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<document id="9F91E22D98DA2914001BE2E085F1FA7B" ID-CLB-Dataset="27348" ID-DOI="10.11646/zootaxa.4581.1.1" ID-GBIF-Dataset="5b259853-01f6-4307-8bef-58c98ec73bd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2633842" ID-ZooBank="20A9776D-AE5F-41BC-A35B-0C5E42EDFE48" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554804587469" checkinUser="plazi" docAuthor="Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian &amp; Wang, Yongjie" docDate="2019" docId="03C47176FF9F8D287AD203ADFB879502" docLanguage="en" docName="zootaxa.4581.1.1.pdf.imd" docOrigin="Zootaxa 4581 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Kempynosmylus Makarkin 2014" docType="treatment" docVersion="16" lastPageNumber="29" masterDocId="FFFD090EFF838D347A450740FFA0923D" masterDocTitle="Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera" masterLastPageNumber="99" masterPageNumber="1" pageNumber="29" updateTime="1698719428342" updateUser="ExternalLinkService">
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<mods:title id="6CAD9D9235615EFFFB17D1796C99245B">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:namePart id="17A8EE1A145A64F950E5008E2F046307">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="4D91CC748F89D60A3748612E1599C5AD">Martins, Caleb Califre</mods:namePart>
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<mods:namePart id="9820CCA14A89C9EDFC6A9F6ADBAB2869">Makarkin, Vladimir</mods:namePart>
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<mods:namePart id="D4138579D8F7840CFBE88BA6A968EB05">Ardila-Camacho, Adrian</mods:namePart>
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<treatment id="03C47176FF9F8D287AD203ADFB879502" ID-DOI="http://doi.org/10.5281/zenodo.5631447" ID-GBIF-Taxon="156146921" ID-Zenodo-Dep="5631447" LSID="urn:lsid:plazi:treatment:03C47176FF9F8D287AD203ADFB879502" httpUri="http://treatment.plazi.org/id/03C47176FF9F8D287AD203ADFB879502" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="C37793EBFF9F8D287AD203ADFD8C973A" box="[151,556,1261,1287]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BD2C060FF9F8D287AD203ADFD8C973A" blockId="28.[151,556,1261,1320]" box="[151,556,1261,1287]" pageId="28" pageNumber="29">
<heading id="D09A770CFF9F8D287AD203ADFD8C973A" bold="true" box="[151,556,1261,1287]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<taxonomicName id="4C6DBBE3FF9F8D287AD203ADFD8C973A" ID-CoL="637WH" authority="Makarkin, 2014" authorityName="Makarkin" authorityYear="2014" box="[151,556,1261,1287]" class="Insecta" family="Osmylidae" genus="Kempynosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF9F8D287AD203ADFD8C973A" bold="true" box="[151,556,1261,1287]" pageId="28" pageNumber="29">
<emphasis id="B9191C72FF9F8D287AD203ADFEFB973A" bold="true" box="[151,347,1261,1287]" italics="true" pageId="28" pageNumber="29">Kempynosmylus</emphasis>
<bibRefCitation id="EFFCBD91FF9F8D287B2603ADFD8C973A" author="Makarkin" box="[355,556,1261,1287]" firstAuthor="Makarkin" pageId="28" pageNumber="29" pagination="8 - 12" refId="ref50661" refString="Makarkin, V. N. (2014) A new fossil genus of Osmylidae (Neuroptera) from the Early Cretaceous of Baissa, Transbaikalia. Far Eastern Entomologist, 278, 8 - 12." type="journal article" year="2014">Makarkin, 2014</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C37793EBFF9F8D287AD2024FFEAB9715" box="[151,267,1295,1320]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BD2C060FF9F8D287AD2024FFEAB9715" blockId="28.[151,556,1261,1320]" box="[151,267,1295,1320]" pageId="28" pageNumber="29">
(
<figureCitation id="1356DCE5FF9F8D287ADA024FFEA29715" box="[159,258,1295,1320]" captionStart="FIGURE 11" captionStartId="26.[151,250,1390,1412]" captionTargetBox="[413,1100,192,1333]" captionTargetId="figure@26.[363,1224,139,1405]" captionText="FIGURE 11. Kempyninae fossils. A. Jurakempynus sinensis Wang et al. (from Wang et al., 2011a); B. Cretosmylus sibiricus Makarkin (from Makarkin, 1990) (forewing); C. Kempynosmylus zherikhini Makarkin (from Makarkin, 2014) (forewing) (figures not to scale)." httpUri="https://zenodo.org/record/2644606/files/figure.png" pageId="28" pageNumber="29">Fig. 11C</figureCitation>
)
</paragraph>
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<subSubSection id="C37793EBFF9F8D287AD20218FBA39753" box="[151,1027,1368,1390]" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BD2C060FF9F8D287AD20218FBA39753" blockId="28.[151,1027,1368,1390]" box="[151,1027,1368,1390]" pageId="28" pageNumber="29">
<emphasis id="B9191C72FF9F8D287AD20218FE829753" bold="true" box="[151,290,1368,1390]" pageId="28" pageNumber="29">
<typeStatus id="54D67EC2FF9F8D287AD20218FF6C9753" box="[151,204,1368,1390]" pageId="28" pageNumber="29">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6DBBE3FF9F8D287B6C0218FD5B9753" authority="Makarkin, 2014: 9" authorityName="Makarkin" authorityPageNumber="9" authorityYear="2014" box="[297,763,1368,1390]" class="Insecta" family="Osmylidae" genus="Kempynosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="zherikhini">
<emphasis id="B9191C72FF9F8D287B6C0218FD999753" box="[297,569,1368,1390]" italics="true" pageId="28" pageNumber="29">Kempynosmylus zherikhini</emphasis>
<bibRefCitation id="EFFCBD91FF9F8D2878050218FD459753" author="Makarkin" box="[576,741,1368,1390]" firstAuthor="Makarkin" pageId="28" pageNumber="29" pagination="8 - 12" refId="ref50661" refString="Makarkin, V. N. (2014) A new fossil genus of Osmylidae (Neuroptera) from the Early Cretaceous of Baissa, Transbaikalia. Far Eastern Entomologist, 278, 8 - 12." type="journal article" year="2014">Makarkin, 2014</bibRefCitation>
: 9
</taxonomicName>
(by original designation).
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<subSubSection id="C37793EBFF9F8D287AD202DAFDBB945A" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BD2C060FF9F8D287AD202DAFDBB945A" blockId="28.[151,1437,1434,1855]" pageId="28" pageNumber="29">
<emphasis id="B9191C72FF9F8D287AD202DAFEB1978E" bold="true" box="[151,273,1434,1459]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
Forewing ovoid; RP with numerous branches (&gt;16), strongly sinuate distally; numerous regularly arranged crossveins in basal half of wing, outer gradate series apparently absent; FW M forked nearly level with origin of RP2, approximately at level of A2 termination, MA few branched, MP pectinately branched (56 branches); FW CuA (34 branches) and CuP (1112 branches) both pectinately branched; A1 elongate, strongly pectinate (≥13 branches) with some irregular end-twigging; A2 strongly pectinate (7 branches); A3 with a few pectinate branches; HW unknown.
</paragraph>
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<subSubSection id="C37793EBFF9F8D287A820132FB879502" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BD2C060FF9F8D287A820132FB879502" blockId="28.[151,1437,1434,1855]" pageId="28" pageNumber="29">
<emphasis id="B9191C72FF9F8D287A820132FEEF94B6" bold="true" box="[199,335,1650,1675]" pageId="28" pageNumber="29">Comments.</emphasis>
Based on an incomplete forewing fragment, the apparent kempynine affinity of this genus (
<emphasis id="B9191C72FF9F8D287FC40133FA3D94B6" box="[1409,1437,1651,1675]" italics="true" pageId="28" pageNumber="29">cf.</emphasis>
<taxonomicName id="4C6DBBE3FF9F8D287AD201D6FEB29492" authorityName="Leach" authorityYear="1815" box="[151,274,1686,1711]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Osmylinae">Osmylinae</taxonomicName>
) remains tentative. Characters such as the numerous basal crossveins, position of the medial fork and sinuate distal branches of RP suggest that is should be placed in
<taxonomicName id="4C6DBBE3FF9F8D28792501FAFC4F94EE" authorityName="Carpenter" authorityYear="1943" box="[864,1007,1722,1747]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Kempyninae">Kempyninae</taxonomicName>
but not conclusively and placement in
<taxonomicName id="4C6DBBE3FF9F8D287AD2019EFEB094CA" authorityName="Leach" authorityYear="1815" box="[151,272,1758,1783]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Osmylinae">Osmylinae</taxonomicName>
cannot be ruled out. The wing venation of
<taxonomicName id="4C6DBBE3FF9F8D287945019EFC1694CA" authorityName="Makarkin" authorityYear="2014" box="[768,950,1758,1783]" class="Insecta" family="Osmylidae" genus="Kempynosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF9F8D287945019EFC1694CA" box="[768,950,1758,1783]" italics="true" pageId="28" pageNumber="29">Kempynosmylus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C6DBBE3FF9F8D287E08019EFB1A94CA" authorityName="Khramov" authorityYear="2014" box="[1101,1210,1758,1783]" class="Insecta" family="Osmylidae" genus="Arbusella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF9F8D287E08019EFB1A94CA" box="[1101,1210,1758,1783]" italics="true" pageId="28" pageNumber="29">Arbusella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DBBE3FF9F8D287EB6019EFA3594CA" box="[1267,1429,1758,1783]" class="Insecta" family="Osmylidae" genus="Jurakempynus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FF9F8D287EB6019EFA3594CA" box="[1267,1429,1758,1783]" italics="true" pageId="28" pageNumber="29">Jurakempynus</emphasis>
</taxonomicName>
, genera similarly lacking the outer gradate series. The status of the genus relative to these genera (and its subfamilial placement) will be clearer once additional, more complete material is discovered.
</paragraph>
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</treatment>
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