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<document id="410B5D036E356E3F4C29B4F59F592EB8" ID-DOI="10.11646/zootaxa.4000.1.1" ID-GBIF-Dataset="507a3784-35f5-46f5-9492-e0884a502f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="289219" ID-ZooBank="00769C76-A2D4-4792-A5DD-5F2B7EA352CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461184674989" checkinUser="plazi" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2015" docId="039087C7FFA8FF81E28C4BE234A66C3C" docLanguage="en" docName="zt04000p048.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blennidus (Agraphoderus) crassus Straneo 1993" docType="treatment" docVersion="6" lastPageNumber="8" masterDocId="FFA9FFBFFFAFFF86E21B4B7535296F7A" masterDocTitle="Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="8" updateTime="1698599231025" updateUser="plazi">
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<mods:title id="BFE67E6ED2AA93913FA64BFB85B1AAEE">Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)</mods:title>
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<mods:namePart id="910866407FAE4676C52663B63207D963">Allegro, Gianni</mods:namePart>
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<mods:namePart id="20D5A947AC1F23E9727E843C30E3C156">Giachino, Pier Mauro</mods:namePart>
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<mods:identifier id="D54EC7538ED0D3FB629A622E5AECF80C" type="GBIF-Dataset">507a3784-35f5-46f5-9492-e0884a502f40</mods:identifier>
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<treatment id="039087C7FFA8FF81E28C4BE234A66C3C" ID-DOI="http://doi.org/10.5281/zenodo.6096022" ID-GBIF-Taxon="127692746" ID-Zenodo-Dep="6096022" LSID="urn:lsid:plazi:treatment:039087C7FFA8FF81E28C4BE234A66C3C" httpUri="http://treatment.plazi.org/id/039087C7FFA8FF81E28C4BE234A66C3C" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="C323655AFFA8FF81E28C4BE234076FA9" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B8636D1FFA8FF81E28C4BE237D66FC8" blockId="7.[151,767,151,211]" box="[151,767,151,178]" pageId="7" pageNumber="8">
<heading id="D0CE81BDFFA8FF81E28C4BE237D66FC8" bold="true" box="[151,767,151,178]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="4C394D52FFA8FF81E28C4BE237D66FC8" ID-CoL="M53S" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,767,151,178]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassus" subGenus="Agraphoderus">
<emphasis id="B94DEAC3FFA8FF81E28C4BE237D66FC8" bold="true" box="[151,767,151,178]" pageId="7" pageNumber="8">
<emphasis id="B94DEAC3FFA8FF81E28C4BE2343A6FCB" bold="true" box="[151,275,151,177]" italics="true" pageId="7" pageNumber="8">Blennidus</emphasis>
(
<emphasis id="B94DEAC3FFA8FF81E33F4BE234F96FCB" bold="true" box="[292,464,151,177]" italics="true" pageId="7" pageNumber="8">Agraphoderus</emphasis>
)
<emphasis id="B94DEAC3FFA8FF81E3FA4BE237126FCB" bold="true" box="[481,571,151,177]" italics="true" pageId="7" pageNumber="8">crassus</emphasis>
(
<bibRefCitation id="EFA84B20FFA8FF81E0514BED37DE6FC8" author="Straneo" box="[586,759,152,178]" pageId="7" pageNumber="8" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8636D1FFA8FF81E28C4BCF34076FA9" blockId="7.[151,767,151,211]" box="[151,302,186,211]" pageId="7" pageNumber="8">
(
<figureCitation id="13022A54FFA8FF81E2844BCF35D56FA9" box="[159,252,186,211]" captionStart="FIGURES 16 19" captionStartId="28.[151,264,1953,1976]" captionTargetBox="[207,1382,198,1913]" captionTargetId="figure@28.[185,1402,193,1932]" captionTargetPageId="28" captionText="FIGURES 16 19. Habitus of Blennidus: bellesi male HT (16); bombonensis n. sp. male HT (17); crassus female HT (18); curtatus male HT (19)." httpUri="https://zenodo.org/record/289224/files/figure.png" pageId="7" pageNumber="8">Figs. 18</figureCitation>
, 56)
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA8FF81E28C4A7736616E0E" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8B8636D1FFA8FF81E28C4A7737716E62" blockId="7.[151,840,258,373]" box="[151,600,258,280]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB2911FFFFA8FF81E28C4A7737716E62" box="[151,600,258,280]" pageId="7" pageNumber="8">
<taxonomicName id="4C394D52FFA8FF81E28C4A77370C6E62" ID-CoL="6SJG4" authority="Straneo, 1993: 389" authorityName="Straneo" authorityPageNumber="389" authorityYear="1993" box="[151,549,258,280]" class="Insecta" family="Carabidae" genus="Ogmopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="B94DEAC3FFA8FF81E28C4A7734766E62" box="[151,351,258,280]" italics="true" pageId="7" pageNumber="8">Ogmopleura crassa</emphasis>
<treatmentCitation id="0A9810C0FFA8FF81E37E4A77370C6E62" author="Straneo" box="[357,549,258,280]" page="389" pageId="7" pageNumber="8" year="1993">
<bibRefCitation id="EFA84B20FFA8FF81E37E4A7734DC6E62" author="Straneo" box="[357,501,258,280]" pageId="7" pageNumber="8" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
: 389
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391
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</paragraph>
<paragraph id="8B8636D1FFA8FF81E28C4A5436616E0E" blockId="7.[151,840,258,373]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB2911FFFFA8FF81E28C4A5436616E0E" pageId="7" pageNumber="8">
<taxonomicName id="4C394D52FFA8FF81E28C4A5434DF6E4D" ID-CoL="M53S" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,502,289,311]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA8FF81E28C4A5434676E4D" box="[151,334,289,311]" italics="true" pageId="7" pageNumber="8">Blennidus crassus</emphasis>
(
<bibRefCitation id="EFA84B20FFA8FF81E3404A5434C76E4D" author="Straneo" box="[347,494,289,311]" pageId="7" pageNumber="8" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA8FF81E3E44A5737ED6E4D" author="Lorenz" box="[511,708,289,311]" page="262" pageId="7" pageNumber="8" year="2005">
<bibRefCitation id="EFA84B20FFA8FF81E3E44A5737BF6E4D" author="Lorenz" box="[511,662,289,311]" pageId="7" pageNumber="8" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005" yearSuffix="a">Lorenz, 2005a</bibRefCitation>
: 262
</treatmentCitation>
<taxonomicName id="4C394D52FFA8FF81E28C4A3534DF6E2C" ID-CoL="M53S" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,502,320,342]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA8FF81E28C4A3534676E2C" box="[151,334,320,342]" italics="true" pageId="7" pageNumber="8">Blennidus crassus</emphasis>
(
<bibRefCitation id="EFA84B20FFA8FF81E3404A3534C76E2C" author="Straneo" box="[347,494,320,342]" pageId="7" pageNumber="8" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA8FF81E3E44A3437EF6E2C" author="Lorenz" box="[511,710,320,342]" page="358" pageId="7" pageNumber="8" year="2005">
<bibRefCitation id="EFA84B20FFA8FF81E3E44A3437BF6E2C" author="Lorenz" box="[511,662,320,342]" pageId="7" pageNumber="8" refString="Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp." type="book" year="2005" yearSuffix="b">Lorenz, 2005b</bibRefCitation>
: 358
</treatmentCitation>
<taxonomicName id="4C394D52FFA8FF81E28C4A2B34DF6E0E" ID-CoL="M53S" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,502,350,373]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA8FF81E28C4A2B34676E0F" box="[151,334,350,373]" italics="true" pageId="7" pageNumber="8">Blennidus crassus</emphasis>
(
<bibRefCitation id="EFA84B20FFA8FF81E3404A2B34C76E0E" author="Straneo" box="[347,494,350,372]" pageId="7" pageNumber="8" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA8FF81E3E44A2B36616E0E" author="Allegro" box="[511,840,350,372]" page="173" pageId="7" pageNumber="8" year="2011">
<bibRefCitation id="EFA84B20FFA8FF81E3E44A2B36316E0E" author="Allegro" box="[511,792,350,372]" pageId="7" pageNumber="8" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino, 2011b</bibRefCitation>
: 173
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA8FF81E28C4AD437456D78" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8B8636D1FFA8FF81E28C4AD437E46EC3" blockId="7.[151,1436,417,838]" box="[151,717,417,442]" pageId="7" pageNumber="8">
<emphasis id="B94DEAC3FFA8FF81E28C4AD4341C6EC0" bold="true" box="[151,309,417,442]" pageId="7" pageNumber="8">
<typeStatus id="54828873FFA8FF81E28C4AD435FB6EC0" box="[151,210,417,442]" pageId="7" pageNumber="8">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32E7641FFA8FF81E3274AD7345B6EC3" box="[316,370,418,441]" name="Peru" pageId="7" pageNumber="8">Peru</collectingCountry>
, Cordillera Cumullca,
<quantity id="4CC19B34FFA8FF81E06A4AD437E06EC3" box="[625,713,417,442]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="8" unit="m" value="3600.0">3600 m</quantity>
.
</paragraph>
<paragraph id="8B8636D1FFA8FF81E2DC4AB036C76EA4" blockId="7.[151,1436,417,838]" box="[199,1006,453,478]" pageId="7" pageNumber="8">
<emphasis id="B94DEAC3FFA8FF81E2DC4AB0349A6EA4" bold="true" box="[199,435,453,478]" pageId="7" pageNumber="8">Examined material.</emphasis>
HT ♀ and 1 PT ♀ (CSt at
<collectionCode id="ED28AE14FFA8FF81E0FB4AB3361E6EA4" box="[736,823,454,478]" pageId="7" pageNumber="8">MSNM</collectionCode>
) (complete TS).
</paragraph>
<paragraph id="8B8636D1FFA8FF81E2DC4A9C37456D78" blockId="7.[151,1436,417,838]" box="[199,620,489,514]" pageId="7" pageNumber="8">
<emphasis id="B94DEAC3FFA8FF81E2DC4A9C342F6D78" bold="true" box="[199,262,489,514]" pageId="7" pageNumber="8">Note.</emphasis>
No male specimens are known.
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA8FF81E2DC497834A66C3C" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8B8636D1FFA8FF81E2DC4978342D6D84" blockId="7.[151,1436,417,838]" pageId="7" pageNumber="8">
<emphasis id="B94DEAC3FFA8FF81E2DC497834496D5C" bold="true" box="[199,352,525,550]" pageId="7" pageNumber="8">Distribution.</emphasis>
The two recorded specimens are from the Andes of Northern
<collectingCountry id="F32E7641FFA8FF81E655497B31AD6D5F" box="[1102,1156,526,549]" name="Peru" pageId="7" pageNumber="8">Peru</collectingCountry>
, Dept. Cajamarca (legit Weyroud 1947). Weyroud probably collected these specimens together with the HT of
<taxonomicName id="4C394D52FFA8FF81E644494731EA6D33" box="[1119,1219,562,585]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="B94DEAC3FFA8FF81E644494731EA6D33" box="[1119,1219,562,585]" italics="true" pageId="7" pageNumber="8">B. negrei</emphasis>
</taxonomicName>
, as the locality and the altitude recorded on the labels are the same.
<taxonomicName id="4C394D52FFA8FF81E0DC492336146D17" box="[711,829,598,621]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA8FF81E0DC492336146D17" box="[711,829,598,621]" italics="true" pageId="7" pageNumber="8">B. crassus</emphasis>
</taxonomicName>
does not seem closely related, based on the external morphology, to
<taxonomicName id="4C394D52FFA8FF81E354490F349E6DEB" box="[335,439,634,657]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="B94DEAC3FFA8FF81E354490F349E6DEB" box="[335,439,634,657]" italics="true" pageId="7" pageNumber="8">B. negrei</emphasis>
</taxonomicName>
; because of the similar habitus, this species is tentatively included in the group of
<taxonomicName id="4C394D52FFA8FF81E79F490F342B6DCF" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA8FF81E79F490F342B6DCF" italics="true" pageId="7" pageNumber="8">B. orbicollis</emphasis>
</taxonomicName>
, which is restricted to the same geographic area (
<bibRefCitation id="EFA84B20FFA8FF81E12549E8315F6DCC" author="Allegro" box="[830,1142,669,694]" pageId="7" pageNumber="8" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino 2011b</bibRefCitation>
), although its aedeagus is still unknown and the attribution to this species group will be therefore confirmed only when male specimens are collected.
</paragraph>
<paragraph id="8B8636D1FFA8FF81E2DC487C34A66C3C" blockId="7.[151,1436,417,838]" pageId="7" pageNumber="8">
<emphasis id="B94DEAC3FFA8FF81E2DC487C34036C58" bold="true" box="[199,298,777,802]" pageId="7" pageNumber="8">Habitat.</emphasis>
The data reported on the labels of the TS specimens (
<quantity id="4CC19B34FFA8FF81E19B487C36F76C58" box="[896,990,777,802]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="8" unit="m" value="3600.0">3,600 m</quantity>
) suggest a preference of this species for the Andean grassland.
</paragraph>
</subSubSection>
</treatment>
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