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<document id="BF907D9E7E5A9163BBF29DFD9F512C70" ID-DOI="10.5281/zenodo.179477" ID-GBIF-Dataset="3ff8b316-c411-4d6b-af4c-e7bb7a663759" ID-ISSN="1175-5326" ID-Zenodo-Dep="179477" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459973316064" checkinUser="plazi" docAuthor="Wasmund, Anne M. &amp; Holzenthal, Ralph W." docDate="2007" docId="039087E8FFEFF61FFF09F8AC655D11EA" docLanguage="en" docName="zt01634p059.pdf" docOrigin="Zootaxa 1634" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Rhyacopsyche Muller" docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="FFA9FF90FFEBF61AFF9EFFC36707123E" masterDocTitle="A revision of the Neotropical caddisfly genus Rhyacopsyche, with the description of 13 new species (Trichoptera: Hydroptilidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="5" updateTime="1698231588944" updateUser="plazi">
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<mods:title id="87CB444D87C80B98AD5B9B1F284F3759">A revision of the Neotropical caddisfly genus Rhyacopsyche, with the description of 13 new species (Trichoptera: Hydroptilidae)</mods:title>
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<mods:namePart id="1DFCB2EA62270CC530F08A7A4833CA68">Wasmund, Anne M.</mods:namePart>
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<subSubSection id="C3236575FFEFF61EFF09F8AC66A615B7" box="[151,417,1903,1929]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8B8636FEFFEFF61EFF09F8AC66A615B7" blockId="4.[151,417,1903,1929]" box="[151,417,1903,1929]" pageId="4" pageNumber="5">
<heading id="D0CE8192FFEFF61EFF09F8AC66A615B7" bold="true" box="[151,417,1903,1929]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C394D7DFFEFF61EFF09F8AC66A615B7" ID-CoL="642FF" authority="Muller" authorityName="Muller" box="[151,417,1903,1929]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAECFFEFF61EFF09F8AC66A615B7" bold="true" box="[151,417,1903,1929]" pageId="4" pageNumber="5">
<emphasis id="B94DEAECFFEFF61EFF09F8AC664315B7" bold="true" box="[151,324,1903,1929]" italics="true" pageId="4" pageNumber="5">Rhyacopsyche</emphasis>
Müller
</emphasis>
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<paragraph id="8B8636FEFFEFF61EFF09F87664D015D2" blockId="4.[151,1436,1973,2028]" pageId="4" pageNumber="5">
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<taxonomicName id="4C394D7DFFEFF61EFF09F87666FC15F2" ID-CoL="642FF" authority="Muller, 1879a:40" authorityName="Muller" authorityPageNumber="40" authorityYear="1879" box="[151,507,1973,1996]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAECFFEFF61EFF09F876663715F2" box="[151,304,1973,1996]" italics="true" pageId="4" pageNumber="5">Rhyacopsyche</emphasis>
<treatmentCitation id="0A9810EFFFEFF61EFEA2F87666FC15F2" author="Muller" box="[316,507,1973,1996]" page="40" pageId="4" pageNumber="5" year="1879">
<bibRefCitation id="EFA84B0FFFEFF61EFEA2F87666DC15F2" author="Muller" box="[316,475,1973,1996]" pageId="4" pageNumber="28" refString="Muller, F. (1879 a) Uber Phryganiden. Zoologischer Anzeiger, 2, 38 - 40, 180 - 182, 283 - 284, 405 - 407." type="journal article" year="1879" yearSuffix="a">Müller, 1879a</bibRefCitation>
:40
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[nomen nudum]; 1879b:143 [
<typeStatus id="5482885CFFEFF61EFCCDF876648C15F2" box="[851,907,1973,1996]" pageId="4" pageNumber="5">Type</typeStatus>
species:
<taxonomicName id="4C394D7DFFEFF61EFC64F876628D15F2" ID-CoL="4SP92" authority="Muller 1879" authorityName="Muller" authorityYear="1879" box="[1018,1418,1973,1996]" class="Insecta" family="Hydroptilidae" genus="Rhyacopsyche" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hagenii">
<emphasis id="B94DEAECFFEFF61EFC64F87663F715F2" box="[1018,1264,1973,1996]" italics="true" pageId="4" pageNumber="5">Rhyacopsyche hagenii</emphasis>
<bibRefCitation id="EFA84B0FFFEFF61EFB63F876628D15F2" author="Muller" box="[1277,1418,1973,1996]" pageId="4" pageNumber="28" refString="Muller, F. (1879 b) Notes on the cases of some south Brazilian Trichoptera. Transactions of the Entomological Society of London, 1879, 131 - 144." type="journal article" year="1879" yearSuffix="b">Müller 1879</bibRefCitation>
</taxonomicName>
b, monobasic]; 1880a:121; 1880b:72. —
<treatmentCitation id="0A9810EFFFEFF61EFDC1F81665FD15D2" author="Flint" box="[607,762,2005,2028]" page="516" pageId="4" pageNumber="5" year="1971">
<bibRefCitation id="EFA84B0FFFEFF61EFDC1F81665CA15D2" author="Flint" box="[607,717,2005,2028]" pageId="4" pageNumber="27" refString="Flint, O. S., Jr. (1971) Studies of Neotropical caddisflies, XI: the genus Rhyacopsyche in Central America (Hydroptilidae). Proceedings of the Biological Society of Washington, 83, 515 - 526." type="journal article" year="1971">Flint,1971</bibRefCitation>
:516
</treatmentCitation>
[definition, review].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8636FEFFEEF61FFF09FF5467D9136A" blockId="5.[151,1436,151,980]" pageId="5" pageNumber="6">
<emphasis id="B94DEAECFFEEF61FFF09FF54669D128F" bold="true" box="[151,410,151,177]" pageId="5" pageNumber="6">Generic description.</emphasis>
Adult. Forewing length 2.0–
<quantity id="4CC19B1BFFEEF61FFC9CFF5B6463128C" box="[770,868,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="6" unit="mm" value="4.9">4.9 mm</quantity>
; wings narrow, acuminate, mottled with light, medium, and/or dark brown hairs; forewing veins R1-R3, M1 and M2 thickened at apex, not reaching wing margin (
<figureCitation id="13022A7BFFEEF61FFF4EFF29661C133A" box="[208,283,234,260]" captionStart="FIGURE 3" captionStartId="30.[151,255,1062,1086]" captionTargetBox="[246,1340,233,987]" captionTargetId="figure@30.[246,1340,229,1004]" captionTargetPageId="30" captionText="FIGURE 3. Rhyacopsyche andina Flint. Wings: A—forewing; B—hind wing." httpUri="https://zenodo.org/record/179480/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
); hind wing with long fringe of hairs along ventral margin. Unmodified head and antennae; 3 ocelli. Mesoscutellum with transverse suture; metascutellum usually pentagonal in shape (
<figureCitation id="13022A7BFFEEF61FFB2BFED163FB1312" box="[1205,1276,274,300]" captionStart="FIGURE 2" captionStartId="29.[151,255,1376,1400]" captionTargetBox="[151,1435,214,1352]" captionTargetId="figure@29.[151,1436,156,1353]" captionTargetPageId="29" captionText="FIGURE 2. Rhyacopsyche hagenii Müller. Adult: head and thorax, dorsal." httpUri="https://zenodo.org/record/179479/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
). Tibial spurs 1-3-4.
</paragraph>
</subSubSection>
<subSubSection id="C3236575FFEEF61FFF58FEA1655D11EA" pageId="5" pageNumber="6" type="description">
<paragraph id="8B8636FEFFEEF61FFF58FEA16431107A" blockId="5.[151,1436,151,980]" pageId="5" pageNumber="6">
Male genitalia. Segment IX produced into dorsolateral lobes, often bearing 1 or more dark, peglike setae (
<figureCitation id="13022A7BFFEEF61FFF01FE4967E0139A" box="[159,231,394,420]" captionStart="FIGURE 4" captionStartId="31.[151,255,1819,1843]" captionTargetBox="[218,1368,194,1744]" captionTargetId="figure@31.[218,1368,194,1761]" captionTargetPageId="31" captionText="FIGURE 4. Rhyacopsyche hagenii Müller. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal; Eplumose seta." httpUri="https://zenodo.org/record/179481/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
A). Tergum X weakly developed and often retracted partially (
<figureCitation id="13022A7BFFEEF61FFC53FE496314139A" box="[973,1043,394,420]" captionStart="FIGURE 9" captionStartId="36.[151,255,1732,1756]" captionTargetBox="[237,1353,186,1678]" captionTargetId="figure@36.[237,1355,186,1695]" captionTargetPageId="36" captionText="FIGURE 9. Rhyacopsyche chichotla Bueno &amp; Hamilton. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal." httpUri="https://zenodo.org/record/179486/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
A) or completely within segment IX (
<figureCitation id="13022A7BFFEEF61FFF58FE71660A13F2" box="[198,269,434,460]" captionStart="FIGURE 4" captionStartId="31.[151,255,1819,1843]" captionTargetBox="[218,1368,194,1744]" captionTargetId="figure@31.[218,1368,194,1761]" captionTargetPageId="31" captionText="FIGURE 4. Rhyacopsyche hagenii Müller. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal; Eplumose seta." httpUri="https://zenodo.org/record/179481/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
A). Inferior appendage large and distinct, elongate (
<figureCitation id="13022A7BFFEEF61FFCF2FE7164B513F2" box="[876,946,434,460]" captionStart="FIGURE 7" captionStartId="34.[151,255,1831,1855]" captionTargetBox="[220,1363,194,1755]" captionTargetId="figure@34.[220,1367,194,1791]" captionTargetPageId="34" captionText="FIGURE 7. Rhyacopsyche benwa, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal." httpUri="https://zenodo.org/record/179484/files/figure.png" pageId="5" pageNumber="6">Fig. 7</figureCitation>
A) or tall (
<figureCitation id="13022A7BFFEEF61FFBB1FE71638313F2" box="[1071,1156,434,460]" captionStart="FIGURE 12" captionStartId="39.[151,255,1946,1970]" captionText="FIGURE 12. Rhyacopsyche colubrinosa, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal." pageId="5" pageNumber="6">Fig. 12</figureCitation>
A), usually bearing 1 or more large modified setae. Intermediate appendage present, often reduced (
<figureCitation id="13022A7BFFEEF61FFB8FFE19636413CA" box="[1041,1123,474,500]" captionStart="FIGURE 9" captionStartId="36.[151,255,1732,1756]" captionTargetBox="[237,1353,186,1678]" captionTargetId="figure@36.[237,1355,186,1695]" captionTargetPageId="36" captionText="FIGURE 9. Rhyacopsyche chichotla Bueno &amp; Hamilton. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal." httpUri="https://zenodo.org/record/179486/files/figure.png" pageId="5" pageNumber="6">Figs. 9</figureCitation>
A, 9C). Phallus with tubular, frequently inflated basal half, apical half with central tubule and lateral process; often surrounded with a thin membranous sheath covered with spicules (
<figureCitation id="13022A7BFFEEF61FFD54FDE96416107A" box="[714,785,554,580]" captionStart="FIGURE 4" captionStartId="31.[151,255,1819,1843]" captionTargetBox="[218,1368,194,1744]" captionTargetId="figure@31.[218,1368,194,1761]" captionTargetPageId="31" captionText="FIGURE 4. Rhyacopsyche hagenii Müller. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, dorsal; Eplumose seta." httpUri="https://zenodo.org/record/179481/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
D).
</paragraph>
<paragraph id="8B8636FEFFEEF61FFF58FD91637810AA" blockId="5.[151,1436,151,980]" pageId="5" pageNumber="6">
Female genitalia. Simple oviscapt. Tergum VIII bearing a setate lobe (or lobes) from posterior margin (
<figureCitation id="13022A7BFFEEF61FFF01FDB967E210AA" box="[159,229,634,660]" captionStart="FIGURE 5" captionStartId="32.[151,255,1691,1715]" captionTargetBox="[224,1362,187,1636]" captionTargetId="figure@32.[224,1364,187,1652]" captionTargetPageId="32" captionText="FIGURE 5. Rhyacopsyche hagenii Müller. Female genitalia: A—ventral; B—dorsal; C—lateral." httpUri="https://zenodo.org/record/179482/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
B). Segments VII, VIII and IX with long internal rod-like apodemes (
<figureCitation id="13022A7BFFEEF61FFB8DFDB9635E10AA" box="[1043,1113,634,660]" captionStart="FIGURE 5" captionStartId="32.[151,255,1691,1715]" captionTargetBox="[224,1362,187,1636]" captionTargetId="figure@32.[224,1364,187,1652]" captionTargetPageId="32" captionText="FIGURE 5. Rhyacopsyche hagenii Müller. Female genitalia: A—ventral; B—dorsal; C—lateral." httpUri="https://zenodo.org/record/179482/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
A).
</paragraph>
<paragraph id="8B8636FEFFEEF61FFF58FD6166FC110A" blockId="5.[151,1436,151,980]" pageId="5" pageNumber="6">
Larva. All 3 thoracic nota divided mesally (
<figureCitation id="13022A7BFFEEF61FFD41FD6164221082" box="[735,805,674,700]" captionStart="FIGURE 1" captionStartId="28.[151,255,1974,1998]" captionTargetBox="[192,1418,189,1917]" captionTargetId="figure@28.[192,1419,189,1919]" captionTargetPageId="28" captionText="FIGURE 1. Rhyacopsyche sp. Larva: A—lateral; B—abdominal segment, dorsal; C—head and thorax, dorsal; D—pro-, meso-, and metathoracic legs, lateral; E—case; F—anal claw." httpUri="https://zenodo.org/record/179478/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
C); legs stout and similar in structure, apex of tibia expanded bearing several enlarged setae; tarsus broad with 1 or 2 enlarged, platelike setae; claw basally inflated (
<figureCitation id="13022A7BFFEEF61FFE9FFD3166441132" box="[257,323,754,780]" captionStart="FIGURE 1" captionStartId="28.[151,255,1974,1998]" captionTargetBox="[192,1418,189,1917]" captionTargetId="figure@28.[192,1419,189,1919]" captionTargetPageId="28" captionText="FIGURE 1. Rhyacopsyche sp. Larva: A—lateral; B—abdominal segment, dorsal; C—head and thorax, dorsal; D—pro-, meso-, and metathoracic legs, lateral; E—case; F—anal claw." httpUri="https://zenodo.org/record/179478/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
D). Abdominal segments IV–VII slightly swollen (
<figureCitation id="13022A7BFFEEF61FFC03FD3164D81132" box="[925,991,754,780]" captionStart="FIGURE 1" captionStartId="28.[151,255,1974,1998]" captionTargetBox="[192,1418,189,1917]" captionTargetId="figure@28.[192,1419,189,1919]" captionTargetPageId="28" captionText="FIGURE 1. Rhyacopsyche sp. Larva: A—lateral; B—abdominal segment, dorsal; C—head and thorax, dorsal; D—pro-, meso-, and metathoracic legs, lateral; E—case; F—anal claw." httpUri="https://zenodo.org/record/179478/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
A); tergites absent; anal prolegs compressed to sides of segment X.
</paragraph>
<paragraph id="8B8636FEFFEEF61FFF58FC81655D11EA" blockId="5.[151,1436,151,980]" pageId="5" pageNumber="6">
Larval case. Bivalve case elongate, tubular and fusiform; constructed of silk only or covered in sand grains; anterior end often with long, silken stalk used in attachment to the substrate (
<figureCitation id="13022A7BFFEEF61FFBEAFCA963B211BA" box="[1140,1205,874,900]" captionStart="FIGURE 1" captionStartId="28.[151,255,1974,1998]" captionTargetBox="[192,1418,189,1917]" captionTargetId="figure@28.[192,1419,189,1919]" captionTargetPageId="28" captionText="FIGURE 1. Rhyacopsyche sp. Larva: A—lateral; B—abdominal segment, dorsal; C—head and thorax, dorsal; D—pro-, meso-, and metathoracic legs, lateral; E—case; F—anal claw." httpUri="https://zenodo.org/record/179478/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
E). Pupal case with oval anterior and posterior openings sealed with silk; when stalk is present, it is shortened and thickened for more rigid attachment to the substrate.
</paragraph>
</subSubSection>
</treatment>
</document>
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