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<document id="6C219BB85E2D06C476184ED01D778A0A" ID-CLB-Dataset="55837" ID-DOI="10.11646/zootaxa.5128.4.3" ID-GBIF-Dataset="bb39ebf4-a5cd-452a-afcc-841459b7274e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6479987" ID-ZooBank="DFF17C52-A983-4F73-84A7-CE5889D62C13" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650722170275" checkinUser="plazi" docAuthor="Yáñez-Rivera, Beatriz &amp; Salazar-Vallejo, Sergio I." docDate="2022" docId="03913D582816FF848CE7B343FC9CFE72" docLanguage="en" docName="zootaxa.5128.4.3.pdf" docOrigin="Zootaxa 5128 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chloeia flava" docType="treatment" docVersion="6" lastPageNumber="517" masterDocId="FFA84520281BFF8A8C70B759FFDFFFDF" masterDocTitle="Revision of Chloeia Savigny in Lamarck, 1818 from tropical American seas (Annelida, Amphinomidae)" masterLastPageNumber="537" masterPageNumber="503" pageNumber="516" updateTime="1699329117022" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4E31E9456B031FE01B454D14A6C82E67">Revision of Chloeia Savigny in Lamarck, 1818 from tropical American seas (Annelida, Amphinomidae)</mods:title>
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<mods:namePart id="7CC0F4306F64F205BFD897E958B245CD">Yáñez-Rivera, Beatriz</mods:namePart>
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<treatment id="03913D582816FF848CE7B343FC9CFE72" ID-DOI="http://doi.org/10.5281/zenodo.6479983" ID-GBIF-Taxon="194851607" ID-Zenodo-Dep="6479983" LSID="urn:lsid:plazi:treatment:03913D582816FF848CE7B343FC9CFE72" httpUri="http://treatment.plazi.org/id/03913D582816FF848CE7B343FC9CFE72" lastPageId="14" lastPageNumber="517" pageId="13" pageNumber="516">
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<heading id="D0CF3B222816FF878CE7B343FE37FBEB" bold="true" box="[151,488,1050,1077]" fontSize="11" level="1" pageId="13" pageNumber="516" reason="1">
<taxonomicName id="4C38F7CD2816FF878CE7B343FE37FBEB" ID-CoL="69WW9" authority="(Pallas, 1766)" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[151,488,1050,1077]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="13" pageNumber="516" phylum="Annelida" rank="species" species="flava">
<emphasis id="B94C505C2816FF878CE7B343FE37FBEB" bold="true" box="[151,488,1050,1077]" pageId="13" pageNumber="516">
<emphasis id="B94C505C2816FF878CE7B343FEE6FBEA" bold="true" box="[151,313,1050,1077]" italics="true" pageId="13" pageNumber="516">Chloeia flava</emphasis>
(
<bibRefCitation id="EFA9F1BF2816FF878D39B343FE3FFBEB" author="Pallas, P. S." box="[329,480,1050,1076]" pageId="13" pageNumber="516" refId="ref24974" refString="Pallas, P. S. (1766) Miscellanea Zoologica. Petrum van Cleef, Hagae Comitum, 224 pp., 14 pls." type="book" year="1766">Pallas, 1766</bibRefCitation>
)
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<paragraph id="8B878C4E2816FF878CE7B366FF06FB86" blockId="13.[151,488,1050,1113]" box="[151,217,1087,1113]" pageId="13" pageNumber="516">
<figureCitation id="130390CB2816FF878CE7B366FF06FB86" box="[151,217,1087,1113]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[151,1436,381,1881]" captionTargetId="figure-116@12.[151,1436,381,1881]" captionTargetPageId="12" captionText="FIGURE 6. Chloeia flava (Pallas, 1766). A. Illustration of complete specimen from Pallas (1766). B. Anterior end, dorsal view. C. Pigmentation pattern from median segments (Photo: Elizabeth Borda). D. Neurochaeta from median chaetigers. EG. Notochaetae, black arrows indicate position of spurs (Li: lips, LL: lateral (plicated basal) lobes, MA: median antenna, ML: median (plicate) lobe, Pa: palps, PA: paired antennae). Scale bars: B, 2 mm; C, 1 mm; DF, 20 μm; G, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6480001" httpUri="https://zenodo.org/record/6480001/files/figure.png" pageId="13" pageNumber="516">Fig. 6</figureCitation>
</paragraph>
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<subSubSection id="C322DFC52816FF878CE7B3DAFB46FB65" pageId="13" pageNumber="516" type="reference_group">
<paragraph id="8B878C4E2816FF878CE7B3DAFD1BFB44" blockId="13.[151,1177,1155,1210]" box="[151,708,1155,1179]" pageId="13" pageNumber="516">
<treatmentCitationGroup id="AB28AB602816FF878CE7B3DAFD1BFB44" box="[151,708,1155,1179]" pageId="13" pageNumber="516">
<treatmentCitation id="0A99AA5F2816FF878CE7B3DAFDCBFB44" author="Pallas, P. S." box="[151,532,1155,1179]" page="102" pageId="13" pageNumber="516" year="1766">
<taxonomicName id="4C38F7CD2816FF878CE7B3DAFDCBFB44" authority="Pallas, 1766: 97 - 102" authorityName="Pallas" authorityPageNumber="97 - 102" authorityYear="1766" box="[151,532,1155,1179]" class="Polychaeta" family="Aphroditidae" genus="Aphrodita" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="516" phylum="Annelida" rank="species" species="flava">
<emphasis id="B94C505C2816FF878CE7B3DAFEE9FB44" box="[151,310,1155,1179]" italics="true" pageId="13" pageNumber="516">Aphrodita flava</emphasis>
<bibRefCitation id="EFA9F1BF2816FF878D4CB3DAFDCBFB44" author="Pallas, P. S." box="[316,532,1155,1179]" pageId="13" pageNumber="516" refId="ref24974" refString="Pallas, P. S. (1766) Miscellanea Zoologica. Petrum van Cleef, Hagae Comitum, 224 pp., 14 pls." type="book" year="1766">Pallas, 1766: 97102</bibRefCitation>
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, Pl. 8,
<figureCitation id="130390CB2816FF878E29B3DAFD1FFB44" box="[601,704,1155,1179]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStartId-0="14.[151,250,1602,1626]" captionStartId-1="16.[151,250,1156,1180]" captionStartId-2="17.[151,250,1302,1326]" captionStartId-3="18.[151,250,1570,1594]" captionStartId-4="20.[151,250,1790,1814]" captionTargetBox-0="[151,1436,453,1579]" captionTargetBox-1="[152,1436,345,1133]" captionTargetBox-2="[151,1434,525,1279]" captionTargetBox-3="[151,1434,346,1547]" captionTargetBox-4="[151,1436,957,1767]" captionTargetId-0="figure-134@14.[151,1436,453,1579]" captionTargetId-1="figure-97@16.[152,1436,345,1133]" captionTargetId-2="figure-151@17.[151,1436,525,1279]" captionTargetId-3="figure-92@18.[151,1436,345,1547]" captionTargetId-4="figure-372@20.[151,1436,957,1767]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="20" captionText-0="FIGURE 7. Chloeia nuriae sp. n., holotype (EMU 13145). A. Dorsal view, recently collected. B. Anterior end, dorsal view. C. Prostomium and caruncle. D. Neurochaeta from first chaetiger. E. Neurochaeta from same chaetiger 10. F. Notochaetae from first chaetiger. G. Notochaeta from chaetiger 10. H. Pigmentation pattern from median segments. I. Posterior end with cirriform lobes. A, DI. Holotype; B, C. Paratypes (ADC: accessory cirriform branchiae, Br: bipinnate branchiae, DC: dorsal cirri, Li: lips, LL: lateral lobes, MA: median antenna, ML: median lobe, Pa: palps, PA: paired antennae). Scale bars: A, 5 mm; B, C, 1 mm; D, 20 μm; E, 15 μm; F, G, 50 μm; H, 2 mm; I, 0.5 mm." captionText-1="FIGURE 8. Chloeia paulayi sp. n. A. Holotype (UF 493), dorsal view, depressed by glass slide, after Methyl Green staining. B. Same, anterior region, dorsal view. C. Same, left lateral view. D. Same, ventral view. E. Paratype (UF 493b), anterior region, dorsal view. F. Same, close-up of anterior end. Scale bars: A, E 2.2 mm; B, F, 1.3 mm; C, 0.9 mm; D, 1.8 mm." captionText-2="FIGURE 9. Chloeia paulayi sp. n. A. Holotype (UF 493), chaetiger 1, notochaetal tips. B. Paratype (UF 493b), chaetiger 16, notochaetae. C. Holotype, chaetiger 4, neurochaetae. Scale bars: A: 45 µm, B: 90 µm, C: 30 µm." captionText-3="FIGURE 10. Chloeia pinnata Moore, 1911, holotype (USNM 16842).A,Anteriorend, dorsal view. B. Segment 3. C.Prostomium and caruncle. D. Segments 3 and 4. E. Pigmentation pattern from median segments. F. Neurochaeta from first chaetiger. G. Long neurochaeta from chaetiger 10. H. Short neurochaeta from same chaetiger. I. Notochaeta from first chaetiger, insert shows the serrated inner border. J. Notochaeta from chaetiger 10. C. Paratype, all other holotype; B, C. White arrows show branchiae (Li: lips, LL: lateral lobes, MA: median antenna, ML: median lobe, Pa: palps, PA: paired antennae). Scale bars: A, C, D, 1 mm; B, E, 0.5 mm; FH, 10 μm; I, 40 μm (inset 25 μm); J, 50 μm." captionText-4="FIGURE 11. Chloeia pseudeuglochis Augener, 1922, holotype (ZMH V-7466). A. Dorsal view. B. Oblique, dorsal view. C. Anterior end, dorsal view. D. Labels. Scale bars: A, 0.5 mm; B, 0.4 mm; C, 0.7 mm (Photos: Alexandra Kerbl)." figureDoi-0="http://doi.org/10.5281/zenodo.6480003" figureDoi-1="http://doi.org/10.5281/zenodo.6480005" figureDoi-2="http://doi.org/10.5281/zenodo.6480007" figureDoi-3="http://doi.org/10.5281/zenodo.6480009" figureDoi-4="http://doi.org/10.5281/zenodo.6480011" httpUri-0="https://zenodo.org/record/6480003/files/figure.png" httpUri-1="https://zenodo.org/record/6480005/files/figure.png" httpUri-2="https://zenodo.org/record/6480007/files/figure.png" httpUri-3="https://zenodo.org/record/6480009/files/figure.png" httpUri-4="https://zenodo.org/record/6480011/files/figure.png" pageId="13" pageNumber="516">Figs 711</figureCitation>
.
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</paragraph>
<paragraph id="8B878C4E2816FF878CE7B3FBFB46FB65" blockId="13.[151,1177,1155,1210]" box="[151,1177,1186,1210]" pageId="13" pageNumber="516">
<treatmentCitationGroup id="AB28AB602816FF878CE7B3FBFB46FB65" box="[151,1177,1186,1210]" pageId="13" pageNumber="516">
<taxonomicName id="4C38F7CD2816FF878CE7B3FBFEFEFB65" ID-CoL="69WW9" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[151,289,1186,1210]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="13" pageNumber="516" phylum="Annelida" rank="species" species="flava">
<emphasis id="B94C505C2816FF878CE7B3FBFEFEFB65" box="[151,289,1186,1210]" italics="true" pageId="13" pageNumber="516">Chloeia flava</emphasis>
</taxonomicName>
.
<treatmentCitation id="0A99AA5F2816FF878D4AB3FBFE26FB65" author="Horst, R." box="[314,505,1186,1210]" page="18" pageId="13" pageNumber="516" year="1912">
<bibRefCitation id="EFA9F1BF2816FF878D4AB3FBFE26FB65" author="Horst, R." box="[314,505,1186,1210]" pageId="13" pageNumber="516" pagination="1 - 43" refId="ref23777" refString="Horst, R. (1912) Polychaeta Errantia of the Siboga Expedition, Part 1. Amphinomidae. Siboga-Expeditie Leyden, 24 a, 1 - 43." type="book chapter" year="1912">Horst 1912: 1819</bibRefCitation>
</treatmentCitation>
, Pl. 7,
<figureCitation id="130390CB2816FF878E30B3FBFDA2FB65" box="[576,637,1186,1210]" captionStart="FIGURE 2" captionStartId="4.[151,250,1526,1550]" captionTargetBox="[152,1436,347,1499]" captionTargetId="figure-108@4.[151,1436,345,1502]" captionTargetPageId="4" captionText="FIGURE 2. Chloeia pinnata Moore, 1911, non-type specimens (LACM 992-39), epitokes, anterior end after cropping chaetae of right parapodia of chaetigers 13, after Methyl Green staining for caruncle.A. Smallest specimen (11 mm long), dorsal view. B. Same, lateral view. C. Same, after Metyhl Green staining. D. Medium-sized specimen (18 mm long), dorsal view. E. Same, lateral view. F. Same, after Methyl Green staining. G. Largest specimen (23 mm long), dorsal view. H. Same, lateral view. I. Same, after Methyl green. Scale bars: A, 0.25 mm; B, 0.33 mm; C, 0.26 mm; D, G, 0.29; E, 0.44 mm; F, 0.35 mm." figureDoi="http://doi.org/10.5281/zenodo.6479993" httpUri="https://zenodo.org/record/6479993/files/figure.png" pageId="13" pageNumber="516">Fig. 2</figureCitation>
;
<treatmentCitation id="0A99AA5F2816FF878EF8B3FBFCF0FB65" author="Fauvel, P." box="[648,815,1186,1210]" page="96" pageId="13" pageNumber="516" year="1953">
<bibRefCitation id="EFA9F1BF2816FF878EF8B3FBFCF0FB65" author="Fauvel, P." box="[648,815,1186,1210]" pageId="13" pageNumber="516" refId="ref23035" refString="Fauvel, P. (1953) The Fauna of India including Pakistan, Ceylon, Burma and Malaya. Annelida Polychaeta. Indian Press, Allahabad, 507 pp." type="book" year="1953">Fauvel 1953: 96</bibRefCitation>
</treatmentCitation>
, Fig. 46a;
<treatmentCitation id="0A99AA5F2816FF878FEAB3FBFBEDFB65" author="Day, J. H." box="[922,1074,1186,1210]" page="124" pageId="13" pageNumber="516" year="1967">
<bibRefCitation id="EFA9F1BF2816FF878FEAB3FBFBEDFB65" author="Day, J. H." box="[922,1074,1186,1210]" pageId="13" pageNumber="516" pagination="1 - 878" refId="ref22273" refString="Day, J. H. (1967) A Monograph on the Polychaeta of Southern Africa. British Museum (Natural History) Publications, 656, 1 - 878. https: // doi. org / 10.5962 / bhl. title. 8596" type="journal article" year="1967">Day 1967: 124</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="130390CB2816FF87884DB3FBFB49FB65" box="[1085,1174,1186,1210]" captionStart="FIGURE 3" captionStartId="7.[151,250,1747,1771]" captionTargetBox="[151,1436,184,1724]" captionTargetId="figure-17@7.[151,1436,181,1724]" captionTargetPageId="7" captionText="FIGURE 3. Chloeia entypa Chamberlin, 1919. A. Holotype (USNM 19346), ventral view; B. Prostomium and caruncle (Li: lips; MA: median antenna; ML: medial lobe; Pa: palps; PA: paired antenna). C. Neurochaeta from first chaetiger. D. Neurochaeta from chaetiger 10. E. Notochaeta from first chaetiger. F. Notochaeta from chaetiger 10. G. Holotype and original label (photo: Elizabeth Borda). Scale bars: A, 2 mm; B, 0.3 mm; C, D, 5 μm; E, 2 μm; F, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6479995" httpUri="https://zenodo.org/record/6479995/files/figure.png" pageId="13" pageNumber="516">Fig. 3.1r</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C322DFC52816FF878CE7B3BAFB67FA9A" pageId="13" pageNumber="516" type="materials_examined">
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<emphasis id="B94C505C2816FF878CE7B3BAFEE1FB23" bold="true" box="[151,318,1250,1277]" pageId="13" pageNumber="516">
<typeStatus id="548332EC2816FF878CE7B3BAFF0DFB22" box="[151,210,1251,1277]" pageId="13" pageNumber="516">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548332EC2816FF878D3BB3BAFE5CFB22" box="[331,387,1251,1277]" pageId="13" pageNumber="516">Type</typeStatus>
specimens were not deposited.
</paragraph>
<paragraph id="8B878C4E2816FF878CB7B25FFB67FA9A" blockId="13.[151,1437,1250,2033]" pageId="13" pageNumber="516">
<emphasis id="B94C505C2816FF878CB7B25FFE64FAFF" bold="true" box="[199,443,1286,1312]" pageId="13" pageNumber="516">Additional material.</emphasis>
[
<materialsCitation id="3B5086132816FF878DB9B25EFB00FAFE" ID-GBIF-Occurrence="3757330359" box="[457,1247,1286,1313]" collectingDate="1970-07-12" collectionCode="USNM" collectorName="D. K. Massey" country="India" location="India" pageId="13" pageNumber="516" specimenCode="USNM 46277" specimenCount="1">
<specimenCode id="DB9E24352816FF878DB9B25EFDB2FAFE" box="[457,621,1286,1313]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="516" type="Museum">USNM 46277</specimenCode>
] Kottayam,
<collectingCountry id="F32FCCDE2816FF878E88B25EFCEBFAFE" box="[760,820,1287,1313]" name="India" pageId="13" pageNumber="516">India</collectingCountry>
,
<date id="FF86AA8E2816FF878F31B25EFC10FAFE" box="[833,975,1287,1313]" pageId="13" pageNumber="516" value="1970-07-12">
<collectingDate id="EFC253662816FF878F31B25EFC10FAFE" box="[833,975,1287,1313]" pageId="13" pageNumber="516" value="1970-07-12">12 Jul. 1970</collectingDate>
</date>
,
<collectorName id="26CDE9982816FF878FABB25EFBA8FAFE" box="[987,1143,1287,1313]" pageId="13" pageNumber="516">D. K. Massey</collectorName>
, coll. (5)
</materialsCitation>
.
<materialsCitation id="3B5086132816FF87889BB25EFB56FA9A" ID-GBIF-Occurrence="3757330312" collectingDate="1908-02-25" collectionCode="USMN" collectorName="Tumindao Island, R &amp; V Albatross" country="Philippines" location="Tumindao Island" pageId="13" pageNumber="516" specimenCode="USMN 17480" specimenCount="1" stateProvince="Sulu">
[
<specimenCode id="DB9E24352816FF878881B25EFA49FAFE" box="[1265,1430,1286,1313]" collectionCode="USMN" pageId="13" pageNumber="516">USMN 17480</specimenCode>
]
<collectingRegion id="49FC42AC2816FF878CE7B272FF15FA9A" box="[151,202,1323,1349]" country="Philippines" name="Sulu" pageId="13" pageNumber="516">Sulu</collectingRegion>
Archipielago,
<collectorName id="26CDE9982816FF878D00B272FD92FA9A" box="[368,589,1323,1349]" pageId="13" pageNumber="516">
<location id="8EE7DA952816FF878D00B272FDEFFA9A" LSID="urn:lsid:plazi:treatment:03913D582816FF848CE7B343FC9CFE72:8EE7DA952816FF878D00B272FDEFFA9A" box="[368,560,1323,1349]" country="Philippines" name="Tumindao Island" pageId="13" pageNumber="516" stateProvince="Sulu">Tumindao Island</location>
, R
</collectorName>
/
<collectorName id="26CDE9982816FF878E26B272FD02FA9A" box="[598,733,1323,1349]" pageId="13" pageNumber="516">V Albatross</collectorName>
,
<collectingCountry id="F32FCCDE2816FF878E98B272FCB9FA9A" box="[744,870,1323,1349]" name="Philippines" pageId="13" pageNumber="516">Philippines</collectingCountry>
Expedition,
<date id="FF86AA8E2816FF878F86B273FB56FA9A" box="[1014,1161,1322,1349]" pageId="13" pageNumber="516" value="1908-02-25">
<collectingDate id="EFC253662816FF878F86B273FB56FA9A" box="[1014,1161,1322,1349]" pageId="13" pageNumber="516" value="1908-02-25">25 Feb. 1908</collectingDate>
</date>
</materialsCitation>
(2).
</paragraph>
</subSubSection>
<subSubSection id="C322DFC52816FF878CB7B217FA5FFA52" pageId="13" pageNumber="516" type="diagnosis">
<paragraph id="8B878C4E2816FF878CB7B217FA5FFA52" blockId="13.[151,1437,1250,2033]" pageId="13" pageNumber="516">
<emphasis id="B94C505C2816FF878CB7B217FE9DFAB7" bold="true" box="[199,322,1358,1384]" pageId="13" pageNumber="516">Diagnosis.</emphasis>
<taxonomicName id="4C38F7CD2816FF878D3BB216FE7BFAB6" authorityName="Savigny" authorityYear="1818" box="[331,420,1359,1385]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="13" pageNumber="516" phylum="Annelida" rank="genus">
<emphasis id="B94C505C2816FF878D3BB216FE7BFAB6" box="[331,420,1359,1385]" italics="true" pageId="13" pageNumber="516">Chloeia</emphasis>
</taxonomicName>
with ventral cirri of similar size throughout body; bipinnate branchiae from chaetiger 4; dorsum with a round, blackish spot surrounded by pale areas per segment; chaetal bundles homogeneously pale.
</paragraph>
</subSubSection>
<subSubSection id="C322DFC52816FF878CB7B2CFFD60F875" pageId="13" pageNumber="516" type="description">
<paragraph id="8B878C4E2816FF878CB7B2CFFC0EF9C2" blockId="13.[151,1437,1250,2033]" pageId="13" pageNumber="516">
<emphasis id="B94C505C2816FF878CB7B2CFFE86FA6F" bold="true" box="[199,345,1430,1456]" pageId="13" pageNumber="516">Description.</emphasis>
Best preserved specimen (USNM 46277) complete,
<quantity id="4CC021AB2816FF878FECB2CFFC35FA6E" box="[924,1002,1430,1457]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="516" unit="mm" value="85.0">85 mm</quantity>
long,
<quantity id="4CC021AB2816FF87885CB2CFFBA6FA6E" box="[1068,1145,1430,1457]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="516" unit="mm" value="15.0">15 mm</quantity>
wide, 38 chaetigers. Body fusiform; dorsal pigmentation pattern consists of a central dark brown circle or round spot per segment positioned towards posterior border of each segment, and brown pigmentation around the lateral side under branchiae and notochaetal lobes of prededent segments (
<figureCitation id="130390CB2816FF878E1DB15AFD38F9C2" box="[621,743,1539,1565]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[151,1436,381,1881]" captionTargetId="figure-116@12.[151,1436,381,1881]" captionTargetPageId="12" captionText="FIGURE 6. Chloeia flava (Pallas, 1766). A. Illustration of complete specimen from Pallas (1766). B. Anterior end, dorsal view. C. Pigmentation pattern from median segments (Photo: Elizabeth Borda). D. Neurochaeta from median chaetigers. EG. Notochaetae, black arrows indicate position of spurs (Li: lips, LL: lateral (plicated basal) lobes, MA: median antenna, ML: median (plicate) lobe, Pa: palps, PA: paired antennae). Scale bars: B, 2 mm; C, 1 mm; DF, 20 μm; G, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6480001" httpUri="https://zenodo.org/record/6480001/files/figure.png" pageId="13" pageNumber="516">Fig. 6A, C</figureCitation>
). Chaetae yellowish.
</paragraph>
<paragraph id="8B878C4E2816FF878CB7B17EFE88F90E" blockId="13.[151,1437,1250,2033]" pageId="13" pageNumber="516">
Prostomium semicircular with four black eyes, anterior eyes 23× larger than posterior ones (
<figureCitation id="130390CB2816FF8788ABB17EFAECF99D" box="[1243,1331,1575,1602]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[151,1436,381,1881]" captionTargetId="figure-116@12.[151,1436,381,1881]" captionTargetPageId="12" captionText="FIGURE 6. Chloeia flava (Pallas, 1766). A. Illustration of complete specimen from Pallas (1766). B. Anterior end, dorsal view. C. Pigmentation pattern from median segments (Photo: Elizabeth Borda). D. Neurochaeta from median chaetigers. EG. Notochaetae, black arrows indicate position of spurs (Li: lips, LL: lateral (plicated basal) lobes, MA: median antenna, ML: median (plicate) lobe, Pa: palps, PA: paired antennae). Scale bars: B, 2 mm; C, 1 mm; DF, 20 μm; G, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6480001" httpUri="https://zenodo.org/record/6480001/files/figure.png" pageId="13" pageNumber="516">Fig. 6B</figureCitation>
). Median antenna arising from anterior caruncle region, longer than lateral antennae and palps. Caruncle with a V-shaped posterior margin (
<quantity id="4CC021AB2816FF878D12B137FE64F956" box="[354,443,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="13" pageNumber="516" unit="mm" value="5.5">5.5 mm</quantity>
long); median lobe with about 30 folds. Lateral basal lobes spread, each with a similar number of folds, without pigmentation. Mouth ventral on chaetiger 3. Bipinnate branchiae from chaetiger 4, present throughout body.
</paragraph>
<paragraph id="8B878C4E2816FF878CB7B182FC71F8E2" blockId="13.[151,1437,1250,2033]" pageId="13" pageNumber="516">Parapodia biramous. Notopodia with cirriform branchiae along chaetigers 14, progressively decreasing in size, subsequent chaetigers with branchiae bipinnate. Cirriform branchiae present in chaetiger 4 smaller than preceding ones. Neuropodia with second ventral cirri as long as contiguous ones.</paragraph>
<paragraph id="8B878C4E2816FF878CB7B01EFD60F875" blockId="13.[151,1437,1250,2033]" pageId="13" pageNumber="516">
Neurochaetae furcates; relative length of tines 1:31:4 (
<figureCitation id="130390CB2816FF878F36B01EFC7DF8BD" box="[838,930,1863,1890]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[151,1436,381,1881]" captionTargetId="figure-116@12.[151,1436,381,1881]" captionTargetPageId="12" captionText="FIGURE 6. Chloeia flava (Pallas, 1766). A. Illustration of complete specimen from Pallas (1766). B. Anterior end, dorsal view. C. Pigmentation pattern from median segments (Photo: Elizabeth Borda). D. Neurochaeta from median chaetigers. EG. Notochaetae, black arrows indicate position of spurs (Li: lips, LL: lateral (plicated basal) lobes, MA: median antenna, ML: median (plicate) lobe, Pa: palps, PA: paired antennae). Scale bars: B, 2 mm; C, 1 mm; DF, 20 μm; G, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6480001" httpUri="https://zenodo.org/record/6480001/files/figure.png" pageId="13" pageNumber="516">Fig. 6D</figureCitation>
). Notochaetae with serrated external border, short tine very short, like a spur (
<figureCitation id="130390CB2816FF878D8AB032FDACF859" box="[506,627,1899,1926]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[151,1436,381,1881]" captionTargetId="figure-116@12.[151,1436,381,1881]" captionTargetPageId="12" captionText="FIGURE 6. Chloeia flava (Pallas, 1766). A. Illustration of complete specimen from Pallas (1766). B. Anterior end, dorsal view. C. Pigmentation pattern from median segments (Photo: Elizabeth Borda). D. Neurochaeta from median chaetigers. EG. Notochaetae, black arrows indicate position of spurs (Li: lips, LL: lateral (plicated basal) lobes, MA: median antenna, ML: median (plicate) lobe, Pa: palps, PA: paired antennae). Scale bars: B, 2 mm; C, 1 mm; DF, 20 μm; G, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6480001" httpUri="https://zenodo.org/record/6480001/files/figure.png" pageId="13" pageNumber="516">Fig. 6EG</figureCitation>
). Serrated external border consists of few teeth (38) or border completely serrated with more than 20 denticles (
<figureCitation id="130390CB2816FF878E4DB0D6FD6CF875" box="[573,691,1935,1962]" captionStart="FIGURE 6" captionStartId="12.[151,250,1904,1928]" captionTargetBox="[151,1436,381,1881]" captionTargetId="figure-116@12.[151,1436,381,1881]" captionTargetPageId="12" captionText="FIGURE 6. Chloeia flava (Pallas, 1766). A. Illustration of complete specimen from Pallas (1766). B. Anterior end, dorsal view. C. Pigmentation pattern from median segments (Photo: Elizabeth Borda). D. Neurochaeta from median chaetigers. EG. Notochaetae, black arrows indicate position of spurs (Li: lips, LL: lateral (plicated basal) lobes, MA: median antenna, ML: median (plicate) lobe, Pa: palps, PA: paired antennae). Scale bars: B, 2 mm; C, 1 mm; DF, 20 μm; G, 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6480001" httpUri="https://zenodo.org/record/6480001/files/figure.png" pageId="13" pageNumber="516">Fig. 6EF</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C322DFC52816FF848CB7B0EAFD1BFE55" lastPageId="14" lastPageNumber="517" pageId="13" pageNumber="516" type="discussion">
<paragraph id="8B878C4E2816FF848CB7B0EAFD1BFE55" blockId="13.[151,1437,1250,2033]" lastBlockId="14.[151,1437,150,429]" lastPageId="14" lastPageNumber="517" pageId="13" pageNumber="516">
<emphasis id="B94C505C2816FF878CB7B0EAFEE4F812" bold="true" box="[199,315,1971,1997]" pageId="13" pageNumber="516">Remarks.</emphasis>
The original description by
<bibRefCitation id="EFA9F1BF2816FF878EF1B0EAFCC8F811" author="Pallas, P. S." box="[641,791,1971,1998]" pageId="13" pageNumber="516" refId="ref24974" refString="Pallas, P. S. (1766) Miscellanea Zoologica. Petrum van Cleef, Hagae Comitum, 224 pp., 14 pls." type="book" year="1766">Pallas (1766)</bibRefCitation>
is adequate with good illustrations, but does not include chaetal features, probably due to lack of microscopes without spheric aberrations. However, chaetal features were illustrated later by
<bibRefCitation id="EFA9F1BF2815FF848D04B7CEFDEAFF6E" author="M'Intosh, W. C." box="[372,565,150,177]" pageId="14" pageNumber="517" pagination="1 - 554" refId="ref24351" refString="M'Intosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 76. Challenger Reports, 12, 1 - 554." type="journal article" year="1885">MIntosh (1885)</bibRefCitation>
. Thus, the delimitation of this species is rather settled, and several authors provided complete characterizations (
<bibRefCitation id="EFA9F1BF2815FF848E38B7E2FD11FF0A" author="Horst, R." box="[584,718,187,213]" pageId="14" pageNumber="517" pagination="1 - 43" refId="ref23777" refString="Horst, R. (1912) Polychaeta Errantia of the Siboga Expedition, Part 1. Amphinomidae. Siboga-Expeditie Leyden, 24 a, 1 - 43." type="book chapter" year="1912">Horst 1912</bibRefCitation>
;
<bibRefCitation id="EFA9F1BF2815FF848EAEB7E2FCAFFF0A" author="Fauvel, P." box="[734,880,186,213]" pageId="14" pageNumber="517" refId="ref23035" refString="Fauvel, P. (1953) The Fauna of India including Pakistan, Ceylon, Burma and Malaya. Annelida Polychaeta. Indian Press, Allahabad, 507 pp." type="book" year="1953">Fauvel 1953</bibRefCitation>
;
<bibRefCitation id="EFA9F1BF2815FF848FF0B7E2FC29FF09" author="Day, J. H." box="[896,1014,187,214]" pageId="14" pageNumber="517" pagination="1 - 878" refId="ref22273" refString="Day, J. H. (1967) A Monograph on the Polychaeta of Southern Africa. British Museum (Natural History) Publications, 656, 1 - 878. https: // doi. org / 10.5962 / bhl. title. 8596" type="journal article" year="1967">Day 1967</bibRefCitation>
). However, the record by
<bibRefCitation id="EFA9F1BF2815FF84895CB7E2FF3DFF26" author="Treadwell, A. L." pageId="14" pageNumber="517" pagination="1 - 11" refId="ref25681" refString="Treadwell, A. L. (1923) Polychaetous annelids from lower California with descriptions of new species. American Museum Novitates, 74, 1 - 11." type="journal article" year="1923">Treadwell (1923)</bibRefCitation>
from San Francisquito Bay,
<collectingRegion id="49FC42AC2815FF848E47B786FD35FF26" box="[567,746,223,249]" country="Mexico" name="Baja California" pageId="14" pageNumber="517">Baja California</collectingRegion>
, in the Gulf of
<collectingRegion id="49FC42AC2815FF848FD8B786FBC3FF26" box="[936,1052,223,249]" country="United States of America" name="California" pageId="14" pageNumber="517">California</collectingRegion>
, is questionable because he only provided species name, without any comment about its appearance. Consequently, some illustrations of this species are herein provided to facilitate its identification and to avoid future misidentifications in Eastern Pacific waters. We think Treadwell might have found
<taxonomicName id="4C38F7CD2815FF848E32B612FC63FEBA" authority="Augener, 1922" authorityName="Augener" authorityYear="1922" box="[578,956,331,357]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="14" pageNumber="517" phylum="Annelida" rank="species" species="pseudeuglochis">
<emphasis id="B94C505C2815FF848E32B612FD8AFEBA" box="[578,597,331,357]" italics="true" pageId="14" pageNumber="517">C</emphasis>
.
<emphasis id="B94C505C2815FF848E12B612FCD0FEBA" box="[610,783,331,357]" italics="true" pageId="14" pageNumber="517">pseudeuglochis</emphasis>
<bibRefCitation id="EFA9F1BF2815FF848F65B612FC63FEBA" author="Augener, H." box="[789,956,331,357]" pageId="14" pageNumber="517" pagination="38 - 53" refId="ref21238" refString="Augener, H. (1922) Ueber litorale Polychaeten von Westindien. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1922 (3 - 5), 38 - 53." type="journal article" year="1922">Augener, 1922</bibRefCitation>
</taxonomicName>
, which was described some time before his contribution was sent for publication (see below).
</paragraph>
</subSubSection>
<subSubSection id="C322DFC52815FF848CB7B6CBFC9CFE72" box="[199,835,402,429]" pageId="14" pageNumber="517" type="distribution">
<paragraph id="8B878C4E2815FF848CB7B6CBFC9CFE72" blockId="14.[151,1437,150,429]" box="[199,835,402,429]" pageId="14" pageNumber="517">
<emphasis id="B94C505C2815FF848CB7B6CBFEBEFE73" bold="true" box="[199,353,402,428]" pageId="14" pageNumber="517">Distribution.</emphasis>
Indian Ocean, in shallow water sediments.
</paragraph>
</subSubSection>
</treatment>
</document>