308 lines
51 KiB
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308 lines
51 KiB
XML
<document id="43DF5CBC31D9486342CEAD22A3BE26A1" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B094E3C4C57F94C3D26F839" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria tholiformis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov." docType="treatment" docUuid="C083E997-420A-4CAA-8A67-088C02177B83" docUuidSource="ZooBank" docVersion="15" lastPageNumber="38" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="37" updateTime="1716316918745" updateUser="carolina">
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<mods:title id="F608D3E0208FAF81D6CE34E3F10401E7">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:roleTerm id="6242D57CEFAF4ECD2FDAA3D76D2D855A">Author</mods:roleTerm>
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<mods:namePart id="EE7A1D14D610641689C9EF6525CD0DB0">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="5CAFC163A83985699399629053041534">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
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<mods:nameIdentifier id="40025E31FD1AF6F25345AA6901AE5F2A" type="email">ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="425D330422557E517C47BC8B8F7964FA">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="B73BA741B55135476ECE65D91BC9F4EE">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="4D5E0069FBA0E7BFFDE1E3BAB8806B83" type="email">mneiber@hotmail.de</mods:nameIdentifier>
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<mods:namePart id="9C8165389908986F4D2EF414716FAC54">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="DB9C39C1806A576AC736F96B122F8337">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:nameIdentifier id="F0DC817082AE143F3FA0309D135E60E4" type="email">hausdorf@zoologie.uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="7B79A9E31B6D8C9A4CAA1DC20F32943B">European Journal of Taxonomy</mods:title>
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<mods:date id="7D3461D9C53DE3B54918D0989BC6953F">2019</mods:date>
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<mods:number id="E9337497391B57570FC9981B6399A51F">2019-05-21</mods:number>
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<mods:number id="DBA958537367BB132A536FAE29CA2E8D">526</mods:number>
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<mods:identifier id="96865065F895FB23431C82747BB81A72" type="DOI">10.5852/ejt.2019.526</mods:identifier>
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<treatment id="03B1DB4C8B094E3C4C57F94C3D26F839" ID-GBIF-Taxon="157239599" LSID="urn:lsid:zoobank.org:act:C083E997-420A-4CAA-8A67-088C02177B83" httpUri="http://treatment.plazi.org/id/03B1DB4C8B094E3C4C57F94C3D26F839" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
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<subSubSection id="C30239D18B094E3D4C57F94C3FDFF910" box="[610,978,1712,1739]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BA76A5A8B094E3D4C57F94C3FDFF910" blockId="36.[368,1219,1712,1810]" box="[610,978,1712,1739]" pageId="36" pageNumber="37">
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<heading id="D0EFDD368B094E3D4C57F94C3FDFF910" box="[610,978,1712,1739]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
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<taxonomicName id="4C1811D98B094E3D4C57F94C3F7DF910" ID-CoL="7RTJC" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[610,880,1712,1739]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
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<emphasis id="B96CB6488B094E3D4C57F94C3F7DF910" bold="true" box="[610,880,1712,1739]" italics="true" pageId="36" pageNumber="37">Landouria tholiformis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F0B338B094E3D4D42F94D3FDFF910" box="[887,978,1713,1739]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30239D18B094E3D4F45F92F3FD0F8CA" pageId="36" pageNumber="37" type="description">
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<paragraph id="8BA76A5A8B094E3D4F45F92F38CEF936" blockId="36.[368,1219,1712,1810]" box="[368,1219,1747,1773]" pageId="36" pageNumber="37">
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<uri id="FF8966588B094E3D4F45F92F38CEF936" box="[368,1219,1747,1773]" pageId="36" pageNumber="37">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBE508F8B094E3D4CBBF92F38CEF936" box="[654,1219,1747,1773]" pageId="36" pageNumber="37">C083E997-420A-4CAA-8A67-088C02177B83</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BA76A5A8B094E3D4C63F90B3FD0F8CA" blockId="36.[368,1219,1712,1810]" box="[598,989,1783,1810]" pageId="36" pageNumber="37">
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<figureCitation id="132376DF8B094E3D4C63F90B3EACF8CA" box="[598,673,1783,1809]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="36" pageNumber="37">Figs 7</figureCitation>
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,
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<figureCitation id="132376DF8B094E3D4C98F90B3EC0F8CA" box="[685,717,1783,1809]" captionStart="Figs 15–24" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 15–24. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="36" pageNumber="37">20</figureCitation>
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,
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<figureCitation id="132376DF8B094E3D4CECF90B3EF4F8CA" box="[729,761,1783,1809]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="36" pageNumber="37">34</figureCitation>
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,
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<figureCitation id="132376DF8B094E3D4D30F90B3F5FF8C9" box="[773,850,1783,1810]" captionStart-0="Figs 48–51" captionStart-1="Figs 52–55" captionStartId-0="31.[189,241,1824,1850]" captionStartId-1="33.[189,241,1717,1743]" captionTargetBox-0="[280,1308,265,2019]" captionTargetBox-1="[189,1398,265,1993]" captionTargetId-0="figure@32.[280,1308,265,2019]" captionTargetId-1="figure@34.[189,1398,265,1993]" captionTargetPageId-0="32" captionTargetPageId-1="34" captionText-0="Figs 48–51 (opposite page). Shells of species of Landouria from Java. 48. L. parahyangensis sp. nov., West Java, Karangnunggal, surroundings of Pasir Taraje Cave (holotype, MZB 20429). 49. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 50. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 51. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). Scale bar: 5 mm." captionText-1="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri-0="https://zenodo.org/record/3152259/files/figure.png" httpUri-1="https://zenodo.org/record/3152265/files/figure.png" pageId="36" pageNumber="37">51–52</figureCitation>
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;
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<tableCitation id="C69A5FE18B094E3D4D6BF90B3FD0F8CA" box="[862,989,1783,1809]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="36" pageNumber="37" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30239D18B094E3D4E88F8BC3F83F814" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="8BA76A5A8B094E3D4E88F8BC3D3BF881" blockId="36.[189,310,1856,1882]" box="[189,310,1856,1882]" pageId="36" pageNumber="37">
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<heading id="D0EFDD368B094E3D4E88F8BC3D3BF881" bold="true" box="[189,310,1856,1882]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
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<emphasis id="B96CB6488B094E3D4E88F8BC3D3BF881" bold="true" box="[189,310,1856,1882]" pageId="36" pageNumber="37">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA76A5A8B094E3D4E88F8923F83F814" blockId="36.[189,1398,1902,1999]" pageId="36" pageNumber="37">
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<taxonomicName id="4C1811D98B094E3D4E88F8923DCCF853" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,449,1902,1928]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
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<emphasis id="B96CB6488B094E3D4E88F8923DCCF853" box="[189,449,1902,1928]" italics="true" pageId="36" pageNumber="37">Landouria tholiformis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F0B338B094E3D4FFDF8933E2FF852" box="[456,546,1903,1929]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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is characterized by a brownish-corneous, sharply keeled shell with scaly processes all over the shell, with a bulge at the proximal end of the penis forming a short coecum, and a crenated flagellum, which tapers abruptly at its proximal end.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30239D18B084E3C4E88FEF73E3BFEAC" pageId="37" pageNumber="38" type="etymology">
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<paragraph id="8BA76A5A8B084E3C4E88FEF73D49FEFE" blockId="37.[189,324,267,293]" box="[189,324,267,293]" pageId="37" pageNumber="38">
|
||
<heading id="D0EFDD368B084E3C4E88FEF73D49FEFE" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B96CB6488B084E3C4E88FEF73D49FEFE" bold="true" box="[189,324,267,293]" pageId="37" pageNumber="38">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88FEC53E3BFEAC" blockId="37.[189,1399,313,375]" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C1811D98B084E3C4E88FEC53DCFFE88" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,450,313,339]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4E88FEC53DCFFE88" box="[189,450,313,339]" italics="true" pageId="37" pageNumber="38">Landouria tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4FFFFEC63E28FE8F" box="[458,549,314,340]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is named after its dome-shaped shell (Latin: ʻtholusʼ = ʻdomeʼ, ʻformaʼ = ʻshapeʼ; used as an adjective).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B084E3C4E88FE623E94FD0A" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8BA76A5A8B084E3C4E88FE623DA7FE63" blockId="37.[189,426,414,440]" box="[189,426,414,440]" pageId="37" pageNumber="38">
|
||
<heading id="D0EFDD368B084E3C4E88FE623DA7FE63" bold="true" box="[189,426,414,440]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B96CB6488B084E3C4E88FE623DA7FE63" bold="true" box="[189,426,414,440]" pageId="37" pageNumber="38">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B7060078B084E3C4EDFFE313F5AFD8B" ID-GBIF-Occurrence="2243137373" collectedFrom="agroforestry" collectingDate="2014-08-09" collectionCode="MZB" collectorName="A. S. Nurinsiyah & F. L. H. Irsyad & F. J. Sari & E. Nurlaela" country="Indonesia" county="Poktunggal Beach" elevation="137" latitude="-8.1449995" location="Gunung Kidul" longLatPrecision="21" longitude="110.63111" municipality="Tepus" pageId="37" pageNumber="38" specimenCode="17999" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BA76A5A8B084E3C4EDFFE313D51FE3C" blockId="37.[189,1398,461,593]" box="[234,348,461,487]" pageId="37" pageNumber="38">
|
||
<typeStatus id="54A3D4F88B084E3C4EDFFE313D51FE3C" box="[234,348,461,487]" pageId="37" pageNumber="38" type="holotype">
|
||
<emphasis id="B96CB6488B084E3C4EDFFE313D51FE3C" bold="true" box="[234,348,461,487]" pageId="37" pageNumber="38">Holotype</emphasis>
|
||
</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88FE0C3F5AFD8B" blockId="37.[189,1398,461,593]" pageId="37" pageNumber="38">
|
||
<collectingCountry id="F30F2ACA8B084E3C4E88FE0C3D56FDD1" box="[189,347,496,522]" name="Indonesia" pageId="37" pageNumber="38">INDONESIA</collectingCountry>
|
||
• holotype (det. anat.);
|
||
<location id="8EC73C818B084E3C4CBAFE0C3F4CFDD1" LSID="urn:lsid:plazi:treatment:03B1DB4C8B094E3C4C57F94C3D26F839:8EC73C818B084E3C4CBAFE0C3F4CFDD1" box="[655,833,496,522]" country="Indonesia" county="Poktunggal Beach" latitude="-8.1449995" longLatPrecision="21" longitude="110.63111" municipality="Tepus" name="Gunung Kidul" pageId="37" pageNumber="38">Gunung Kidul</location>
|
||
,
|
||
<collectingMunicipality id="6BC3F0208B084E3C4D66FE0C3F91FDD1" box="[851,924,496,522]" pageId="37" pageNumber="38">Tepus</collectingMunicipality>
|
||
, near
|
||
<collectingCounty id="62C612D68B084E3C4DD8FE0C38C0FDD1" box="[1005,1229,496,522]" pageId="37" pageNumber="38">Poktunggal Beach</collectingCounty>
|
||
,
|
||
<collectedFrom id="380429A78B084E3C4AEAFE0C397FFDD1" box="[1247,1394,496,522]" pageId="37" pageNumber="38">agroforestry</collectedFrom>
|
||
;
|
||
<geoCoordinate id="EE2C0C9D8B084E3C4E88FDED3D4FFDF6" box="[189,322,529,557]" degrees="8" direction="south" minutes="08" orientation="latitude" pageId="37" pageNumber="38" precision="15" seconds="42" value="-8.1449995">8°08′42″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C0C9D8B084E3C4F66FDED3DF5FDF6" box="[339,504,529,557]" degrees="110" direction="east" minutes="37" orientation="longitude" pageId="37" pageNumber="38" precision="15" seconds="52" value="110.63111">110°37′52″ E</geoCoordinate>
|
||
;
|
||
<elevation id="00358D698B084E3C4C3DFDEF3E97FDF6" box="[520,666,531,557]" metricMagnitude="2" metricUnit="m" metricValue="1.3699999999999999" pageId="37" pageNumber="38" unit="m" value="137.0">
|
||
<quantity id="4CE0C7BF8B084E3C4C3DFDEF3E5AFDF6" box="[520,599,531,557]" metricMagnitude="2" metricUnit="m" metricValue="1.3699999999999999" pageId="37" pageNumber="38" unit="m" value="137.0">137 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectingDate id="EFE2B5728B084E3C4C99FDEF3F45FDF6" box="[684,840,531,557]" pageId="37" pageNumber="38" value="2014-08-09">
|
||
<date id="FFA64C9A8B084E3C4C99FDEF3F45FDF6" box="[684,840,531,557]" pageId="37" pageNumber="38" value="2014-08-09">9 Aug. 2014</date>
|
||
</collectingDate>
|
||
;
|
||
<collectorName id="26ED0F8C8B084E3C4D62FDEF3816FDF6" box="[855,1051,531,557]" pageId="37" pageNumber="38">A.S. Nurinsiyah</collectorName>
|
||
,
|
||
<collectorName id="26ED0F8C8B084E3C4A1EFDEF38C3FDF6" box="[1067,1230,531,557]" pageId="37" pageNumber="38">F.L.H. Irsyad</collectorName>
|
||
,
|
||
<collectorName id="26ED0F8C8B084E3C4AEBFDEF394DFDF6" box="[1246,1344,531,557]" pageId="37" pageNumber="38">F.J. Sari</collectorName>
|
||
and
|
||
<collectorName id="26ED0F8C8B084E3C4E88FDCB3D4EFD8B" box="[189,323,566,593]" pageId="37" pageNumber="38">E. Nurlaela</collectorName>
|
||
leg.; D =
|
||
<quantity id="4CE0C7BF8B084E3C4F8CFDCA3E1BFD8A" box="[441,534,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="37" pageNumber="38" unit="mm" value="8.7">8.7 mm</quantity>
|
||
, H =
|
||
<quantity id="4CE0C7BF8B084E3C4C63FDCA3EBEFD8A" box="[598,691,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="37" pageNumber="38" unit="mm" value="5.2">5.2 mm</quantity>
|
||
;
|
||
<collectionCode id="ED09F29F8B084E3C4C8AFDCB3EF2FD8A" box="[703,767,567,593]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="37" pageNumber="38" type="Museum">MZB</collectionCode>
|
||
<specimenCode id="DBBEC2218B084E3C4D33FDCA3F5EFD8B" box="[774,851,566,592]" pageId="37" pageNumber="38">17999</specimenCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BA76A5A8B084E3C4EDFFD8C3E94FD0A" blockId="37.[189,1399,624,721]" pageId="37" pageNumber="38">
|
||
<materialsCitation id="3B7060078B084E3C4EDFFD8C3E98FD0A" ID-GBIF-Occurrence="2243137491" collectedFrom="agroforestry" collectionCode="MZB" country="Indonesia" county="Gunung Kidul" elevation="230" latitude="-8.102778" location="Pego Hill" longLatPrecision="21" longitude="110.630554" municipality="Tepus" pageId="37" pageNumber="38" specimenCount="8" stateProvince="Yogyakarta" typeStatus="paratype">
|
||
<typeStatus id="54A3D4F88B084E3C4EDFFD8C3D6AFD51" box="[234,359,624,650]" pageId="37" pageNumber="38" type="paratype">
|
||
<emphasis id="B96CB6488B084E3C4EDFFD8C3D6AFD51" bold="true" box="[234,359,624,650]" pageId="37" pageNumber="38">Paratypes</emphasis>
|
||
</typeStatus>
|
||
<collectingCountry id="F30F2ACA8B084E3C4E88FD6F3D56FD76" box="[189,347,659,685]" name="Indonesia" pageId="37" pageNumber="38">INDONESIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D1EA1D38B084E3C4FB1FD6F3DECFD75" box="[388,481,659,686]" count="8" pageId="37" pageNumber="38" type="generic">8 spec.</specimenCount>
|
||
;
|
||
<collectingRegion id="49DCA4B88B084E3C4FC1FD6F3E70FD76" box="[500,637,659,685]" country="Indonesia" name="Yogyakarta" pageId="37" pageNumber="38">Yogyakarta</collectingRegion>
|
||
,
|
||
<collectingCounty id="62C612D68B084E3C4CA5FD6F3F49FD76" box="[656,836,659,685]" pageId="37" pageNumber="38">Gunung Kidul</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BC3F0208B084E3C4D62FD683FADFD75" box="[855,928,660,686]" pageId="37" pageNumber="38">Tepus</collectingMunicipality>
|
||
,
|
||
<location id="8EC73C818B084E3C4D86FD683820FD76" LSID="urn:lsid:plazi:treatment:03B1DB4C8B094E3C4C57F94C3D26F839:8EC73C818B084E3C4D86FD683820FD76" box="[947,1069,659,686]" country="Indonesia" county="Gunung Kidul" latitude="-8.102778" longLatPrecision="21" longitude="110.630554" municipality="Tepus" name="Pego Hill" pageId="37" pageNumber="38" stateProvince="Yogyakarta">Pego Hill</location>
|
||
,
|
||
<collectedFrom id="380429A78B084E3C4A74FD6F38D9FD76" box="[1089,1236,659,685]" pageId="37" pageNumber="38">agroforestry</collectedFrom>
|
||
;
|
||
<geoCoordinate id="EE2C0C9D8B084E3C4ADDFD6D397CFD76" box="[1256,1393,657,685]" degrees="8" direction="south" minutes="06" orientation="latitude" pageId="37" pageNumber="38" precision="15" seconds="10" value="-8.102778">8°06′10″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C0C9D8B084E3C4E88FD493D50FD0A" box="[189,349,693,721]" degrees="110" direction="east" minutes="37" orientation="longitude" pageId="37" pageNumber="38" precision="15" seconds="50" value="110.630554">110°37′50″ E</geoCoordinate>
|
||
;
|
||
<elevation id="00358D698B084E3C4F5FFD4B3DF9FD0B" box="[362,500,694,721]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="37" pageNumber="38" unit="m" value="230.0">
|
||
<quantity id="4CE0C7BF8B084E3C4F5FFD4B3DB9FD0A" box="[362,436,695,721]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="37" pageNumber="38" unit="m" value="230.0">230 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<collectionCode id="ED09F29F8B084E3C4C34FD4B3E4CFD0A" box="[513,577,695,721]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="37" pageNumber="38" type="Museum">MZB</collectionCode>
|
||
19299
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B084E3C4E88FD0438E5FA9F" pageId="37" pageNumber="38" type="description">
|
||
<paragraph id="8BA76A5A8B084E3C4E88FD043D43FCC9" blockId="37.[189,334,760,786]" box="[189,334,760,786]" pageId="37" pageNumber="38">
|
||
<heading id="D0EFDD368B084E3C4E88FD043D43FCC9" bold="true" box="[189,334,760,786]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B96CB6488B084E3C4E88FD043D43FCC9" bold="true" box="[189,334,760,786]" pageId="37" pageNumber="38">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88FCDB3CF1FBCF" blockId="37.[189,1398,806,1044]" pageId="37" pageNumber="38">
|
||
SHELL (
|
||
<figureCitation id="132376DF8B084E3C4F23FCDA3D6BFC9A" box="[278,358,806,833]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="37" pageNumber="38">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="132376DF8B084E3C4F42FCDA3D9AFC9A" box="[375,407,806,833]" captionStart="Figs 48–51" captionStartId="31.[189,241,1824,1850]" captionTargetBox="[280,1308,265,2019]" captionTargetId="figure@32.[280,1308,265,2019]" captionTargetPageId="32" captionText="Figs 48–51 (opposite page). Shells of species of Landouria from Java. 48. L. parahyangensis sp. nov., West Java, Karangnunggal, surroundings of Pasir Taraje Cave (holotype, MZB 20429). 49. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 50. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 51. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152259/files/figure.png" pageId="37" pageNumber="38">51</figureCitation>
|
||
;
|
||
<tableCitation id="C69A5FE18B084E3C4F9DFCDA3E22FC9A" box="[424,559,806,833]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="37" pageNumber="38" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
|
||
). Depressed conical, with a dome-shaped spire, with 6–6.25 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over shell; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, wide, comprising 32–40% of shell diameter, hardly obscured by columellar edge.
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88FBC838E5FA9F" blockId="37.[189,1399,1075,1348]" pageId="37" pageNumber="38">
|
||
GENITALIA (
|
||
<figureCitation id="132376DF8B084E3C4F74FBCF3D90FB95" box="[321,413,1075,1102]" captionStart="Figs 15–24" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 15–24. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="37" pageNumber="38">Figs 20</figureCitation>
|
||
,
|
||
<figureCitation id="132376DF8B084E3C4F9CFBC83DC7FB95" box="[425,458,1076,1102]" captionStart="Figs 29–47" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="37" pageNumber="38">34</figureCitation>
|
||
;
|
||
<tableCitation id="C69A5FE18B084E3C4FE3FBCF3E3CFB95" box="[470,561,1075,1102]" captionStart="Table 4" captionStartId="22.[188,257,551,579]" captionTargetBox="[205,1929,661,1732]" captionTargetPageId="22" captionText="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="37" pageNumber="38" tableUuid="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Table 4</tableCitation>
|
||
). Atrium short; penis long, becoming slightly broader proximally, with a bulge at proximal end forming a short coecum; membranaceous sheath around distal part of penis connected with epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum moderately long, broadest in its middle part, crenated, without node, abruptly tapering at its proximal end; vas deferens narrow and long; vagina moderately long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, almost cylindrical distal part that abruptly passes into a narrower and longer proximal part that ends in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B084E3C4E88FA903E47F895" pageId="37" pageNumber="38" type="discussion">
|
||
<paragraph id="8BA76A5A8B084E3C4E88FA903D22FA5D" blockId="37.[189,303,1388,1414]" box="[189,303,1388,1414]" pageId="37" pageNumber="38">
|
||
<heading id="D0EFDD368B084E3C4E88FA903D22FA5D" bold="true" box="[189,303,1388,1414]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B96CB6488B084E3C4E88FA903D22FA5D" bold="true" box="[189,303,1388,1414]" pageId="37" pageNumber="38">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88FA663E38F999" blockId="37.[189,1399,1434,1602]" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C1811D98B084E3C4E88FA663DCCFA6F" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,449,1434,1460]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4E88FA663DCCFA6F" box="[189,449,1434,1460]" italics="true" pageId="37" pageNumber="38">Landouria tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4FFDFA663E2FFA6F" box="[456,546,1434,1460]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
clusters in the phylogeny (
|
||
<figureCitation id="132376DF8B084E3C4D68FA663FADFA6F" box="[861,928,1434,1460]" captionStart="Fig" captionStartId="7.[189,232,1718,1744]" captionTargetBox="[189,1398,270,1663]" captionTargetId="figure@7.[189,1398,270,1663]" captionTargetPageId="7" captionText="Fig. 1. Bayesian 50% majority rule consensus tree based on mitochondrial 16S rDNA sequences of species of Landouria Godwin-Austen, 1918 with Mandarina suenoae Minato, 1978 and Euhadra peliomphala (Pfeiffer, 1850) as outgroups. All species of Landouria in this tree are from Java except L. omphalostoma Páll-Gergely & Hunyadi, 2013 from China and L. timorensis Köhler et al. in press, L. montana Köhler et al. in press and Landouria sp. nov. (= L. winteriana (non Pfeiffer, 1842) in Köhler et al. (in press)) from Timor. Values at the nodes represent Bayesian posterior probabilities (left) and maximum likelihood bootstrap values (right). Numbers after taxon names correspond to MZB and ZMH catalogue numbers, AM catalogue numbers or GenBank accession numbers (see Table 1)." httpUri="https://zenodo.org/record/3152219/files/figure.png" pageId="37" pageNumber="38">Fig. 1</figureCitation>
|
||
) with
|
||
<taxonomicName id="4C1811D98B084E3C4DDBFA673885FA6F" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[1006,1160,1434,1460]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4DDBFA673885FA6F" box="[1006,1160,1434,1460]" italics="true" pageId="37" pageNumber="38">L. abdidalem</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4ABAFA6638E4FA6F" box="[1167,1257,1434,1460]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Fig.
|
||
<figureCitation id="132376DF8B084E3C4B1AFA66395DFA6E" box="[1327,1360,1434,1461]" captionStart="Figs 56–59" captionStartId="40.[189,241,1859,1885]" captionTargetBox="[189,1399,265,2004]" captionTargetId="figure@39.[189,1399,265,2004]" captionTargetPageId="39" captionText="Figs 56–59 (opposite page). Shells of species of Landouria from Java. 56. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 57. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (holotype MZB 21432). 58. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 59. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152267/files/figure.png" pageId="37" pageNumber="38">56</figureCitation>
|
||
). It differs from that species in the more depressed (D/H 1.77–1.88 vs
|
||
<quantity id="4CE0C7BF8B084E3C4DF8FA41386CFA0C" box="[973,1121,1469,1495]" metricMagnitude="-2" metricUnit="m" metricValue="5.1689" metricValueMax="5.4356" metricValueMin="4.9022" pageId="37" pageNumber="38" unit="in" value="2.035" valueMax="2.14" valueMin="1.93">1.93–2.14 in</quantity>
|
||
<taxonomicName id="4C1811D98B084E3C4A5DFA42390EFA0C" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[1128,1283,1469,1495]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4A5DFA42390EFA0C" box="[1128,1283,1469,1495]" italics="true" pageId="37" pageNumber="38">L. abdidalem</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4B3EFA423964FA03" box="[1291,1385,1470,1496]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), less sharply keeled shell with scaly processes all over the teleoconch (
|
||
<figureCitation id="132376DF8B084E3C4DC5FA1C3835FA20" box="[1008,1080,1504,1531]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="37" pageNumber="38">Fig. 7</figureCitation>
|
||
), in the flagellum, which is slightly longer than the short epiphallus, and in having a bulge at the proximal end of the penis (perhaps this is an individual abnormity).
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88F99D3E47F895" blockId="37.[189,1399,1632,1870]" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C1811D98B084E3C4E88F99D3DCCF9A0" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,449,1633,1659]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4E88F99D3DCCF9A0" box="[189,449,1633,1659]" italics="true" pageId="37" pageNumber="38">Landouria tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4FFCF99D3E29F9A0" box="[457,548,1633,1659]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
occurs sympatrically with
|
||
<taxonomicName id="4C1811D98B084E3C4D57F99D3FFBF9A1" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[866,1014,1633,1658]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="sewuensis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4D57F99D3FFBF9A1" box="[866,1014,1633,1658]" italics="true" pageId="37" pageNumber="38">L. sewuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4DCBF99D3854F9A0" box="[1022,1113,1633,1659]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Fig. 58), from which it differs in the less sharply keeled shell with a dome-shaped spire, a more distinct granular microsculpture, with scaly processes all over the shell (
|
||
<figureCitation id="132376DF8B084E3C4CACF95B3EEFF91A" box="[665,738,1703,1729]" captionStart="Figs 6–11" captionStartId="17.[189,241,1787,1813]" captionTargetBox="[189,1399,501,1756]" captionTargetId="figure@17.[189,1399,501,1756]" captionTargetPageId="17" captionText="Figs 6–11. Shell sculpture of species of Landouria from Java (umbilical side of shell in 6; apical side in 7–11). 6. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 7. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 8. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 9. L. pacitanensis sp. nov., East Java, Pacitan, Srau Beach (paratype, ZMH 148510). 10. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 11. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152225/files/figure.png" pageId="37" pageNumber="38">Fig. 7</figureCitation>
|
||
) and less distinct spiral lines at the umbilical side, the bulge at the proximal end of the penis that forms a short coecum, and a narrow, crenated flagellum. The flagellum of
|
||
<taxonomicName id="4C1811D98B084E3C4F61F9123DE5F8DC" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[340,488,1774,1799]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="sewuensis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4F61F9123DE5F8DC" box="[340,488,1774,1799]" italics="true" pageId="37" pageNumber="38">L. sewuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4FC4F9123E41F8D3" box="[497,588,1774,1800]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is broader than that of
|
||
<taxonomicName id="4C1811D98B084E3C4D6BF9123FF2F8DC" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[862,1023,1773,1799]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4D6BF9123FF2F8DC" box="[862,1023,1773,1799]" italics="true" pageId="37" pageNumber="38">L. tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4A33F912386CF8D3" box="[1030,1121,1774,1800]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and equipped with two rows of tubercles. There is a globular bulge in the distal part of the vagina in
|
||
<taxonomicName id="4C1811D98B084E3C4A03F8ED38C4F8F1" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[1078,1225,1809,1834]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="sewuensis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4A03F8ED38C4F8F1" box="[1078,1225,1809,1834]" italics="true" pageId="37" pageNumber="38">L. sewuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4AFAF8ED3924F8F0" box="[1231,1321,1809,1835]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
which is absent in
|
||
<taxonomicName id="4C1811D98B084E3C4F7DF8C93DE5F895" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[328,488,1844,1870]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4F7DF8C93DE5F895" box="[328,488,1844,1870]" italics="true" pageId="37" pageNumber="38">L. tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4FDAF8C83E47F895" box="[495,586,1844,1870]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30239D18B084E3C4E88F8893D26F839" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="8BA76A5A8B084E3C4E88F8893D5BF854" blockId="37.[189,342,1909,1935]" box="[189,342,1909,1935]" pageId="37" pageNumber="38">
|
||
<heading id="D0EFDD368B084E3C4E88F8893D5BF854" bold="true" box="[189,342,1909,1935]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B96CB6488B084E3C4E88F8893D5BF854" bold="true" box="[189,342,1909,1935]" pageId="37" pageNumber="38">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA76A5A8B084E3C4E88F8583D26F839" blockId="37.[189,1399,1956,2018]" pageId="37" pageNumber="38">
|
||
<taxonomicName id="4C1811D98B084E3C4E88F8583DCFF865" authority="Nurinsiyah & Neiber & Hausdorf, 2019" authorityName="Nurinsiyah & Neiber & Hausdorf" authorityYear="2019" box="[189,450,1956,1982]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
|
||
<emphasis id="B96CB6488B084E3C4E88F8583DCFF865" box="[189,450,1956,1982]" italics="true" pageId="37" pageNumber="38">Landouria tholiformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F0B338B084E3C4FFCF8583E29F865" box="[457,548,1956,1982]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is endemic to the Gunung Sewu karst region in Gunung Kidul Regency (Fig. 52).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |