143 lines
19 KiB
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143 lines
19 KiB
XML
<document id="1B9BE6B01BDE3E607CCB6D6D0F97CBBE" ID-DOI="10.11646/zootaxa.4808.3.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="3933267" ID-ZooBank="01C982CD-56F1-4847-89AD-D6DD727D3172" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594121354116" checkinUser="plazi" docAuthor="Bogan, Sergio & Agnolin, Federico L." docDate="2020" docId="03D5261DFFD2DF06FF5AF886AD22FEFD" docLanguage="en" docName="zootaxa..4808.3.11.pdf" docOrigin="Zootaxa 4808 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthicus Agassiz 1829" docType="treatment" docVersion="3" lastPageNumber="575" masterDocId="FFEC5E65FFD3DF02FFCDFFF4AE7AFFE2" masterDocTitle="First fossil of the Giant Armored Catfish Acanthicus (Siluriformes, Loricariidae) from the Late Miocene of Entre Ríos Province, Argentina" masterLastPageNumber="581" masterPageNumber="571" pageNumber="572" updateTime="1700277237756" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="307BB05223E4905C0FB540E1D2B07FE7">
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<mods:title id="4DD9DDF6545723E68E66B6981C172642">First fossil of the Giant Armored Catfish Acanthicus (Siluriformes, Loricariidae) from the Late Miocene of Entre Ríos Province, Argentina</mods:title>
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<mods:name id="8D21560B0ACD23A377B8F0EB1DDB2B55" type="personal">
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<mods:roleTerm id="63F27991B554BCCF7E6AF6161B198D10">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E39F6D8BD8A52AAB81EA791C89037EA">Bogan, Sergio</mods:namePart>
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<mods:affiliation id="4F45ED7BBEC4BE06C94D513471136D71">Fundación de Historia Natural “ Félix de Azara ”. Centro de Ciencias Naturales, Ambientales y Antropológicas, Universidad Maimónides, Hidalgo 775 piso 7 (1405 BDB). Buenos Aires, Argentina.</mods:affiliation>
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<mods:name id="CD1A078AA14EAED51F0FE43A2AEFAAEF" type="personal">
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<mods:namePart id="1A58284969B7D9C7EA41BF0ACF978028">Agnolin, Federico L.</mods:namePart>
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<mods:affiliation id="F74D73D8D259B50D5B6DC6164B748A07">Fundación de Historia Natural “ Félix de Azara ”. Centro de Ciencias Naturales, Ambientales y Antropológicas, Universidad Maimónides, Hidalgo 775 piso 7 (1405 BDB). Buenos Aires, Argentina. & Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Av. Ángel Gallardo 470 (1405). Buenos Aires, Argentina.</mods:affiliation>
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<mods:typeOfResource id="FB40033CDC65818DF15AF9AF2E0EC120">text</mods:typeOfResource>
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<mods:title id="ABE75DF0079F741CD45628EAD1FF906B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B66013D115FC46AEB142EE1B13E4055C">
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<mods:date id="D21943EC12B824F6FA3FA52DB67C234D">2020</mods:date>
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<mods:number id="3D4D815EDD43805EC7C65C5B8139072F">2020-07-03</mods:number>
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<mods:number id="181BFDE7DE98FEE3F51360C3747862ED">4808</mods:number>
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<mods:detail id="745B0E05C1506CA63722508D131250E0" type="issue">
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<mods:number id="6D5DE68CCFFB3592CCDED67AB829A968">3</mods:number>
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<mods:start id="EF82F91C730AC20DF10BACD9DD447E35">571</mods:start>
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<mods:classification id="6E5CB5CD1453A9F6B1FBED6B3B956EE5">journal article</mods:classification>
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<mods:identifier id="8E0C3B24F8807FC4C2BA525F1AFBC5E2" type="DOI">10.11646/zootaxa.4808.3.11</mods:identifier>
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<mods:identifier id="A4756C081B0BAAAB31B6E54F8ADDCDE6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D5261DFFD2DF06FF5AF886AD22FEFD" LSID="urn:lsid:plazi:treatment:03D5261DFFD2DF06FF5AF886AD22FEFD" httpUri="http://treatment.plazi.org/id/03D5261DFFD2DF06FF5AF886AD22FEFD" lastPageId="4" lastPageNumber="575" pageId="1" pageNumber="572">
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<subSubSection id="C366C480FFD2DF03FF5AF886AF31F86F" box="[151,331,1906,1933]" pageId="1" pageNumber="572" type="nomenclature">
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<paragraph id="8BC3970BFFD2DF03FF5AF886AF31F86F" blockId="1.[151,331,1906,1933]" box="[151,331,1906,1933]" pageId="1" pageNumber="572">
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<heading id="D08B2067FFD2DF03FF5AF886AF31F86F" bold="true" box="[151,331,1906,1933]" fontSize="11" level="1" pageId="1" pageNumber="572" reason="1">
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<emphasis id="B9084B19FFD2DF03FF5AF886AF31F86F" bold="true" box="[151,331,1906,1933]" pageId="1" pageNumber="572">
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<taxonomicName id="4C7CEC88FFD2DF03FF5AF886AF5AF86F" ID-CoL="LZY" ID-ENA="337731" authorityName="Agassiz" authorityYear="1829" box="[151,288,1906,1933]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B9084B19FFD2DF03FF5AF886AF5AF86F" bold="true" box="[151,288,1906,1933]" italics="true" pageId="1" pageNumber="572">Acanthicus</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C366C480FFD2DF00FF5AF84DAAFBFF35" lastPageId="2" lastPageNumber="573" pageId="1" pageNumber="572" type="materials_examined">
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<paragraph id="8BC3970BFFD2DF03FF5AF84DABE6F81A" blockId="1.[151,1436,1977,2041]" pageId="1" pageNumber="572">
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<emphasis id="B9084B19FFD2DF03FF5AF84DAF0DF836" bold="true" box="[151,375,1977,2004]" pageId="1" pageNumber="572">Referred material.</emphasis>
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<collectionCode id="ED6D0FCEFFD2DF03FEB2F84EAFA9F836" box="[383,467,1978,2004]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="1" pageNumber="572" type="Museum">MACN</collectionCode>
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Pv-16105, partial skull table composed of the parieto-supraoccipital, and incomplete left and right sphenotics and frontals (
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<figureCitation id="13478B8EFFD2DF03FDFBF82AACEDF81A" box="[566,663,2014,2040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1368,1393]" captionTargetBox="[158,1429,661,1341]" captionTargetId="figure@3.[151,1436,654,1351]" captionTargetPageId="3" captionText="FIGURE 2. Acanthicus sp. MACN Pv-16105, partial skull table composed of the parieto-supraoccipital, and incomplete sphenotic and frontal bones in A, B, dorsal, C, ventral, D, left lateral, and E, right lateral views. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3933271" httpUri="https://zenodo.org/record/3933271/files/figure.png" pageId="1" pageNumber="572">Figure 2</figureCitation>
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);
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<collectionCode id="ED6D0FCEFFD2DF03FD66F82AAC85F81A" box="[683,767,2014,2040]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="1" pageNumber="572" type="Museum">MACN</collectionCode>
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Pv-13307, posterior half of a parieto-supraoccipital;
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<collectionCode id="ED6D0FCEFFD2DF03FA85F82AABE6F81A" box="[1352,1436,2014,2040]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="1" pageNumber="572" type="Museum">MACN</collectionCode>
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</paragraph>
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<paragraph id="8BC3970BFFD1DF00FF5AFF6DAAFBFF35" blockId="2.[151,1436,153,215]" pageId="2" pageNumber="573">
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Pv-16070, three incomplete compound pterotics, and right frontal; based on size congruence and differential preservation, it is very probable that these elements belong to different individuals (
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<figureCitation id="13478B8EFFD1DF00FBDCFF49AA09FF35" box="[1041,1139,189,215]" captionStart="FIGURE 3" captionStartId="4.[151,250,1038,1063]" captionTargetBox="[155,1431,315,1007]" captionTargetId="figure@4.[151,1436,309,1013]" captionTargetPageId="4" captionText="FIGURE 3. Acanthicus sp. MACN Pv-13307, posterior half of a parieto-supraoccipital bone in A dorsal, B ventral, and C anterior views; D-G, MACN Pv-16070, three incomplete compound pterotic bones D, indeterminate side, E,F, left side, and G, right side in D-E,G, dorsal, and F, ventral views; H-I, right frontal in H dorsal, and I ventral views. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3933273" httpUri="https://zenodo.org/record/3933273/files/figure.png" pageId="2" pageNumber="573">Figure 3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF03C783FFD1DF00FF5AF843ADC6F816" ID-DOI="http://doi.org/10.5281/zenodo.3933269" ID-Zenodo-Dep="3933269" httpUri="https://zenodo.org/record/3933269/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,250,1975,2000]" targetBox="[166,1421,238,1950]" targetPageId="2">
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<paragraph id="8BC3970BFFD1DF00FF5AF843ADC6F816" blockId="2.[151,1436,1975,2036]" pageId="2" pageNumber="573">
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<emphasis id="B9084B19FFD1DF00FF5AF843AF69F832" bold="true" box="[151,275,1975,2000]" pageId="2" pageNumber="573">FIGURE 1.</emphasis>
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Map showing the distribution of extant species of the genus
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<taxonomicName id="4C7CEC88FFD1DF00FCB8F84CAD9CF832" authorityName="Agassiz" authorityYear="1829" box="[885,998,1976,2000]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="573" phylum="Chordata" rank="genus">
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<emphasis id="B9084B19FFD1DF00FCB8F84CAD9CF832" box="[885,998,1976,2000]" italics="true" pageId="2" pageNumber="573">Acanthicus</emphasis>
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</taxonomicName>
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. The star indicates the fossiliferous locality of Paraná, Entre Ríos province, Argentina; Ituzaingó Formation, Late Miocene.
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</paragraph>
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</caption>
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<subSubSection id="C366C480FFD0DF06FF0AFF6CAD22FEFD" lastPageId="4" lastPageNumber="575" pageId="3" pageNumber="574" type="description">
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<paragraph id="8BC3970BFFD0DF01FF0AFF6CAB46FF35" blockId="3.[151,1437,152,647]" pageId="3" pageNumber="574">
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<emphasis id="B9084B19FFD0DF01FF0AFF6CAF23FF51" bold="true" box="[199,345,152,179]" pageId="3" pageNumber="574">Description.</emphasis>
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The skull roof is relatively thick and dorsally flat, showing a depressed area between the contact of the parieto-supraoccipital and frontals. It is dorsally ornamented by strong annular bases for odontodes.
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</paragraph>
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<paragraph id="8BC3970BFFD0DF01FF0AFF15AA9FFE31" blockId="3.[151,1437,152,647]" pageId="3" pageNumber="574">Parieto-supraoccipital bone is very elongate anteroposteriorly (its length representing more than two and a half times its maximum width). It is roughly subrectangular in contour, with its maximum transverse width located at the level of the articulation with the sphenotics. Lateral margins of the bone are nearly straight. Posteriorly, gradually decreases in transverse width. Posterior rear of the bone shows a rounded and almost smooth posterior process that is centrally located. The process is laterally delimited by shallow concavities for articulation with bony plates. The anterior margin of the bone shows the articulation for sphenotics and frontal. The parieto-supraoccipital shows an anterior process that is subtriangular in contour and extends as a wedge between the frontal bones.</paragraph>
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<paragraph id="8BC3970BFFD0DF01FF0AFE29AA54FD65" blockId="3.[151,1437,152,647]" pageId="3" pageNumber="574">
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The dorsal surface of the parieto-supraoccipital shows a particular ornamentation. From the center of the element radiate several grooves and lines of annular bases of odontodes that are roughly anteroposteriorly oriented. It shows anterior and posterior grooves and odontoid rows that result in a hourglass-shaped pattern, that is attenutaed towards the middle portion of the bone. Towards the lateral sides, the ornamentation is shallower and the rows of grooves and odontodes are more separate from each other (
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<figureCitation id="13478B8EFFD0DF01FCE0FD99ADBCFD65" box="[813,966,621,647]" captionStart="FIGURE 2" captionStartId="3.[151,250,1368,1393]" captionTargetBox="[158,1429,661,1341]" captionTargetId="figure@3.[151,1436,654,1351]" captionTargetPageId="3" captionText="FIGURE 2. Acanthicus sp. MACN Pv-16105, partial skull table composed of the parieto-supraoccipital, and incomplete sphenotic and frontal bones in A, B, dorsal, C, ventral, D, left lateral, and E, right lateral views. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3933271" httpUri="https://zenodo.org/record/3933271/files/figure.png" pageId="3" pageNumber="574">Figures 2A,B</figureCitation>
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and
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<figureCitation id="13478B8EFFD0DF01FC31FD99AA5BFD65" box="[1020,1057,621,647]" captionStart="FIGURE 3" captionStartId="4.[151,250,1038,1063]" captionTargetBox="[155,1431,315,1007]" captionTargetId="figure@4.[151,1436,309,1013]" captionTargetPageId="4" captionText="FIGURE 3. Acanthicus sp. MACN Pv-13307, posterior half of a parieto-supraoccipital bone in A dorsal, B ventral, and C anterior views; D-G, MACN Pv-16070, three incomplete compound pterotic bones D, indeterminate side, E,F, left side, and G, right side in D-E,G, dorsal, and F, ventral views; H-I, right frontal in H dorsal, and I ventral views. Scale bar equals 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3933273" httpUri="https://zenodo.org/record/3933273/files/figure.png" pageId="3" pageNumber="574">3A</figureCitation>
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).
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</paragraph>
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<caption id="DF03C783FFD0DF01FF5AFAACAB6AFA77" ID-DOI="http://doi.org/10.5281/zenodo.3933271" ID-Zenodo-Dep="3933271" httpUri="https://zenodo.org/record/3933271/files/figure.png" pageId="3" pageNumber="574" startId="3.[151,250,1368,1393]" targetBox="[158,1429,661,1341]" targetPageId="3">
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<paragraph id="8BC3970BFFD0DF01FF5AFAACAB6AFA77" blockId="3.[151,1437,1368,1429]" pageId="3" pageNumber="574">
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<emphasis id="B9084B19FFD0DF01FF5AFAACAF68FA93" bold="true" box="[151,274,1368,1393]" pageId="3" pageNumber="574">FIGURE 2.</emphasis>
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<taxonomicName id="4C7CEC88FFD0DF01FED5FAADAFF3FA93" authorityName="Agassiz" authorityYear="1829" box="[280,393,1369,1393]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="574" phylum="Chordata" rank="genus">
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<emphasis id="B9084B19FFD0DF01FED5FAADAFF3FA93" box="[280,393,1369,1393]" italics="true" pageId="3" pageNumber="574">Acanthicus</emphasis>
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</taxonomicName>
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sp. MACN Pv-16105, partial skull table composed of the parieto-supraoccipital, and incomplete sphenotic and frontal bones in
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<emphasis id="B9084B19FFD0DF01FE59FA88AFDCFA77" bold="true" box="[404,422,1404,1429]" pageId="3" pageNumber="574">A</emphasis>
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,
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<emphasis id="B9084B19FFD0DF01FE7EFA89AFBEFA77" bold="true" box="[435,452,1405,1429]" pageId="3" pageNumber="574">B</emphasis>
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, dorsal,
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<emphasis id="B9084B19FFD0DF01FDD6FA88AC57FA77" bold="true" box="[539,557,1404,1429]" pageId="3" pageNumber="574">C</emphasis>
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, ventral,
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<emphasis id="B9084B19FFD0DF01FD40FA89ACE5FA77" bold="true" box="[653,671,1405,1429]" pageId="3" pageNumber="574">D</emphasis>
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, left lateral, and
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<emphasis id="B9084B19FFD0DF01FC86FA89AD26FA77" bold="true" box="[843,860,1405,1429]" pageId="3" pageNumber="574">E</emphasis>
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, right lateral views. Scale bar equals 1 cm.
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</paragraph>
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</caption>
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<paragraph id="8BC3970BFFD0DF01FF0AFA37AA67F9C4" blockId="3.[151,1437,1475,2041]" pageId="3" pageNumber="574">The sphenotics are poorly preserved and show eroded lateral margins. They articulate between the frontal and the parieto-supraoccipital anteriorly and forming a deep and concave notch that shows a homogeneously convex margin. The bone is relatively anteroposteriorly elongate and is dorsally convex.</paragraph>
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<paragraph id="8BC3970BFFD0DF01FF0AF9DBAFABF973" blockId="3.[151,1437,1475,2041]" pageId="3" pageNumber="574">The suture between the frontals and with the parieto-supraoccipital is somewhat curly. From the central portion of the element radiate several strong lines of annular bases of odontodes that orientate towards the posterior and medial margins of the bone.</paragraph>
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<paragraph id="8BC3970BFFD0DF01FF5AF968ADEEF8A7" blockId="3.[151,1437,1475,2041]" pageId="3" pageNumber="574">
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A more complete frontal bone (MACN Pv-16070) is available. The frontal is roughly
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<emphasis id="B9084B19FFD0DF01FB86F968AA21F957" box="[1099,1115,1692,1717]" italics="true" pageId="3" pageNumber="574">T</emphasis>
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-shaped and shows an orbital process that forms part of the orbital margin. The orbital process is well-developed and subrectangular in contour, with the anterior and posterior margins gradually decreasing their width near the orbital margin. The orbital surface is slightly concave and forms a smooth step. The ornamentation of the orbital process is represented by notable odontodes that are randonly distributed over the surface of the bone.
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</paragraph>
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<paragraph id="8BC3970BFFD0DF01FF0AF8BBAF40F81B" blockId="3.[151,1437,1475,2041]" pageId="3" pageNumber="574">The posterior process shows a convex posterior margin and is laterally delimited by a smooth surface for articulation with the sphenotic. The compound pterotic is represented by three incomplete specimens (MACN Pv-16070). It is a relatively large element and in ventral view is very complex. It shows the ossified transcapular ligament and a free posterior margin with a rounded lateral margin. There is a large and posterolaterally located opening for the postotic canal.</paragraph>
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<paragraph id="8BC3970BFFD7DF06FF0AFF6DAD22FEFD" blockId="4.[151,1436,153,287]" pageId="4" pageNumber="575">
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It is dorsally covered by odontode bases and a large number of pits that become more evident towards the anteromedial margin of the bone resulting in a honeycomb aspect. This fenestrate condition was regarded by
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<bibRefCitation id="EFEDEAFAFFD7DF06FAF0FF49AE9BFF19" author="Chamon, C. C." pageId="4" pageNumber="575" refId="ref5542" refString="Chamon, C. C. (2011) Revisao taxonomica e relacoes filogeneticas do grupo Acanthicus (Siluriformes, Loricariidae). Doctoral dissertation, Universidade de Sao Paulo, Sao Paulo, 270 pp." type="book" year="2011">Chamon (2011)</bibRefCitation>
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as typical for
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<taxonomicName id="4C7CEC88FFD7DF06FE5CFF15AC77FF19" authorityName="Agassiz" authorityYear="1829" box="[401,525,225,251]" class="Actinopterygii" family="Loricariidae" genus="Acanthicus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="575" phylum="Chordata" rank="genus">
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<emphasis id="B9084B19FFD7DF06FE5CFF15AC77FF19" box="[401,525,225,251]" italics="true" pageId="4" pageNumber="575">Acanthicus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7CEC88FFD7DF06FD81FF15AC80FF19" authorityName="Isbrucker" authorityYear="1980" box="[588,762,225,251]" class="Actinopterygii" family="Loricariidae" genus="Megalancistrus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="575" phylum="Chordata" rank="genus">
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<emphasis id="B9084B19FFD7DF06FD81FF15AC80FF19" box="[588,762,225,251]" italics="true" pageId="4" pageNumber="575">Megalancistrus</emphasis>
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</taxonomicName>
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. Posteriorly, the bone shows ornamentation composed by strong crests with well-developed annular bases for odontodes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |