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<document id="3AA159753944E1B30969E92294A6B6EA" ID-DOI="10.5281/zenodo.184620" ID-GBIF-Dataset="1d4a10eb-c628-4525-b1f1-5d92412e6346" ID-ISSN="1175-5326" ID-Zenodo-Dep="184620" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459991941659" checkinUser="plazi" docAuthor="Ocampo, Federico C. &amp; Vaz-De-Mello, Fernando Z." docDate="2008" docId="03CADA28F71A713AFF5AF973FD27772F" docLanguage="en" docName="zt01916p062.pdf" docOrigin="Zootaxa 1916" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Xenaclopus borneensis Arrow" docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="FFF3A250F71B7139FFCDFF99FFA87545" masterDocTitle="The genus Xenaclopus Arrow (Coleoptera: Scarabaeidae): redescription and removal from the Aclopinae, with systematic notes" masterLastPageNumber="62" masterPageNumber="57" pageNumber="58" updateTime="1698232666687" updateUser="plazi">
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<mods:title id="5F485AB331716C5247C5D820777D9263">The genus Xenaclopus Arrow (Coleoptera: Scarabaeidae): redescription and removal from the Aclopinae, with systematic notes</mods:title>
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<mods:namePart id="8A462AA93C020A3017F3EB7E77C658A7">Ocampo, Federico C.</mods:namePart>
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<mods:namePart id="5C028897893DDE4D516D1C45F3871025">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date id="8169B8D3972719CB554A37CD57862584">2008</mods:date>
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<mods:number id="00402E4452BC50E7C116FE591D6A1681">1916</mods:number>
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<mods:identifier id="F36FDD9582F9B20E8EEB64DE52C8CE38" type="DOI">10.5281/zenodo.184620</mods:identifier>
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<mods:identifier id="DC8A9BDE710FDB994B160985612AE5EF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="553476ACE2C527745A5CA34C7F9E1978" type="Zenodo-Dep">184620</mods:identifier>
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<treatment id="03CADA28F71A713AFF5AF973FD27772F" ID-DOI="http://doi.org/10.5281/zenodo.6231976" ID-GBIF-Taxon="119380019" ID-Zenodo-Dep="6231976" LSID="urn:lsid:plazi:treatment:03CADA28F71A713AFF5AF973FD27772F" httpUri="http://treatment.plazi.org/id/03CADA28F71A713AFF5AF973FD27772F" lastPageId="3" lastPageNumber="60" pageId="1" pageNumber="58">
<subSubSection id="C37938B5F71A713AFF5AF973FED57707" lastPageId="3" lastPageNumber="60" pageId="1" pageNumber="58" type="nomenclature">
<paragraph id="8BDC6B3EF71A7138FF5AF973FDAF7241" blockId="1.[151,519,1770,1796]" box="[151,519,1770,1796]" pageId="1" pageNumber="58">
<heading id="D094DC52F71A7138FF5AF973FDAF7241" bold="true" box="[151,519,1770,1796]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
<taxonomicName id="4C6310BDF71A7138FF5AF973FDAF7241" authority="Arrow" authorityName="Arrow" box="[151,519,1770,1796]" class="Insecta" family="Melolonthidae" genus="Xenaclopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="B917B72CF71A7138FF5AF973FDAF7241" bold="true" box="[151,519,1770,1796]" pageId="1" pageNumber="58">
<emphasis id="B917B72CF71A7138FF5AF973FE067241" bold="true" box="[151,430,1770,1796]" italics="true" pageId="1" pageNumber="58">Xenaclopus borneensis</emphasis>
Arrow
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDC6B3EF71A713AFF5AF8ACFED57707" blockId="1.[151,1437,1845,2031]" lastBlockId="3.[151,1437,152,618]" lastPageId="3" lastPageNumber="60" pageId="1" pageNumber="58">
<emphasis id="B917B72CF71A7138FF5AF8ACFE36720A" bold="true" box="[151,414,1845,1871]" pageId="1" pageNumber="58">Redescription: Male.</emphasis>
Length
<quantity id="4C9BC6DBF71A7138FDCFF8ACFD18720A" box="[514,688,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="3.965" metricValueMax="4.11" metricValueMin="3.82" pageId="1" pageNumber="58" unit="mm" value="3.965" valueMax="4.11" valueMin="3.82">3.82–4.11 mm</quantity>
, width
<quantity id="4C9BC6DBF71A7138FCC5F8ACFC1F720A" box="[776,951,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="2.505" metricValueMax="2.75" metricValueMin="2.26" pageId="1" pageNumber="58" unit="mm" value="2.505" valueMax="2.75" valueMin="2.26">2.26–2.75 mm</quantity>
. Color: Head, pronotum, elytra brown to dark-brown; head progressively darker from clypeus to frons, pronotum with 2 darker areas at middle, elytral margins and suture darker than disc.
<emphasis id="B917B72CF71A7138FD82F81CFD2672DB" box="[591,654,1925,1950]" italics="true" pageId="1" pageNumber="58">Head</emphasis>
(
<figureCitation id="135877BBF71A7138FD6DF81CFD4E72DA" box="[672,742,1925,1951]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1329,1353]" captionTargetBox="[278,1294,193,1281]" captionTargetId="figure@2.[269,1317,171,1325]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Xenaclopus borneensis. 1. head, dorsal view; 2. left mandible, dorsal view; 3. labium, ventral view; right maxilla, ventral view; 4. left antenna, dorsal view." httpUri="https://zenodo.org/record/184621/files/figure.png" pageId="1" pageNumber="58">Fig. 1</figureCitation>
): Eye canthus well developed, marginate with moderately long setae on anterior margin (note: setae not shown in illustration). Eyes well developed, rounded, not entirely divided by eye canthus. Frons slightly convex, surface sparsely punctate, sparsely setose; setae long, slender. Clypeus transverse; lateral margins oblique; anterior margin straight, slightly reflexed; frontoclypeal suture obsolete. Labrum protruding beyond clypeal margin, shape pentagonal with round apex with 5–6 long slender setae (setae not shown in illustration). Mandibles protruding beyond clypeal margin, broadly rounded externally, slightly concave dorsoventrally, lacking incisory teeth, molar area poorly developed (
<figureCitation id="135877BBF719713BFAF2F988FA25736E" box="[1343,1421,1553,1579]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1329,1353]" captionTargetBox="[278,1294,193,1281]" captionTargetId="figure@2.[269,1317,171,1325]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Xenaclopus borneensis. 1. head, dorsal view; 2. left mandible, dorsal view; 3. labium, ventral view; right maxilla, ventral view; 4. left antenna, dorsal view." httpUri="https://zenodo.org/record/184621/files/figure.png" pageId="2" pageNumber="59">Fig. 2</figureCitation>
). Labium (
<figureCitation id="135877BBF719713BFEC9F9A0FEE47316" box="[260,332,1593,1619]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1329,1353]" captionTargetBox="[278,1294,193,1281]" captionTargetId="figure@2.[269,1317,171,1325]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Xenaclopus borneensis. 1. head, dorsal view; 2. left mandible, dorsal view; 3. labium, ventral view; right maxilla, ventral view; 4. left antenna, dorsal view." httpUri="https://zenodo.org/record/184621/files/figure.png" pageId="2" pageNumber="59">Fig. 3</figureCitation>
) longer than wide, labial palpi with 3 palpomeres; surface setose, setae dense to moderately dense, long, slender. Maxillae poorly developed, maxillary palpus with 4 palpomeres (
<figureCitation id="135877BBF719713BFB40F9F8FB7E733E" box="[1165,1238,1633,1659]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1329,1353]" captionTargetBox="[278,1294,193,1281]" captionTargetId="figure@2.[269,1317,171,1325]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Xenaclopus borneensis. 1. head, dorsal view; 2. left mandible, dorsal view; 3. labium, ventral view; right maxilla, ventral view; 4. left antenna, dorsal view." httpUri="https://zenodo.org/record/184621/files/figure.png" pageId="2" pageNumber="59">Fig. 4</figureCitation>
). Antennae (
<figureCitation id="135877BBF719713BFAA1F9F8FF0E73E6" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1329,1353]" captionTargetBox="[278,1294,193,1281]" captionTargetId="figure@2.[269,1317,171,1325]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Xenaclopus borneensis. 1. head, dorsal view; 2. left mandible, dorsal view; 3. labium, ventral view; right maxilla, ventral view; 4. left antenna, dorsal view." httpUri="https://zenodo.org/record/184621/files/figure.png" pageId="2" pageNumber="59">Fig. 5</figureCitation>
) with 9 antennomeres, antennal club with 3 antennomeres; sixth antennomere greatly reduced, antennal club longer than antennomeres 1–6 combined.
<emphasis id="B917B72CF719713BFD4CF928FD5D738F" box="[641,757,1713,1738]" italics="true" pageId="2" pageNumber="59">Pronotum</emphasis>
(
<figureCitation id="135877BBF719713BFCC8F928FCE5738E" box="[773,845,1713,1739]" captionStart="FIGURES 6 – 7" captionStartId="3.[151,269,1267,1291]" captionTargetBox="[155,1417,691,1236]" captionTargetId="figure@3.[155,821,677,1243]" captionTargetPageId="3" captionText="FIGURES 6 – 7. Xenaclopus borneensis habitus. 6. dorsal view; 7. ventral view." httpUri="https://zenodo.org/record/184622/files/figure.png" pageId="2" pageNumber="59">Fig. 6</figureCitation>
): Convex, wider than long. Surface sparsely punctate, setose; setae long, slender. Marginal bead complete; anterior margin concave, with membrane; lateral margins broadly rounded; posterior margin slightly sinuous. Anterior angles nearly right-angled; posterior angles broadly rounded.
<taxonomicName id="4C6310BDF719713BFE0EF8B0FD9F7207" box="[451,567,1833,1858]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B917B72CF719713BFE0EF8B0FD9F7207" box="[451,567,1833,1858]" italics="true" pageId="2" pageNumber="59">Scutellum</emphasis>
</taxonomicName>
(
<figureCitation id="135877BBF719713BFD87F8B0FD3E7206" box="[586,662,1833,1859]" captionStart="FIGURES 6 – 7" captionStartId="3.[151,269,1267,1291]" captionTargetBox="[155,1417,691,1236]" captionTargetId="figure@3.[155,821,677,1243]" captionTargetPageId="3" captionText="FIGURES 6 – 7. Xenaclopus borneensis habitus. 6. dorsal view; 7. ventral view." httpUri="https://zenodo.org/record/184622/files/figure.png" pageId="2" pageNumber="59">Fig. 6</figureCitation>
): Large, triangular apex acute, surface sparsely punctate.
<emphasis id="B917B72CF719713BFA9EF8B0FA347207" box="[1363,1436,1833,1858]" italics="true" pageId="2" pageNumber="59">Elytra</emphasis>
(
<figureCitation id="135877BBF719713BFF52F8C8FF43722E" box="[159,235,1873,1899]" captionStart="FIGURES 6 – 7" captionStartId="3.[151,269,1267,1291]" captionTargetBox="[155,1417,691,1236]" captionTargetId="figure@3.[155,821,677,1243]" captionTargetPageId="3" captionText="FIGURES 6 – 7. Xenaclopus borneensis habitus. 6. dorsal view; 7. ventral view." httpUri="https://zenodo.org/record/184622/files/figure.png" pageId="2" pageNumber="59">Fig. 6</figureCitation>
.): Convex, globose, lateral margins broadly rounded. Elytral striae absent sparsely punctate, sparsely setose; setae moderately long; pseudoepipleura poorly developed.
<emphasis id="B917B72CF719713BFC65F8E0FC5872D7" box="[936,1008,1913,1938]" italics="true" pageId="2" pageNumber="59">Ve n te r</emphasis>
(
<figureCitation id="135877BBF719713BFBC9F8E0FBE672D6" box="[1028,1102,1913,1939]" captionStart="FIGURES 6 – 7" captionStartId="3.[151,269,1267,1291]" captionTargetBox="[155,1417,691,1236]" captionTargetId="figure@3.[155,821,677,1243]" captionTargetPageId="3" captionText="FIGURES 6 – 7. Xenaclopus borneensis habitus. 6. dorsal view; 7. ventral view." httpUri="https://zenodo.org/record/184622/files/figure.png" pageId="2" pageNumber="59">Fig. 7</figureCitation>
): Metasternum narrow, 0.51 times as long as wide; 6 ventrites exposed medially; penultimate ventrite as long as previous three ventrites combined (at middle), membranous on apical margin. Pygidium not exposed beyond elytral margin, wider than long, surface sparsely setose; setae long, slender. Abdominal spiracles dorsal to sternotergal suture.
<emphasis id="B917B72CF718713AFA91FF01FA3B75F4" box="[1372,1427,152,177]" italics="true" pageId="3" pageNumber="60">Legs</emphasis>
: Protibiae with 3 teeth (
<figureCitation id="135877BBF718713AFE67FF59FE5B759F" box="[426,499,192,218]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Fig. 8</figureCitation>
), protibial spur slightly curved. Mesotibiae and metatibiae (
<figureCitation id="135877BBF718713AFB0FFF59FAA2759F" box="[1218,1290,192,218]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Fig. 9</figureCitation>
) with short, medial, transverse carinae. Mesotibial and metatibial (
<figureCitation id="135877BBF718713AFCD8FF71FCC27447" box="[789,874,232,258]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Fig. 10</figureCitation>
) apex with fringe of thick setae; mesotibial and metatibial spurs contiguous, metatibial spurs (
<figureCitation id="135877BBF718713AFD79FE89FC9B746F" box="[692,819,272,298]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Figs. 9–10</figureCitation>
) both set below tarsal articulation; inner spur longer than outer spur. Metatibial tarsal insertion simple, without notch; metatibial apical margin entire. Tarsus approximately as long as tibiae. Tarsomeres 1 and 5 each longer than 2, 3, 4 individually; all tarsi with long, apical setae (
<figureCitation id="135877BBF718713AFEE3FE11FEDD74E7" box="[302,373,392,418]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Fig. 9</figureCitation>
). Protarsal claws curved, outer claw bifid (split) (
<figureCitation id="135877BBF718713AFC7BFE11FBA474E7" box="[950,1036,392,418]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Fig. 11</figureCitation>
). Mesotarsal and metatarsal claws simple (not bifid), symmetrical (
<figureCitation id="135877BBF718713AFDFDFE29FD22748F" box="[560,650,432,458]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Figs. 9</figureCitation>
).
<emphasis id="B917B72CF718713AFD6AFE29FCC9748C" box="[679,865,432,457]" italics="true" pageId="3" pageNumber="60">Male Genitalia</emphasis>
: phallobase dilated basally, elongate apically, entirely open ventrally; parameres simple, symmetrical, elongate; paramere apex acute, angled in relation to phallobase (
<figureCitation id="135877BBF718713AFEEAFD99FE11775F" box="[295,441,512,538]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Figs. 12–13</figureCitation>
), genital segment (spiculum gastrale) poorly sclerotized, Y-shaped with very wide basal arm (
<figureCitation id="135877BBF718713AFED5FDB1FEC77707" box="[280,367,552,578]" captionStart="FIGURES 8 – 14" captionStartId="4.[151,269,1818,1842]" captionTargetBox="[169,1433,200,1753]" captionTargetId="figure@4.[153,1433,194,1794]" captionTargetPageId="4" captionText="FIGURES 8 – 14. Xenaclopus borneensis. 8. left protibia, dorsal view; 9. right metatibia, dorsal view; 10. right metatibial apex, apical view; 11. left proclaws, ventral view; 12. aedeagus, lateral view; 13. parameres, apical view; 14. genital sclerite (spiculum gastrale), ventral view." httpUri="https://zenodo.org/record/184623/files/figure.png" pageId="3" pageNumber="60">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1C3BB6F719713BFF5AFAA8FD05702E" httpUri="https://zenodo.org/record/184621/files/figure.png" pageId="2" pageNumber="59" targetBox="[278,1294,193,1281]" targetPageId="2">
<paragraph id="8BDC6B3EF719713BFF5AFAA8FD05702E" blockId="2.[151,1436,1329,1387]" pageId="2" pageNumber="59">
<emphasis id="B917B72CF719713BFF5AFAA8FEEA700C" bold="true" box="[151,322,1329,1353]" pageId="2" pageNumber="59">FIGURES 1–5.</emphasis>
<taxonomicName id="4C6310BDF719713BFE85FAABFD95700C" box="[328,573,1330,1353]" class="Insecta" family="Melolonthidae" genus="Xenaclopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="B917B72CF719713BFE85FAABFD95700C" box="[328,573,1330,1353]" italics="true" pageId="2" pageNumber="59">Xenaclopus borneensis</emphasis>
</taxonomicName>
. 1. head, dorsal view; 2. left mandible, dorsal view; 3. labium, ventral view; right maxilla, ventral view; 4. left antenna, dorsal view.
</paragraph>
</caption>
<subSubSection id="C37938B5F718713AFF0BFDD6FD27772F" box="[198,655,591,618]" pageId="3" pageNumber="60" type="discussion">
<paragraph id="8BDC6B3EF718713AFF0BFDD6FD27772F" blockId="3.[151,1437,152,618]" box="[198,655,591,618]" pageId="3" pageNumber="60">
<emphasis id="B917B72CF718713AFF0BFDD6FEE8772C" bold="true" box="[198,320,591,617]" pageId="3" pageNumber="60">Remarks.</emphasis>
Females are unknown to us.
</paragraph>
</subSubSection>
</treatment>
</document>
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