. A., Wesselingh, F. P., Wilson, M. E. J. & Pandolfi, J. M. (2008) Hopping hotspots: global shifts in marine biodiversity. Science, 321, 654 - 657. http: // dx. doi. org / 10.1126 / science. 1155674" type="journal article">
|
||
Renema
|
||
<emphasis id="B9067A33FF8DFFB4FEEB43B4674D6CE7" box="[280,328,404,429]" italics="true" pageId="5" pageNumber="392">et al</emphasis>
|
||
. (2008)
|
||
</bibRefCitation>
|
||
, it is difficult to accept it uncritically for such a heterogeneous taxon as
|
||
<taxonomicName id="4C72DDA2FF8DFFB4FB3943B563306CE7" box="[1226,1333,404,429]" class="Malacostraca" family="Leucosiidae" genus="Typilobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="392" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9067A33FF8DFFB4FB3943B563306CE7" box="[1226,1333,404,429]" italics="true" pageId="5" pageNumber="392">Typilobus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |