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<mods:title id="A06A13AF17EB0F89F0202C1DE6629945">Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)</mods:title>
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<mods:namePart id="74FBB5C799E6C3FC5311857FD728D9CF">Hunyadi, András</mods:namePart>
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<emphasis id="B959EAE22F14240EFF12B7C8FE1CFD29" bold="true" box="[151,391,726,753]" italics="true" pageId="52" pageNumber="53">Metalycaeus heudei</emphasis>
(Bavay &amp; Dautzenberg, 1900)
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<figureCitation id="13162A752F14240EFF12B7E5FF7CFCCD" box="[151,231,763,789]" captionStart="FIGURE 41" captionStartId="53.[151,250,1943,1967]" captionTargetBox="[170,1418,181,1920]" captionTargetId="figure-18@53.[170,1418,181,1920]" captionTargetPageId="53" captionText="FIGURE 41. Metalycaeus heudei (Bavay &amp; Dautzenberg, 1900). A: from Kayah State, Myanmar (site JG102); B: holotype of Alycaeus kengtungensis Godwin-Austen, 1914 (NHMUK 1903.7.1.3037)." figureDoi="http://doi.org/10.5281/zenodo.4772049" httpUri="https://zenodo.org/record/4772049/files/figure.png" pageId="52" pageNumber="53">Fig. 41</figureCitation>
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<paragraph id="8B9236F02F14240EFF12B621FCDBFC8F" blockId="52.[151,1436,831,1162]" box="[151,832,831,856]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="AB3D11DE2F14240EFF12B621FCDBFC8F" box="[151,832,831,856]" pageId="52" pageNumber="53">
<treatmentCitation id="0A8C10E12F14240EFF12B621FCA7FC8F" author="Bavay, A. &amp; Dautzenberg, P." box="[151,828,831,856]" page="122" pageId="52" pageNumber="53" year="1900">
<taxonomicName id="4C2D4D732F14240EFF12B621FCA7FC8F" authority="Bavay &amp; Dautzenberg, 1900 a: 121 - 122" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="121 - 122" authorityYear="1900" box="[151,828,831,856]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei" subGenus="Charax">
<emphasis id="B959EAE22F14240EFF12B621FF69FC8F" box="[151,242,831,855]" italics="true" pageId="52" pageNumber="53">Alycaeus</emphasis>
(
<emphasis id="B959EAE22F14240EFE84B621FED6FC8F" box="[257,333,831,855]" italics="true" pageId="52" pageNumber="53">Charax</emphasis>
) 
<emphasis id="B959EAE22F14240EFED9B621FE3BFC8F" box="[348,416,831,855]" italics="true" pageId="52" pageNumber="53">heudei</emphasis>
<bibRefCitation id="EFBC4B012F14240EFE23B65EFCA7FC8F" author="Bavay, A. &amp; Dautzenberg, P." box="[422,828,831,856]" pageId="52" pageNumber="53" pagination="108 - 122" refId="ref24401" refString="Bavay, A. &amp; Dautzenberg, P. (1900 a) Diagnoses de coquilles nouvelles de l'Indo-Chine. Journal de Conchyliologie, 48, 108 - 122." type="journal article" year="1900">Bavay &amp; Dautzenberg, 1900a: 121–122</bibRefCitation>
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.
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<taxonomicName id="4C2D4D732F14240EFF12B640FE3BFCAE" authorityName="Bavay &amp; Dautzenberg" authorityYear="1900" box="[151,416,862,886]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei" subGenus="Charax">
<emphasis id="B959EAE22F14240EFF12B640FF69FCAE" box="[151,242,862,886]" italics="true" pageId="52" pageNumber="53">Alycaeus</emphasis>
(
<emphasis id="B959EAE22F14240EFE84B640FED6FCAE" box="[257,333,862,886]" italics="true" pageId="52" pageNumber="53">Charax</emphasis>
) 
<emphasis id="B959EAE22F14240EFED9B640FE3BFCAE" box="[348,416,862,886]" italics="true" pageId="52" pageNumber="53">heudei</emphasis>
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— 
<treatmentCitation id="0A8C10E12F14240EFE43B640FCCDFCAE" author="Bavay, A. &amp; Dautzenberg, P." box="[454,854,862,886]" page="458" pageId="52" pageNumber="53" year="1900">
<bibRefCitation id="EFBC4B012F14240EFE43B640FCCDFCAE" author="Bavay, A. &amp; Dautzenberg, P." box="[454,854,862,886]" pageId="52" pageNumber="53" pagination="435 - 460" refId="ref24431" refString="Bavay, A. &amp; Dautzenberg, P. (1900 b) Description de coquilles nouvelles de l'Indo-Chine. Journal de Conchyliologie, 48, 435 - 460." type="journal article" year="1900">Bavay &amp; Dautzenberg 1900b: 458–459</bibRefCitation>
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, plate 11, figs 15–18.
</paragraph>
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<treatmentCitation id="0A8C10E12F14240EFF12B662FD35FC4C" author="Bavay, A. &amp; Dautzenberg, P." box="[151,686,892,917]" page="51" pageId="52" pageNumber="53" year="1912">
<taxonomicName id="4C2D4D732F14240EFF12B662FD35FC4C" authority="Bavay &amp; Dautzenberg, 1912: 50 - 51" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="50 - 51" authorityYear="1912" box="[151,686,892,917]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="paviei">
<emphasis id="B959EAE22F14240EFF12B662FEACFC4C" box="[151,311,892,916]" italics="true" pageId="52" pageNumber="53">Alycaeus paviei</emphasis>
<bibRefCitation id="EFBC4B012F14240EFEB8B663FD35FC4C" author="Bavay, A. &amp; Dautzenberg, P." box="[317,686,892,917]" pageId="52" pageNumber="53" pagination="1 - 54" refId="ref24461" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de coquilles nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912: 50–51</bibRefCitation>
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, pl. 4, figs 5–8.
</paragraph>
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<treatmentCitation id="0A8C10E12F14240EFF12B685FD6EFC6B" author="Bavay, A. &amp; Dautzenberg, P." box="[151,757,923,948]" page="51" pageId="52" pageNumber="53" year="1912">
<taxonomicName id="4C2D4D732F14240EFF12B685FD6EFC6B" authority="Bavay &amp; Dautzenberg, 1912: 51" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="51" authorityYear="1912" box="[151,757,923,948]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="variety" species="paviei" variety="minor">
<emphasis id="B959EAE22F14240EFF12B685FEACFC6B" box="[151,311,923,947]" italics="true" pageId="52" pageNumber="53">Alycaeus paviei</emphasis>
var. 
<emphasis id="B959EAE22F14240EFEECB682FE3DFC6B" box="[361,422,924,947]" italics="true" pageId="52" pageNumber="53">minor</emphasis>
<bibRefCitation id="EFBC4B012F14240EFE29B682FD6EFC6B" author="Bavay, A. &amp; Dautzenberg, P." box="[428,757,923,948]" pageId="52" pageNumber="53" pagination="1 - 54" refId="ref24461" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de coquilles nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912: 51</bibRefCitation>
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, pl. 4, fig. 9.
</paragraph>
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<treatmentCitation id="0A8C10E12F14240EFF12B6A4FCB2FC0A" author="Zilch, A." box="[151,809,954,978]" page="146" pageId="52" pageNumber="53" year="1957">
<taxonomicName id="4C2D4D732F14240EFF12B6A4FCB2FC0A" authority="Zilch, 1957: 145 - 146" authorityName="Zilch" authorityPageNumber="145 - 146" authorityYear="1957" box="[151,809,954,978]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="compressicosta" subGenus="Dicharax">
<emphasis id="B959EAE22F14240EFF12B6A4FEB7FC0A" box="[151,300,954,978]" italics="true" pageId="52" pageNumber="53">Chamalycaeus</emphasis>
(
<emphasis id="B959EAE22F14240EFEBEB6A4FE00FC0A" box="[315,411,954,978]" italics="true" pageId="52" pageNumber="53">Dicharax</emphasis>
) 
<emphasis id="B959EAE22F14240EFE2FB6A4FDDEFC09" box="[426,581,954,977]" italics="true" pageId="52" pageNumber="53">compressicosta</emphasis>
<bibRefCitation id="EFBC4B012F14240EFDCEB6A4FCB2FC0A" author="Zilch, A." box="[587,809,954,978]" pageId="52" pageNumber="53" pagination="141 - 150" refId="ref26585" refString="Zilch, A. (1957) Die Typen und Typoide des Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fur Molluskenkunde, 86 (4 / 6), 141 - 150." type="journal article" year="1957">Zilch, 1957: 145–146</bibRefCitation>
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, fig. 33.
</paragraph>
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<treatmentCitation id="0A8C10E12F14240EFF12B6C6FD43FC28" author="Varga, A." box="[151,728,984,1009]" page="167" pageId="52" pageNumber="53" year="1974">
<taxonomicName id="4C2D4D732F14240EFF12B6C6FD43FC28" authority="Varga, 1974: 165 - 167" authorityName="Varga" authorityPageNumber="165 - 167" authorityYear="1974" box="[151,728,984,1009]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="fractus" subGenus="Dicharax">
<emphasis id="B959EAE22F14240EFF12B6C6FEB7FC28" box="[151,300,984,1008]" italics="true" pageId="52" pageNumber="53">Chamalycaeus</emphasis>
(
<emphasis id="B959EAE22F14240EFEBEB6C6FE00FC28" box="[315,411,984,1008]" italics="true" pageId="52" pageNumber="53">Dicharax</emphasis>
) 
<emphasis id="B959EAE22F14240EFE2FB6C7FE6BFC29" box="[426,496,985,1009]" italics="true" pageId="52" pageNumber="53">fractus</emphasis>
<bibRefCitation id="EFBC4B012F14240EFE73B6C7FD43FC28" author="Varga, A." box="[502,728,984,1009]" pageId="52" pageNumber="53" pagination="165 - 167" refId="ref26510" refString="Varga, A. (1974) Eine neue Chamalycaeus - Art aus Vietnam (Gastropoda: Cyclophoridae, Alycaeninae). Folia Historico Naturalia Musei Matraensis, 2, 165 - 167." type="journal article" year="1974">Varga, 1974: 165–167</bibRefCitation>
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, figs 1–5.
</paragraph>
<paragraph id="8B9236F02F14240EFF12B6E9FC85FBD7" blockId="52.[151,1436,831,1162]" box="[151,798,1015,1040]" pageId="52" pageNumber="53">
<treatmentCitation id="0A8C10E12F14240EFF12B6E9FD43FBD7" author="Yang, H. &amp; Fan, Z. &amp; Qiao, D. &amp; He, J." box="[151,728,1015,1040]" page="32" pageId="52" pageNumber="53" year="2012">
<taxonomicName id="4C2D4D732F14240EFF12B6E9FD43FBD7" authority="Yang, Fan, Qiao &amp; He, 2012: 32" authorityName="Yang, Fan, Qiao &amp; He" authorityPageNumber="32" authorityYear="2012" box="[151,728,1015,1040]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="zhuangiyucuii">
<emphasis id="B959EAE22F14240EFF12B6E9FE1CFBD7" box="[151,391,1015,1039]" italics="true" pageId="52" pageNumber="53">Alycaeus zhuangiyucuii</emphasis>
<bibRefCitation id="EFBC4B012F14240EFE09B6E6FD43FBD7" author="Yang, H. &amp; Fan, Z. &amp; Qiao, D. &amp; He, J." box="[396,728,1015,1040]" pageId="52" pageNumber="53" pagination="32 - 33" refId="ref26544" refString="Yang, H., Fan, Z., Qiao, D. &amp; He, J. (2012) Description of four landsnails from China. Shell Discoveries, 1 (1), 32 - 33." type="journal article" year="2012">Yang, Fan, Qiao &amp; He, 2012: 32</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 2.
</paragraph>
<paragraph id="8B9236F02F14240EFF12B108FCEFFBB3" blockId="52.[151,1436,831,1162]" pageId="52" pageNumber="53">
<taxonomicName id="4C2D4D732F14240EFF12B108FEFBFBF6" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[151,352,1046,1070]" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="B959EAE22F14240EFF12B108FEFBFBF6" box="[151,352,1046,1070]" italics="true" pageId="52" pageNumber="53">Metalycaeus heudei</emphasis>
</taxonomicName>
— Páll-Gergely 
<emphasis id="B959EAE22F14240EFD8FB109FDAEFBF6" box="[522,565,1046,1070]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. 2017: 74–84, figs 49C–D, 50–52, 53C–D. (
<taxonomicName id="4C2D4D732F14240EFC77B108FACEFBF6" authority="Bavay &amp; Dautzenberg, 1912" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[1010,1365,1046,1070]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="paviei">
<emphasis id="B959EAE22F14240EFC77B108FBABFBF5" box="[1010,1072,1046,1069]" italics="true" pageId="52" pageNumber="53">paviei</emphasis>
<bibRefCitation id="EFBC4B012F14240EFBB0B108FACEFBF6" author="Bavay, A. &amp; Dautzenberg, P." box="[1077,1365,1046,1070]" pageId="52" pageNumber="53" pagination="1 - 54" refId="ref24461" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de coquilles nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2D4D732F14240EFADBB108FDF8FB95" authority="Bavay &amp; Dautzenberg, 1912" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="variety" species="paviei" variety="minor">
<emphasis id="B959EAE22F14240EFADBB108FA07FBF5" box="[1374,1436,1046,1069]" italics="true" pageId="52" pageNumber="53">paviei</emphasis>
var. 
<emphasis id="B959EAE22F14240EFF70B12BFEA9FB94" box="[245,306,1077,1100]" italics="true" pageId="52" pageNumber="53">minor</emphasis>
<bibRefCitation id="EFBC4B012F14240EFEBFB12BFDF8FB95" author="Bavay, A. &amp; Dautzenberg, P." box="[314,611,1076,1101]" pageId="52" pageNumber="53" pagination="1 - 54" refId="ref24461" refString="Bavay, A. &amp; Dautzenberg, P. (1912) Description de coquilles nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60, 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2D4D732F14240EFDEAB12BFC17FB94" authority="Zilch, 1957" authorityName="Zilch" authorityYear="1957" box="[623,908,1076,1100]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="compressicosta">
<emphasis id="B959EAE22F14240EFDEAB12BFC91FB94" box="[623,778,1077,1100]" italics="true" pageId="52" pageNumber="53">compressicosta</emphasis>
<bibRefCitation id="EFBC4B012F14240EFC96B12AFC17FB94" author="Zilch, A." box="[787,908,1076,1100]" pageId="52" pageNumber="53" pagination="141 - 150" refId="ref26585" refString="Zilch, A. (1957) Die Typen und Typoide des Natur-Museums Senckenberg, 20: Mollusca, Cyclophoridae, Alycaeinae. Archiv fur Molluskenkunde, 86 (4 / 6), 141 - 150." type="journal article" year="1957">Zilch, 1957</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2D4D732F14240EFC1DB12BFBF8FB95" authority="Varga, 1974" authorityName="Varga" authorityYear="1974" box="[920,1123,1077,1101]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="fractus">
<emphasis id="B959EAE22F14240EFC1DB12BFC45FB95" box="[920,990,1077,1101]" italics="true" pageId="52" pageNumber="53">fractus</emphasis>
<bibRefCitation id="EFBC4B012F14240EFC63B12BFBF8FB95" author="Varga, A." box="[998,1123,1077,1101]" pageId="52" pageNumber="53" pagination="165 - 167" refId="ref26510" refString="Varga, A. (1974) Eine neue Chamalycaeus - Art aus Vietnam (Gastropoda: Cyclophoridae, Alycaeninae). Folia Historico Naturalia Musei Matraensis, 2, 165 - 167." type="journal article" year="1974">Varga, 1974</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C2D4D732F14240EFB12B12AFEEDFBB3" authority="Yang, Fan, Qiao &amp; He 2012" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="subSpecies" species="heudei" subSpecies="zhuangiyucuii">
<emphasis id="B959EAE22F14240EFB12B12AFABEFB94" box="[1175,1317,1076,1100]" italics="true" pageId="52" pageNumber="53">zhuangiyucuii</emphasis>
<bibRefCitation id="EFBC4B012F14240EFAA8B12BFA07FB94" author="Yang, H. &amp; Fan, Z. &amp; Qiao, D. &amp; He, J." box="[1325,1436,1077,1101]" pageId="52" pageNumber="53" pagination="32 - 33" refId="ref26544" refString="Yang, H., Fan, Z., Qiao, D. &amp; He, J. (2012) Description of four landsnails from China. Shell Discoveries, 1 (1), 32 - 33." type="journal article" year="2012">Yang, Fan,</bibRefCitation>
Qiao &amp; He, 2012
</taxonomicName>
are synonyms; 
<bibRefCitation id="EFBC4B012F14240EFD90B14DFC87FBB3" author="Inkhavilay, K. &amp; Sutcharit, C. &amp; Bantaowong, U. &amp; Chanabun, R. &amp; Siriwut, W. &amp; Srisonchai, R. &amp; Pholyotha, A. &amp; Jirapatrasilp, P. &amp; Panha, S." box="[533,796,1107,1131]" pageId="52" pageNumber="53" pagination="1 - 166" refId="ref25340" refString="Inkhavilay, K., Sutcharit, C., Bantaowong, U., Chanabun, R., Siriwut, W., Srisonchai, R., Pholyotha, A., Jirapatrasilp, P. &amp; Panha, S. (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834, 1 - 166. https: // doi. org / 10.3897 / zookeys. 834.28800" type="journal article" year="2019">
Inkhavilay 
<emphasis id="B959EAE22F14240EFD0DB14AFD2EFBB3" box="[648,693,1107,1131]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. 2019: 16
</bibRefCitation>
, fig. 6C.
</paragraph>
<paragraph id="8B9236F02F14240EFF12B16CFB8BFB52" blockId="52.[151,1436,831,1162]" box="[151,1040,1138,1162]" pageId="52" pageNumber="53">
<taxonomicName id="4C2D4D732F14240EFF12B16CFD05FB52" authority="Godwin-Austen, 1914: 409" authorityName="Godwin-Austen" authorityPageNumber="409" authorityYear="1914" box="[151,670,1138,1162]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="kengtungensis">
<emphasis id="B959EAE22F14240EFF12B16CFE19FB52" box="[151,386,1138,1162]" italics="true" pageId="52" pageNumber="53">Alycaeus kengtungensis</emphasis>
Godwin-Austen, 1914: 409
</taxonomicName>
, plate 139, figs 6, 6a. 
<emphasis id="B959EAE22F14240EFCF8B16CFB8BFB52" bold="true" box="[893,1040,1138,1162]" pageId="52" pageNumber="53">new synonym</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F14240EFF12B1A6FAB7F9CE" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8B9236F02F14240EFF12B1A6FAB7F9CE" blockId="52.[151,1437,1208,1846]" pageId="52" pageNumber="53">
<emphasis id="B959EAE22F14240EFF12B1A6FE1FFB0A" bold="true" box="[151,388,1208,1234]" pageId="52" pageNumber="53">Material examined.</emphasis>
<materialsCitation id="3B453CAD2F14240EFE0AB1A6FE92FAC2" ID-GBIF-Occurrence="3128117331" collectionCode="NHMUK" collectorName="Woodthorpe" country="Myanmar" location="Shan State" pageId="52" pageNumber="53" specimenCode="NHMUK 1903.7" specimenCount="2" stateProvince="Shan" typeStatus="holotype">
<emphasis id="B959EAE22F14240EFE0AB1A6FD9EFB0A" bold="true" box="[399,517,1208,1234]" pageId="52" pageNumber="53">
<typeStatus id="549688522F14240EFE0AB1A6FD9BFB0A" box="[399,512,1208,1234]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F33A76602F14240EFD95B1A7FDE4FB0B" box="[528,639,1209,1235]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
, 
<collectingRegion id="49E9F8122F14240EFD0BB1A6FC97FB0A" box="[654,780,1208,1234]" country="Myanmar" name="Shan" pageId="52" pageNumber="53">Shan State</collectingRegion>
, “Kengtung, Siam Frontier, leg. 
<collectorName id="26D853262F14240EFB0FB1A6FA81FB0A" box="[1162,1306,1208,1234]" pageId="52" pageNumber="53">Woodthorpe</collectorName>
”, 
<specimenCode id="DB8B9E8B2F14240EFAB7B1A7FF7DFB2E" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="52" pageNumber="53" type="Museum">NHMUK 1903.7</specimenCode>
.1.3037 (
<typeStatus id="549688522F14240EFECDB1C2FE37FB2E" box="[328,428,1244,1270]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C2D4D732F14240EFE56B1C2FD09FB2E" authorityName="Godwin-Austen" authorityYear="1914" box="[467,658,1244,1270]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="kengtungensis">
<emphasis id="B959EAE22F14240EFE56B1C2FD09FB2E" box="[467,658,1244,1270]" italics="true" pageId="52" pageNumber="53">A. kengtungensis</emphasis>
</taxonomicName>
). For the examined type specimens of other taxa see Páll-Gergely 
<emphasis id="B959EAE22F14240EFA0DB1C3FF36FAC2" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. (2017)
</materialsCitation>
. 
<emphasis id="B959EAE22F14240EFE96B01EFE4BFAC2" bold="true" box="[275,464,1280,1306]" pageId="52" pageNumber="53">Other material:</emphasis>
<materialsCitation id="3B453CAD2F14240EFE53B01EFBFEFAE6" ID-GBIF-Occurrence="3128117339" collectingDate="2019-02-10" collectionCode="JG" collectorName="J. Grego" country="Myanmar" county="Hpruso Distr." latitude="19.352016" location="Bo Pha Gu Cave" longLatPrecision="1" longitude="97.06265" municipality="Maw Thi Do Village" pageId="52" pageNumber="53" specimenCode="JG10" specimenCount="1" stateProvince="Kayah">
4 shells, 
<collectingCountry id="F33A76602F14240EFDBCB01FFD33FAC3" box="[569,680,1281,1307]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
, 
<collectingRegion id="49E9F8122F14240EFD37B01EFCA6FAC2" box="[690,829,1280,1306]" country="Myanmar" name="Kayah" pageId="52" pageNumber="53">Kayah State</collectingRegion>
, 
<collectingCounty id="62F34E7C2F14240EFCCDB01FFC79FAC2" box="[840,994,1280,1307]" pageId="52" pageNumber="53">Hpruso Distr.</collectingCounty>
, 
<collectingMunicipality id="6BF6AC8A2F14240EFC68B01FFB4CFAC2" box="[1005,1239,1280,1307]" pageId="52" pageNumber="53">Maw Thi Do Village</collectingMunicipality>
, 
<location id="8EF2602B2F14240EFB64B01FFA06FAC2" LSID="urn:lsid:plazi:treatment:038487E62F14240EFF12B7C8FED8F8EE:8EF2602B2F14240EFB64B01FFA06FAC2" box="[1249,1437,1280,1307]" country="Myanmar" county="Hpruso Distr." latitude="19.352016" longLatPrecision="1" longitude="97.06265" municipality="Maw Thi Do Village" name="Bo Pha Gu Cave" pageId="52" pageNumber="53" stateProvince="Kayah">Bo Pha Gu Cave</location>
sink, 
<geoCoordinate id="EE1950372F14240EFF50B03AFEF2FAE6" box="[213,361,1316,1342]" degrees="19" direction="north" minutes="21.121" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="19.352016">19°21.121’N</geoCoordinate>
, 
<geoCoordinate id="EE1950372F14240EFEF3B03AFE6CFAE7" box="[374,503,1316,1343]" degrees="97" direction="east" minutes="3.759" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="97.06265">97°3.759’E</geoCoordinate>
(locality code: 
<specimenCode id="DB8B9E8B2F14240EFD2FB03AFD72FAE6" box="[682,745,1316,1342]" collectionCode="JG" pageId="52" pageNumber="53">JG10</specimenCode>
), 
<date id="FF9310302F14240EFD78B03AFC0DFAE6" box="[765,918,1316,1342]" pageId="52" pageNumber="53" value="2019-02-10">
<collectingDate id="EFD7E9D82F14240EFD78B03AFC0DFAE6" box="[765,918,1316,1342]" pageId="52" pageNumber="53" value="2019-02-10">10 Feb. 2019</collectingDate>
</date>
, 
<collectorName id="26D853262F14240EFC27B03BFB98FAE6" box="[930,1027,1316,1343]" pageId="52" pageNumber="53">J. Grego</collectorName>
leg., 
<collectionCode id="ED3CAE352F14240EFBC6B03AFBFEFAE6" box="[1091,1125,1316,1342]" pageId="52" pageNumber="53">JG</collectionCode>
</materialsCitation>
; 
<materialsCitation id="3B453CAD2F14240EFBF6B03AFC0BFA5E" ID-GBIF-Occurrence="3128117353" collectingDate="2019-12-12" collectionCode="JG" collectorName="J. Grego" country="Myanmar" county="Hpruso Distr." latitude="19.382767" location="Han Li Village" longLatPrecision="1" longitude="97.03588" municipality="Maw Thi Do" pageId="52" pageNumber="53" specimenCode="JG100" specimenCount="1" stateProvince="Kayah">
2 shells, 
<collectingCountry id="F33A76602F14240EFB5DB03BFADCFAE7" box="[1240,1351,1317,1343]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
, 
<collectingRegion id="49E9F8122F14240EFAD6B03AFF49FABA" country="Myanmar" name="Kayah" pageId="52" pageNumber="53">Kayah State</collectingRegion>
, 
<collectingCounty id="62F34E7C2F14240EFF65B057FEE7FABA" box="[224,380,1352,1379]" pageId="52" pageNumber="53">Hpruso Distr.</collectingCounty>
, 
<collectingMunicipality id="6BF6AC8A2F14240EFE0CB057FDBAFABB" box="[393,545,1352,1379]" pageId="52" pageNumber="53">Maw Thi Do</collectingMunicipality>
, road towards 
<location id="8EF2602B2F14240EFD49B057FCE2FABA" LSID="urn:lsid:plazi:treatment:038487E62F14240EFF12B7C8FED8F8EE:8EF2602B2F14240EFD49B057FCE2FABA" box="[716,889,1352,1379]" country="Myanmar" county="Hpruso Distr." latitude="19.382767" longLatPrecision="1" longitude="97.03588" municipality="Maw Thi Do" name="Han Li Village" pageId="52" pageNumber="53" stateProvince="Kayah">Han Li Village</location>
, under bridge over 
<location id="8EF2602B2F14240EFBD9B056FB6DFABA" LSID="urn:lsid:plazi:treatment:038487E62F14240EFF12B7C8FED8F8EE:8EF2602B2F14240EFBD9B056FB6DFABA" box="[1116,1270,1352,1378]" country="Myanmar" county="Hpruso Distr." latitude="19.382767" longLatPrecision="1" longitude="97.03588" municipality="Maw Thi Do" name="Phruno River" pageId="52" pageNumber="53" stateProvince="Kayah">Phruno River</location>
, 
<geoCoordinate id="EE1950372F14240EFA81B056FA03FABA" box="[1284,1432,1352,1378]" degrees="19" direction="north" minutes="22.966" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="19.382767">19°22.966’N</geoCoordinate>
, 
<geoCoordinate id="EE1950372F14240EFF12B072FEBDFA5F" box="[151,294,1388,1415]" degrees="97" direction="east" minutes="02.153" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="97.03588">97°02.153’E</geoCoordinate>
(locality code: 
<specimenCode id="DB8B9E8B2F14240EFE57B072FD84FA5E" box="[466,543,1388,1414]" collectionCode="JG" pageId="52" pageNumber="53">JG100</specimenCode>
), 
<date id="FF9310302F14240EFDB7B072FD52FA5E" box="[562,713,1388,1415]" pageId="52" pageNumber="53" value="2019-12-12">
<collectingDate id="EFD7E9D82F14240EFDB7B072FD52FA5E" box="[562,713,1388,1415]" pageId="52" pageNumber="53" value="2019-12-12">12 Dec. 2019</collectingDate>
</date>
, 
<collectorName id="26D853262F14240EFD51B073FCA9FA5E" box="[724,818,1388,1415]" pageId="52" pageNumber="53">J. Grego</collectorName>
leg., 
<collectionCode id="ED3CAE352F14240EFCEBB072FC0BFA5E" box="[878,912,1388,1414]" pageId="52" pageNumber="53">JG</collectionCode>
</materialsCitation>
; 
<materialsCitation id="3B453CAD2F14240EFC1EB072FD70FA16" ID-GBIF-Occurrence="3128117397" collectingDate="2019-12-12" collectionCode="JG" collectorName="J. Grego" country="Myanmar" county="Hpruso Distr." latitude="19.383516" location="Han Li Village" longLatPrecision="1" longitude="97.03513" municipality="Maw Thi Do" pageId="52" pageNumber="53" specimenCode="JG101" specimenCount="1" stateProvince="Kayah">
1 shell, 
<collectingCountry id="F33A76602F14240EFC77B073FBFAFA5F" box="[1010,1121,1389,1415]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
, 
<collectingRegion id="49E9F8122F14240EFBEEB072FB6EFA5E" box="[1131,1269,1388,1414]" country="Myanmar" name="Kayah" pageId="52" pageNumber="53">Kayah State</collectingRegion>
, 
<collectingCounty id="62F34E7C2F14240EFB7AB073FA03FA5E" box="[1279,1432,1388,1415]" pageId="52" pageNumber="53">Hpruso Distr.</collectingCounty>
, 
<collectingMunicipality id="6BF6AC8A2F14240EFF12B08FFEB0FA73" box="[151,299,1424,1451]" pageId="52" pageNumber="53">Maw Thi Do</collectingMunicipality>
, road towards 
<location id="8EF2602B2F14240EFE55B08FFDE2FA72" LSID="urn:lsid:plazi:treatment:038487E62F14240EFF12B7C8FED8F8EE:8EF2602B2F14240EFE55B08FFDE2FA72" box="[464,633,1424,1451]" country="Myanmar" county="Hpruso Distr." latitude="19.383516" longLatPrecision="1" longitude="97.03513" municipality="Maw Thi Do" name="Han Li Village" pageId="52" pageNumber="53" stateProvince="Kayah">Han Li Village</location>
, rocks above bridge over 
<location id="8EF2602B2F14240EFC1EB08EFBA8FA72" LSID="urn:lsid:plazi:treatment:038487E62F14240EFF12B7C8FED8F8EE:8EF2602B2F14240EFC1EB08EFBA8FA72" box="[923,1075,1424,1450]" country="Myanmar" county="Hpruso Distr." latitude="19.383516" longLatPrecision="1" longitude="97.03513" municipality="Maw Thi Do" name="Phruno River" pageId="52" pageNumber="53" stateProvince="Kayah">Phruno River</location>
, 
<geoCoordinate id="EE1950372F14240EFBBAB08EFB49FA72" box="[1087,1234,1424,1450]" degrees="19" direction="north" minutes="23.011" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="19.383516">19°23.011’N</geoCoordinate>
, 
<geoCoordinate id="EE1950372F14240EFB5BB08EFAF7FA72" box="[1246,1388,1424,1450]" degrees="97" direction="east" minutes="02.108" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="97.03513">97°02.108’E</geoCoordinate>
(locality code: 
<specimenCode id="DB8B9E8B2F14240EFEA7B0AAFEF5FA16" box="[290,366,1460,1486]" collectionCode="JG" pageId="52" pageNumber="53">JG101</specimenCode>
), 
<date id="FF9310302F14240EFE06B0AAFD85FA16" box="[387,542,1460,1487]" pageId="52" pageNumber="53" value="2019-12-12">
<collectingDate id="EFD7E9D82F14240EFE06B0AAFD85FA16" box="[387,542,1460,1487]" pageId="52" pageNumber="53" value="2019-12-12">12 Dec. 2019</collectingDate>
</date>
, 
<collectorName id="26D853262F14240EFDAFB0ABFD12FA16" box="[554,649,1460,1487]" pageId="52" pageNumber="53">J. Grego</collectorName>
leg., 
<collectionCode id="ED3CAE352F14240EFD4CB0AAFD70FA16" box="[713,747,1460,1486]" pageId="52" pageNumber="53">JG</collectionCode>
</materialsCitation>
; 
<materialsCitation id="3B453CAD2F14240EFD7DB0AAFCE2F9CF" ID-GBIF-Occurrence="3128117388" collectingDate="2019-12-12" collectionCode="JG, HNHM" collectorName="J. Grego" country="Myanmar" county="Hpruso Distr." latitude="19.379066" location="Entrance of Phruno River Cave" longLatPrecision="1" longitude="97.04283" municipality="Maw Thi Do Village" pageId="52" pageNumber="53" specimenCode="JG102, HNHM 104867" specimenCount="1" stateProvince="Kayah">
1 shell, 
<collectingCountry id="F33A76602F14240EFCD4B0ABFC5BFA17" box="[849,960,1461,1487]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
, 
<collectingRegion id="49E9F8122F14240EFC49B0AAFBC3FA16" box="[972,1112,1460,1486]" country="Myanmar" name="Kayah" pageId="52" pageNumber="53">Kayah State</collectingRegion>
, 
<collectingCounty id="62F34E7C2F14240EFBE1B0ABFB64FA16" box="[1124,1279,1460,1487]" pageId="52" pageNumber="53">Hpruso Distr.</collectingCounty>
, 
<collectingMunicipality id="6BF6AC8A2F14240EFA8FB0ABFF70FA2A" pageId="52" pageNumber="53">Maw Thi Do Village</collectingMunicipality>
, 
<location id="8EF2602B2F14240EFF70B0C7FDCEFA2A" LSID="urn:lsid:plazi:treatment:038487E62F14240EFF12B7C8FED8F8EE:8EF2602B2F14240EFF70B0C7FDCEFA2A" box="[245,597,1496,1523]" country="Myanmar" county="Hpruso Distr." latitude="19.379066" longLatPrecision="1" longitude="97.04283" municipality="Maw Thi Do Village" name="Entrance of Phruno River Cave" pageId="52" pageNumber="53" stateProvince="Kayah">Entrance of Phruno River Cave</location>
, 
<geoCoordinate id="EE1950372F14240EFDE5B0C6FD6FFA2A" box="[608,756,1496,1522]" degrees="19" direction="north" minutes="22.744" orientation="latitude" pageId="52" pageNumber="53" precision="1" value="19.379066">19°22.744’N</geoCoordinate>
, 
<geoCoordinate id="EE1950372F14240EFD7AB0C6FC15FA2B" box="[767,910,1496,1523]" degrees="97" direction="east" minutes="02.570" orientation="longitude" pageId="52" pageNumber="53" precision="1" value="97.04283">97°02.570’E</geoCoordinate>
(locality code: 
<specimenCode id="DB8B9E8B2F14240EFBBEB0C6FB12FA2A" box="[1083,1161,1496,1522]" collectionCode="JG" pageId="52" pageNumber="53">JG102</specimenCode>
), 
<date id="FF9310302F14240EFB1EB0C6FAA8FA2A" box="[1179,1331,1496,1523]" pageId="52" pageNumber="53" value="2019-12-12">
<collectingDate id="EFD7E9D82F14240EFB1EB0C6FAA8FA2A" box="[1179,1331,1496,1523]" pageId="52" pageNumber="53" value="2019-12-12">12 Dec. 2019</collectingDate>
</date>
, 
<collectorName id="26D853262F14240EFABBB0C7FA07FA2A" box="[1342,1436,1496,1523]" pageId="52" pageNumber="53">J. Grego</collectorName>
leg., 
<specimenCode id="DB8B9E8B2F14240EFF4BB0E3FEE4F9CE" box="[206,383,1532,1559]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="52" pageNumber="53">HNHM 104867</specimenCode>
(photographed shell: 
<quantity id="4CD59B152F14240EFDF1B0E2FD41F9CF" box="[628,730,1532,1559]" metricMagnitude="-3" metricUnit="m" metricValue="4.77" pageId="52" pageNumber="53" unit="mm" value="4.77">4.77 mm</quantity>
, H: 
<quantity id="4CD59B152F14240EFC8DB0E2FCEBF9CF" box="[776,880,1532,1559]" metricMagnitude="-3" metricUnit="m" metricValue="3.66" pageId="52" pageNumber="53" unit="mm" value="3.66">3.66 mm</quantity>
)
</materialsCitation>
; 
<materialsCitation id="3B453CAD2F14240EFC01B0E2FAB2F9CE" ID-GBIF-Occurrence="3128117348" box="[900,1321,1532,1559]" collectingDate="2019-12-12" collectionCode="JG" collectorName="J. Grego" country="Myanmar" county="Hpruso Distr." latitude="19.379066" location="Entrance of Phruno River Cave" longLatPrecision="1" longitude="97.04283" municipality="Maw Thi Do Village" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Kayah">
17 shells, same data as preceding, 
<collectionCode id="ED3CAE352F14240EFA83B0E2FAB2F9CE" box="[1286,1321,1532,1558]" pageId="52" pageNumber="53">JG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F14240EFF42B33EFA88F987" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8B9236F02F14240EFF42B33EFA88F987" blockId="52.[151,1437,1208,1846]" pageId="52" pageNumber="53">
<emphasis id="B959EAE22F14240EFF42B33EFD38F9E2" bold="true" box="[199,675,1568,1594]" pageId="52" pageNumber="53">
Diagnosis of specimens from 
<collectingCountry id="F33A76602F14240EFDA2B33EFD05F9E2" box="[551,670,1568,1594]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
.
</emphasis>
A medium-sized to large species with very long R2 having typical 
<taxonomicName id="4C2D4D732F14240EFF12B35AFEBFF986" authorityName="Pilsbry" authorityYear="1900" box="[151,292,1604,1630]" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F14240EFF12B35AFEBFF986" box="[151,292,1604,1630]" italics="true" pageId="52" pageNumber="53">Metalycaeus</emphasis>
</taxonomicName>
- 
<typeStatus id="549688522F14240EFEA9B35BFEC5F987" box="[300,350,1605,1631]" pageId="52" pageNumber="53">type</typeStatus>
elevated, thin, widely spaced ribs, a narrow swelling on R3, and a rounded aperture.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F14240EFF42B376FBF0F95B" box="[199,1131,1640,1667]" pageId="52" pageNumber="53" type="description">
<paragraph id="8B9236F02F14240EFF42B376FBF0F95B" blockId="52.[151,1437,1208,1846]" box="[199,1131,1640,1667]" pageId="52" pageNumber="53">
<emphasis id="B959EAE22F14240EFF42B376FEE5F95A" bold="true" box="[199,382,1640,1666]" pageId="52" pageNumber="53">Measurements.</emphasis>
D: 
<quantity id="4CD59B152F14240EFE22B376FDD7F95B" box="[423,588,1640,1667]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.05" metricValueMin="4.65" pageId="52" pageNumber="53" unit="mm" value="4.85" valueMax="5.05" valueMin="4.65">4.65–5.05 mm</quantity>
, H: 
<quantity id="4CD59B152F14240EFDFEB376FC85F95B" box="[635,798,1640,1667]" metricMagnitude="-3" metricUnit="m" metricValue="3.685" metricValueMax="3.71" metricValueMin="3.66" pageId="52" pageNumber="53" unit="mm" value="3.685" valueMax="3.71" valueMin="3.66">3.66–3.71 mm</quantity>
(newly collected specimens).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F14240EFF42B392FED8F8EE" pageId="52" pageNumber="53" type="discussion">
<paragraph id="8B9236F02F14240EFF42B392FED8F8EE" blockId="52.[151,1437,1208,1846]" pageId="52" pageNumber="53">
<emphasis id="B959EAE22F14240EFF42B392FEA0F97E" bold="true" box="[199,315,1676,1702]" pageId="52" pageNumber="53">Remarks.</emphasis>
<taxonomicName id="4C2D4D732F14240EFEC4B392FD84F97E" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[321,543,1676,1702]" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="B959EAE22F14240EFEC4B392FD84F97E" box="[321,543,1676,1702]" italics="true" pageId="52" pageNumber="53">Metalycaeus heudei</emphasis>
</taxonomicName>
is a widely distributed, variable species reported from 
<collectingCountry id="F33A76602F14240EFBF8B392FB58F97E" box="[1149,1219,1676,1702]" name="China" pageId="52" pageNumber="53">China</collectingCountry>
, 
<collectingCountry id="F33A76602F14240EFB49B392FAB0F97E" box="[1228,1323,1676,1702]" name="Vietnam" pageId="52" pageNumber="53">Vietnam</collectingCountry>
and 
<collectingCountry id="F33A76602F14240EFADAB393FA02F97F" box="[1375,1433,1677,1703]" name="Laos" pageId="52" pageNumber="53">Laos</collectingCountry>
. 
<taxonomicName id="4C2D4D732F14240EFF12B3AEFE39F912" authorityName="Godwin-Austen" authorityYear="1914" box="[151,418,1712,1738]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="kengtungensis">
<emphasis id="B959EAE22F14240EFF12B3AEFE39F912" box="[151,418,1712,1738]" italics="true" pageId="52" pageNumber="53">Alycaeus kengtungensis</emphasis>
</taxonomicName>
(
<figureCitation id="13162A752F14240EFE34B3AEFD8EF912" box="[433,533,1712,1738]" captionStart="FIGURE 41" captionStartId="53.[151,250,1943,1967]" captionTargetBox="[170,1418,181,1920]" captionTargetId="figure-18@53.[170,1418,181,1920]" captionTargetPageId="53" captionText="FIGURE 41. Metalycaeus heudei (Bavay &amp; Dautzenberg, 1900). A: from Kayah State, Myanmar (site JG102); B: holotype of Alycaeus kengtungensis Godwin-Austen, 1914 (NHMUK 1903.7.1.3037)." figureDoi="http://doi.org/10.5281/zenodo.4772049" httpUri="https://zenodo.org/record/4772049/files/figure.png" pageId="52" pageNumber="53">Fig. 41B</figureCitation>
) from the eastern border of 
<collectingRegion id="49E9F8122F14240EFCCAB3AEFC53F912" box="[847,968,1712,1738]" country="Myanmar" name="Shan" pageId="52" pageNumber="53">Shan State</collectingRegion>
, 
<collectingCountry id="F33A76602F14240EFC56B3AFFBD9F913" box="[979,1090,1713,1739]" name="Myanmar" pageId="52" pageNumber="53">Myanmar</collectingCountry>
, and the newly collected shells from 
<collectingRegion id="49E9F8122F14240EFF52B3CAFEFAF936" box="[215,353,1748,1774]" country="Myanmar" name="Kayah" pageId="52" pageNumber="53">Kayah State</collectingRegion>
(
<figureCitation id="13162A752F14240EFEF7B3CAFE2CF937" box="[370,439,1748,1775]" captionStart="FIGURE 5" captionStartId="9.[151,250,1831,1855]" captionTargetBox="[164,1423,181,1808]" captionTargetId="figure-18@9.[164,1423,181,1808]" captionTargetPageId="9" captionText="FIGURE 5. Distribution of alycaeid species. Inverted empty triangle: Dicharax cyclostoma Páll-Gergely, n. sp., empty triangle: Chamalycaeus spiratus Páll-Gergely, n. sp., filled triangle: Metalycaeus bicarinatus Páll-Gergely &amp; Hunyadi, n. sp., inverted filled triangles: Chamalycaeus erawan Páll-Gergely &amp; Hunyadi n. sp., empty circles: Metalycaeus aries Páll-Gergely &amp; Auffenberg, n. sp., filled circles: Metalycaeus heudei (Bavay &amp; Dautzenberg, 1900) (the one in the north indicates the type locality of Alycaeus kengtungensis Godwin-Austen, 1914), square: Metalycaeus pygmachos Páll-Gergely &amp; Hunyadi, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4771970" httpUri="https://zenodo.org/record/4771970/files/figure.png" pageId="52" pageNumber="53">Fig. 5</figureCitation>
) fall within the known morphological variation of 
<taxonomicName id="4C2D4D732F14240EFB86B3CAFB7EF936" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[1027,1253,1748,1774]" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="B959EAE22F14240EFB86B3CAFB7EF936" box="[1027,1253,1748,1774]" italics="true" pageId="52" pageNumber="53">Metalycaeus heudei</emphasis>
</taxonomicName>
. Therefore, we consider 
<taxonomicName id="4C2D4D732F14240EFF79B3E6FE22F8CA" authorityName="Godwin-Austen" authorityYear="1914" box="[252,441,1784,1810]" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="kengtungensis">
<emphasis id="B959EAE22F14240EFF79B3E6FE22F8CA" box="[252,441,1784,1810]" italics="true" pageId="52" pageNumber="53">A. kengtungensis</emphasis>
</taxonomicName>
a new synonym of 
<taxonomicName id="4C2D4D732F14240EFD0AB3E7FD64F8CA" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[655,767,1784,1810]" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="B959EAE22F14240EFD0AB3E7FD64F8CA" box="[655,767,1784,1810]" italics="true" pageId="52" pageNumber="53">M. heudei</emphasis>
</taxonomicName>
. With this, 
<taxonomicName id="4C2D4D732F14240EFCF8B3E7FC77F8CA" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[893,1004,1784,1810]" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="heudei">
<emphasis id="B959EAE22F14240EFCF8B3E7FC77F8CA" box="[893,1004,1784,1810]" italics="true" pageId="52" pageNumber="53">M. heudei</emphasis>
</taxonomicName>
is the most widely distributed alycaeid species known.
</paragraph>
</subSubSection>
</treatment>
</document>