284 lines
35 KiB
XML
284 lines
35 KiB
XML
<document id="0ADC07DFC87672BB86313CD194D89E74" ID-CLB-Dataset="26030" ID-DOI="10.11646/zootaxa.4656.1.7" ID-GBIF-Dataset="67dbf573-5358-4a7f-80a6-fd07df93fdb1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3366805" ID-ZooBank="C63683B3-5CE3-4B71-A15B-1B886E80E6DE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565681870283" checkinUser="plazi" docAuthor="Belousov, Igor A., Kabak, Ilya I. & Liang, Hongbin" docDate="2019" docId="039087E40668FFF040F63B9AFEED37F7" docLanguage="en" docName="zootaxa.4656.1.7.pdf" docOrigin="Zootaxa 4656 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protrechiama Belousov & Kabak 2003" docType="treatment" docVersion="9" lastPageNumber="152" masterDocId="FFA9FF9C066FFFF940613B00FFC53314" masterDocTitle="New species of the tribe Trechini from China (Coleoptera: Carabidae)" masterLastPageNumber="152" masterPageNumber="143" pageNumber="150" updateTime="1698750868340" updateUser="ExternalLinkService">
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<mods:titleInfo id="DCEE7C9D9D11280722EFD59F24F32112">
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<mods:title id="040882F792D8E80E3D17899AE4B5E2F2">New species of the tribe Trechini from China (Coleoptera: Carabidae)</mods:title>
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<mods:name id="8B9965A0E7E7F93A88E5C60628F51C76" type="personal">
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<mods:namePart id="9CAEA6FE64C822505507BEC7A0494684">Belousov, Igor A.</mods:namePart>
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<mods:namePart id="3CA539AD06F600332FF6EB91A115AE20">Kabak, Ilya I.</mods:namePart>
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<mods:namePart id="BAAA713FF8E3DA749075134DCF0E2CFA">Liang, Hongbin</mods:namePart>
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<mods:date id="F2B2AA9CA5B72646346396E4D8824670">2019</mods:date>
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<treatment id="039087E40668FFF040F63B9AFEED37F7" ID-DOI="http://doi.org/10.5281/zenodo.5944199" ID-GBIF-Taxon="159126060" ID-Zenodo-Dep="5944199" LSID="urn:lsid:plazi:treatment:039087E40668FFF040F63B9AFEED37F7" httpUri="http://treatment.plazi.org/id/039087E40668FFF040F63B9AFEED37F7" lastPageId="9" lastPageNumber="152" pageId="7" pageNumber="150">
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<subSubSection id="C32365790668FFFE40F63B9AFD9233A0" box="[151,599,153,181]" pageId="7" pageNumber="150" type="nomenclature">
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<paragraph id="8B8636F20668FFFE40F63B9AFD9233A0" blockId="7.[151,599,153,217]" box="[151,599,153,181]" pageId="7" pageNumber="150">
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<heading id="D0CE819E0668FFFE40F63B9AFD9233A0" bold="true" box="[151,599,153,181]" fontSize="11" level="1" pageId="7" pageNumber="150" reason="1">
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<emphasis id="B94DEAE00668FFFE40F63B9AFD9233A0" bold="true" box="[151,599,153,181]" pageId="7" pageNumber="150">
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<emphasis id="B94DEAE00668FFFE40F63B9AFE7733A0" bold="true" box="[151,434,154,181]" italics="true" pageId="7" pageNumber="150">
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<taxonomicName id="4C394D710668FFFE40F63B9AFEF833A1" ID-CoL="8KYKV" authorityName="Belousov & Kabak" authorityYear="2003" box="[151,317,154,181]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="150" phylum="Arthropoda" rank="genus">Protrechiama</taxonomicName>
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сrassipes
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</emphasis>
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(
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<bibRefCitation id="EFA84B030668FFFE41A03B99FD8A33A0" author="Ueno, S. - I." box="[449,591,153,180]" pageId="7" pageNumber="150" pagination="37 - 42" refId="ref5944" refString="Ueno, S. - I. (1997) Occurrence of Trechiama (Coleoptera, Trechinae) in Southwest China. Journal of the Speleological Society of Japan, 22, 37 - 42." type="journal article" year="1997">Uéno, 1997</bibRefCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32365790668FFFE40F63BBEFF2933CD" box="[151,236,190,217]" pageId="7" pageNumber="150" type="description">
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<paragraph id="8B8636F20668FFFE40F63BBEFF2933CD" blockId="7.[151,599,153,217]" box="[151,236,190,217]" pageId="7" pageNumber="150">
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(
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<figureCitation id="13022A770668FFFE40C13BBEFF2633CD" box="[160,227,190,217]" captionStart="FIGURE 6" captionStartId="8.[151,250,2000,2025]" captionTargetId="figure@8.[266,1321,164,1978]" captionTargetPageId="8" captionText="FIGURE 6. Protrechiama crassipes (Uéno, 1997) female, habitus." figureDoi="http://doi.org/10.5281/zenodo.3443676" httpUri="https://zenodo.org/record/3443676/files/figure.png" pageId="7" pageNumber="150">Fig. 6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32365790668FFFE40F63A02FDD4322D" pageId="7" pageNumber="150" type="reference_group">
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<paragraph id="8B8636F20668FFFE40F63A02FDD4322D" blockId="7.[151,1354,257,313]" pageId="7" pageNumber="150">
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<treatmentCitationGroup id="AB2911DC0668FFFE40F63A02FDD4322D" pageId="7" pageNumber="150">
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<taxonomicName id="4C394D710668FFFE40F63A02FDC8320E" ID-CoL="7CNWG" authority="Ueno, 1997: 38" authorityName="Ueno" authorityPageNumber="38" authorityYear="1997" box="[151,525,258,282]" class="Insecta" family="Carabidae" genus="Trechiama" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="150" phylum="Arthropoda" rank="species" species="crassipes">
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<emphasis id="B94DEAE00668FFFE40F63A02FEA2320E" box="[151,359,258,282]" italics="true" pageId="7" pageNumber="150">Trechiama crassipes</emphasis>
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<treatmentCitation id="0A9810E30668FFFE410F3A02FDC8320E" author="Ueno, S. - I." box="[366,525,258,282]" page="38" pageId="7" pageNumber="150" year="1997">
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<bibRefCitation id="EFA84B030668FFFE410F3A02FDC8320E" author="Ueno, S. - I." box="[366,525,258,282]" pageId="7" pageNumber="150" pagination="37 - 42" refId="ref5944" refString="Ueno, S. - I. (1997) Occurrence of Trechiama (Coleoptera, Trechinae) in Southwest China. Journal of the Speleological Society of Japan, 22, 37 - 42." type="journal article" year="1997">Uéno, 1997: 38</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="548288500668FFFE42763A03FD8C320F" box="[535,585,259,283]" pageId="7" pageNumber="150">Type</typeStatus>
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locality: “Mt Dafengding, Wahei Xiang of Meigu Xian, southern
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<collectingRegion id="49FDF8100668FFFE44873A01FAF8320E" box="[1254,1341,257,282]" country="China" name="Sichuan" pageId="7" pageNumber="150">Sichuan</collectingRegion>
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”.
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<taxonomicName id="4C394D710668FFFE40F63A21FDCB322D" ID-CoL="4N7RM" authority=": Ueno, 2006" authorityName="Ueno" authorityYear="2006" box="[151,526,288,313]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="150" phylum="Arthropoda" rank="species" species="crassipes">
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<emphasis id="B94DEAE00668FFFE40F63A21FE43322D" box="[151,390,289,313]" italics="true" pageId="7" pageNumber="150">Protrechiama crassipes</emphasis>
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:
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<bibRefCitation id="EFA84B030668FFFE41F53A21FDCB322D" author="Ueno, S. - I." box="[404,526,288,313]" pageId="7" pageNumber="150" pagination="13 - 32" refId="ref5980" refString="Ueno, S. - I. (2006) The trechine beetles (Coleoptera, Trechinae) from Mt. Wawu Shan in central Sichuan, southwest China. Elytra, 34, 13 - 32." type="journal article" year="2006">Uéno, 2006</bibRefCitation>
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</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32365790668FFF040F63A66FEED37F7" lastPageId="9" lastPageNumber="152" pageId="7" pageNumber="150" type="materials_examined">
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<paragraph id="8B8636F20668FFFE40F63A66FBAC32DD" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">
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<emphasis id="B94DEAE00668FFFE40F63A66FE463295" bold="true" box="[151,387,358,385]" pageId="7" pageNumber="150">Material examined:</emphasis>
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<materialsCitation id="3B513CAF0668FFFE41EB3A66FBA132DD" ID-GBIF-Occurrence="2332229317" collectingDate="2012-06-12" collectionCode="CAS, IOZ" collectorName="Liu Ye" country="China" county="Meigu County" elevation="2923" latitude="28.67477" location="Yizi Pass" longLatPrecision="1" longitude="103.05248" municipality="China" pageId="7" pageNumber="150" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan">
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||
<emphasis id="B94DEAE00668FFFE41EB3A66FE343295" bold="true" box="[394,497,358,385]" pageId="7" pageNumber="150">
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||
<collectingCountry id="F32E76620668FFFE41EB3A66FE293295" box="[394,492,358,385]" name="China" pageId="7" pageNumber="150">CHINA</collectingCountry>
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:
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</emphasis>
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||
<specimenCount id="9D3FFD7B0668FFFE41993A67FDDE3295" box="[504,539,359,385]" pageId="7" pageNumber="150" type="female">1♀</specimenCount>
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, with label data “
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<collectingMunicipality id="6BE2AC880668FFFE42813A67FCED3295" box="[736,808,359,385]" pageId="7" pageNumber="150">China</collectingMunicipality>
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,
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<collectingRegion id="49FDF8100668FFFE43553A67FC143296" box="[820,977,359,386]" country="China" name="Sichuan" pageId="7" pageNumber="150">Sichuan prov.</collectingRegion>
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,
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<location id="8EE660290668FFFE43BA3A67FB863296" LSID="urn:lsid:plazi:treatment:039087E40668FFF040F63B9AFEED37F7:8EE660290668FFFE43BA3A67FB863296" box="[987,1091,359,386]" country="China" county="Meigu County" latitude="28.67477" longLatPrecision="1" longitude="103.05248" municipality="China" name="Yizi Pass" pageId="7" pageNumber="150" stateProvince="Sichuan">Yizi Pass</location>
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btw.
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<collectingCounty id="62E74E7E0668FFFE44E03A67FAE63295" box="[1153,1315,359,385]" pageId="7" pageNumber="150">Meigu County</collectingCounty>
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and
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<location id="8EE660290668FFFE453A3A67FF2E32B1" LSID="urn:lsid:plazi:treatment:039087E40668FFF040F63B9AFEED37F7:8EE660290668FFFE453A3A67FF2E32B1" country="China" county="Meigu County" latitude="28.67477" longLatPrecision="1" longitude="103.05248" municipality="China" name="Ebian County" pageId="7" pageNumber="150" stateProvince="Sichuan">Ebian County</location>
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, mixed forest,
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<quantity id="4CC19B170668FFFE41F53A8BFE2832B2" box="[404,493,395,422]" metricMagnitude="3" metricUnit="m" metricValue="2.923" pageId="7" pageNumber="150" unit="m" value="2923.0">
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<elevation id="0014D1C10668FFFE41F53A8BFE2832B2" box="[404,493,395,422]" metricMagnitude="3" metricUnit="m" metricValue="2.923" pageId="7" pageNumber="150" unit="m" value="2923.0">2923 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE0D50350668FFFE419B3A8CFD4732B1" box="[506,642,395,422]" degrees="28.67477" direction="north" orientation="latitude" pageId="7" pageNumber="150" precision="1" value="28.67477">N 28.67477</geoCoordinate>
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,
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<geoCoordinate id="EE0D50350668FFFE42EF3A8CFCE032B1" box="[654,805,394,422]" degrees="103.05248" direction="east" orientation="longitude" pageId="7" pageNumber="150" precision="1" value="103.05248">E 103.05248</geoCoordinate>
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”; “
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<date id="FF8710320668FFFE43273A8BFC0C32B1" box="[838,969,395,421]" pageId="7" pageNumber="150" value="2012-06-12">
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<collectingDate id="EFC3E9DA0668FFFE43273A8BFC0C32B1" box="[838,969,395,421]" pageId="7" pageNumber="150" value="2012-06-12">2012.VI.12</collectingDate>
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</date>
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, in dead log, Shi Hongliang,
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<collectorName id="26CC53240668FFFE45793A8BFAA132B2" box="[1304,1380,395,422]" pageId="7" pageNumber="150">Liu Ye</collectorName>
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leg.,
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<location id="8EE660290668FFFE40F63AAFFEBD32DD" LSID="urn:lsid:plazi:treatment:039087E40668FFF040F63B9AFEED37F7:8EE660290668FFFE40F63AAFFEBD32DD" box="[151,376,431,457]" country="China" county="Meigu County" latitude="28.67477" longLatPrecision="1" longitude="103.05248" municipality="China" name="Institute of Zoology" pageId="7" pageNumber="150" stateProvince="Sichuan">Institute of Zoology</location>
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,
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<collectionCode id="ED28AE370668FFFE41E23AAFFE7832DD" box="[387,445,431,457]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="150" type="Museum">CAS</collectionCode>
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||
. [In Chinese]: Meigu Xian—Ebian Xian, Yizi Yakou” (
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<collectionCode id="ED28AE370668FFFE44493AAFFB9932DD" box="[1064,1116,431,457]" pageId="7" pageNumber="150">IOZ</collectionCode>
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)
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</materialsCitation>
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.
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||
</paragraph>
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||
<paragraph id="8B8636F20668FFFE40A63AD3FC29314E" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">
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||
Although the species was adequately and thoroughly described by S.–I.
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<bibRefCitation id="EFA84B030668FFFE438F3AD3FBBA32F9" author="Ueno, S. - I." box="[1006,1151,467,493]" pageId="7" pageNumber="150" pagination="37 - 42" refId="ref5944" refString="Ueno, S. - I. (1997) Occurrence of Trechiama (Coleoptera, Trechinae) in Southwest China. Journal of the Speleological Society of Japan, 22, 37 - 42." type="journal article" year="1997">Uéno (1997)</bibRefCitation>
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||
, we give here its description based on newly available female specimen from another collecting site in order to illustrate variation in some characters and to draw attention to some others. In addition, the photograph will help in more accurate understanding of external morphology, in particular concerning the shape of pronotum.
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||
</paragraph>
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<paragraph id="8B8636F20668FFFE40A63962FBDF31B6" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">
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||
<emphasis id="B94DEAE00668FFFE40A63962FE943169" bold="true" box="[199,337,610,637]" pageId="7" pageNumber="150">Description</emphasis>
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||
. The smallest species of the genus, body length
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<quantity id="4CC19B170668FFFE43E33963FC18316A" box="[898,989,611,638]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="7" pageNumber="150" unit="mm" value="4.8">4.8 mm</quantity>
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. Habitus relatively narrow, subparallel, depressed, with rather thick (compared to congeners) antennae and legs (
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<figureCitation id="13022A770668FFFE43A63987FBC931B6" box="[967,1036,647,674]" captionStart="FIGURE 6" captionStartId="8.[151,250,2000,2025]" captionTargetId="figure@8.[266,1321,164,1978]" captionTargetPageId="8" captionText="FIGURE 6. Protrechiama crassipes (Uéno, 1997) female, habitus." figureDoi="http://doi.org/10.5281/zenodo.3443676" httpUri="https://zenodo.org/record/3443676/files/figure.png" pageId="7" pageNumber="150">Fig. 6</figureCitation>
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||
).
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</paragraph>
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<paragraph id="8B8636F20668FFFE40A639ABFE2731FD" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">Color of dorsum amber brownish, with slightly darker posterior portion of head and disc of pronotum and elytra. Appendages pale reddish.</paragraph>
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<paragraph id="8B8636F20668FFFE40A639F3FE4E3041" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">Microsculpture fine, consisting of well-engraved rather small isodiametric meshes on head, very shallow irregular transverse meshes on pronotum and fine transverse lines on elytra. All upper surface finely micropunctured and faintly iridescent.</paragraph>
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<paragraph id="8B8636F20668FFFE40A6385FFAED37F5" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">
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Head somewhat trapezoid (
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<collectionCode id="ED28AE370668FFFE4263385FFDE8306D" box="[514,557,863,889]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="150">PW</collectionCode>
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/
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||
<collectionCode id="ED28AE370668FFFE4257385FFDA0306D" box="[566,613,863,889]" pageId="7" pageNumber="150">HW</collectionCode>
|
||
1.34), eyes medium-sized (YL/TL 1.42, L3/YL 1.12), slightly convex; tempora glabrous, slightly convex anteriorly, much more markedly in posterior portion. Frontal furrows rather deep, subparallel in median portion, more deeply impressed and divergent anteriorly, abruptly divergent posteriorly. Parietal transverse impression distinct. Supraorbital setiferous pores located in lines clearly convergent posteriad, anterior one deeply, posterior one moderately foveolate. Anterior margin of labrum concave. Mandibles relatively slender, evenly curved. Tooth on right mandible not large, with long basis, tridentate, median denticle being shortest, while the distal one—longest. Labial tooth distinctly cleft at apex. Labial suture very fine but traceable. Six submental setae. Segments of maxillary palpi of medium length, fusiform, glabrous. Penultimate segment of the labial palpi quadrisetose. Ligula obtusangular, blunt at apex, with two long apical setae and three lateral setae on each side. Paraglossae moderately long, slender, arcuate and distinctly pubescent.Antennae average in length, rather thick, AL/EL 1.04, middle antennomeres approximately twice as long as wide, L3/
|
||
<geoCoordinate id="EE0D50350668FFFE44553FC7FB5D37F5" box="[1076,1176,1223,1249]" degrees="3" direction="west" minutes="2.60" orientation="longitude" pageId="7" pageNumber="150" precision="9" value="-3.0433333">W3 2.60</geoCoordinate>
|
||
, L3/L2 1.46.
|
||
</paragraph>
|
||
<paragraph id="8B8636F20668FFFE40A63FECFB9B3531" blockId="7.[151,1437,358,2042]" pageId="7" pageNumber="150">
|
||
Pronotum transverse (
|
||
<collectionCode id="ED28AE370668FFFE41DF3FEBFE2D3611" box="[446,488,1259,1285]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="150">PW</collectionCode>
|
||
/
|
||
<collectionCode id="ED28AE370668FFFE41903FEBFDD73611" box="[497,530,1259,1285]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="150" type="Museum">PL</collectionCode>
|
||
1.27), markedly contracted to base (
|
||
<collectionCode id="ED28AE370668FFFE43D13FEBFC193611" box="[944,988,1259,1285]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="150">PW</collectionCode>
|
||
/
|
||
<collectionCode id="ED28AE370668FFFE43853FEBFBC33611" box="[996,1030,1259,1285]" pageId="7" pageNumber="150">PB</collectionCode>
|
||
1.39), slightly wider at base than at apex (
|
||
<collectionCode id="ED28AE370668FFFE40BB3E0FFF3F363D" box="[218,250,1295,1321]" pageId="7" pageNumber="150">PB</collectionCode>
|
||
/PA 1.14). Lateral margins convex nearly for all of their length, except for a slight and shallow emargination just before hind angles, markedly reflexed, especially in their posterior portion. Hind angles rather small and slightly obtuse, with blunt apices. Anterior margin concave, front angles rounded, slightly but distinctly produced anteriad. Posterior margin straight medially, obliquely emarginate laterally. Disc of pronotum slightly convex, depressed along suture, basal foveae large and deep. Prebasal transverse impression not sharply impressed. Base of pronotum irregularly wrinkled. Apical transverse impression distinct, triangular-shaped. Median line distinct but rather shallow. Lateral groove rather wide, its inner margin slightly waved. Discal foveae shallow. Anterior lateral seta located in anterior third of pronotum (PSa 34%), posterior one just at hind angles.
|
||
</paragraph>
|
||
<paragraph id="8B8636F20668FFF040A63D2FFBD3320A" blockId="7.[151,1437,358,2042]" lastBlockId="9.[151,1437,153,1252]" lastPageId="9" lastPageNumber="152" pageId="7" pageNumber="150">
|
||
Elytra (
|
||
<collectionCode id="ED28AE370668FFFE417B3D2FFEF9355D" box="[282,316,1583,1609]" pageId="7" pageNumber="150">EL</collectionCode>
|
||
/
|
||
<collectionCode id="ED28AE370668FFFE41243D2FFEA3355D" box="[325,358,1583,1609]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="150" type="Museum">PL</collectionCode>
|
||
2.70; EW/
|
||
<collectionCode id="ED28AE370668FFFE41803D2FFDCE355D" box="[481,523,1583,1609]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="150">PW</collectionCode>
|
||
1.39;
|
||
<collectionCode id="ED28AE370668FFFE42333D2FFDB8355D" box="[594,637,1583,1609]" pageId="7" pageNumber="150">EW</collectionCode>
|
||
/HW 1.86) rather narrow (
|
||
<collectionCode id="ED28AE370668FFFE43C53D2FFC03355D" box="[932,966,1583,1609]" pageId="7" pageNumber="150">EL</collectionCode>
|
||
/
|
||
<collectionCode id="ED28AE370668FFFE43AE3D2FFC3E355D" box="[975,1019,1583,1609]" pageId="7" pageNumber="150">EW</collectionCode>
|
||
1.53), subparallel, depressed on disk (
|
||
<collectionCode id="ED28AE370668FFFE40FE3D53FF073579" box="[159,194,1619,1645]" pageId="7" pageNumber="150">EL</collectionCode>
|
||
/BH 2.44), their sides only marginally convex in their median portion; maximum width slightly behind middle, apex truncate. Humeri distinct though rounded. Marginal groove narrow. Elytral striation distinct: inner striae deep and coarsely punctured, outer striae shallower but traceable, finely punctured. Striolation on the apical slope distinctly engraved. Stria 8 shallow in anterior portion, becoming deeper on the apical slope and angularly curved near umbilicate pore 7, interval between stria 8 and the apical striole wide and convex, narrowing to the elytral apex, the neighbouring outer interval, on the contrary, plane. Inner interspaces subconvex, outer interspaces flat. Parascutellar striole long and deep, scutellar pore present. Apical striole nearly straight in anterior portion, abruptly terminated anteriorly at level slightly behind umbilicate pores 7; apical carinula well-developed and sharply delimited. On each elytron, 10 foveolate setiferous pores arranged in two irregular longitudinal series. The inner series, apart from the preapical pore, consists of four setae, of which the anterior one located in stria 2 and the remaining three pores in stria 3. The outer series consists of five pores located in stria 5. Preapical pore situated in the apical cross of striae 2 and 3 markedly behind the anterior termination of the apical striole, DP 91.9%. All pores of the apical triangle present, the inner side of the apical triangle is divergent from the elytral suture posteriad, since the angulo-apical pore shifted outward and located much closer to the exterior pore than to the elytral suture. All umbilicate pores well-aggregated within their groups, equidistant, except for pores 7 and 8 which are markedly more spaced. Umbilicate formula U1–8: 7.9%, 13.8%, 19.8%, 25.2%, 64.2%, 69.6%, 84.9%, 92.8%.
|
||
</paragraph>
|
||
<caption id="DF46667A0667FFF140F63CD0FCFA34FD" ID-DOI="http://doi.org/10.5281/zenodo.3443676" ID-Zenodo-Dep="3443676" box="[151,831,2000,2025]" httpUri="https://zenodo.org/record/3443676/files/figure.png" pageId="8" pageNumber="151" targetBox="[333,1300,184,1971]" targetPageId="8">
|
||
<paragraph id="8B8636F20667FFF140F63CD0FCFA34FD" blockId="8.[151,831,2000,2025]" box="[151,831,2000,2025]" pageId="8" pageNumber="151">
|
||
<emphasis id="B94DEAE00667FFF140F63CD0FED634FD" bold="true" box="[151,275,2000,2025]" pageId="8" pageNumber="151">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C394D710667FFF141783CD1FD5334FD" authority="(Ueno, 1997)" authorityName="Ueno" authorityYear="1997" baseAuthorityName="Ueno" baseAuthorityYear="1997" box="[281,662,2001,2025]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="151" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B94DEAE00667FFF141783CD1FDCC34FC" box="[281,521,2001,2025]" italics="true" pageId="8" pageNumber="151">Protrechiama crassipes</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA84B030667FFF142773CD1FD4A34FD" author="Ueno, S. - I." box="[534,655,2001,2025]" pageId="8" pageNumber="151" pagination="37 - 42" refId="ref5944" refString="Ueno, S. - I. (1997) Occurrence of Trechiama (Coleoptera, Trechinae) in Southwest China. Journal of the Speleological Society of Japan, 22, 37 - 42." type="journal article" year="1997">Uéno, 1997</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
female, habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8636F20666FFF040A63A28FE71329D" blockId="9.[151,1437,153,1252]" pageId="9" pageNumber="152">Ventral surface: metepisternites clearly longer than wide. Abdominal sternite 2 without setae, sternite 3 with one pair of paramedian setae, sternites 4 and 5 with 2–3 (usually 2) pair of setae. Anal sternite quadrisetose in one female specimen studied.</paragraph>
|
||
<paragraph id="8B8636F20666FFF040A63A94FB3D32C5" blockId="9.[151,1437,153,1252]" pageId="9" pageNumber="152">Legs rather thick, foretibiae sharply grooved and carinate on exterior surface, nearly glabrous, with only a few hairs just near the tibia apex on their anterior surface. Hind tibia ratios: EL/HTi 1.93; HTi/HTa 1.31.</paragraph>
|
||
<paragraph id="8B8636F20666FFF040A63ADAFCCF3007" blockId="9.[151,1437,153,1252]" pageId="9" pageNumber="152">
|
||
<emphasis id="B94DEAE00666FFF040A63ADAFE7532E1" bold="true" box="[199,432,474,501]" pageId="9" pageNumber="152">Comparative notes.</emphasis>
|
||
As it was indicated by
|
||
<bibRefCitation id="EFA84B030666FFF042D53ADBFC8132E1" author="Ueno, S. - I." box="[692,836,475,501]" pageId="9" pageNumber="152" pagination="37 - 42" refId="ref5944" refString="Ueno, S. - I. (1997) Occurrence of Trechiama (Coleoptera, Trechinae) in Southwest China. Journal of the Speleological Society of Japan, 22, 37 - 42." type="journal article" year="1997">Uéno (1997)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C394D710666FFF0432E3ADBFC1632E0" baseAuthorityName="Ueno" baseAuthorityYear="1997" box="[847,979,475,500]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B94DEAE00666FFF0432E3ADBFC1632E0" box="[847,979,475,500]" italics="true" pageId="9" pageNumber="152">P. crassipes</emphasis>
|
||
</taxonomicName>
|
||
is the most isolated species of the genus, although it shares all major characters of other congeners: the number and disposition of elytral setiferous pores (closer to the lower limit of variation in
|
||
<taxonomicName id="4C394D710666FFF0420D3923FBFD3128" authority="Belousov & Kabak, 2003" authorityName="Belousov & Kabak" authorityYear="2003" box="[620,1080,546,572]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="species" species="glabricollis">
|
||
<emphasis id="B94DEAE00666FFF0420D3923FCCE3128" box="[620,779,546,572]" italics="true" pageId="9" pageNumber="152">P. glabricollis</emphasis>
|
||
<bibRefCitation id="EFA84B030666FFF043753922FBFD3128" author="Belousov, I. & Kabak, I." box="[788,1080,546,572]" pageId="9" pageNumber="152" pagination="15 - 86" refId="ref5869" refString="Belousov, I. & Kabak, I. (2003) New Trechini from China (Coleoptera, Carabidae) (Coleoptera, Carabidae). Tethys Entomological Research, 8, 15 - 86." type="journal article" year="2003">Belousov & Kabak, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D710666FFF044153923FEEF3174" authority="Belousov & Kabak, 2003" authorityName="Belousov & Kabak" authorityYear="2003" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="species" species="marginalis">
|
||
<emphasis id="B94DEAE00666FFF044153923FACC3128" box="[1140,1289,546,572]" italics="true" pageId="9" pageNumber="152">P. marginalis</emphasis>
|
||
<bibRefCitation id="EFA84B030666FFF045723922FEEF3174" author="Belousov, I. & Kabak, I." pageId="9" pageNumber="152" pagination="15 - 86" refId="ref5869" refString="Belousov, I. & Kabak, I. (2003) New Trechini from China (Coleoptera, Carabidae) (Coleoptera, Carabidae). Tethys Entomological Research, 8, 15 - 86." type="journal article" year="2003">Belousov & Kabak, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
), the anterior discal setiferous pore located in stria 2, the absence of setae on the pronotal disc and the broad lateral groove of the pronotum. However,
|
||
<taxonomicName id="4C394D710666FFF042D8396AFCF83197" baseAuthorityName="Ueno" baseAuthorityYear="1997" box="[697,829,618,643]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B94DEAE00666FFF042D8396AFCF83197" box="[697,829,618,643]" italics="true" pageId="9" pageNumber="152">P. crassipes</emphasis>
|
||
</taxonomicName>
|
||
differs readily from all other currently known
|
||
<taxonomicName id="4C394D710666FFF0452D396AFF2C31B3" authorityName="Belousov & Kabak" authorityYear="2003" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAE00666FFF0452D396AFF2C31B3" italics="true" pageId="9" pageNumber="152">Protrechiama</emphasis>
|
||
</taxonomicName>
|
||
in much smaller size (
|
||
<quantity id="4CC19B170666FFF0418D398CFDBA31BC" box="[492,639,652,680]" metricMagnitude="-3" metricUnit="m" metricValue="5.025" metricValueMax="5.25" metricValueMin="4.8" pageId="9" pageNumber="152" unit="mm" value="5.025" valueMax="5.25" valueMin="4.8">4.8-5.25 mm</quantity>
|
||
from anterior margin of labrum for all specimens currently known vs. more than
|
||
<quantity id="4CC19B170666FFF0417039B0FEB031DF" box="[273,373,688,715]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" pageId="9" pageNumber="152" unit="mm" value="6.65">6.65 mm</quantity>
|
||
in other species), short and flat elytra (vs. long and convex, oblong-ovate elytra) and, specifically, much stouter legs and antennae with middle antennomeres only marginally more than twice as long as wide (vs. more than 3 times as long as wide in other species).
|
||
</paragraph>
|
||
<paragraph id="8B8636F20666FFF040A6381CFBC23719" blockId="9.[151,1437,153,1252]" pageId="9" pageNumber="152">
|
||
The female from the Yizi Pass differs from the
|
||
<typeStatus id="548288500666FFF04288381DFCDC3023" box="[745,793,797,823]" pageId="9" pageNumber="152">type</typeStatus>
|
||
series in the following characters: 5 setiferous pores attached to stria 5 vs.
|
||
<quantity id="4CC19B170666FFF0411A3840FE06304E" box="[379,451,832,858]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="9" pageNumber="152" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
|
||
the
|
||
<typeStatus id="548288500666FFF041943840FDE0304E" box="[501,549,832,858]" pageId="9" pageNumber="152">type</typeStatus>
|
||
specimens; shape of pronotum with basal margin straight medially vs. salient, nearly arcuate in the
|
||
<typeStatus id="548288500666FFF041E33864FE78306A" box="[386,445,868,894]" pageId="9" pageNumber="152">types</typeStatus>
|
||
specimens and tempora more evenly convex in anterior portion vs. nearly straight in the
|
||
<typeStatus id="548288500666FFF040F63888FF0230B6" box="[151,199,904,930]" pageId="9" pageNumber="152">type</typeStatus>
|
||
specimens. These differences do not seem to be of great taxonomic importance and, for the moment, we prefer to consider the female from the Yizi Pass to be conspecific with the
|
||
<typeStatus id="548288500666FFF043EE38ACFC7A30D2" box="[911,959,940,966]" pageId="9" pageNumber="152">type</typeStatus>
|
||
series. Additionally, it is worth noting that figure
|
||
<quantity id="4CC19B170666FFF0408138CFFECF30FD" box="[224,266,975,1001]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="152" unit="in" value="1.0">1 in</quantity>
|
||
the original description is slightly misleading with regard to, at least, basal foveae of pronotum which are large according to the paper text and nearly missing in the figure (
|
||
<bibRefCitation id="EFA84B030666FFF0431838F3FC3E3719" author="Ueno, S. - I." box="[889,1019,1011,1037]" pageId="9" pageNumber="152" pagination="37 - 42" refId="ref5944" refString="Ueno, S. - I. (1997) Occurrence of Trechiama (Coleoptera, Trechinae) in Southwest China. Journal of the Speleological Society of Japan, 22, 37 - 42." type="journal article" year="1997">Uéno 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636F20666FFF040A63F15FCA0378F" blockId="9.[151,1437,153,1252]" pageId="9" pageNumber="152">
|
||
<emphasis id="B94DEAE00666FFF040A63F15FE9C3724" bold="true" box="[199,345,1045,1072]" pageId="9" pageNumber="152">Distribution</emphasis>
|
||
. All of the known members of the genus
|
||
<taxonomicName id="4C394D710666FFF043433F16FC7E3724" authorityName="Belousov & Kabak" authorityYear="2003" box="[802,955,1046,1072]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAE00666FFF043433F16FC7E3724" box="[802,955,1046,1072]" italics="true" pageId="9" pageNumber="152">Protrechiama</emphasis>
|
||
</taxonomicName>
|
||
are confined to a rather small mountainous area between the Niuri River in the west, the Dadu River in the north and the Yangtze River in the east.
|
||
<taxonomicName id="4C394D710666FFF045783F3BFA593740" baseAuthorityName="Ueno" baseAuthorityYear="1997" box="[1305,1436,1083,1108]" class="Insecta" family="Carabidae" genus="Protrechiama" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="152" phylum="Arthropoda" rank="species" species="crassipes">
|
||
<emphasis id="B94DEAE00666FFF045783F3BFA593740" box="[1305,1436,1083,1108]" italics="true" pageId="9" pageNumber="152">P. crassipes</emphasis>
|
||
</taxonomicName>
|
||
is not an exception, two known collecting sites are located less than
|
||
<quantity id="4CC19B170666FFF043FF3F5EFC23376C" box="[926,998,1118,1144]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="152" unit="km" value="10.0">10 km</quantity>
|
||
from each other in the north of Meigu County and in the Yizi Pass between Meigu and Ebian counties.
|
||
</paragraph>
|
||
<paragraph id="8B8636F20666FFF040A63FA4FEED37F7" blockId="9.[151,1437,153,1252]" pageId="9" pageNumber="152">
|
||
<emphasis id="B94DEAE00666FFF040A63FA4FE8E37AB" bold="true" box="[199,331,1188,1215]" pageId="9" pageNumber="152">Bionomics.</emphasis>
|
||
The species was found in a mixed forest at rather low elevations from 2300 to slightly exceeding
|
||
<quantity id="4CC19B170666FFF040F63FC9FF2937F0" box="[151,236,1225,1252]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="152" unit="m" value="2900.0">2900 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |