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<mods:title id="2FFD3A626EBEC3C503B86AED3E624106">Crinoids of the stalked family Hyocrinidae (Echinodermata) collected by the USNS Eltanin in the Southern Ocean</mods:title>
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<taxonomicName id="4C5667B0FFD4FFF75483FF4DA9EFFF17" ID-CoL="8GHQM" authority="Vaney" authorityName="Vaney" box="[151,707,152,178]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="14" pageNumber="395" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFD4FFF75483FF4DA943FF17" bold="true" box="[151,623,152,178]" italics="true" pageId="14" pageNumber="395">Ptilocrinus (Chambersaecrinus) brucei</emphasis>
Vaney
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<bibRefCitation id="EFC761C2FFD4FFF756FEFF4DA8A6FF17" author="Wilton" box="[746,906,152,178]" pageId="14" pageNumber="395" refString="Wilton, D. W. (1908) Resultats de l'Expedition nationale Antarctique du S. Y. Scotia, 1902 - 1904, 4, Zoologie, 1, 1 - 133." type="journal article" year="1908">Wilton, 1908</bibRefCitation>
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<paragraph id="8BE91C33FFD4FFF75483FF6FAAEBFF76" blockId="14.[151,906,152,211]" box="[151,455,186,211]" pageId="14" pageNumber="395">
<figureCitation id="136D00B6FFD4FFF75483FF6FAA00FF76" box="[151,300,186,211]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStart-5="FIGURE 14" captionStart-6="FIGURE 15" captionStart-7="FIGURE 16" captionStartId-0="14.[151,250,1495,1517]" captionStartId-1="16.[151,250,1533,1555]" captionStartId-2="17.[151,250,1593,1615]" captionStartId-3="18.[151,250,1272,1294]" captionStartId-4="19.[151,250,1743,1765]" captionStartId-5="20.[151,250,1738,1760]" captionStartId-6="21.[151,250,1868,1890]" captionStartId-7="22.[151,250,1830,1852]" captionTargetBox-0="[295,1291,921,1473]" captionTargetBox-1="[151,1436,589,1511]" captionTargetBox-2="[215,1372,193,1568]" captionTargetBox-3="[336,1245,373,1243]" captionTargetBox-4="[151,1436,193,1720]" captionTargetBox-5="[177,1409,913,1716]" captionTargetBox-6="[242,1344,565,1845]" captionTargetBox-7="[151,1436,913,1808]" captionTargetId-0="figure@14.[295,1291,921,1474]" captionTargetId-1="figure@16.[151,1436,589,1511]" captionTargetId-2="figure@17.[215,1372,193,1569]" captionTargetId-3="figure@18.[336,1245,373,1243]" captionTargetId-4="figure@19.[151,1436,193,1722]" captionTargetId-5="figure@20.[177,1410,913,1717]" captionTargetId-6="figure@21.[242,1345,565,1846]" captionTargetId-7="figure@22.[151,1436,913,1809]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionTargetPageId-5="20" captionTargetPageId-6="21" captionTargetPageId-7="22" captionText-0="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." captionText-1="FIGURE 10. Brachial muscular synarthries in Ptilocrinus brucei. A – D: specimen 1005438; E – F: specimen 1005439; A: episynostosial brachial; B – D: free brachial, B: side view showing pinnule socket, C – D: oblique adoral view, D: detail of adoral process; E – F: transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea." captionText-2="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." captionText-3="FIGURE 12. Proximal stalk symplexies in large specimens of Ptilocrinus brucei. A: specimen USNM 1005453, B – D: specimen USNM 1005439; A: proximalmost columnal; B: at a distance of 5.0 mm from aboral cup; C: at a distance of 6.0 mm from aboral cup; D: at a distance of 22.0 mm from aboral cup." captionText-4="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." captionText-5="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." captionText-6="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." captionText-7="FIGURE 16. Columnal articulations in mid and distal stalk of Ptilocrinus brucei paratype. A: symplexy in proximal mesistele, B, D: transition between symplexy and syzygy; B: in distal mesistele; D: in proximal dististele; C, E: in distalmost stalk at a distance of 36 mm from D." httpUri-0="https://zenodo.org/record/243058/files/figure.png" httpUri-1="https://zenodo.org/record/243059/files/figure.png" httpUri-2="https://zenodo.org/record/243060/files/figure.png" httpUri-3="https://zenodo.org/record/243061/files/figure.png" httpUri-4="https://zenodo.org/record/243062/files/figure.png" httpUri-5="https://zenodo.org/record/243063/files/figure.png" httpUri-6="https://zenodo.org/record/243064/files/figure.png" httpUri-7="https://zenodo.org/record/243065/files/figure.png" pageId="14" pageNumber="395">Figures 9–16</figureCitation>
, 
<tableCitation id="C6D42988FFD4FFF7552CFF6FAAEFFF76" box="[312,451,186,211]" captionStart-0="TABLE 9" captionStart-1="TABLE 10" captionStart-2="TABLE 11" captionStart-3="TABLE 12" captionStart-4="TABLE 13" captionStart-5="TABLE 14" captionStart-6="TABLE 15" captionStartId-0="15.[151,239,584,607]" captionStartId-1="15.[151,239,1174,1197]" captionStartId-2="18.[151,239,1401,1424]" captionStartId-3="18.[151,239,1665,1688]" captionStartId-4="21.[151,239,260,283]" captionStartId-5="23.[151,239,1052,1075]" captionStartId-6="24.[151,239,440,463]" captionTargetBox-0="[155,1397,764,1106]" captionTargetBox-1="[158,1405,1257,1568]" captionTargetBox-2="[159,1363,1452,1598]" captionTargetBox-3="[154,1393,1780,1967]" captionTargetBox-4="[159,1335,342,488]" captionTargetBox-5="[159,1320,1134,1322]" captionTargetPageId-0="15" captionTargetPageId-1="15" captionTargetPageId-2="18" captionTargetPageId-3="18" captionTargetPageId-4="21" captionTargetPageId-5="23" captionText-0="TABLE 9. Ptilocrinus (Chambersaecrinus) brucei. Main morphological variables in USNM specimens and type series. Wb: Br 1 width; Wr-b: radial aboral width; Wr-t: radial adoral width; Hc: aboral cup height; Hr-r: radial axial height; Hri: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: adoral diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm. Dp and Dm are used as growth index (in yellow)." captionText-1="TABLE 10. Ptilocrinus (Chambersaecrinus) brucei. Variation of main morphological ratios in USNM specimens and type series. Abbreviations and growth index as in Table 9." captionText-2="TABLE 11. Ptilocrinus (Chambersaecrinus) brucei USNM 1005438. Quantitative characters of proximal brachials." captionText-3="TABLE 12. Ptilocrinus (Chambersaecrinus) brucei USNM 1005438. Quantitative characters of columnals. Columnals numbered from proximal to distal. Dal: axial lumen diameter, Dcol: columnal diameter, Hcol: maximum columnal height. Values in mm, except ratios." captionText-4="TABLE 13. Ptilocrinus (Chambersaecrinus) brucei USNM 1005433 - B. Quantitative characters of columnals. Columnals numbered from proximal to distal. D: maximum columnal diameter, H: columnal height. Values in mm, except H / D." captionText-5="TABLE 14. Ptilocrinus (Chambersaecrinus) brucei, R / V Dmitri Mendeleev sta. 4094. Stalk measurements. D: maximum columnal diameter, H: columnal height. Values in mm, except H / D." captionText-6="TABLE 15. Ptilocrinus (Chambersaecrinus) brucei. Variations in proximal arm pattern. * holotype, ** paratype." httpUri-0="http://table.plazi.org/id/DF294CBBFFD5FFF65483FD9DA86AFD45" httpUri-1="http://table.plazi.org/id/DF294CBBFFD5FFF65483FB43A82EFB68" httpUri-2="http://table.plazi.org/id/DF294CBBFFC8FFEB5483FAACAE41FA35" httpUri-3="http://table.plazi.org/id/DF294CBBFFC8FFEB5483F954A939F97D" httpUri-4="http://table.plazi.org/id/DF294CBBFFCFFFEC5483FED1AE4DFE9E" httpUri-5="http://table.plazi.org/id/DF294CBBFFCDFFEE5483FBC9A8F5FBF6" pageId="14" pageNumber="395" tableUuid-0="DF294CBBFFD5FFF65483FD9DA86AFD45" tableUuid-1="DF294CBBFFD5FFF65483FB43A82EFB68" tableUuid-2="DF294CBBFFC8FFEB5483FAACAE41FA35" tableUuid-3="DF294CBBFFC8FFEB5483F954A939F97D" tableUuid-4="DF294CBBFFCFFFEC5483FED1AE4DFE9E" tableUuid-5="DF294CBBFFCDFFEE5483FBC9A8F5FBF6">Tables 9–15</tableCitation>
.
</paragraph>
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Synonymy: 
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<emphasis id="B922C021FFD4FFF75507FED7AAE0FEBD" box="[275,460,258,280]" italics="true" pageId="14" pageNumber="395">Ptilocrinus brucei</emphasis>
Vaney
</taxonomicName>
<emphasis id="B922C021FFD4FFF7560DFED6A901FEBD" box="[537,557,259,280]" italics="true" pageId="14" pageNumber="395">in</emphasis>
<bibRefCitation id="EFC761C2FFD4FFF75620FED7A991FEBD" author="Wilton" box="[564,701,258,280]" pageId="14" pageNumber="395" refString="Wilton, D. W. (1908) Resultats de l'Expedition nationale Antarctique du S. Y. Scotia, 1902 - 1904, 4, Zoologie, 1, 1 - 133." type="journal article" year="1908">Wilton, 1908</bibRefCitation>
: fig. 70, pl. 23; 
<bibRefCitation id="EFC761C2FFD4FFF75776FED6AF1BFEBD" author="Vaney" box="[866,1079,258,280]" pageId="14" pageNumber="395" refString="Vaney, C. &amp; John, D. D. (1939) Scientific results of the voyage of S. Y. Scotia, 1902 - 04: The Crinoidea. Transactions of the Royal Society of Edinburg, 59, 661 - 672. http: // dx. doi. org / 10.1017 / S 0080456800017385" type="journal article" year="1939">Vaney &amp; John, 1939</bibRefCitation>
: 661–664, 1 pl.; 
<bibRefCitation id="EFC761C2FFD4FFF750F0FED6AEB4FEBD" author="Clark" box="[1252,1432,258,280]" pageId="14" pageNumber="395" refString="Clark, A. H. (1915) Die crinoiden der Antarktis. In: Deutsche Sudpolar-Expedition 1901 - 1903. Druck und Verlag von Georg Reimer, Berlin, 101 - 209." type="book chapter" year="1915">A.H. Clark, 1915</bibRefCitation>
: 161; 
<bibRefCitation id="EFC761C2FFD4FFF754EEFEF4AAFDFE92" author="Smaldon" box="[250,465,289,311]" pageId="14" pageNumber="395" refString="Smaldon, G., Heppell, D. &amp; Watt, K. R. (1976) Type specimens of invertebrates (excluding insects) held at the Royal Scottish Museum, Edinburg. Royal Scottish Museum Information Series, Natural History, 4, 1 - 118." type="journal article" year="1976">
Smaldon 
<emphasis id="B922C021FFD4FFF7554EFEF7AAA1FE92" box="[346,397,289,311]" italics="true" pageId="14" pageNumber="395">et al.</emphasis>
, 1976
</bibRefCitation>
: 74; 
<bibRefCitation id="EFC761C2FFD4FFF75614FEF7A9ABFE92" author="Roux" box="[512,647,289,311]" pageId="14" pageNumber="395" refString="Roux, M. (1980 b) Les articulations du pedoncule des Hyocrinidae (Echinodermes, Crinoides pedoncules): interet systematique et consequences. Bulletin du Museum national d'Histoire naturelle, Paris, Serie 4 e, 2 (Section A), 1, 31 - 57." type="journal article" year="1980" yearSuffix="b">Roux, 1980b</bibRefCitation>
: 36, 41, 53, pl. 3 (figs. 5–6); Roux 
<emphasis id="B922C021FFD4FFF757FEFEF7AF31FE92" box="[1002,1053,289,311]" italics="true" pageId="14" pageNumber="395">et al.</emphasis>
, 2002: 822. 
<taxonomicName id="4C5667B0FFD4FFF75483FE94A823FEF3" authority="Mironov &amp; Sorokina, 1998: 52" authorityName="Mironov &amp; Sorokina" authorityPageNumber="52" authorityYear="1998" box="[151,783,320,342]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="14" pageNumber="395" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFD4FFF75483FE94AAEBFEF3" box="[151,455,320,342]" italics="true" pageId="14" pageNumber="395">P. (Chambersaecrinus) brucei</emphasis>
: 
<bibRefCitation id="EFC761C2FFD4FFF755C0FE95A9C6FEF3" author="Mironov" box="[468,746,320,342]" pageId="14" pageNumber="395" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina, 1998</bibRefCitation>
: 52
</taxonomicName>
–55, fig. 19–20, pl. 8 (figs. 4–5); 
<bibRefCitation id="EFC761C2FFD4FFF7504AFE94AE61FEF3" author="Roux" box="[1118,1357,320,342]" pageId="14" pageNumber="395" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert, 2011</bibRefCitation>
: 45, 48, 51; Eléaume 
<emphasis id="B922C021FFD4FFF75558FEB5AA53FED1" box="[332,383,350,373]" italics="true" pageId="14" pageNumber="395">et al.</emphasis>
, 2014: 210–211.
</paragraph>
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<treatmentCitationGroup id="AB463B1DFFD4FFF75483FEABAF07FE36" box="[151,1067,381,403]" pageId="14" pageNumber="395">
<taxonomicName id="4C5667B0FFD4FFF75483FEABA832FE36" ID-CoL="8GHR5" authority="Mironov &amp; Sorokina, 1998: 55" authorityName="Mironov &amp; Sorokina" authorityPageNumber="55" authorityYear="1998" box="[151,798,381,403]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="14" pageNumber="395" phylum="Echinodermata" rank="species" species="flexibilis" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFD4FFF75483FEABAAF1FE36" box="[151,477,381,403]" italics="true" pageId="14" pageNumber="395">P. (Chambersaecrinus) flexibilis</emphasis>
<treatmentCitation id="0AF73A22FFD4FFF755F0FEA8A832FE36" author="Mironov" box="[484,798,381,403]" page="55" pageId="14" pageNumber="395" year="1998">
<bibRefCitation id="EFC761C2FFD4FFF755F0FEA8A9D6FE36" author="Mironov" box="[484,762,381,403]" pageId="14" pageNumber="395" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina, 1998</bibRefCitation>
: 55
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</taxonomicName>
–57, fig. 21, pl. 11 (fig. 2).
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</paragraph>
</subSubSection>
<subSubSection id="C34C4FB8FFD4FFF65483FE6AA92BFE08" lastPageId="15" lastPageNumber="396" pageId="14" pageNumber="395" type="materials_examined">
<paragraph id="8BE91C33FFD4FFF75483FE6AABFDFE58" blockId="14.[151,1436,447,869]" pageId="14" pageNumber="395">
<emphasis id="B922C021FFD4FFF75483FE6AAA27FE7D" bold="true" box="[151,267,447,472]" pageId="14" pageNumber="395">
<typeStatus id="54EDA291FFD4FFF75483FE6AAA2AFE7D" box="[151,262,447,472]" pageId="14" pageNumber="395" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED4784F6FFD4FFF75506FE15AA63FE7D" box="[274,335,448,472]" httpUri="http://grbio.org/cool/h8hj-4rxx" name="National Museum of Scotland - Natural Sciences" pageId="14" pageNumber="395">NMS</collectionCode>
1921.143.1743, S. 
<collectingCountry id="F3415CA3FFD4FFF7560BFE15A91AFE7D" box="[543,566,448,472]" name="Italy" pageId="14" pageNumber="395">Y</collectingCountry>
. 
<taxonomicName id="4C5667B0FFD4FFF75656FE15A9A4FE72" box="[578,648,448,471]" genus="Scotia" kingdom="Animalia" pageId="14" pageNumber="395" phylum="Nemertea" rank="genus">
<collectingCountry id="F3415CA3FFD4FFF75656FE15A9A4FE72" box="[578,648,448,471]" name="United Kingdom" pageId="14" pageNumber="395">
<emphasis id="B922C021FFD4FFF75656FE15A9A4FE72" box="[578,648,448,471]" italics="true" pageId="14" pageNumber="395">Scotia</emphasis>
</collectingCountry>
</taxonomicName>
, sta. 301, NW Weddell Sea, 
<geoCoordinate id="EE627AF4FFD4FFF757DAFE6AAF09FE7D" box="[974,1061,447,472]" direction="south" orientation="latitude" pageId="14" pageNumber="395" precision="925" value="-64.8">64°48’S</geoCoordinate>
– 
<geoCoordinate id="EE627AF4FFD4FFF75022FE6AAFB3FE7D" box="[1078,1183,447,472]" direction="west" orientation="longitude" pageId="14" pageNumber="395" precision="925" value="-44.433334">44°26’W</geoCoordinate>
, depth 
<quantity id="4CAEB1D6FFD4FFF750E4FE6AAE61FE72" box="[1264,1357,447,472]" metricMagnitude="3" metricUnit="m" metricValue="4.585" pageId="14" pageNumber="395" unit="m" value="4585.0">4,585 m</quantity>
(2,485 fms).
</paragraph>
<paragraph id="8BE91C33FFD4FFF754D3FDD2AEA2FCC0" blockId="14.[151,1436,447,869]" pageId="14" pageNumber="395">
<emphasis id="B922C021FFD4FFF754D3FDD2AA98FD85" bold="true" box="[199,436,519,544]" pageId="14" pageNumber="395">Material examined.</emphasis>
NHM 1939.5.13.1 (
<typeStatus id="54EDA291FFD4FFF756B7FDDDA825FD85" box="[675,777,520,544]" pageId="14" pageNumber="395" type="paratype">paratype</typeStatus>
), S. 
<collectingCountry id="F3415CA3FFD4FFF75727FDDDA866FD85" box="[819,842,520,544]" name="Italy" pageId="14" pageNumber="395">Y</collectingCountry>
. 
<taxonomicName id="4C5667B0FFD4FFF7574EFDD2A88FFD85" box="[858,931,519,544]" genus="Scotia" kingdom="Animalia" pageId="14" pageNumber="395" phylum="Nemertea" rank="genus">
<collectingCountry id="F3415CA3FFD4FFF7574EFDD2A88FFD85" box="[858,931,519,544]" name="United Kingdom" pageId="14" pageNumber="395">Scotia</collectingCountry>
</taxonomicName>
, trawl 3426, off South Orcade Islands, 
<geoCoordinate id="EE627AF4FFD4FFF75161FDD2ABE1FDE1" direction="south" orientation="latitude" pageId="14" pageNumber="395" precision="925" value="-62.166668">62° 10’S</geoCoordinate>
, 
<geoCoordinate id="EE627AF4FFD4FFF754CFFDF9AA7FFDE1" box="[219,339,556,580]" direction="west" orientation="longitude" pageId="14" pageNumber="395" precision="925" value="-41.333332">41° 20’ W</geoCoordinate>
, depth 
<quantity id="4CAEB1D6FFD4FFF755B2FDF9A92FFDE1" box="[422,515,556,581]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="14" pageNumber="395" unit="m" value="3245.0">3,245 m</quantity>
(includes note by D. John in 1938: specimen was “broken into small species but among them some individual plates, radials, etc.. are recognizable”.); 
<collectionCode id="ED4784F6FFD4FFF757A1FD85AF2AFDCD" LSID="urn:lsid:biocol.org:col:34871" box="[949,1030,592,616]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="395">USNM</collectionCode>
1005433, South East Pacific basin, 63°0.3’to 
<geoCoordinate id="EE627AF4FFD4FFF7551CFDA1AA55FD28" box="[264,377,628,653]" direction="south" orientation="latitude" pageId="14" pageNumber="395" precision="92" value="-63.2">63°12.0’S</geoCoordinate>
–128°12’ to 
<geoCoordinate id="EE627AF4FFD4FFF7561DFDA1A950FD29" box="[521,636,628,652]" direction="west" orientation="longitude" pageId="14" pageNumber="395" precision="925" value="-128.18333">128°11’W</geoCoordinate>
, depth 
<quantity id="4CAEB1D6FFD4FFF756C4FDA1A802FD29" box="[720,814,628,653]" metricMagnitude="3" metricUnit="m" metricValue="4.682" pageId="14" pageNumber="395" unit="m" value="4682.0">4,682 m</quantity>
(
<specimenCount id="9D50D7BAFFD4FFF7572AFDA1A8E2FD28" box="[830,974,628,653]" pageId="14" pageNumber="395" type="generic">2 specimens</specimenCount>
; 1 with theca, proximal arm and stalk; 
<specimenCount id="9D50D7BAFFD4FFF7519AFDA1ABDDFD15" pageId="14" pageNumber="395" type="juvenile">1 juvenile</specimenCount>
with complete stalk, basal ring and fragments of proximal arms and radials); 
<collectionCode id="ED4784F6FFD4FFF7509FFD4DAFF0FD15" LSID="urn:lsid:biocol.org:col:34871" box="[1163,1244,664,688]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="395">USNM</collectionCode>
1005438, South Atlantic Ocean, 
<taxonomicName id="4C5667B0FFD4FFF75541FD69AAFDFD71" authority="Sea" authorityName="Sea" box="[341,465,700,725]" genus="Scotia" kingdom="Animalia" pageId="14" pageNumber="395" phylum="Nemertea" rank="genus">
<collectingCountry id="F3415CA3FFD4FFF75541FD69AAB7FD70" box="[341,411,700,725]" name="United Kingdom" pageId="14" pageNumber="395">Scotia</collectingCountry>
Sea
</taxonomicName>
, 
<collectingCountry id="F3415CA3FFD4FFF755F5FD69A9ABFD70" box="[481,647,700,725]" name="South Georgia and the South Sandwich Islands" pageId="14" pageNumber="395">South Georgia</collectingCountry>
<collectingCountry id="F3415CA3FFD4FFF75685FD69A9F4FD70" box="[657,728,700,725]" name="Iceland" pageId="14" pageNumber="395">Island</collectingCountry>
, 54°00’– 
<geoCoordinate id="EE627AF4FFD4FFF75750FD69A8B2FD70" box="[836,926,700,725]" direction="south" orientation="latitude" pageId="14" pageNumber="395" precision="925" value="-54.083332">54°05’S</geoCoordinate>
, 33°40’– 
<geoCoordinate id="EE627AF4FFD4FFF75001FD69AF50FD71" box="[1045,1148,700,724]" direction="west" orientation="longitude" pageId="14" pageNumber="395" precision="925" value="-33.716667">33°43’W</geoCoordinate>
, depth 
<quantity id="4CAEB1D6FFD4FFF750C0FD69AE5FFD71" box="[1236,1395,700,725]" metricMagnitude="3" metricUnit="m" metricValue="2.6905" metricValueMax="2.718" metricValueMin="2.663" pageId="14" pageNumber="395" unit="m" value="2690.5" valueMax="2718.0" valueMin="2663.0">2663–2718 m</quantity>
(1, partly fragmented, theca poorly preserved, fragments of proximal arms with radials); 
<collectionCode id="ED4784F6FFD4FFF7509DFD35AFF6FD5D" LSID="urn:lsid:biocol.org:col:34871" box="[1161,1242,736,760]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="395">USNM</collectionCode>
1005439, South Atlantic, 
<taxonomicName id="4C5667B0FFD4FFF75515FCD1AA56FCB9" authority="Sea" authorityName="Sea" box="[257,378,772,797]" genus="Scotia" kingdom="Animalia" pageId="14" pageNumber="395" phylum="Nemertea" rank="genus">
<collectingCountry id="F3415CA3FFD4FFF75515FCD1AA6BFCB8" box="[257,327,772,797]" name="United Kingdom" pageId="14" pageNumber="395">Scotia</collectingCountry>
Sea
</taxonomicName>
, NE of South 
<collectingCountry id="F3415CA3FFD4FFF75634FCD1A9F1FCB8" box="[544,733,772,797]" name="United Kingdom" pageId="14" pageNumber="395">Shetland Islands</collectingCountry>
, 59°S– 
<geoCoordinate id="EE627AF4FFD4FFF75739FCD1A8AAFCB8" box="[813,902,772,797]" direction="west" orientation="longitude" pageId="14" pageNumber="395" precision="5555" value="-51.9">51.9°W</geoCoordinate>
, depth 
<quantity id="4CAEB1D6FFD4FFF757CEFCD1AFA8FCB9" box="[986,1156,772,797]" metricMagnitude="3" metricUnit="m" metricValue="3.2600000000000002" metricValueMax="3.5100000000000002" metricValueMin="3.0100000000000002" pageId="14" pageNumber="395" unit="m" value="3260.0" valueMax="3510.0" valueMin="3010.0">3,010–3,510 m</quantity>
(1 theca, proximal arms and stalk fragments); 
<collectionCode id="ED4784F6FFD4FFF755B6FCFDAADFFCE5" LSID="urn:lsid:biocol.org:col:34871" box="[418,499,808,832]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="395">USNM</collectionCode>
1105453-A South 
<collectingCountry id="F3415CA3FFD4FFF756CBFCF2A8B8FCE5" box="[735,916,807,832]" name="United Kingdom" pageId="14" pageNumber="395">Orkney Islands</collectingCountry>
, 
<taxonomicName id="4C5667B0FFD4FFF757B3FCF2AF6DFCE5" authority="Ridge" authorityName="Ridge" box="[935,1089,807,832]" genus="Scotia" kingdom="Animalia" pageId="14" pageNumber="395" phylum="Nemertea" rank="genus">
<collectingCountry id="F3415CA3FFD4FFF757B3FCF2A8C1FCE5" box="[935,1005,807,832]" name="United Kingdom" pageId="14" pageNumber="395">Scotia</collectingCountry>
Ridge
</taxonomicName>
, 62°09’ à 
<geoCoordinate id="EE627AF4FFD4FFF750DFFCF2AE0BFCE5" box="[1227,1319,807,832]" direction="south" orientation="latitude" pageId="14" pageNumber="395" precision="925" value="-62.266666">62°16’S</geoCoordinate>
–38°11’ à 
<geoCoordinate id="EE627AF4FFD4FFF75483FC99ABD2FCC1" box="[151,254,844,868]" direction="west" orientation="longitude" pageId="14" pageNumber="395" precision="925" value="-38.4">38°24’W</geoCoordinate>
, depth 
<quantity id="4CAEB1D6FFD4FFF7555AFC99AADBFCC1" box="[334,503,844,869]" metricMagnitude="3" metricUnit="m" metricValue="3.2575" metricValueMax="3.4589999999999996" metricValueMin="3.056" pageId="14" pageNumber="395" unit="m" value="3257.5" valueMax="3459.0" valueMin="3056.0">3,056–3,459 m</quantity>
(dissociated radials, basal ring, proximal brachials and proximalmost columnals).
</paragraph>
<caption id="DF294CBBFFD4FFF75483FA02A88EF9A9" httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="14" pageNumber="395" targetBox="[295,1291,921,1473]" targetPageId="14">
<paragraph id="8BE91C33FFD4FFF75483FA02A88EF9A9" blockId="14.[151,1435,1495,1548]" pageId="14" pageNumber="395">
<emphasis id="B922C021FFD4FFF75483FA02AA3FFA49" bold="true" box="[151,275,1495,1517]" pageId="14" pageNumber="395">FIGURE 9.</emphasis>
Theca and proximal arms of 
<taxonomicName id="4C5667B0FFD4FFF75650FA02A9D1FA48" box="[580,765,1495,1517]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="14" pageNumber="395" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFD4FFF75650FA02A9D1FA48" box="[580,765,1495,1517]" italics="true" pageId="14" pageNumber="395">Ptilocrinus brucei</emphasis>
</taxonomicName>
. A–B: specimen 1005439; C: specimen 1005433-A; A: view of anal interray; B: view of A–B interray; C: anterior view; as: anal sac; o: oral.
</paragraph>
</caption>
<paragraph id="8BE91C33FFD4FFF654D3F9EDAE36FEB8" blockId="14.[151,1437,1592,2014]" lastBlockId="15.[151,1437,151,536]" lastPageId="15" lastPageNumber="396" pageId="14" pageNumber="395">
<emphasis id="B922C021FFD4FFF754D3F9EDAA95F9F4" bold="true" box="[199,441,1592,1617]" pageId="14" pageNumber="395">Emended diagnosis.</emphasis>
A species of 
<taxonomicName id="4C5667B0FFD4FFF7564BF9EDA8E0F9F4" authority="(Chambersaecrinus)" baseAuthorityName="Chambersaecrinus" box="[607,972,1592,1617]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="14" pageNumber="395" phylum="Echinodermata" rank="genus">
<emphasis id="B922C021FFD4FFF7564BF9EDA8E0F9F4" box="[607,972,1592,1617]" italics="true" pageId="14" pageNumber="395">Ptilocrinus (Chambersaecrinus)</emphasis>
</taxonomicName>
reaching large size; proximalmost stalk diameter to 
<quantity id="4CAEB1D6FFD4FFF75535F989AA50F9D0" box="[289,380,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="395" unit="mm" value="4.5">4.5 mm</quantity>
, arm length to 
<quantity id="4CAEB1D6FFD4FFF7563AF988A9A1F9D0" box="[558,653,1629,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="395" unit="mm" value="130.0">130 mm</quantity>
with about 40 pinnules on each side, and pinnule length to 
<quantity id="4CAEB1D6FFD4FFF75150F989AEB5F9D0" box="[1348,1433,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="395" unit="mm" value="55.0">55 mm</quantity>
. External morphology variable; ratio of cup height to adoral diameter 0.97–1.36; ratio of cup to radial height 1.01– 1.65 (usually 1.20–1.34); ratio of radial to first brachial width 1.45–1.97; ratio of adoral cup to proximalmost stalk diameter 3.25 (small specimen) to 4.27 (usually 3.97–4.20); arm deeply inserted on adoral radial edge; basal cup and proximalmost stalk diameter usually equal; maximum ratio of columnal height to diameter in a given stalk from 
<quantity id="4CAEB1D6FFD4FFF754C2F8C4AA09F88C" box="[214,293,1808,1833]" metricMagnitude="-2" metricUnit="m" metricValue="3.6322" pageId="14" pageNumber="395" unit="in" value="1.43">1.43 in</quantity>
smallest specimen to 
<quantity id="4CAEB1D6FFD4FFF75636F8C4A95EF88C" box="[546,626,1808,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.0413999999999999" pageId="14" pageNumber="395" unit="in" value="0.41">0.41 in</quantity>
large specimens. Tegmen inflated and rounded; anal cone significantly taller than oral cone and reaching level of Br8 to Br 
<quantity id="4CAEB1D6FFD4FFF756C8F8E0A838F8EB" box="[732,788,1845,1870]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="14" pageNumber="395" unit="in" value="14.0">14 in</quantity>
large specimens; base of anal cone peripheral on tegmen; upper part of cone oriented toward tegmen center; orals smooth and concave with rounded top; as many as 20 large polygonal tegminal plates sometimes with a regular median arrangement. Arm pattern variable, from chiefly alternating free brachial and brachial pairs to long series (more than 20) of successive free brachials beyond the third brachial pair; proximalmost arm pattern 1+2 3 4 5+6 7 (75.9%), 1+2 3 4 5+6 7+8 (13.8%); first pinnule usually on Br4 (94.7%), rarely on Br5; most frequent brachial pairs beyond Br7 at 8+9 (44.0%), 10+11 (22.7%) and 18+19 (27.3%). Brachial muscular synarthry with conspicuous process prolonging adoral groove. Pinnule inserted on distal adoral side of brachial; proximal part with asymmetrical genital inflation; great number of additional small proximal plates decreasing distally; additional plates imbricated or irregularly aligned.
</paragraph>
<paragraph id="8BE91C33FFD5FFF654D3FEF2A92BFE08" blockId="15.[151,1437,151,536]" pageId="15" pageNumber="396">Stalk symplexies with 6 (young stage) to 14 (usually 8–12) crenular units of 1 crenula of various lengths; up to 3 crenulae or subsidiary crenulae developed between units in large specimens; axial canal in proximal mesistele enlarges substantially during ontogeny; lumen bordered by coarsely-meshed galleried stereom; distal syzygies with thin irregular radial crenularium.</paragraph>
</subSubSection>
<subSubSection id="C34C4FB8FFD5FFEF54D3FE62A8F7FE75" lastPageId="22" lastPageNumber="403" pageId="15" pageNumber="396" type="description">
<paragraph id="8BE91C33FFD5FFF654D3FE62A820FDBD" blockId="15.[151,1437,151,536]" pageId="15" pageNumber="396">
<emphasis id="B922C021FFD5FFF654D3FE62AA75FE75" bold="true" box="[199,345,439,464]" pageId="15" pageNumber="396">Description.</emphasis>
<tableCitation id="C6D42988FFD5FFF6554BFE62AA90FE75" box="[351,444,439,464]" captionStart="TABLE 9" captionStartId="15.[151,239,584,607]" captionTargetBox="[155,1397,764,1106]" captionTargetPageId="15" captionText="TABLE 9. Ptilocrinus (Chambersaecrinus) brucei. Main morphological variables in USNM specimens and type series. Wb: Br 1 width; Wr-b: radial aboral width; Wr-t: radial adoral width; Hc: aboral cup height; Hr-r: radial axial height; Hri: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: adoral diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm. Dp and Dm are used as growth index (in yellow)." httpUri="http://table.plazi.org/id/DF294CBBFFD5FFF65483FD9DA86AFD45" pageId="15" pageNumber="396" tableUuid="DF294CBBFFD5FFF65483FD9DA86AFD45">Tables 9</tableCitation>
and 
<tableCitation id="C6D42988FFD5FFF655E7FE6DA923FE75" box="[499,527,440,464]" captionStart="TABLE 10" captionStartId="15.[151,239,1174,1197]" captionTargetBox="[158,1405,1257,1568]" captionTargetPageId="15" captionText="TABLE 10. Ptilocrinus (Chambersaecrinus) brucei. Variation of main morphological ratios in USNM specimens and type series. Abbreviations and growth index as in Table 9." httpUri="http://table.plazi.org/id/DF294CBBFFD5FFF65483FB43A82EFB68" pageId="15" pageNumber="396" tableUuid="DF294CBBFFD5FFF65483FB43A82EFB68">10</tableCitation>
list biometric parameters, including 
<typeStatus id="54EDA291FFD5FFF657BFFE62AF21FE75" box="[939,1037,439,464]" pageId="15" pageNumber="396" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54EDA291FFD5FFF65057FE6DAF8BFE75" box="[1091,1191,440,464]" pageId="15" pageNumber="396" type="paratype">paratype</typeStatus>
. The small number of specimens and the wide range of character variations preclude obtaining robust results from statistical analyses on the main changes in quantitative ratios during ontogeny.
</paragraph>
<caption id="DF294CBBFFD5FFF65483FD9DA86AFD45" ID-Table-UUID="DF294CBBFFD5FFF65483FD9DA86AFD45" httpUri="http://table.plazi.org/id/DF294CBBFFD5FFF65483FD9DA86AFD45" pageId="15" pageNumber="396" targetBox="[155,1397,764,1106]" targetIsTable="true" targetPageId="15">
<paragraph id="8BE91C33FFD5FFF65483FD9DA86AFD45" blockId="15.[151,1436,584,736]" pageId="15" pageNumber="396">
<emphasis id="B922C021FFD5FFF65483FD9DAA27FDFA" bold="true" box="[151,267,584,607]" pageId="15" pageNumber="396">TABLE 9.</emphasis>
<emphasis id="B922C021FFD5FFF65506FD9DA992FDFA" box="[274,702,584,607]" italics="true" pageId="15" pageNumber="396">
<taxonomicName id="4C5667B0FFD5FFF65506FD9DA995FDFA" box="[274,697,584,607]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="15" pageNumber="396" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">Ptilocrinus (Chambersaecrinus) brucei</taxonomicName>
.
</emphasis>
Main morphological variables in USNM specimens and type series. Wb: Br1 width; Wr-b: radial aboral width; Wr-t: radial adoral width; Hc: aboral cup height; Hr-r: radial axial height; Hri: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: adoral diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm. Dp and Dm are used as growth index (in yellow).
</paragraph>
</caption>
<paragraph id="8BE91C33FFD5FFF6548FFD29AE4BFBF7" pageId="15" pageNumber="396">
<table id="F956EE93FFD50006548FFD29AE59FBF7" box="[155,1397,764,1106]" gridcols="13" gridrows="8" pageId="15" pageNumber="396">
<tr id="35661E71FFD50006548FFD29AE59FCB7" box="[155,1397,764,786]" gridrow="0" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFD29AA22FCB7" box="[155,270,764,786]" gridcol="0" gridrow="0" pageId="15" pageNumber="396">Specimen</th>
<th id="76B7770DFFD500065555FD29AA41FCB7" box="[321,365,764,786]" gridcol="1" gridrow="0" pageId="15" pageNumber="396">Wb</th>
<th id="76B7770DFFD50006558AFD29AAE6FCB7" box="[414,458,764,786]" gridcol="2" gridrow="0" pageId="15" pageNumber="396">Wrb</th>
<th id="76B7770DFFD5000655E8FD29A904FCB7" box="[508,552,764,786]" gridcol="3" gridrow="0" pageId="15" pageNumber="396">Wrt</th>
<th id="76B7770DFFD50006564DFD29A9BEFCB7" box="[601,658,764,786]" gridcol="4" gridrow="0" pageId="15" pageNumber="396">Hc</th>
<th id="76B7770DFFD5000656ACFD29A9C8FCB7" box="[696,740,764,786]" gridcol="5" gridrow="0" pageId="15" pageNumber="396">Hrr</th>
<th id="76B7770DFFD500065701FD29A86DFCB7" box="[789,833,764,786]" gridcol="6" gridrow="0" pageId="15" pageNumber="396">Hri</th>
<th id="76B7770DFFD500065765FD29A88EFCB7" box="[881,930,764,786]" gridcol="7" gridrow="0" pageId="15" pageNumber="396">Htga</th>
<th id="76B7770DFFD5000657C5FD29AF28FCB7" box="[977,1028,764,786]" gridcol="8" gridrow="0" pageId="15" pageNumber="396">Htgo</th>
<th id="76B7770DFFD50006503BFD29AF4AFCB7" box="[1071,1126,764,786]" gridcol="9" gridrow="0" pageId="15" pageNumber="396">Dc</th>
<th id="76B7770DFFD500065099FD29AF95FCB7" box="[1165,1209,764,786]" gridcol="10" gridrow="0" pageId="15" pageNumber="396">Drb</th>
<th id="76B7770DFFD5000650FDFD29AE3BFCB7" box="[1257,1303,764,786]" gridcol="11" gridrow="0" pageId="15" pageNumber="396">Dp</th>
<th id="76B7770DFFD500065153FD29AE59FCB7" box="[1351,1397,764,786]" gridcol="12" gridrow="0" pageId="15" pageNumber="396">Dm</th>
</tr>
<tr id="35661E71FFD50006548FFCF0AE59FC9E" box="[155,1397,805,827]" gridrow="1" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFCF0AA22FC9E" box="[155,270,805,827]" gridcol="0" gridrow="1" pageId="15" pageNumber="396">1005433-B</th>
<td id="76B7770DFFD500065555FCF0AA41FC9E" box="[321,365,805,827]" gridcol="1" gridrow="1" pageId="15" pageNumber="396">2.1</td>
<td id="76B7770DFFD50006558AFCF0AAE6FC9E" box="[414,458,805,827]" gridcol="2" gridrow="1" pageId="15" pageNumber="396">1.9</td>
<td id="76B7770DFFD5000655E8FCF0A904FC9E" box="[508,552,805,827]" gridcol="3" gridrow="1" pageId="15" pageNumber="396">3.7</td>
<td id="76B7770DFFD50006564DFCF0A9BEFC9E" box="[601,658,805,827]" gridcol="4" gridrow="1" pageId="15" pageNumber="396">7.45</td>
<td id="76B7770DFFD5000656ACFCF0A9C8FC9E" box="[696,740,805,827]" gridcol="5" gridrow="1" pageId="15" pageNumber="396">3.8</td>
<td id="76B7770DFFD500065701FCF0A86DFC9E" box="[789,833,805,827]" gridcol="6" gridrow="1" pageId="15" pageNumber="396">4.5</td>
<td id="76B7770DFFD500065765FCF0A88EFC9E" box="[881,930,805,827]" gridcol="7" gridrow="1" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657C5FCF0AF28FC9E" box="[977,1028,805,827]" gridcol="8" gridrow="1" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006503BFCF0AF4AFC9E" box="[1071,1126,805,827]" gridcol="9" gridrow="1" pageId="15" pageNumber="396">6.2</td>
<td id="76B7770DFFD500065099FCF0AF95FC9E" box="[1165,1209,805,827]" gridcol="10" gridrow="1" pageId="15" pageNumber="396">3.4</td>
<td id="76B7770DFFD5000650FDFCF0AE3BFC9E" box="[1257,1303,805,827]" gridcol="11" gridrow="1" pageId="15" pageNumber="396">1.9</td>
<td id="76B7770DFFD500065153FCF0AE59FC9E" box="[1351,1397,805,827]" gridcol="12" gridrow="1" pageId="15" pageNumber="396">1.4</td>
</tr>
<tr id="35661E71FFD50006548FFC9BAE59FCC1" box="[155,1397,846,868]" gridrow="2" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFC9BAA22FCC1" box="[155,270,846,868]" gridcol="0" gridrow="2" pageId="15" pageNumber="396">paratype</th>
<td id="76B7770DFFD500065555FC9BAA41FCC1" box="[321,365,846,868]" gridcol="1" gridrow="2" pageId="15" pageNumber="396">3.4</td>
<td id="76B7770DFFD50006558AFC9BAAE6FCC1" box="[414,458,846,868]" gridcol="2" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000655E8FC9BA904FCC1" box="[508,552,846,868]" gridcol="3" gridrow="2" pageId="15" pageNumber="396">6.7</td>
<td id="76B7770DFFD50006564DFC9BA9BEFCC1" box="[601,658,846,868]" gridcol="4" gridrow="2" pageId="15" pageNumber="396">11.55</td>
<td id="76B7770DFFD5000656ACFC9BA9C8FCC1" box="[696,740,846,868]" gridcol="5" gridrow="2" pageId="15" pageNumber="396">6.6</td>
<td id="76B7770DFFD500065701FC9BA86DFCC1" box="[789,833,846,868]" gridcol="6" gridrow="2" pageId="15" pageNumber="396">6.75</td>
<td id="76B7770DFFD500065765FC9BA88EFCC1" box="[881,930,846,868]" gridcol="7" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657C5FC9BAF28FCC1" box="[977,1028,846,868]" gridcol="8" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006503BFC9BAF4AFCC1" box="[1071,1126,846,868]" gridcol="9" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065099FC9BAF95FCC1" box="[1165,1209,846,868]" gridcol="10" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000650FDFC9BAE3BFCC1" box="[1257,1303,846,868]" gridcol="11" gridrow="2" pageId="15" pageNumber="396">~2.5</td>
<td id="76B7770DFFD500065153FC9BAE59FCC1" box="[1351,1397,846,868]" gridcol="12" gridrow="2" pageId="15" pageNumber="396">1.95</td>
</tr>
<tr id="35661E71FFD50006548FFCADAE59FC2B" box="[155,1397,888,910]" gridrow="3" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFCADAA22FC2B" box="[155,270,888,910]" gridcol="0" gridrow="3" pageId="15" pageNumber="396">1005453-A</th>
<td id="76B7770DFFD500065555FCADAA41FC2B" box="[321,365,888,910]" gridcol="1" gridrow="3" pageId="15" pageNumber="396">3.4</td>
<td id="76B7770DFFD50006558AFCADAAE6FC2B" box="[414,458,888,910]" gridcol="2" gridrow="3" pageId="15" pageNumber="396">3.75</td>
<td id="76B7770DFFD5000655E8FCADA904FC2B" box="[508,552,888,910]" gridcol="3" gridrow="3" pageId="15" pageNumber="396">5.6</td>
<td id="76B7770DFFD50006564DFCADA9BEFC2B" box="[601,658,888,910]" gridcol="4" gridrow="3" pageId="15" pageNumber="396">12.6</td>
<td id="76B7770DFFD5000656ACFCADA9C8FC2B" box="[696,740,888,910]" gridcol="5" gridrow="3" pageId="15" pageNumber="396">6.6</td>
<td id="76B7770DFFD500065701FCADA86DFC2B" box="[789,833,888,910]" gridcol="6" gridrow="3" pageId="15" pageNumber="396">7.5</td>
<td id="76B7770DFFD500065765FCADA88EFC2B" box="[881,930,888,910]" gridcol="7" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657C5FCADAF28FC2B" box="[977,1028,888,910]" gridcol="8" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006503BFCADAF4AFC2B" box="[1071,1126,888,910]" gridcol="9" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065099FCADAF95FC2B" box="[1165,1209,888,910]" gridcol="10" gridrow="3" pageId="15" pageNumber="396">6.25</td>
<td id="76B7770DFFD5000650FDFCADAE3BFC2B" box="[1257,1303,888,910]" gridcol="11" gridrow="3" pageId="15" pageNumber="396">2.75</td>
<td id="76B7770DFFD500065153FCADAE59FC2B" box="[1351,1397,888,910]" gridcol="12" gridrow="3" pageId="15" pageNumber="396">-</td>
</tr>
<tr id="35661E71FFD50006548FFC74AE59FC73" box="[155,1397,929,982]" gridrow="4" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFC74AA22FC73" box="[155,270,929,982]" gridcol="0" gridrow="4" pageId="15" pageNumber="396">1005433-A</th>
<td id="76B7770DFFD500065555FC74AA41FC73" box="[321,365,929,982]" gridcol="1" gridrow="4" pageId="15" pageNumber="396">4.1 4.7</td>
<td id="76B7770DFFD50006558AFC74AAE6FC73" box="[414,458,929,982]" gridcol="2" gridrow="4" pageId="15" pageNumber="396">4.1</td>
<td id="76B7770DFFD5000655E8FC74A904FC73" box="[508,552,929,982]" gridcol="3" gridrow="4" pageId="15" pageNumber="396">6.3 6.8</td>
<td id="76B7770DFFD50006564DFC74A9BEFC73" box="[601,658,929,982]" gridcol="4" gridrow="4" pageId="15" pageNumber="396">16.15 15.2</td>
<td id="76B7770DFFD5000656ACFC74A9C8FC73" box="[696,740,929,982]" gridcol="5" gridrow="4" pageId="15" pageNumber="396">8.8</td>
<td id="76B7770DFFD500065701FC74A86DFC73" box="[789,833,929,982]" gridcol="6" gridrow="4" pageId="15" pageNumber="396">10.5</td>
<td id="76B7770DFFD500065765FC74A88EFC73" box="[881,930,929,982]" gridcol="7" gridrow="4" pageId="15" pageNumber="396">&gt;9.0</td>
<td id="76B7770DFFD5000657C5FC74AF28FC73" box="[977,1028,929,982]" gridcol="8" gridrow="4" pageId="15" pageNumber="396">5.7</td>
<td id="76B7770DFFD50006503BFC74AF4AFC73" box="[1071,1126,929,982]" gridcol="9" gridrow="4" pageId="15" pageNumber="396">11.7 11.85</td>
<td id="76B7770DFFD500065099FC74AF95FC73" box="[1165,1209,929,982]" gridcol="10" gridrow="4" pageId="15" pageNumber="396">7.15</td>
<td id="76B7770DFFD5000650FDFC74AE3BFC73" box="[1257,1303,929,982]" gridcol="11" gridrow="4" pageId="15" pageNumber="396">2.8</td>
<td id="76B7770DFFD500065153FC74AE59FC73" box="[1351,1397,929,982]" gridcol="12" gridrow="4" pageId="15" pageNumber="396">2.4</td>
</tr>
<tr id="35661E71FFD50006548FFC3CAE59FC5A" box="[155,1397,1001,1023]" gridrow="5" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFC3CAA22FC5A" box="[155,270,1001,1023]" gridcol="0" gridrow="5" pageId="15" pageNumber="396">1005439</th>
<td id="76B7770DFFD500065555FC3CAA41FC5A" box="[321,365,1001,1023]" gridcol="1" gridrow="5" pageId="15" pageNumber="396">4.05</td>
<td id="76B7770DFFD50006558AFC3CAAE6FC5A" box="[414,458,1001,1023]" gridcol="2" gridrow="5" pageId="15" pageNumber="396">4.35</td>
<td id="76B7770DFFD5000655E8FC3CA904FC5A" box="[508,552,1001,1023]" gridcol="3" gridrow="5" pageId="15" pageNumber="396">7.45</td>
<td id="76B7770DFFD50006564DFC3CA9BEFC5A" box="[601,658,1001,1023]" gridcol="4" gridrow="5" pageId="15" pageNumber="396">15.05</td>
<td id="76B7770DFFD5000656ACFC3CA9C8FC5A" box="[696,740,1001,1023]" gridcol="5" gridrow="5" pageId="15" pageNumber="396">7.85</td>
<td id="76B7770DFFD500065701FC3CA86DFC5A" box="[789,833,1001,1023]" gridcol="6" gridrow="5" pageId="15" pageNumber="396">9.05</td>
<td id="76B7770DFFD500065765FC3CA88EFC5A" box="[881,930,1001,1023]" gridcol="7" gridrow="5" pageId="15" pageNumber="396">16.5</td>
<td id="76B7770DFFD5000657C5FC3CAF28FC5A" box="[977,1028,1001,1023]" gridcol="8" gridrow="5" pageId="15" pageNumber="396">9.8</td>
<td id="76B7770DFFD50006503BFC3CAF4AFC5A" box="[1071,1126,1001,1023]" gridcol="9" gridrow="5" pageId="15" pageNumber="396">13.7</td>
<td id="76B7770DFFD500065099FC3CAF95FC5A" box="[1165,1209,1001,1023]" gridcol="10" gridrow="5" pageId="15" pageNumber="396">7.85</td>
<td id="76B7770DFFD5000650FDFC3CAE3BFC5A" box="[1257,1303,1001,1023]" gridcol="11" gridrow="5" pageId="15" pageNumber="396">3.4</td>
<td id="76B7770DFFD500065153FC3CAE59FC5A" box="[1351,1397,1001,1023]" gridcol="12" gridrow="5" pageId="15" pageNumber="396">2.8</td>
</tr>
<tr id="35661E71FFD50006548FFBC7AE59FB8D" box="[155,1397,1042,1064]" gridrow="6" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFBC7AA22FB8D" box="[155,270,1042,1064]" gridcol="0" gridrow="6" pageId="15" pageNumber="396">holotype</th>
<td id="76B7770DFFD500065555FBC7AA41FB8D" box="[321,365,1042,1064]" gridcol="1" gridrow="6" pageId="15" pageNumber="396">4.8</td>
<td id="76B7770DFFD50006558AFBC7AAE6FB8D" box="[414,458,1042,1064]" gridcol="2" gridrow="6" pageId="15" pageNumber="396">4.4</td>
<td id="76B7770DFFD5000655E8FBC7A904FB8D" box="[508,552,1042,1064]" gridcol="3" gridrow="6" pageId="15" pageNumber="396">8.0</td>
<td id="76B7770DFFD50006564DFBC7A9BEFB8D" box="[601,658,1042,1064]" gridcol="4" gridrow="6" pageId="15" pageNumber="396">13.6</td>
<td id="76B7770DFFD5000656ACFBC7A9C8FB8D" box="[696,740,1042,1064]" gridcol="5" gridrow="6" pageId="15" pageNumber="396">8.4</td>
<td id="76B7770DFFD500065701FBC7A86DFB8D" box="[789,833,1042,1064]" gridcol="6" gridrow="6" pageId="15" pageNumber="396">9.4</td>
<td id="76B7770DFFD500065765FBC7A88EFB8D" box="[881,930,1042,1064]" gridcol="7" gridrow="6" pageId="15" pageNumber="396">12.0</td>
<td id="76B7770DFFD5000657C5FBC7AF28FB8D" box="[977,1028,1042,1064]" gridcol="8" gridrow="6" pageId="15" pageNumber="396">~6.0</td>
<td id="76B7770DFFD50006503BFBC7AF4AFB8D" box="[1071,1126,1042,1064]" gridcol="9" gridrow="6" pageId="15" pageNumber="396">14.0</td>
<td id="76B7770DFFD500065099FBC7AF95FB8D" box="[1165,1209,1042,1064]" gridcol="10" gridrow="6" pageId="15" pageNumber="396">7.4</td>
<td id="76B7770DFFD5000650FDFBC7AE3BFB8D" box="[1257,1303,1042,1064]" gridcol="11" gridrow="6" pageId="15" pageNumber="396">3.5</td>
<td id="76B7770DFFD500065153FBC7AE59FB8D" box="[1351,1397,1042,1064]" gridcol="12" gridrow="6" pageId="15" pageNumber="396">&lt;3.0</td>
</tr>
<tr id="35661E71FFD50006548FFBE9AE59FBF7" box="[155,1397,1084,1106]" gridrow="7" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548FFBE9AA22FBF7" box="[155,270,1084,1106]" gridcol="0" gridrow="7" pageId="15" pageNumber="396">1005438</th>
<td id="76B7770DFFD500065555FBE9AA41FBF7" box="[321,365,1084,1106]" gridcol="1" gridrow="7" pageId="15" pageNumber="396">5.5</td>
<td id="76B7770DFFD50006558AFBE9AAE6FBF7" box="[414,458,1084,1106]" gridcol="2" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000655E8FBE9A904FBF7" box="[508,552,1084,1106]" gridcol="3" gridrow="7" pageId="15" pageNumber="396">8.9</td>
<td id="76B7770DFFD50006564DFBE9A9BEFBF7" box="[601,658,1084,1106]" gridcol="4" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000656ACFBE9A9C8FBF7" box="[696,740,1084,1106]" gridcol="5" gridrow="7" pageId="15" pageNumber="396">9.25</td>
<td id="76B7770DFFD500065701FBE9A86DFBF7" box="[789,833,1084,1106]" gridcol="6" gridrow="7" pageId="15" pageNumber="396">12.0</td>
<td id="76B7770DFFD500065765FBE9A88EFBF7" box="[881,930,1084,1106]" gridcol="7" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657C5FBE9AF28FBF7" box="[977,1028,1084,1106]" gridcol="8" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006503BFBE9AF4AFBF7" box="[1071,1126,1084,1106]" gridcol="9" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065099FBE9AF95FBF7" box="[1165,1209,1084,1106]" gridcol="10" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000650FDFBE9AE3BFBF7" box="[1257,1303,1084,1106]" gridcol="11" gridrow="7" pageId="15" pageNumber="396">&gt;4.2</td>
<td id="76B7770DFFD500065153FBE9AE59FBF7" box="[1351,1397,1084,1106]" gridcol="12" gridrow="7" pageId="15" pageNumber="396">3.8</td>
</tr>
</table>
</paragraph>
<caption id="DF294CBBFFD5FFF65483FB43A82EFB68" ID-Table-UUID="DF294CBBFFD5FFF65483FB43A82EFB68" httpUri="http://table.plazi.org/id/DF294CBBFFD5FFF65483FB43A82EFB68" pageId="15" pageNumber="396" targetBox="[158,1405,1257,1568]" targetIsTable="true" targetPageId="15">
<paragraph id="8BE91C33FFD5FFF65483FB43A82EFB68" blockId="15.[151,1436,1174,1229]" pageId="15" pageNumber="396">
<emphasis id="B922C021FFD5FFF65483FB43AA35FB08" bold="true" box="[151,281,1174,1197]" pageId="15" pageNumber="396">TABLE 10.</emphasis>
<emphasis id="B922C021FFD5FFF65536FB43A9FDFB08" box="[290,721,1174,1197]" italics="true" pageId="15" pageNumber="396">
<taxonomicName id="4C5667B0FFD5FFF65536FB43A9E1FB08" box="[290,717,1174,1197]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="15" pageNumber="396" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">Ptilocrinus (Chambersaecrinus) brucei</taxonomicName>
.
</emphasis>
Variation of main morphological ratios in USNM specimens and type series. Abbreviations and growth index as in Table 9.
</paragraph>
</caption>
<paragraph id="8BE91C33FFD5FFF6548AFB3CAE51F985" pageId="15" pageNumber="396">
<table id="F956EE93FFD50006548AFB3CAE51F985" box="[158,1405,1257,1568]" gridcols="11" gridrows="8" pageId="15" pageNumber="396">
<tr id="35661E71FFD50006548AFB3CAE51FB5A" box="[158,1405,1257,1279]" gridrow="0" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFB3CAA3EFB5A" box="[158,274,1257,1279]" gridcol="0" gridrow="0" pageId="15" pageNumber="396">Specimen</th>
<th id="76B7770DFFD50006555DFB3CAAB1FB5A" box="[329,413,1257,1279]" gridcol="1" gridrow="0" pageId="15" pageNumber="396">Hc/Dc</th>
<th id="76B7770DFFD5000655D4FB3CAAD3FB5A" box="[448,511,1257,1279]" gridcol="2" gridrow="0" pageId="15" pageNumber="396">Hr/Hc</th>
<th id="76B7770DFFD500065634FB3CA946FB5A" box="[544,618,1257,1279]" gridcol="3" gridrow="0" pageId="15" pageNumber="396">Htg/Hc</th>
<th id="76B7770DFFD50006569EFB3CA9F3FB5A" box="[650,735,1257,1279]" gridcol="4" gridrow="0" pageId="15" pageNumber="396">Hr/Wr</th>
<th id="76B7770DFFD5000656EFFB3CA863FB5A" box="[763,847,1257,1279]" gridcol="5" gridrow="0" pageId="15" pageNumber="396">Wr/ Wb</th>
<th id="76B7770DFFD500065767FB3CA8F9FB5A" box="[883,981,1257,1279]" gridcol="6" gridrow="0" pageId="15" pageNumber="396">Dc/Dp</th>
<th id="76B7770DFFD5000657E0FB3CAF65FB5A" box="[1012,1097,1257,1279]" gridcol="7" gridrow="0" pageId="15" pageNumber="396">Dc/Drb</th>
<th id="76B7770DFFD500065061FB3CAFEEFB5A" box="[1141,1218,1257,1279]" gridcol="8" gridrow="0" pageId="15" pageNumber="396">Drb/Dp</th>
<th id="76B7770DFFD5000650F0FB3CAE1CFB5A" box="[1252,1328,1257,1279]" gridcol="9" gridrow="0" pageId="15" pageNumber="396">Dp/Dm</th>
<th id="76B7770DFFD50006515BFB3CAE51FB5A" box="[1359,1405,1257,1279]" gridcol="10" gridrow="0" pageId="15" pageNumber="396">Dp</th>
</tr>
<tr id="35661E71FFD50006548AFAC7AE51FA8D" box="[158,1405,1298,1320]" gridrow="1" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFAC7AA3EFA8D" box="[158,274,1298,1320]" gridcol="0" gridrow="1" pageId="15" pageNumber="396">1005433-B</th>
<td id="76B7770DFFD50006555DFAC7AAB1FA8D" box="[329,413,1298,1320]" gridcol="1" gridrow="1" pageId="15" pageNumber="396">1.20</td>
<td id="76B7770DFFD5000655D4FAC7AAD3FA8D" box="[448,511,1298,1320]" gridcol="2" gridrow="1" pageId="15" pageNumber="396">0.60</td>
<td id="76B7770DFFD500065634FAC7A946FA8D" box="[544,618,1298,1320]" gridcol="3" gridrow="1" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006569EFAC7A9F3FA8D" box="[650,735,1298,1320]" gridcol="4" gridrow="1" pageId="15" pageNumber="396">1.21</td>
<td id="76B7770DFFD5000656EFFAC7A863FA8D" box="[763,847,1298,1320]" gridcol="5" gridrow="1" pageId="15" pageNumber="396">1.73</td>
<td id="76B7770DFFD500065767FAC7A8F9FA8D" box="[883,981,1298,1320]" gridcol="6" gridrow="1" pageId="15" pageNumber="396">3.25</td>
<td id="76B7770DFFD5000657E0FAC7AF65FA8D" box="[1012,1097,1298,1320]" gridcol="7" gridrow="1" pageId="15" pageNumber="396">1.81</td>
<td id="76B7770DFFD500065061FAC7AFEEFA8D" box="[1141,1218,1298,1320]" gridcol="8" gridrow="1" pageId="15" pageNumber="396">1.80</td>
<td id="76B7770DFFD5000650F0FAC7AE1CFA8D" box="[1252,1328,1298,1320]" gridcol="9" gridrow="1" pageId="15" pageNumber="396">1.40</td>
<td id="76B7770DFFD50006515BFAC7AE51FA8D" box="[1359,1405,1298,1320]" gridcol="10" gridrow="1" pageId="15" pageNumber="396">1.9</td>
</tr>
<tr id="35661E71FFD50006548AFAE9AE51FAF7" box="[158,1405,1340,1362]" gridrow="2" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFAE9AA3EFAF7" box="[158,274,1340,1362]" gridcol="0" gridrow="2" pageId="15" pageNumber="396">paratype</th>
<td id="76B7770DFFD50006555DFAE9AAB1FAF7" box="[329,413,1340,1362]" gridcol="1" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000655D4FAE9AAD3FAF7" box="[448,511,1340,1362]" gridcol="2" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065634FAE9A946FAF7" box="[544,618,1340,1362]" gridcol="3" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006569EFAE9A9F3FAF7" box="[650,735,1340,1362]" gridcol="4" gridrow="2" pageId="15" pageNumber="396">1.01</td>
<td id="76B7770DFFD5000656EFFAE9A863FAF7" box="[763,847,1340,1362]" gridcol="5" gridrow="2" pageId="15" pageNumber="396">1.97</td>
<td id="76B7770DFFD500065767FAE9A8F9FAF7" box="[883,981,1340,1362]" gridcol="6" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657E0FAE9AF65FAF7" box="[1012,1097,1340,1362]" gridcol="7" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065061FAE9AFEEFAF7" box="[1141,1218,1340,1362]" gridcol="8" gridrow="2" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000650F0FAE9AE1CFAF7" box="[1252,1328,1340,1362]" gridcol="9" gridrow="2" pageId="15" pageNumber="396">~1.28</td>
<td id="76B7770DFFD50006515BFAE9AE51FAF7" box="[1359,1405,1340,1362]" gridcol="10" gridrow="2" pageId="15" pageNumber="396">~2.5</td>
</tr>
<tr id="35661E71FFD50006548AFAB0AE51FADE" box="[158,1405,1381,1403]" gridrow="3" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFAB0AA3EFADE" box="[158,274,1381,1403]" gridcol="0" gridrow="3" pageId="15" pageNumber="396">1005453-A</th>
<td id="76B7770DFFD50006555DFAB0AAB1FADE" box="[329,413,1381,1403]" gridcol="1" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000655D4FAB0AAD3FADE" box="[448,511,1381,1403]" gridcol="2" gridrow="3" pageId="15" pageNumber="396">0.60</td>
<td id="76B7770DFFD500065634FAB0A946FADE" box="[544,618,1381,1403]" gridcol="3" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006569EFAB0A9F3FADE" box="[650,735,1381,1403]" gridcol="4" gridrow="3" pageId="15" pageNumber="396">1.34</td>
<td id="76B7770DFFD5000656EFFAB0A863FADE" box="[763,847,1381,1403]" gridcol="5" gridrow="3" pageId="15" pageNumber="396">1.69</td>
<td id="76B7770DFFD500065767FAB0A8F9FADE" box="[883,981,1381,1403]" gridcol="6" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657E0FAB0AF65FADE" box="[1012,1097,1381,1403]" gridcol="7" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065061FAB0AFEEFADE" box="[1141,1218,1381,1403]" gridcol="8" gridrow="3" pageId="15" pageNumber="396">2.13</td>
<td id="76B7770DFFD5000650F0FAB0AE1CFADE" box="[1252,1328,1381,1403]" gridcol="9" gridrow="3" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006515BFAB0AE51FADE" box="[1359,1405,1381,1403]" gridcol="10" gridrow="3" pageId="15" pageNumber="396">2.75</td>
</tr>
<tr id="35661E71FFD50006548AFA5BAE51FA01" box="[158,1405,1422,1444]" gridrow="4" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFA5BAA3EFA01" box="[158,274,1422,1444]" gridcol="0" gridrow="4" pageId="15" pageNumber="396">1005433-A</th>
<td id="76B7770DFFD50006555DFA5BAAB1FA01" box="[329,413,1422,1444]" gridcol="1" gridrow="4" pageId="15" pageNumber="396">1.3-1.36</td>
<td id="76B7770DFFD5000655D4FA5BAAD3FA01" box="[448,511,1422,1444]" gridcol="2" gridrow="4" pageId="15" pageNumber="396">0.65</td>
<td id="76B7770DFFD500065634FA5BA946FA01" box="[544,618,1422,1444]" gridcol="3" gridrow="4" pageId="15" pageNumber="396">0.38</td>
<td id="76B7770DFFD50006569EFA5BA9F3FA01" box="[650,735,1422,1444]" gridcol="4" gridrow="4" pageId="15" pageNumber="396">1.54-.65</td>
<td id="76B7770DFFD5000656EFFA5BA863FA01" box="[763,847,1422,1444]" gridcol="5" gridrow="4" pageId="15" pageNumber="396">1.45-.55</td>
<td id="76B7770DFFD500065767FA5BA8F9FA01" box="[883,981,1422,1444]" gridcol="6" gridrow="4" pageId="15" pageNumber="396">4.2-4.27</td>
<td id="76B7770DFFD5000657E0FA5BAF65FA01" box="[1012,1097,1422,1444]" gridcol="7" gridrow="4" pageId="15" pageNumber="396">1.64-.66</td>
<td id="76B7770DFFD500065061FA5BAFEEFA01" box="[1141,1218,1422,1444]" gridcol="8" gridrow="4" pageId="15" pageNumber="396">2.57</td>
<td id="76B7770DFFD5000650F0FA5BAE1CFA01" box="[1252,1328,1422,1444]" gridcol="9" gridrow="4" pageId="15" pageNumber="396">1.17</td>
<td id="76B7770DFFD50006515BFA5BAE51FA01" box="[1359,1405,1422,1444]" gridcol="10" gridrow="4" pageId="15" pageNumber="396">2.8</td>
</tr>
<tr id="35661E71FFD50006548AFA6DAE51FA6B" box="[158,1405,1464,1486]" gridrow="5" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFA6DAA3EFA6B" box="[158,274,1464,1486]" gridcol="0" gridrow="5" pageId="15" pageNumber="396">1005439</th>
<td id="76B7770DFFD50006555DFA6DAAB1FA6B" box="[329,413,1464,1486]" gridcol="1" gridrow="5" pageId="15" pageNumber="396">1.1-1.12</td>
<td id="76B7770DFFD5000655D4FA6DAAD3FA6B" box="[448,511,1464,1486]" gridcol="2" gridrow="5" pageId="15" pageNumber="396">0.60</td>
<td id="76B7770DFFD500065634FA6DA946FA6B" box="[544,618,1464,1486]" gridcol="3" gridrow="5" pageId="15" pageNumber="396">0.65</td>
<td id="76B7770DFFD50006569EFA6DA9F3FA6B" box="[650,735,1464,1486]" gridcol="4" gridrow="5" pageId="15" pageNumber="396">1.21</td>
<td id="76B7770DFFD5000656EFFA6DA863FA6B" box="[763,847,1464,1486]" gridcol="5" gridrow="5" pageId="15" pageNumber="396">1.83</td>
<td id="76B7770DFFD500065767FA6DA8F9FA6B" box="[883,981,1464,1486]" gridcol="6" gridrow="5" pageId="15" pageNumber="396">3.97-4.06</td>
<td id="76B7770DFFD5000657E0FA6DAF65FA6B" box="[1012,1097,1464,1486]" gridcol="7" gridrow="5" pageId="15" pageNumber="396">1.71-.75</td>
<td id="76B7770DFFD500065061FA6DAFEEFA6B" box="[1141,1218,1464,1486]" gridcol="8" gridrow="5" pageId="15" pageNumber="396">2.32</td>
<td id="76B7770DFFD5000650F0FA6DAE1CFA6B" box="[1252,1328,1464,1486]" gridcol="9" gridrow="5" pageId="15" pageNumber="396">1.20</td>
<td id="76B7770DFFD50006515BFA6DAE51FA6B" box="[1359,1405,1464,1486]" gridcol="10" gridrow="5" pageId="15" pageNumber="396">3.4</td>
</tr>
<tr id="35661E71FFD50006548AFA34AE51FA52" box="[158,1405,1505,1527]" gridrow="6" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AFA34AA3EFA52" box="[158,274,1505,1527]" gridcol="0" gridrow="6" pageId="15" pageNumber="396">holotype</th>
<td id="76B7770DFFD50006555DFA34AAB1FA52" box="[329,413,1505,1527]" gridcol="1" gridrow="6" pageId="15" pageNumber="396">0.97</td>
<td id="76B7770DFFD5000655D4FA34AAD3FA52" box="[448,511,1505,1527]" gridcol="2" gridrow="6" pageId="15" pageNumber="396">0.69</td>
<td id="76B7770DFFD500065634FA34A946FA52" box="[544,618,1505,1527]" gridcol="3" gridrow="6" pageId="15" pageNumber="396">0.88</td>
<td id="76B7770DFFD50006569EFA34A9F3FA52" box="[650,735,1505,1527]" gridcol="4" gridrow="6" pageId="15" pageNumber="396">1.17</td>
<td id="76B7770DFFD5000656EFFA34A863FA52" box="[763,847,1505,1527]" gridcol="5" gridrow="6" pageId="15" pageNumber="396">1.67</td>
<td id="76B7770DFFD500065767FA34A8F9FA52" box="[883,981,1505,1527]" gridcol="6" gridrow="6" pageId="15" pageNumber="396">4.0</td>
<td id="76B7770DFFD5000657E0FA34AF65FA52" box="[1012,1097,1505,1527]" gridcol="7" gridrow="6" pageId="15" pageNumber="396">1.89</td>
<td id="76B7770DFFD500065061FA34AFEEFA52" box="[1141,1218,1505,1527]" gridcol="8" gridrow="6" pageId="15" pageNumber="396">2.11</td>
<td id="76B7770DFFD5000650F0FA34AE1CFA52" box="[1252,1328,1505,1527]" gridcol="9" gridrow="6" pageId="15" pageNumber="396">~1.40</td>
<td id="76B7770DFFD50006515BFA34AE51FA52" box="[1359,1405,1505,1527]" gridcol="10" gridrow="6" pageId="15" pageNumber="396">3.5</td>
</tr>
<tr id="35661E71FFD50006548AF9DFAE51F985" box="[158,1405,1546,1568]" gridrow="7" pageId="15" pageNumber="396">
<th id="76B7770DFFD50006548AF9DFAA3EF985" box="[158,274,1546,1568]" gridcol="0" gridrow="7" pageId="15" pageNumber="396">1005438</th>
<td id="76B7770DFFD50006555DF9DFAAB1F985" box="[329,413,1546,1568]" gridcol="1" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000655D4F9DFAAD3F985" box="[448,511,1546,1568]" gridcol="2" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065634F9DFA946F985" box="[544,618,1546,1568]" gridcol="3" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD50006569EF9DFA9F3F985" box="[650,735,1546,1568]" gridcol="4" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000656EFF9DFA863F985" box="[763,847,1546,1568]" gridcol="5" gridrow="7" pageId="15" pageNumber="396">&gt;1.32</td>
<td id="76B7770DFFD500065767F9DFA8F9F985" box="[883,981,1546,1568]" gridcol="6" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000657E0F9DFAF65F985" box="[1012,1097,1546,1568]" gridcol="7" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD500065061F9DFAFEEF985" box="[1141,1218,1546,1568]" gridcol="8" gridrow="7" pageId="15" pageNumber="396">-</td>
<td id="76B7770DFFD5000650F0F9DFAE1CF985" box="[1252,1328,1546,1568]" gridcol="9" gridrow="7" pageId="15" pageNumber="396">&gt;1.11</td>
<td id="76B7770DFFD50006515BF9DFAE51F985" box="[1359,1405,1546,1568]" gridcol="10" gridrow="7" pageId="15" pageNumber="396">&gt;4.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE91C33FFD5FFF654D3F9B7A889F8D3" blockId="15.[151,1437,1633,2018]" pageId="15" pageNumber="396">
<emphasis id="B922C021FFD5FFF654D3F9B7AA53F9DF" box="[199,383,1634,1658]" italics="true" pageId="15" pageNumber="396">USNM 1005439</emphasis>
: Theca well preserved with proximal and mid arms (
<figureCitation id="136D00B6FFD5FFF657FBF9B4AF18F9DF" box="[1007,1076,1633,1658]" captionStart="FIGURE 9" captionStartId="14.[151,250,1495,1517]" captionTargetBox="[295,1291,921,1473]" captionTargetId="figure@14.[295,1291,921,1474]" captionTargetPageId="14" captionText="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="15" pageNumber="396">Fig. 9</figureCitation>
A–B) strikingly resembling the original figure of the 
<typeStatus id="54EDA291FFD5FFF65598F950AAC2F93B" box="[396,494,1669,1694]" pageId="15" pageNumber="396" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFC761C2FFD5FFF655EAF950A9BCF93B" author="Wilton" box="[510,656,1669,1694]" pageId="15" pageNumber="396" refString="Wilton, D. W. (1908) Resultats de l'Expedition nationale Antarctique du S. Y. Scotia, 1902 - 1904, 4, Zoologie, 1, 1 - 133." type="journal article" year="1908">Wilton 1908</bibRefCitation>
; 
<bibRefCitation id="EFC761C2FFD5FFF6568AF953A8A8F93B" author="Vaney" box="[670,900,1669,1694]" pageId="15" pageNumber="396" refString="Vaney, C. &amp; John, D. D. (1939) Scientific results of the voyage of S. Y. Scotia, 1902 - 04: The Crinoidea. Transactions of the Royal Society of Edinburg, 59, 661 - 672. http: // dx. doi. org / 10.1017 / S 0080456800017385" type="journal article" year="1939">Vaney &amp; John 1939</bibRefCitation>
). Tegmen inflated and rounded; high anal cone reaching level of Br14, significantly taller than oral cone; base of anal cone peripheral on tegmen; upper portion of cone oriented toward tegmen center (
<figureCitation id="136D00B6FFD5FFF65623F918A955F943" box="[567,633,1741,1766]" captionStart="FIGURE 9" captionStartId="14.[151,250,1495,1517]" captionTargetBox="[295,1291,921,1473]" captionTargetId="figure@14.[295,1291,921,1474]" captionTargetPageId="14" captionText="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="15" pageNumber="396">Fig. 9</figureCitation>
A–B); orals subvertical, smooth and concave with rounded tip; as many as 20 large polygonal tegminal plates without peculiar median arrangement in inter-rays (
<figureCitation id="136D00B6FFD5FFF65095F924AFE9F8AF" box="[1153,1221,1777,1802]" captionStart="FIGURE 9" captionStartId="14.[151,250,1495,1517]" captionTargetBox="[295,1291,921,1473]" captionTargetId="figure@14.[295,1291,921,1474]" captionTargetPageId="14" captionText="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="15" pageNumber="396">Fig. 9</figureCitation>
B); inter-rays A–B, C–D, A–E wider than the two other inter-rays; hydropores on adoral quarter of tegmen; conspicuous ambulacral “bridges” between Br10–11 and top of oral cone with cover plates in a gathered arrangement; outer ramifications of “bridges” corresponding to proximal pinnules not attached to tegmen.
</paragraph>
<paragraph id="8BE91C33FFD5FFE954D3F854AEA1FDBD" blockId="15.[151,1437,1633,2018]" lastBlockId="16.[151,1437,151,536]" lastPageId="16" lastPageNumber="397" pageId="15" pageNumber="396">
Proximal arm pattern 1+2 3 4 5+6 7 (5 cases) with first pinnule on Br4; brachial pair at 8+9 (3 cases) and 9+10 (1); number of successive muscular synarthries beyond the last brachial pair and before broken end: 6 (2 cases), 8 (1), 24 (1), and 16 including Br7 (1). Two ossicles of a brachial pair united by flat synostosis. Pinnule socket in distal half of brachial adorally (
<figureCitation id="136D00B6FFCAFFE95617FF42A97AFF15" box="[515,598,151,176]" captionStart="FIGURE 10" captionStartId="16.[151,250,1533,1555]" captionTargetBox="[151,1436,589,1511]" captionTargetId="figure@16.[151,1436,589,1511]" captionTargetPageId="16" captionText="FIGURE 10. Brachial muscular synarthries in Ptilocrinus brucei. A – D: specimen 1005438; E – F: specimen 1005439; A: episynostosial brachial; B – D: free brachial, B: side view showing pinnule socket, C – D: oblique adoral view, D: detail of adoral process; E – F: transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea." httpUri="https://zenodo.org/record/243059/files/figure.png" pageId="16" pageNumber="397">Fig. 10</figureCitation>
A–B); transverse muscular synarthry uniting pinnule to arm with marked fulcral ridge and deep muscular fossa on brachial (
<figureCitation id="136D00B6FFCAFFE956E7FF69A864FF71" box="[755,840,188,213]" captionStart="FIGURE 10" captionStartId="16.[151,250,1533,1555]" captionTargetBox="[151,1436,589,1511]" captionTargetId="figure@16.[151,1436,589,1511]" captionTargetPageId="16" captionText="FIGURE 10. Brachial muscular synarthries in Ptilocrinus brucei. A – D: specimen 1005438; E – F: specimen 1005439; A: episynostosial brachial; B – D: free brachial, B: side view showing pinnule socket, C – D: oblique adoral view, D: detail of adoral process; E – F: transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea." httpUri="https://zenodo.org/record/243059/files/figure.png" pageId="16" pageNumber="397">Fig. 10</figureCitation>
E–F); free brachial with strongly oblique muscular articulations (
<figureCitation id="136D00B6FFCAFFE95520FF0AAAA6FF5D" box="[308,394,223,248]" captionStart="FIGURE 10" captionStartId="16.[151,250,1533,1555]" captionTargetBox="[151,1436,589,1511]" captionTargetId="figure@16.[151,1436,589,1511]" captionTargetPageId="16" captionText="FIGURE 10. Brachial muscular synarthries in Ptilocrinus brucei. A – D: specimen 1005438; E – F: specimen 1005439; A: episynostosial brachial; B – D: free brachial, B: side view showing pinnule socket, C – D: oblique adoral view, D: detail of adoral process; E – F: transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea." httpUri="https://zenodo.org/record/243059/files/figure.png" pageId="16" pageNumber="397">Fig. 10</figureCitation>
B); muscular synarthries with ambulacral groove forming a conspicuous adoral process in mid-articulation (
<figureCitation id="136D00B6FFCAFFE95576FED1AA95FEB9" box="[354,441,260,285]" captionStart="FIGURE 10" captionStartId="16.[151,250,1533,1555]" captionTargetBox="[151,1436,589,1511]" captionTargetId="figure@16.[151,1436,589,1511]" captionTargetPageId="16" captionText="FIGURE 10. Brachial muscular synarthries in Ptilocrinus brucei. A – D: specimen 1005438; E – F: specimen 1005439; A: episynostosial brachial; B – D: free brachial, B: side view showing pinnule socket, C – D: oblique adoral view, D: detail of adoral process; E – F: transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea." httpUri="https://zenodo.org/record/243059/files/figure.png" pageId="16" pageNumber="397">Fig. 10</figureCitation>
C–D). Adoral proximal arm inflated with numerous lateral plates not aligned, and conspicuous festooned cover plates in a gathered arrangement; proximal part of pinnules with proximal genital inflation of asymmetric, multiplated architecture, and cover plates poorly differentiated (
<figureCitation id="136D00B6FFCAFFE95066FE99AF93FEC1" box="[1138,1215,332,357]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="16" pageNumber="397">Fig. 11</figureCitation>
A–B); 3 (maximum 4) irregular plate rows on convex side decreasing to 1 row distally (
<figureCitation id="136D00B6FFCAFFE9578FFEBAA8C1FE2D" box="[923,1005,367,392]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="16" pageNumber="397">Fig. 11</figureCitation>
B); concave side without rows; lateral plates small or disappearing in mid arm with cover plates rounded or elliptical; muscular synarthries of first pinnular with brachial (
<figureCitation id="136D00B6FFCAFFE955B4FE62AADDFE75" box="[416,497,439,464]" captionStart="FIGURE 10" captionStartId="16.[151,250,1533,1555]" captionTargetBox="[151,1436,589,1511]" captionTargetId="figure@16.[151,1436,589,1511]" captionTargetPageId="16" captionText="FIGURE 10. Brachial muscular synarthries in Ptilocrinus brucei. A – D: specimen 1005438; E – F: specimen 1005439; A: episynostosial brachial; B – D: free brachial, B: side view showing pinnule socket, C – D: oblique adoral view, D: detail of adoral process; E – F: transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea." httpUri="https://zenodo.org/record/243059/files/figure.png" pageId="16" pageNumber="397">Fig. 10</figureCitation>
E) and with second pinnular (
<figureCitation id="136D00B6FFCAFFE95754FE62A8A3FE75" box="[832,911,439,464]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="16" pageNumber="397">Fig. 11</figureCitation>
E) having perpendicular fulcral ridge allowing wide amplitude of movements; symmetrical synostosis between second and third pinnular (
<figureCitation id="136D00B6FFCAFFE95082FE09AFC8FE51" box="[1174,1252,476,501]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="16" pageNumber="397">Fig. 11</figureCitation>
F–G); following pinnulars asymmetrical, supporting genital inflation (
<figureCitation id="136D00B6FFCAFFE956FFFE2AA815FDBD" box="[747,825,511,536]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="16" pageNumber="397">Fig. 11</figureCitation>
H–I); pinnulars becoming more symmetrical distally.
</paragraph>
<caption id="DF294CBBFFCAFFE95483FA28A970F928" httpUri="https://zenodo.org/record/243059/files/figure.png" pageId="16" pageNumber="397" targetBox="[151,1436,589,1511]" targetPageId="16">
<paragraph id="8BE91C33FFCAFFE95483FA28A970F928" blockId="16.[151,1436,1532,1677]" pageId="16" pageNumber="397">
<emphasis id="B922C021FFCAFFE95483FA28AA08F9B7" bold="true" box="[151,292,1533,1555]" pageId="16" pageNumber="397">FIGURE 10.</emphasis>
Brachial muscular synarthries in 
<taxonomicName id="4C5667B0FFCAFFE9569AFA29A866F9B7" box="[654,842,1532,1554]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="16" pageNumber="397" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCAFFE9569AFA29A866F9B7" box="[654,842,1532,1554]" italics="true" pageId="16" pageNumber="397">Ptilocrinus brucei</emphasis>
</taxonomicName>
. A–D: specimen 1005438; E–F: specimen 1005439; A: episynostosial brachial; B–D: free brachial, B: side view showing pinnule socket, C–D: oblique adoral view, D: detail of adoral process; E–F:transverse synarthry articulating pinnule on brachial, E: proximal facet of first pinnular, F: oblique adoral view of pinnule socket with roof over muscular fossea broken (arrow); ap: adoral process, il: inner ligament area, ol: outer ligament area, m: muscular area, mf: muscular fossea.
</paragraph>
</caption>
<paragraph id="8BE91C33FFCAFFE954D3F96FAAD5F856" blockId="16.[151,1437,1722,2035]" pageId="16" pageNumber="397">
Length of proximal stalk attached to aboral cup 5.0 mm with 10 columnals; proximalmost diameter 
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; cup aboral diameter 
<quantity id="4CAEB1D6FFCAFFE95595F90AAAF0F953" box="[385,476,1759,1783]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="397" unit="mm" value="3.4">3.4 mm</quantity>
; stalk distal diameter 3.0 mm. Stalk fragment with distal proxistele and transition to mesistele: length 
<quantity id="4CAEB1D6FFCAFFE9554AF8D7AAFEF8BF" box="[350,466,1794,1819]" metricMagnitude="-2" metricUnit="m" metricValue="1.705" pageId="16" pageNumber="397" unit="mm" value="17.05">17.05 mm</quantity>
with 22 columnals; proximal diameter 
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with maximum columnal height 
<quantity id="4CAEB1D6FFCAFFE9516DF8D7ABE9F89B" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="397" unit="mm" value="0.9">0.9 mm</quantity>
, and symplexy with 11 crenular units of 2–3 crenulae (fig. 12 B–C); distal diameter 
<quantity id="4CAEB1D6FFCAFFE95084F8F2AFC4F89B" box="[1168,1256,1831,1855]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="397" unit="mm" value="2.8">2.8 mm</quantity>
with maximum columnal height 
<quantity id="4CAEB1D6FFCAFFE9554CF89FAA9DF8C7" box="[344,433,1866,1891]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="397" unit="mm" value="1.6">1.6 mm</quantity>
and symplexy with 11 crenular units of 1 robust crenula (
<figureCitation id="136D00B6FFCAFFE95042F89FAF85F8C7" box="[1110,1193,1866,1891]" captionStart="FIGURE 12" captionStartId="18.[151,250,1272,1294]" captionTargetBox="[336,1245,373,1243]" captionTargetId="figure@18.[336,1245,373,1243]" captionTargetPageId="18" captionText="FIGURE 12. Proximal stalk symplexies in large specimens of Ptilocrinus brucei. A: specimen USNM 1005453, B – D: specimen USNM 1005439; A: proximalmost columnal; B: at a distance of 5.0 mm from aboral cup; C: at a distance of 6.0 mm from aboral cup; D: at a distance of 22.0 mm from aboral cup." httpUri="https://zenodo.org/record/243061/files/figure.png" pageId="16" pageNumber="397">Fig. 12</figureCitation>
D); inner areola and probably claustrum depressed proximally (diameter 
<quantity id="4CAEB1D6FFCAFFE956F1F8BAA813F822" box="[741,831,1903,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="16" pageNumber="397" unit="mm" value="1.3">1.3 mm</quantity>
), with round lacuna of diameter 
<quantity id="4CAEB1D6FFCAFFE950A4F8BBAE26F823" box="[1200,1290,1902,1927]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="397" unit="mm" value="0.5">0.5 mm</quantity>
; inner areola surrounded by coarsely-meshed stereom; stereom of depressed area progressively resorbed distally, resulting in a larger axial canal (diameter 1.0 mm) (
<figureCitation id="136D00B6FFCAFFE95654F863A9BDF86A" box="[576,657,1974,1999]" captionStart="FIGURE 12" captionStartId="18.[151,250,1272,1294]" captionTargetBox="[336,1245,373,1243]" captionTargetId="figure@18.[336,1245,373,1243]" captionTargetPageId="18" captionText="FIGURE 12. Proximal stalk symplexies in large specimens of Ptilocrinus brucei. A: specimen USNM 1005453, B – D: specimen USNM 1005439; A: proximalmost columnal; B: at a distance of 5.0 mm from aboral cup; C: at a distance of 6.0 mm from aboral cup; D: at a distance of 22.0 mm from aboral cup." httpUri="https://zenodo.org/record/243061/files/figure.png" pageId="16" pageNumber="397">Fig. 12</figureCitation>
B–D); ratio of axial canal to columnal diameter in this fragment from 0.44 (proximal) to 0.36 (distal).
</paragraph>
<caption id="DF294CBBFFCBFFE85483F9ECA94BF94C" httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="17" pageNumber="398" targetBox="[215,1372,193,1568]" targetPageId="17">
<paragraph id="8BE91C33FFCBFFE85483F9ECA94BF94C" blockId="17.[151,1436,1593,1769]" pageId="17" pageNumber="398">
<emphasis id="B922C021FFCBFFE85483F9ECAA33F9EB" bold="true" box="[151,287,1593,1615]" pageId="17" pageNumber="398">FIGURE 11.</emphasis>
Architecture and articulations in 
<taxonomicName id="4C5667B0FFCBFFE85661F9ECA802F9EA" box="[629,814,1593,1615]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="17" pageNumber="398" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCBFFE85661F9ECA802F9EA" box="[629,814,1593,1615]" italics="true" pageId="17" pageNumber="398">Ptilocrinus brucei</emphasis>
</taxonomicName>
pinnules. A–B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C–D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E–I: specimen USNM 1005439; E: proximal muscular synarthry of Pn2; F: aboral distal view of Pn2; G: proximal synostose of Pn3; H: pinnule broken proximally to Pn4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn4 (adoral plates removed, orientation like in H).
</paragraph>
</caption>
<paragraph id="8BE91C33FFCBFFEB54D3F8C3AA5CFEE5" blockId="17.[151,1436,1813,2018]" lastBlockId="18.[151,1436,151,320]" lastPageId="18" lastPageNumber="399" pageId="17" pageNumber="398">
<emphasis id="B922C021FFCBFFE854D3F8C3AAADF88B" box="[199,385,1814,1838]" italics="true" pageId="17" pageNumber="398">USNM 1005438</emphasis>
: Large specimen in fragments, lacking aboral cup: radials attached to arms, fragment of inflated tegmen and tall anal tube, and complete stalk except proximalmost columnals. Proximal arm pattern 1+2 3 4 5+6 7 8 with first pinnule on Br4 (5 cases), brachial pair at 9+10 (1), 10+11 (2), and no other brachial pair (1); fifth arm broken after Br8; mid arm with free brachial only; maximum number of muscular articulations before broken end 21; distal arm fragments rarely with one brachial pair; muscular articulations oblique beyond Br4; Br1 to Br3 with lateral wings. Height and width of brachials moderately decreasing from Br1 to Br3 and very little beyond Br3 (
<tableCitation id="C6D42988FFC8FFEB5521FF42AAB3FF15" box="[309,415,151,176]" captionStart="TABLE 11" captionStartId="18.[151,239,1401,1424]" captionTargetBox="[159,1363,1452,1598]" captionTargetPageId="18" captionText="TABLE 11. Ptilocrinus (Chambersaecrinus) brucei USNM 1005438. Quantitative characters of proximal brachials." httpUri="http://table.plazi.org/id/DF294CBBFFC8FFEB5483FAACAE41FA35" pageId="18" pageNumber="399" tableUuid="DF294CBBFFC8FFEB5483FAACAE41FA35">Table 11</tableCitation>
). Arms attached to tegmen to Br9; adoral face of arm inflated and multiplated, with festooned cover plates in a gathered arrangement; genital inflation of some pinnules with numerous imbricated irregular polygonal plates not in rows (
<figureCitation id="136D00B6FFC8FFEB5642FF0AA98BFF5D" box="[598,679,223,248]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="18" pageNumber="399">Fig. 11</figureCitation>
C), covering exceptionally preserved gonads with oocytes of 100– 150µ (
<figureCitation id="136D00B6FFC8FFEB54F5FED1AA1CFEB9" box="[225,304,260,285]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="18" pageNumber="399">Fig. 11</figureCitation>
D); other pinnules with asymmetric architecture described above; arm and pinnule articulations as in previous specimen.
</paragraph>
<caption id="DF294CBBFFC8FFEB5483FB2DA821FAEE" httpUri="https://zenodo.org/record/243061/files/figure.png" pageId="18" pageNumber="399" targetBox="[336,1245,373,1243]" targetPageId="18">
<paragraph id="8BE91C33FFC8FFEB5483FB2DA821FAEE" blockId="18.[151,1437,1272,1355]" pageId="18" pageNumber="399">
<emphasis id="B922C021FFC8FFEB5483FB2DAA09FAA8" bold="true" box="[151,293,1272,1294]" pageId="18" pageNumber="399">FIGURE 12.</emphasis>
Proximal stalk symplexies in large specimens of 
<taxonomicName id="4C5667B0FFC8FFEB5752FB2DAF2FFAAB" box="[838,1027,1272,1294]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="18" pageNumber="399" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFC8FFEB5752FB2DAF2FFAAB" box="[838,1027,1272,1294]" italics="true" pageId="18" pageNumber="399">Ptilocrinus brucei</emphasis>
</taxonomicName>
. A: specimen USNM 1005453, B–D: specimen USNM 1005439; A: proximalmost columnal; B: at a distance of 5.0 mm from aboral cup; C: at a distance of 6.0 mm from aboral cup; D: at a distance of 22.0 mm from aboral cup.
</paragraph>
</caption>
<caption id="DF294CBBFFC8FFEB5483FAACAE41FA35" ID-Table-UUID="DF294CBBFFC8FFEB5483FAACAE41FA35" box="[151,1389,1401,1424]" httpUri="http://table.plazi.org/id/DF294CBBFFC8FFEB5483FAACAE41FA35" pageId="18" pageNumber="399" targetBox="[159,1363,1452,1598]" targetIsTable="true" targetPageId="18">
<paragraph id="8BE91C33FFC8FFEB5483FAACAE41FA35" blockId="18.[151,1389,1401,1424]" box="[151,1389,1401,1424]" pageId="18" pageNumber="399">
<emphasis id="B922C021FFC8FFEB5483FAACAA3AFA35" bold="true" box="[151,278,1401,1424]" pageId="18" pageNumber="399">TABLE 11.</emphasis>
<taxonomicName id="4C5667B0FFC8FFEB5508FAACA993FA35" box="[284,703,1401,1424]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="18" pageNumber="399" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFC8FFEB5508FAACA993FA35" box="[284,703,1401,1424]" italics="true" pageId="18" pageNumber="399">Ptilocrinus (Chambersaecrinus) brucei</emphasis>
</taxonomicName>
USNM 1005438. Quantitative characters of proximal brachials.
</paragraph>
</caption>
<paragraph id="8BE91C33FFC8FFEB548BFA79AE7FF99B" pageId="18" pageNumber="399">
<table id="F956EE93FFC80006548BFA79AE7FF99B" box="[159,1363,1452,1598]" gridcols="9" gridrows="4" pageId="18" pageNumber="399">
<tr id="35661E71FFC80006548BFA79AE7FFA67" box="[159,1363,1452,1474]" gridrow="0" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548BFA79AA61FA67" box="[159,333,1452,1474]" gridcol="0" gridrow="0" pageId="18" pageNumber="399">Brachial rank</th>
<th id="76B7770DFFC8000655D2FA79AADEFA67" box="[454,498,1452,1474]" gridcol="1" gridrow="0" pageId="18" pageNumber="399">Br1</th>
<th id="76B7770DFFC80006565AFA79A956FA67" box="[590,634,1452,1474]" gridcol="2" gridrow="0" pageId="18" pageNumber="399">Br2</th>
<th id="76B7770DFFC8000656D4FA79A9C0FA67" box="[704,748,1452,1474]" gridcol="3" gridrow="0" pageId="18" pageNumber="399">Br3</th>
<th id="76B7770DFFC800065727FA79A873FA67" box="[819,863,1452,1474]" gridcol="4" gridrow="0" pageId="18" pageNumber="399">Br4</th>
<th id="76B7770DFFC8000657A4FA79A8F0FA67" box="[944,988,1452,1474]" gridcol="5" gridrow="0" pageId="18" pageNumber="399">Br5</th>
<th id="76B7770DFFC800065039FA79AF75FA67" box="[1069,1113,1452,1474]" gridcol="6" gridrow="0" pageId="18" pageNumber="399">Br6</th>
<th id="76B7770DFFC8000650BEFA79AFFAFA67" box="[1194,1238,1452,1474]" gridcol="7" gridrow="0" pageId="18" pageNumber="399">Br7</th>
<th id="76B7770DFFC800065133FA79AE7FFA67" box="[1319,1363,1452,1474]" gridcol="8" gridrow="0" pageId="18" pageNumber="399">Br8</th>
</tr>
<tr id="35661E71FFC80006548BFA00AE7FFA4E" box="[159,1363,1493,1515]" gridrow="1" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548BFA00AA61FA4E" box="[159,333,1493,1515]" gridcol="0" gridrow="1" pageId="18" pageNumber="399">Upper width (W)</th>
<td id="76B7770DFFC8000655D2FA00AADEFA4E" box="[454,498,1493,1515]" gridcol="1" gridrow="1" pageId="18" pageNumber="399">5.2</td>
<td id="76B7770DFFC80006565AFA00A956FA4E" box="[590,634,1493,1515]" gridcol="2" gridrow="1" pageId="18" pageNumber="399">5.3</td>
<td id="76B7770DFFC8000656D4FA00A9C0FA4E" box="[704,748,1493,1515]" gridcol="3" gridrow="1" pageId="18" pageNumber="399">4.1</td>
<td id="76B7770DFFC800065727FA00A873FA4E" box="[819,863,1493,1515]" gridcol="4" gridrow="1" pageId="18" pageNumber="399">3.6</td>
<td id="76B7770DFFC8000657A4FA00A8F0FA4E" box="[944,988,1493,1515]" gridcol="5" gridrow="1" pageId="18" pageNumber="399">3.6</td>
<td id="76B7770DFFC800065039FA00AF75FA4E" box="[1069,1113,1493,1515]" gridcol="6" gridrow="1" pageId="18" pageNumber="399">3.6</td>
<td id="76B7770DFFC8000650BEFA00AFFAFA4E" box="[1194,1238,1493,1515]" gridcol="7" gridrow="1" pageId="18" pageNumber="399">3.5</td>
<td id="76B7770DFFC800065133FA00AE7FFA4E" box="[1319,1363,1493,1515]" gridcol="8" gridrow="1" pageId="18" pageNumber="399">3.5</td>
</tr>
<tr id="35661E71FFC80006548BFA2AAE7FF9B0" box="[159,1363,1535,1557]" gridrow="2" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548BFA2AAA61F9B0" box="[159,333,1535,1557]" gridcol="0" gridrow="2" pageId="18" pageNumber="399">Maxi height (H)</th>
<td id="76B7770DFFC8000655D2FA2AAADEF9B0" box="[454,498,1535,1557]" gridcol="1" gridrow="2" pageId="18" pageNumber="399">4.6</td>
<td id="76B7770DFFC80006565AFA2AA956F9B0" box="[590,634,1535,1557]" gridcol="2" gridrow="2" pageId="18" pageNumber="399">4.0</td>
<td id="76B7770DFFC8000656D4FA2AA9C0F9B0" box="[704,748,1535,1557]" gridcol="3" gridrow="2" pageId="18" pageNumber="399">1.7</td>
<td id="76B7770DFFC800065727FA2AA873F9B0" box="[819,863,1535,1557]" gridcol="4" gridrow="2" pageId="18" pageNumber="399">1.4</td>
<td id="76B7770DFFC8000657A4FA2AA8F0F9B0" box="[944,988,1535,1557]" gridcol="5" gridrow="2" pageId="18" pageNumber="399">1.3</td>
<td id="76B7770DFFC800065039FA2AAF75F9B0" box="[1069,1113,1535,1557]" gridcol="6" gridrow="2" pageId="18" pageNumber="399">1.4</td>
<td id="76B7770DFFC8000650BEFA2AAFFAF9B0" box="[1194,1238,1535,1557]" gridcol="7" gridrow="2" pageId="18" pageNumber="399">1.4</td>
<td id="76B7770DFFC800065133FA2AAE7FF9B0" box="[1319,1363,1535,1557]" gridcol="8" gridrow="2" pageId="18" pageNumber="399">1.3</td>
</tr>
<tr id="35661E71FFC80006548BF9FDAE7FF99B" box="[159,1363,1576,1598]" gridrow="3" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548BF9FDAA61F99B" box="[159,333,1576,1598]" gridcol="0" gridrow="3" pageId="18" pageNumber="399">H/W</th>
<td id="76B7770DFFC8000655D2F9FDAADEF99B" box="[454,498,1576,1598]" gridcol="1" gridrow="3" pageId="18" pageNumber="399">0.88</td>
<td id="76B7770DFFC80006565AF9FDA956F99B" box="[590,634,1576,1598]" gridcol="2" gridrow="3" pageId="18" pageNumber="399">0.75</td>
<td id="76B7770DFFC8000656D4F9FDA9C0F99B" box="[704,748,1576,1598]" gridcol="3" gridrow="3" pageId="18" pageNumber="399">0.41</td>
<td id="76B7770DFFC800065727F9FDA873F99B" box="[819,863,1576,1598]" gridcol="4" gridrow="3" pageId="18" pageNumber="399">0.39</td>
<td id="76B7770DFFC8000657A4F9FDA8F0F99B" box="[944,988,1576,1598]" gridcol="5" gridrow="3" pageId="18" pageNumber="399">0.36</td>
<td id="76B7770DFFC800065039F9FDAF75F99B" box="[1069,1113,1576,1598]" gridcol="6" gridrow="3" pageId="18" pageNumber="399">0.39</td>
<td id="76B7770DFFC8000650BEF9FDAFFAF99B" box="[1194,1238,1576,1598]" gridcol="7" gridrow="3" pageId="18" pageNumber="399">0.40</td>
<td id="76B7770DFFC800065133F9FDAE7FF99B" box="[1319,1363,1576,1598]" gridcol="8" gridrow="3" pageId="18" pageNumber="399">0.37</td>
</tr>
</table>
</paragraph>
<caption id="DF294CBBFFC8FFEB5483F954A939F97D" ID-Table-UUID="DF294CBBFFC8FFEB5483F954A939F97D" httpUri="http://table.plazi.org/id/DF294CBBFFC8FFEB5483F954A939F97D" pageId="18" pageNumber="399" targetBox="[154,1393,1780,1967]" targetIsTable="true" targetPageId="18">
<paragraph id="8BE91C33FFC8FFEB5483F954A939F97D" blockId="18.[151,1435,1665,1752]" pageId="18" pageNumber="399">
<emphasis id="B922C021FFC8FFEB5483F954AA34F93D" bold="true" box="[151,280,1665,1688]" pageId="18" pageNumber="399">TABLE 12.</emphasis>
<taxonomicName id="4C5667B0FFC8FFEB5534F954A9E9F93D" box="[288,709,1665,1688]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="18" pageNumber="399" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFC8FFEB5534F954A9E9F93D" box="[288,709,1665,1688]" italics="true" pageId="18" pageNumber="399">Ptilocrinus (Chambersaecrinus) brucei</emphasis>
</taxonomicName>
USNM 1005438. Quantitative characters of columnals. Columnals numbered from proximal to distal. Dal: axial lumen diameter, Dcol: columnal diameter, Hcol: maximum columnal height. Values in mm, except ratios.
</paragraph>
</caption>
<paragraph id="8BE91C33FFC8FFEB548EF920AE5DF80A" pageId="18" pageNumber="399">
<table id="F956EE93FFC80006548EF921AE5DF80A" box="[154,1393,1780,1967]" gridcols="12" gridrows="5" pageId="18" pageNumber="399">
<tr id="35661E71FFC80006548EF921AE5DF8AF" box="[154,1393,1780,1802]" gridrow="0" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548EF921AA6DF8AF" box="[154,321,1780,1802]" gridcol="0" gridrow="0" pageId="18" pageNumber="399">From cup base</th>
<th id="76B7770DFFC80006556EF921AA84F8AF" box="[378,424,1780,1802]" gridcol="1" gridrow="0" pageId="18" pageNumber="399">~0</th>
<th id="76B7770DFFC8000655C1F921A92FF8AF" box="[469,515,1780,1802]" gridcol="2" gridrow="0" pageId="18" pageNumber="399">15</th>
<th id="76B7770DFFC80006563BF921A977F8AF" box="[559,603,1780,1802]" gridcol="3" gridrow="0" pageId="18" pageNumber="399">40</th>
<th id="76B7770DFFC800065686F921A992F8AF" box="[658,702,1780,1802]" gridcol="4" gridrow="0" pageId="18" pageNumber="399">60</th>
<th id="76B7770DFFC8000656E1F921A80DF8AF" box="[757,801,1780,1802]" gridcol="5" gridrow="0" pageId="18" pageNumber="399">100</th>
<th id="76B7770DFFC80006574CF921A8A8F8AF" box="[856,900,1780,1802]" gridcol="6" gridrow="0" pageId="18" pageNumber="399">150</th>
<th id="76B7770DFFC8000657AEF921A8D8F8AF" box="[954,1012,1780,1802]" gridcol="7" gridrow="0" pageId="18" pageNumber="399">200</th>
<th id="76B7770DFFC800065009F921AF65F8AF" box="[1053,1097,1780,1802]" gridcol="8" gridrow="0" pageId="18" pageNumber="399">250</th>
<th id="76B7770DFFC800065094F921AF80F8AF" box="[1152,1196,1780,1802]" gridcol="9" gridrow="0" pageId="18" pageNumber="399">300</th>
<th id="76B7770DFFC8000650F7F921AE23F8AF" box="[1251,1295,1780,1802]" gridcol="10" gridrow="0" pageId="18" pageNumber="399">350</th>
<th id="76B7770DFFC800065151F921AE5DF8AF" box="[1349,1393,1780,1802]" gridcol="11" gridrow="0" pageId="18" pageNumber="399">386</th>
</tr>
<tr id="35661E71FFC80006548EF8C8AE5DF896" box="[154,1393,1821,1843]" gridrow="1" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548EF8C8AA6DF896" box="[154,321,1821,1843]" gridcol="0" gridrow="1" pageId="18" pageNumber="399">Diameter (Dcol)</th>
<td id="76B7770DFFC80006556EF8C8AA84F896" box="[378,424,1821,1843]" gridcol="1" gridrow="1" pageId="18" pageNumber="399">4.15</td>
<td id="76B7770DFFC8000655C1F8C8A92FF896" box="[469,515,1821,1843]" gridcol="2" gridrow="1" pageId="18" pageNumber="399">4.0</td>
<td id="76B7770DFFC80006563BF8C8A977F896" box="[559,603,1821,1843]" gridcol="3" gridrow="1" pageId="18" pageNumber="399">3.9</td>
<td id="76B7770DFFC800065686F8C8A992F896" box="[658,702,1821,1843]" gridcol="4" gridrow="1" pageId="18" pageNumber="399">3.85</td>
<td id="76B7770DFFC8000656E1F8C8A80DF896" box="[757,801,1821,1843]" gridcol="5" gridrow="1" pageId="18" pageNumber="399">3.75</td>
<td id="76B7770DFFC80006574CF8C8A8A8F896" box="[856,900,1821,1843]" gridcol="6" gridrow="1" pageId="18" pageNumber="399">4.1</td>
<td id="76B7770DFFC8000657AEF8C8A8D8F896" box="[954,1012,1821,1843]" gridcol="7" gridrow="1" pageId="18" pageNumber="399">4.1</td>
<td id="76B7770DFFC800065009F8C8AF65F896" box="[1053,1097,1821,1843]" gridcol="8" gridrow="1" pageId="18" pageNumber="399">3.9</td>
<td id="76B7770DFFC800065094F8C8AF80F896" box="[1152,1196,1821,1843]" gridcol="9" gridrow="1" pageId="18" pageNumber="399">4.1</td>
<td id="76B7770DFFC8000650F7F8C8AE23F896" box="[1251,1295,1821,1843]" gridcol="10" gridrow="1" pageId="18" pageNumber="399">4.6</td>
<td id="76B7770DFFC800065151F8C8AE5DF896" box="[1349,1393,1821,1843]" gridcol="11" gridrow="1" pageId="18" pageNumber="399">5.95</td>
</tr>
<tr id="35661E71FFC80006548EF892AE5DF8F8" box="[154,1393,1863,1885]" gridrow="2" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548EF892AA6DF8F8" box="[154,321,1863,1885]" gridcol="0" gridrow="2" pageId="18" pageNumber="399">Height (Hcol)</th>
<td id="76B7770DFFC80006556EF892AA84F8F8" box="[378,424,1863,1885]" gridcol="1" gridrow="2" pageId="18" pageNumber="399">&lt;07</td>
<td id="76B7770DFFC8000655C1F892A92FF8F8" box="[469,515,1863,1885]" gridcol="2" gridrow="2" pageId="18" pageNumber="399">&lt;07</td>
<td id="76B7770DFFC80006563BF892A977F8F8" box="[559,603,1863,1885]" gridcol="3" gridrow="2" pageId="18" pageNumber="399">1.0</td>
<td id="76B7770DFFC800065686F892A992F8F8" box="[658,702,1863,1885]" gridcol="4" gridrow="2" pageId="18" pageNumber="399">1.5</td>
<td id="76B7770DFFC8000656E1F892A80DF8F8" box="[757,801,1863,1885]" gridcol="5" gridrow="2" pageId="18" pageNumber="399">1.4</td>
<td id="76B7770DFFC80006574CF892A8A8F8F8" box="[856,900,1863,1885]" gridcol="6" gridrow="2" pageId="18" pageNumber="399">1.3</td>
<td id="76B7770DFFC8000657AEF892A8D8F8F8" box="[954,1012,1863,1885]" gridcol="7" gridrow="2" pageId="18" pageNumber="399">1.7</td>
<td id="76B7770DFFC800065009F892AF65F8F8" box="[1053,1097,1863,1885]" gridcol="8" gridrow="2" pageId="18" pageNumber="399">1.6</td>
<td id="76B7770DFFC800065094F892AF80F8F8" box="[1152,1196,1863,1885]" gridcol="9" gridrow="2" pageId="18" pageNumber="399">1.4</td>
<td id="76B7770DFFC8000650F7F892AE23F8F8" box="[1251,1295,1863,1885]" gridcol="10" gridrow="2" pageId="18" pageNumber="399">1.4</td>
<td id="76B7770DFFC800065151F892AE5DF8F8" box="[1349,1393,1863,1885]" gridcol="11" gridrow="2" pageId="18" pageNumber="399">1.1</td>
</tr>
<tr id="35661E71FFC80006548EF8A5AE5DF823" box="[154,1393,1904,1926]" gridrow="3" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548EF8A5AA6DF823" box="[154,321,1904,1926]" gridcol="0" gridrow="3" pageId="18" pageNumber="399">Hcol/Dcol</th>
<td id="76B7770DFFC80006556EF8A5AA84F823" box="[378,424,1904,1926]" gridcol="1" gridrow="3" pageId="18" pageNumber="399">&lt;0.2</td>
<td id="76B7770DFFC8000655C1F8A5A92FF823" box="[469,515,1904,1926]" gridcol="2" gridrow="3" pageId="18" pageNumber="399">&lt;0.2</td>
<td id="76B7770DFFC80006563BF8A5A977F823" box="[559,603,1904,1926]" gridcol="3" gridrow="3" pageId="18" pageNumber="399">0.34</td>
<td id="76B7770DFFC800065686F8A5A992F823" box="[658,702,1904,1926]" gridcol="4" gridrow="3" pageId="18" pageNumber="399">0.39</td>
<td id="76B7770DFFC8000656E1F8A5A80DF823" box="[757,801,1904,1926]" gridcol="5" gridrow="3" pageId="18" pageNumber="399">0.37</td>
<td id="76B7770DFFC80006574CF8A5A8A8F823" box="[856,900,1904,1926]" gridcol="6" gridrow="3" pageId="18" pageNumber="399">0.32</td>
<td id="76B7770DFFC8000657AEF8A5A8D8F823" box="[954,1012,1904,1926]" gridcol="7" gridrow="3" pageId="18" pageNumber="399">0.415</td>
<td id="76B7770DFFC800065009F8A5AF65F823" box="[1053,1097,1904,1926]" gridcol="8" gridrow="3" pageId="18" pageNumber="399">0.41</td>
<td id="76B7770DFFC800065094F8A5AF80F823" box="[1152,1196,1904,1926]" gridcol="9" gridrow="3" pageId="18" pageNumber="399">0.34</td>
<td id="76B7770DFFC8000650F7F8A5AE23F823" box="[1251,1295,1904,1926]" gridcol="10" gridrow="3" pageId="18" pageNumber="399">0.30</td>
<td id="76B7770DFFC800065151F8A5AE5DF823" box="[1349,1393,1904,1926]" gridcol="11" gridrow="3" pageId="18" pageNumber="399">0.19</td>
</tr>
<tr id="35661E71FFC80006548EF84CAE5DF80A" box="[154,1393,1945,1967]" gridrow="4" pageId="18" pageNumber="399">
<th id="76B7770DFFC80006548EF84CAA6DF80A" box="[154,321,1945,1967]" gridcol="0" gridrow="4" pageId="18" pageNumber="399">Dal/Dcol</th>
<td id="76B7770DFFC80006556EF84CAA84F80A" box="[378,424,1945,1967]" gridcol="1" gridrow="4" pageId="18" pageNumber="399">-</td>
<td id="76B7770DFFC8000655C1F84CA92FF80A" box="[469,515,1945,1967]" gridcol="2" gridrow="4" pageId="18" pageNumber="399">0.45</td>
<td id="76B7770DFFC80006563BF84CA977F80A" box="[559,603,1945,1967]" gridcol="3" gridrow="4" pageId="18" pageNumber="399">-</td>
<td id="76B7770DFFC800065686F84CA992F80A" box="[658,702,1945,1967]" gridcol="4" gridrow="4" pageId="18" pageNumber="399">0.45</td>
<td id="76B7770DFFC8000656E1F84CA80DF80A" box="[757,801,1945,1967]" gridcol="5" gridrow="4" pageId="18" pageNumber="399">-</td>
<td id="76B7770DFFC80006574CF84CA8A8F80A" box="[856,900,1945,1967]" gridcol="6" gridrow="4" pageId="18" pageNumber="399">-</td>
<td id="76B7770DFFC8000657AEF84CA8D8F80A" box="[954,1012,1945,1967]" gridcol="7" gridrow="4" pageId="18" pageNumber="399">0.31</td>
<td id="76B7770DFFC800065009F84CAF65F80A" box="[1053,1097,1945,1967]" gridcol="8" gridrow="4" pageId="18" pageNumber="399">-</td>
<td id="76B7770DFFC800065094F84CAF80F80A" box="[1152,1196,1945,1967]" gridcol="9" gridrow="4" pageId="18" pageNumber="399">0.25</td>
<td id="76B7770DFFC8000650F7F84CAE23F80A" box="[1251,1295,1945,1967]" gridcol="10" gridrow="4" pageId="18" pageNumber="399">0.14</td>
<td id="76B7770DFFC800065151F84CAE5DF80A" box="[1349,1393,1945,1967]" gridcol="11" gridrow="4" pageId="18" pageNumber="399">0.09</td>
</tr>
</table>
</paragraph>
<caption id="DF294CBBFFC9FFEA5483F91AAADCF8E4" httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="19" pageNumber="400" targetBox="[151,1436,193,1720]" targetPageId="19">
<paragraph id="8BE91C33FFC9FFEA5483F91AAADCF8E4" blockId="19.[151,1437,1743,1857]" pageId="19" pageNumber="400">
<emphasis id="B922C021FFC9FFEA5483F91AAA33F941" bold="true" box="[151,287,1743,1765]" pageId="19" pageNumber="400">FIGURE 13.</emphasis>
Changes in columnal facet and axial canal along the stalk of 
<taxonomicName id="4C5667B0FFC9FFEA579AF91AAF6AF940" box="[910,1094,1743,1765]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="19" pageNumber="400" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFC9FFEA579AF91AAF6AF940" box="[910,1094,1743,1765]" italics="true" pageId="19" pageNumber="400">Ptilocrinus brucei</emphasis>
</taxonomicName>
specimen USNM 1005438. A–F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D–E: in transition mesistele/dististele; F: in mid dististele; G–L: axial canal; J–L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele.
</paragraph>
</caption>
<paragraph id="8BE91C33FFC9FFED54D3F8BDA9B2FE50" blockId="19.[151,1436,1896,2030]" lastBlockId="20.[151,1437,151,861]" lastPageId="20" lastPageNumber="401" pageId="19" pageNumber="400">
Remaining stalk length 
<quantity id="4CAEB1D6FFC9FFEA55C5F8BDA902F825" box="[465,558,1896,1921]" metricMagnitude="-1" metricUnit="m" metricValue="3.86" pageId="19" pageNumber="400" unit="mm" value="386.0">386 mm</quantity>
(five fragments without gap); 
<tableCitation id="C6D42988FFC9FFEA5790F8BDA8C8F824" box="[900,996,1896,1921]" captionStart="TABLE 12" captionStartId="18.[151,239,1665,1688]" captionTargetBox="[154,1393,1780,1967]" captionTargetPageId="18" captionText="TABLE 12. Ptilocrinus (Chambersaecrinus) brucei USNM 1005438. Quantitative characters of columnals. Columnals numbered from proximal to distal. Dal: axial lumen diameter, Dcol: columnal diameter, Hcol: maximum columnal height. Values in mm, except ratios." httpUri="http://table.plazi.org/id/DF294CBBFFC8FFEB5483F954A939F97D" pageId="19" pageNumber="400" tableUuid="DF294CBBFFC8FFEB5483F954A939F97D">Table 12</tableCitation>
lists quantitative columnal parameters; columnal diameter increasing rapidly at distal end suggesting stalk broken close to attachment disk. Proximalmost columnal with 14 crenular units (poorly preserved); number of crenular units decreasing to 12 of one crenula in proximal mesistele (
<figureCitation id="136D00B6FFC9FFEA5591F800AAF6F848" box="[389,474,2005,2030]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="19" pageNumber="400">Fig. 13</figureCitation>
A), to 11 with numerous subsidiary crenulae in mid mesistele (
<figureCitation id="136D00B6FFC9FFEA50D9F800AE08F848" box="[1229,1316,2005,2030]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="19" pageNumber="400">Fig. 13</figureCitation>
B), and to multiradiate symplexy with initially 10 crenular units observed at center in distal mesistele (
<figureCitation id="136D00B6FFCEFFED50A0FF42AE2BFF15" box="[1204,1287,151,176]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="20" pageNumber="401">Fig. 13</figureCitation>
C); transition symplexy/syzygy (
<figureCitation id="136D00B6FFCEFFED5579FF69AA93FF71" box="[365,447,188,213]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="20" pageNumber="401">Fig. 13</figureCitation>
D) with development of syzygial stereom on facet surface (
<figureCitation id="136D00B6FFCEFFED5066FF69AFE9FF71" box="[1138,1221,188,213]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="20" pageNumber="401">Fig. 13</figureCitation>
E) indicating short dististele (
<quantity id="4CAEB1D6FFCEFFED551EFF35AAAAFF5D" box="[266,390,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="20" pageNumber="401" unit="mm" value="35.0" valueMax="40.0" valueMin="30.0">30–40 mm</quantity>
) with anchylosed articulations; distalmost columnals with facet entirely covered by syzygial stereom, juvenile symplexial pattern of 6 small crenular units of 1 crenula and pentalobate lumen at center (
<figureCitation id="136D00B6FFCEFFED517BFED1AB98FEE5" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="20" pageNumber="401">Fig. 13</figureCitation>
F). Axial lumen diameter to 
<quantity id="4CAEB1D6FFCEFFED55E4FEFDA969FE9A" box="[496,581,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="401" unit="mm" value="1.8">1.8 mm</quantity>
proximally; ratio of lumen to columnal facet diameter 
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proxistele and proximal mesistele, decreasing slowly to 
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distal mesistele and rapidly to 
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dististele. From distal proxistele to mid mesistele, axial canal surrounded by galleried stereom of large meshes bordering small deep ligamentary pits fused with lumen and alternating with crenular units (
<figureCitation id="136D00B6FFCEFFED501AFE41AF49FE09" box="[1038,1125,404,429]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="20" pageNumber="401">Fig. 13</figureCitation>
G–H and J–K); pit floor corresponding to depressed inner areola (see below specimen 1005433B, 
<figureCitation id="136D00B6FFCEFFED57C1FE62AF09FE75" box="[981,1061,439,464]" captionStart="FIGURE 14" captionStartId="20.[151,250,1738,1760]" captionTargetBox="[177,1409,913,1716]" captionTargetId="figure@20.[177,1410,913,1717]" captionTargetPageId="20" captionText="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="20" pageNumber="401">Fig. 14</figureCitation>
D); this character disappearing in distal mesistele (
<figureCitation id="136D00B6FFCEFFED5546FE09AA8DFE51" box="[338,417,476,501]" captionStart="FIGURE 13" captionStartId="19.[151,250,1743,1765]" captionTargetBox="[151,1436,193,1720]" captionTargetId="figure@19.[151,1436,193,1722]" captionTargetPageId="19" captionText="FIGURE 13. Changes in columnal facet and axial canal along the stalk of Ptilocrinus brucei specimen USNM 1005438. A – F: columnal facets; A: in proximal mesistele; B: in mid mesistele; C: in distal mesistele; D – E: in transition mesistele / dististele; F: in mid dististele; G – L: axial canal; J – L: oblique view of G, H, I respectively; G, J: in proximal mesistele, H, K: in mid mesistele, I, L: in distal mesistele." httpUri="https://zenodo.org/record/243062/files/figure.png" pageId="20" pageNumber="401">
Fig. 
<date id="FFE83AF3FFCEFFED5592FE09AA8DFE51" box="[390,417,476,500]" pageId="20" pageNumber="401" value="2013-01">13</date>
</figureCitation>
I and L) and dististele.
</paragraph>
<paragraph id="8BE91C33FFCEFFED54D3FDD5AF19FD68" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="401">
<emphasis id="B922C021FFCEFFED54D3FDD5AABBFDBD" box="[199,407,512,536]" italics="true" pageId="20" pageNumber="401">USNM 1005433-A</emphasis>
: Aboral cup tall; radials more than three times taller than broad (
<figureCitation id="136D00B6FFCEFFED506CFE2AAF97FDBD" box="[1144,1211,511,536]" captionStart="FIGURE 9" captionStartId="14.[151,250,1495,1517]" captionTargetBox="[295,1291,921,1473]" captionTargetId="figure@14.[295,1291,921,1474]" captionTargetPageId="14" captionText="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="20" pageNumber="401">Fig. 9</figureCitation>
C). Tegmen inflated to Br4 or more proximally; about 10 tegminal plates per inter-radius; ambulacral “bridges” between Br8 and mouth with festooned cover plates in a gathered arrangement; oral cone shorter than anal cone; orals smooth and slightly concave; top of oral cone flat. Pattern of proximal arms attached to theca 1+2 3 4 5+6 7 (2 cases) and 1+2 3 4 5+6 7 8+9 10 (3) with first pinnule on Br4; brachial pair following Br10 either 11+12 (1 case) or 15+16 (1). Radial and proximalmost brachial articulations as in USNM 1005453 (see description below).
</paragraph>
<paragraph id="8BE91C33FFCEFFED54D3FD02AE06FCF9" blockId="20.[151,1437,151,861]" pageId="20" pageNumber="401">
Length of preserved stalk 23.0 mm; proximalmost diameter 
<quantity id="4CAEB1D6FFCEFFED579FFD0DA8CBFD4A" box="[907,999,728,752]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="401" unit="mm" value="2.8">2.8 mm</quantity>
, decreasing to 2.4 (ratio of columnal height to diameter 0.42) at distal broken end; proximalmost symplexies with 9 crenular units of 1–2 crenulae (
<figureCitation id="136D00B6FFCEFFED517BFD29AB9FFC9D" captionStart="FIGURE 14" captionStartId="20.[151,250,1738,1760]" captionTargetBox="[177,1409,913,1716]" captionTargetId="figure@20.[177,1410,913,1717]" captionTargetPageId="20" captionText="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="20" pageNumber="401">Fig. 14</figureCitation>
A–C); ratio of claustrum to columnal diameter up to 0.54; tallest columnals (first order in growth sequence) showing formation of a large axial canal with perilumen underlined by large stereom meshes (
<figureCitation id="136D00B6FFCEFFED50A1FC91AE2AFCF9" box="[1205,1286,836,861]" captionStart="FIGURE 14" captionStartId="20.[151,250,1738,1760]" captionTargetBox="[177,1409,913,1716]" captionTargetId="figure@20.[177,1410,913,1717]" captionTargetPageId="20" captionText="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="20" pageNumber="401">Fig. 14</figureCitation>
A).
</paragraph>
<caption id="DF294CBBFFCEFFED5483F91FABD1F8B8" httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="20" pageNumber="401" targetBox="[177,1409,913,1716]" targetPageId="20">
<paragraph id="8BE91C33FFCEFFED5483F91FABD1F8B8" blockId="20.[151,1436,1738,1821]" pageId="20" pageNumber="401">
<emphasis id="B922C021FFCEFFED5483F91FAA33F97A" bold="true" box="[151,287,1738,1760]" pageId="20" pageNumber="401">FIGURE 14.</emphasis>
Stalk articulations in 
<taxonomicName id="4C5667B0FFCEFFED55EEF91FA99DF945" box="[506,689,1738,1760]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="20" pageNumber="401" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCEFFED55EEF91FA99DF945" box="[506,689,1738,1760]" italics="true" pageId="20" pageNumber="401">Ptilocrinus brucei</emphasis>
</taxonomicName>
specimens USNM 1005433. A–C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D–F: columnals from small specimen B; D: proximal mesistele, E–F: distal mesistele.
</paragraph>
</caption>
<paragraph id="8BE91C33FFCEFFEC54D3F89EA859FF71" blockId="20.[151,1437,1866,2035]" lastBlockId="21.[151,1436,151,213]" lastPageId="21" lastPageNumber="402" pageId="20" pageNumber="401">
<emphasis id="B922C021FFCEFFED54D3F89EAAB9F8C6" box="[199,405,1867,1891]" italics="true" pageId="20" pageNumber="401">USNM 1005433-B</emphasis>
: Juvenile specimen of proximal stalk diameter 
<quantity id="4CAEB1D6FFCEFFED57B2F89EA8D1F8C7" box="[934,1021,1867,1891]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="401" unit="mm" value="1.9">1.9 mm</quantity>
, 
<quantity id="4CAEB1D6FFCEFFED501CF89FAF40F8C7" box="[1032,1132,1866,1891]" metricMagnitude="-2" metricUnit="m" metricValue="8.95" pageId="20" pageNumber="401" unit="mm" value="89.5">89.5 mm</quantity>
long, allowing observation of a young stage of stalk ontogeny along stalk length. 
<tableCitation id="C6D42988FFCEFFED5739F8BBA8BFF822" box="[813,915,1902,1927]" captionStart="TABLE 13" captionStartId="21.[151,239,260,283]" captionTargetBox="[159,1335,342,488]" captionTargetPageId="21" captionText="TABLE 13. Ptilocrinus (Chambersaecrinus) brucei USNM 1005433 - B. Quantitative characters of columnals. Columnals numbered from proximal to distal. D: maximum columnal diameter, H: columnal height. Values in mm, except H / D." httpUri="http://table.plazi.org/id/DF294CBBFFCFFFEC5483FED1AE4DFE9E" pageId="20" pageNumber="401" tableUuid="DF294CBBFFCFFFEC5483FED1AE4DFE9E">Table 13</tableCitation>
lists quantitative morphological parameters. Columnal slightly barrel-shaped (
<figureCitation id="136D00B6FFCEFFED561AF847A972F80E" box="[526,606,1938,1963]" captionStart="FIGURE 14" captionStartId="20.[151,250,1738,1760]" captionTargetBox="[177,1409,913,1716]" captionTargetId="figure@20.[177,1410,913,1717]" captionTargetPageId="20" captionText="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="20" pageNumber="401">Fig. 14</figureCitation>
F) having maximum height in mid stalk with ratio of height to diameter up to 1.43. Symplexies with 8 crenular units of 1 short crenula (
<figureCitation id="136D00B6FFCEFFED5756F863A8BEF86A" box="[834,914,1974,1999]" captionStart="FIGURE 14" captionStartId="20.[151,250,1738,1760]" captionTargetBox="[177,1409,913,1716]" captionTargetId="figure@20.[177,1410,913,1717]" captionTargetPageId="20" captionText="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="20" pageNumber="401">Fig. 14</figureCitation>
D–E); inner areola more depressed proximally than distally; ratio of depressed areola to facet diameter 0.46–0.54; axial canal perilumen marked by conspicuous large stereom meshes; claustrum in proximal columnal with pentalobate to multilobate lumen; thin multiradiate syzygial crenularium developing in distal mesistele (
<figureCitation id="136D00B6FFCFFFEC56F1FF69A819FF71" box="[741,821,188,213]" captionStart="FIGURE 14" captionStartId="20.[151,250,1738,1760]" captionTargetBox="[177,1409,913,1716]" captionTargetId="figure@20.[177,1410,913,1717]" captionTargetPageId="20" captionText="FIGURE 14. Stalk articulations in Ptilocrinus brucei specimens USNM 1005433. A – C: proximal columnals in specimen A; A: of first order; B: of second order; C: of third order; D – F: columnals from small specimen B; D: proximal mesistele, E – F: distal mesistele." httpUri="https://zenodo.org/record/243063/files/figure.png" pageId="21" pageNumber="402">Fig. 14</figureCitation>
E–F).
</paragraph>
<caption id="DF294CBBFFCFFFEC5483FED1AE4DFE9E" ID-Table-UUID="DF294CBBFFCFFFEC5483FED1AE4DFE9E" httpUri="http://table.plazi.org/id/DF294CBBFFCFFFEC5483FED1AE4DFE9E" pageId="21" pageNumber="402" targetBox="[159,1335,342,488]" targetIsTable="true" targetPageId="21">
<paragraph id="8BE91C33FFCFFFEC5483FED1AE4DFE9E" blockId="21.[151,1435,260,315]" pageId="21" pageNumber="402">
<emphasis id="B922C021FFCFFFEC5483FED1AA3AFEBE" bold="true" box="[151,278,260,283]" pageId="21" pageNumber="402">TABLE 13.</emphasis>
<taxonomicName id="4C5667B0FFCFFFEC5509FED1A997FEBE" box="[285,699,260,283]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="21" pageNumber="402" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFCFFFEC5509FED1A997FEBE" box="[285,699,260,283]" italics="true" pageId="21" pageNumber="402">Ptilocrinus (Chambersaecrinus) brucei</emphasis>
</taxonomicName>
USNM 1005433-B. Quantitative characters of columnals. Columnals numbered from proximal to distal. D: maximum columnal diameter, H: columnal height. Values in mm, except H/D.
</paragraph>
</caption>
<paragraph id="8BE91C33FFCFFFEC548BFE82AE1BFE4D" pageId="21" pageNumber="402">
<table id="F956EE93FFCF0006548BFE83AE1BFE4D" box="[159,1335,342,488]" gridcols="7" gridrows="4" pageId="21" pageNumber="402">
<tr id="35661E71FFCF0006548BFE83AE1BFEC9" box="[159,1335,342,364]" gridrow="0" pageId="21" pageNumber="402">
<th id="76B7770DFFCF0006548BFE83AA18FEC9" box="[159,308,342,364]" gridcol="0" gridrow="0" pageId="21" pageNumber="402">From cup base</th>
<th id="76B7770DFFCF00065627FE83A94DFEC9" box="[563,609,342,364]" gridcol="1" gridrow="0" pageId="21" pageNumber="402">0</th>
<th id="76B7770DFFCF000656ABFE83A9C1FEC9" box="[703,749,342,364]" gridcol="2" gridrow="0" pageId="21" pageNumber="402">5</th>
<th id="76B7770DFFCF0006575FFE83A854FEC9" box="[843,888,342,364]" gridcol="3" gridrow="0" pageId="21" pageNumber="402">20</th>
<th id="76B7770DFFCF000657CCFE83AF28FEC9" box="[984,1028,342,364]" gridcol="4" gridrow="0" pageId="21" pageNumber="402">35</th>
<th id="76B7770DFFCF00065065FE83AFB1FEC9" box="[1137,1181,342,364]" gridcol="5" gridrow="0" pageId="21" pageNumber="402">50</th>
<th id="76B7770DFFCF0006511EFE83AE1BFEC9" box="[1290,1335,342,364]" gridcol="6" gridrow="0" pageId="21" pageNumber="402">89.5</th>
</tr>
<tr id="35661E71FFCF0006548BFE55AE1BFE33" box="[159,1335,384,406]" gridrow="1" pageId="21" pageNumber="402">
<th id="76B7770DFFCF0006548BFE55AA18FE33" box="[159,308,384,406]" gridcol="0" gridrow="1" pageId="21" pageNumber="402">Diameter (D)</th>
<td id="76B7770DFFCF00065627FE55A94DFE33" box="[563,609,384,406]" gridcol="1" gridrow="1" pageId="21" pageNumber="402">1.9</td>
<td id="76B7770DFFCF000656ABFE55A9C1FE33" box="[703,749,384,406]" gridcol="2" gridrow="1" pageId="21" pageNumber="402">1.5</td>
<td id="76B7770DFFCF0006575FFE55A854FE33" box="[843,888,384,406]" gridcol="3" gridrow="1" pageId="21" pageNumber="402">1.4</td>
<td id="76B7770DFFCF000657CCFE55AF28FE33" box="[984,1028,384,406]" gridcol="4" gridrow="1" pageId="21" pageNumber="402">1.4</td>
<td id="76B7770DFFCF00065065FE55AFB1FE33" box="[1137,1181,384,406]" gridcol="5" gridrow="1" pageId="21" pageNumber="402">1.4</td>
<td id="76B7770DFFCF0006511EFE55AE1BFE33" box="[1290,1335,384,406]" gridcol="6" gridrow="1" pageId="21" pageNumber="402">1.55</td>
</tr>
<tr id="35661E71FFCF0006548BFE7CAE1BFE1A" box="[159,1335,425,447]" gridrow="2" pageId="21" pageNumber="402">
<th id="76B7770DFFCF0006548BFE7CAA18FE1A" box="[159,308,425,447]" gridcol="0" gridrow="2" pageId="21" pageNumber="402">Height (H)</th>
<td id="76B7770DFFCF00065627FE7CA94DFE1A" box="[563,609,425,447]" gridcol="1" gridrow="2" pageId="21" pageNumber="402">&lt;0.4</td>
<td id="76B7770DFFCF000656ABFE7CA9C1FE1A" box="[703,749,425,447]" gridcol="2" gridrow="2" pageId="21" pageNumber="402">&lt;0.4</td>
<td id="76B7770DFFCF0006575FFE7CA854FE1A" box="[843,888,425,447]" gridcol="3" gridrow="2" pageId="21" pageNumber="402">0.8</td>
<td id="76B7770DFFCF000657CCFE7CAF28FE1A" box="[984,1028,425,447]" gridcol="4" gridrow="2" pageId="21" pageNumber="402">1.9</td>
<td id="76B7770DFFCF00065065FE7CAFB1FE1A" box="[1137,1181,425,447]" gridcol="5" gridrow="2" pageId="21" pageNumber="402">2.0</td>
<td id="76B7770DFFCF0006511EFE7CAE1BFE1A" box="[1290,1335,425,447]" gridcol="6" gridrow="2" pageId="21" pageNumber="402">1.85</td>
</tr>
<tr id="35661E71FFCF0006548BFE07AE1BFE4D" box="[159,1335,466,488]" gridrow="3" pageId="21" pageNumber="402">
<th id="76B7770DFFCF0006548BFE07AA18FE4D" box="[159,308,466,488]" gridcol="0" gridrow="3" pageId="21" pageNumber="402">H/D</th>
<td id="76B7770DFFCF00065627FE07A94DFE4D" box="[563,609,466,488]" gridcol="1" gridrow="3" pageId="21" pageNumber="402">0.18</td>
<td id="76B7770DFFCF000656ABFE07A9C1FE4D" box="[703,749,466,488]" gridcol="2" gridrow="3" pageId="21" pageNumber="402">0.22</td>
<td id="76B7770DFFCF0006575FFE07A854FE4D" box="[843,888,466,488]" gridcol="3" gridrow="3" pageId="21" pageNumber="402">0.56</td>
<td id="76B7770DFFCF000657CCFE07AF28FE4D" box="[984,1028,466,488]" gridcol="4" gridrow="3" pageId="21" pageNumber="402">1.36</td>
<td id="76B7770DFFCF00065065FE07AFB1FE4D" box="[1137,1181,466,488]" gridcol="5" gridrow="3" pageId="21" pageNumber="402">1.43</td>
<td id="76B7770DFFCF0006511EFE07AE1BFE4D" box="[1290,1335,466,488]" gridcol="6" gridrow="3" pageId="21" pageNumber="402">1.19</td>
</tr>
</table>
</paragraph>
<caption id="DF294CBBFFCFFFEC5483F899A8F9F83A" httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="21" pageNumber="402" targetBox="[242,1344,565,1845]" targetPageId="21">
<paragraph id="8BE91C33FFCFFFEC5483F899A8F9F83A" blockId="21.[151,1436,1868,1951]" pageId="21" pageNumber="402">
<emphasis id="B922C021FFCFFFEC5483F899AA33F8C4" bold="true" box="[151,287,1868,1890]" pageId="21" pageNumber="402">FIGURE 15.</emphasis>
Radial and proximal brachials in 
<taxonomicName id="4C5667B0FFCFFFEC5667F899A806F8C7" box="[627,810,1868,1890]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="21" pageNumber="402" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCFFFEC5667F899A806F8C7" box="[627,810,1868,1890]" italics="true" pageId="21" pageNumber="402">Ptilocrinus brucei</emphasis>
</taxonomicName>
. A–C, E–F: specimen 1005453–A; D, G: specimen 1005433- A; A–B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br1, D: distal facet ofBr1, E: distal facet of Br2, F: proximal facet of Br2, G: distal facet of Br3.
</paragraph>
</caption>
<paragraph id="8BE91C33FFCCFFEF54D3FF4DA8F7FE75" blockId="22.[151,1437,151,861]" pageId="22" pageNumber="403">
<emphasis id="B922C021FFCCFFEF54D3FF4DAA51FF15" box="[199,381,152,176]" italics="true" pageId="22" pageNumber="403">USNM 1005453</emphasis>
: Badly preserved specimen; all ossicles dissociated; all remaining ossicles similar to those of other specimens. Proximalmost arm pattern 1+2 3 4 with first pinnule on Br4. Radial with deep arm insertion (
<figureCitation id="136D00B6FFCCFFEF517BFF69AB9FFF5D" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
A); inner surface with two sub-parallel conspicuous nervous canals (
<figureCitation id="136D00B6FFCCFFEF57D5FF0AAF3EFF5D" box="[961,1042,223,248]" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
A–B); synarthry joining radial and Br1 with well differentiated inner (adoral) ligament and muscular areas (
<figureCitation id="136D00B6FFCCFFEF57C2FED1AF04FEB8" box="[982,1064,260,285]" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
C), this differentiation becoming less conspicuous in the following muscular synarthries (
<figureCitation id="136D00B6FFCCFFEF571AFEF2A871FEE5" box="[782,861,295,320]" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
E and G); Br3 with deep adoral groove (
<figureCitation id="136D00B6FFCCFFEF5132FEF2AE5BFEE5" box="[1318,1399,295,320]" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
G); synostoses flat; stereom of synostosis 1+2 with small meshes only on Br1 facet (
<figureCitation id="136D00B6FFCCFFEF5045FE99AF89FEC0" box="[1105,1189,332,357]" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
D) and small meshes aborally plus larger ones around lumen and adorally on Br2 adjacent facet (
<figureCitation id="136D00B6FFCCFFEF5022FEBAAFA2FE2D" box="[1078,1166,367,392]" captionStart="FIGURE 15" captionStartId="21.[151,250,1868,1890]" captionTargetBox="[242,1344,565,1845]" captionTargetId="figure@21.[242,1345,565,1846]" captionTargetPageId="21" captionText="FIGURE 15. Radial and proximal brachials in Ptilocrinus brucei. A – C, E – F: specimen 1005453 – A; D, G: specimen 1005433 - A; A – B: inner view of radial, A: oblique view showing arm articulation and nervous canals; C: proximal facet of Br 1, D: distal facet ofBr 1, E: distal facet of Br 2, F: proximal facet of Br 2, G: distal facet of Br 3." httpUri="https://zenodo.org/record/243064/files/figure.png" pageId="22" pageNumber="403">Fig. 15</figureCitation>
F). Only proximalmost columnals remaining; articulations with 14 crenular units of 1 crenula with various lengths and pentalobate lumen (
<figureCitation id="136D00B6FFCCFFEF548BFE62ABDDFE75" box="[159,241,439,464]" captionStart="FIGURE 12" captionStartId="18.[151,250,1272,1294]" captionTargetBox="[336,1245,373,1243]" captionTargetId="figure@18.[336,1245,373,1243]" captionTargetPageId="18" captionText="FIGURE 12. Proximal stalk symplexies in large specimens of Ptilocrinus brucei. A: specimen USNM 1005453, B – D: specimen USNM 1005439; A: proximalmost columnal; B: at a distance of 5.0 mm from aboral cup; C: at a distance of 6.0 mm from aboral cup; D: at a distance of 22.0 mm from aboral cup." httpUri="https://zenodo.org/record/243061/files/figure.png" pageId="22" pageNumber="403">Fig. 12</figureCitation>
A); no stereom differentiation between claustrum and inner areola.
</paragraph>
</subSubSection>
<subSubSection id="C34C4FB8FFCCFFEE54D3FE08A9FEFDBD" lastPageId="23" lastPageNumber="404" pageId="22" pageNumber="403" type="materials_examined">
<paragraph id="8BE91C33FFCCFFEF54D3FE08ABFFFCF8" blockId="22.[151,1437,151,861]" pageId="22" pageNumber="403">
<emphasis id="B922C021FFCCFFEF54D3FE08A903FE56" box="[199,559,476,500]" italics="true" pageId="22" pageNumber="403">
<typeStatus id="54EDA291FFCCFFEF54D3FE08AA79FE51" box="[199,341,477,500]" pageId="22" pageNumber="403" type="holotype">HOLOTYPE</typeStatus>
AND 
<typeStatus id="54EDA291FFCCFFEF55B0FE09A905FE56" box="[420,553,476,499]" pageId="22" pageNumber="403" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
Quantitative parameters of the 
<typeStatus id="54EDA291FFCCFFEF57BFFE09AF21FE50" box="[939,1037,476,501]" pageId="22" pageNumber="403" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54EDA291FFCCFFEF5059FE08AF81FE50" box="[1101,1197,477,501]" pageId="22" pageNumber="403" type="paratype">paratype</typeStatus>
listed by Vaney (
<emphasis id="B922C021FFCCFFEF5192FE08AEB0FE51" box="[1414,1436,477,500]" italics="true" pageId="22" pageNumber="403">in</emphasis>
<bibRefCitation id="EFC761C2FFCCFFEF5483FDD5AA52FDBD" author="Vaney" box="[151,382,511,536]" pageId="22" pageNumber="403" refString="Vaney, C. &amp; John, D. D. (1939) Scientific results of the voyage of S. Y. Scotia, 1902 - 04: The Crinoidea. Transactions of the Royal Society of Edinburg, 59, 661 - 672. http: // dx. doi. org / 10.1017 / S 0080456800017385" type="journal article" year="1939">Vaney &amp; John 1939</bibRefCitation>
) were usually overestimates. 
<bibRefCitation id="EFC761C2FFCCFFEF56C2FE2AAF01FDBD" author="Mironov" box="[726,1069,511,536]" pageId="22" pageNumber="403" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov and Sorokina (1998)</bibRefCitation>
re-examined the large 
<typeStatus id="54EDA291FFCCFFEF512EFE2AAEB0FDBD" box="[1338,1436,511,536]" pageId="22" pageNumber="403" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED4784F6FFCCFFEF548AFDF1ABF0FD99" box="[158,220,548,572]" httpUri="http://grbio.org/cool/h8hj-4rxx" name="National Museum of Scotland - Natural Sciences" pageId="22" pageNumber="403">NMS</collectionCode>
1921.143.1743), which is now in poor condition. I used Vaney &amp; John’s (1939) measurements for calibrating measurements of the large and fine original photograph of the 
<typeStatus id="54EDA291FFCCFFEF57DBFD92AF1DFDC5" box="[975,1073,583,608]" pageId="22" pageNumber="403" type="holotype">holotype</typeStatus>
when it was still well-preserved (Vaney 1908) (
<tableCitation id="C6D42988FFCCFFEF552BFDB9AA8CFD21" box="[319,416,620,645]" captionStart="TABLE 9" captionStartId="15.[151,239,584,607]" captionTargetBox="[155,1397,764,1106]" captionTargetPageId="15" captionText="TABLE 9. Ptilocrinus (Chambersaecrinus) brucei. Main morphological variables in USNM specimens and type series. Wb: Br 1 width; Wr-b: radial aboral width; Wr-t: radial adoral width; Hc: aboral cup height; Hr-r: radial axial height; Hri: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: adoral diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm. Dp and Dm are used as growth index (in yellow)." httpUri="http://table.plazi.org/id/DF294CBBFFD5FFF65483FD9DA86AFD45" pageId="22" pageNumber="403" tableUuid="DF294CBBFFD5FFF65483FD9DA86AFD45">Tables 9</tableCitation>
and 
<tableCitation id="C6D42988FFCCFFEF55CDFDB9AAD4FD21" box="[473,504,620,644]" captionStart="TABLE 10" captionStartId="15.[151,239,1174,1197]" captionTargetBox="[158,1405,1257,1568]" captionTargetPageId="15" captionText="TABLE 10. Ptilocrinus (Chambersaecrinus) brucei. Variation of main morphological ratios in USNM specimens and type series. Abbreviations and growth index as in Table 9." httpUri="http://table.plazi.org/id/DF294CBBFFD5FFF65483FB43A82EFB68" pageId="22" pageNumber="403" tableUuid="DF294CBBFFD5FFF65483FB43A82EFB68">10</tableCitation>
). General 
<typeStatus id="54EDA291FFCCFFEF567AFDB9A9FCFD20" box="[622,720,620,645]" pageId="22" pageNumber="403" type="holotype">holotype</typeStatus>
morphology is as in 
<collectionCode id="ED4784F6FFCCFFEF57D5FDB9AF3EFD21" LSID="urn:lsid:biocol.org:col:34871" box="[961,1042,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="403">USNM</collectionCode>
1005439 (
<figureCitation id="136D00B6FFCCFFEF5099FDB9AFFDFD21" box="[1165,1233,620,645]" captionStart="FIGURE 9" captionStartId="14.[151,250,1495,1517]" captionTargetBox="[295,1291,921,1473]" captionTargetId="figure@14.[295,1291,921,1474]" captionTargetPageId="14" captionText="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="22" pageNumber="403">Fig. 9</figureCitation>
A). Each arm was about 
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long with the distal ends rolled up, about 40 pinnules on each side; maximum pinnule length 
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with 40 pinnulars. Proximal arm pattern 1+2 3 4 5+6 7 with first brachial on Br4 (4 cases); one arm described by Vaney with the following brachial pairs at 9+10, 16+17 24+25, 33+34, 40+41; other arms with brachial pairs at 8+9 (2 cases), 13+14, 14+15, 17+18, 18+19 (1 case each). Stalk length about 
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; proximalmost diameter 
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, decreasing to 3.0 at a distance of 
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from cup; maximum ratio of columnal height to diameter 
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mid stalk.
</paragraph>
<caption id="DF294CBBFFCCFFEF5483F8F3AA85F8DF" httpUri="https://zenodo.org/record/243065/files/figure.png" pageId="22" pageNumber="403" targetBox="[151,1436,913,1808]" targetPageId="22">
<paragraph id="8BE91C33FFCCFFEF5483F8F3AA85F8DF" blockId="22.[151,1436,1830,1914]" pageId="22" pageNumber="403">
<emphasis id="B922C021FFCCFFEF5483F8F3AA33F899" bold="true" box="[151,287,1830,1852]" pageId="22" pageNumber="403">FIGURE 16.</emphasis>
Columnal articulations in mid and distal stalk of 
<taxonomicName id="4C5667B0FFCCFFEF5707F8F3A8E6F899" box="[787,970,1830,1852]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="22" pageNumber="403" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCCFFEF5707F8F3A8E6F899" box="[787,970,1830,1852]" italics="true" pageId="22" pageNumber="403">Ptilocrinus brucei</emphasis>
</taxonomicName>
paratype. A: symplexy in proximal mesistele, B, D: transition between symplexy and syzygy; B: in distal mesistele; D: in proximal dististele; C, E: in distalmost stalk at a distance of 36 mm from D.
</paragraph>
</caption>
<paragraph id="8BE91C33FFCCFFEE54D3F873A9FEFDBD" blockId="22.[151,1437,1958,2019]" lastBlockId="23.[151,1437,151,1005]" lastPageId="23" lastPageNumber="404" pageId="22" pageNumber="403">
The 
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(NHM 1939.5.13.1) is fragmented. However, proximal arm pattern reconstruction is possible: 1+2 3 4 5+6 7+ 
<date id="FFE83AF3FFCCFFEF5557F81EAAA4F846" box="[323,392,1995,2019]" pageId="22" pageNumber="403" value="2010-09-08">8 9 10</date>
+11 12 (3 cases), 1+2 3 4 5+6 7 8+9 
<date id="FFE83AF3FFCCFFEF5734F81EA8ACF846" box="[800,896,1995,2019]" pageId="22" pageNumber="403" value="2012-11-10">10 11 12</date>
+13 (1), 1+2 3 4 5 6 7 8+ 
<date id="FFE83AF3FFCCFFEF5083F81EAFC5F846" box="[1175,1257,1995,2019]" pageId="22" pageNumber="403" value="2011-10-09">9 10 11</date>
+12 (1); usually free brachials alternating with brachial pairs distally. Five radials preserved. Stalk very fragmented; total length of all fragments about 
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; proxistele lost; proximalmost diameter inferred from other stalk fragments about 
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(3.0 mm after Vaney); minimum values in proximal mesistele: columnal diameter 
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, columnal height 
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, ratio of height to diameter 0.86; maximum values in mid and distal mesistele: columnal diameter 
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, columnal height 
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, ratio of height to diameter 1.43; maximum values in distalmost fragments: columnal diameter 
<quantity id="4CAEB1D6FFCDFFEE5514FE99AA75FEC1" box="[256,345,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="23" pageNumber="404" unit="mm" value="2.7">2.7 mm</quantity>
, columnal height 
<quantity id="4CAEB1D6FFCDFFEE5636FE99A956FEC1" box="[546,634,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="404" unit="mm" value="1.5">1.5 mm</quantity>
, ratio of height to diameter 0.56. Symplexies with 10–11 crenular units of 1 crenula in proximalmost fragments and coarsely-meshed stereom in inner areola and perilumen (
<figureCitation id="136D00B6FFCDFFEE5110FEBAAE79FE2D" box="[1284,1365,367,392]" captionStart="FIGURE 16" captionStartId="22.[151,250,1830,1852]" captionTargetBox="[151,1436,913,1808]" captionTargetId="figure@22.[151,1436,913,1809]" captionTargetPageId="22" captionText="FIGURE 16. Columnal articulations in mid and distal stalk of Ptilocrinus brucei paratype. A: symplexy in proximal mesistele, B, D: transition between symplexy and syzygy; B: in distal mesistele; D: in proximal dististele; C, E: in distalmost stalk at a distance of 36 mm from D." httpUri="https://zenodo.org/record/243065/files/figure.png" pageId="23" pageNumber="404">Fig. 16</figureCitation>
A); 8– 9 crenular units of 1 crenula in mid mesistele; 7 crenular units of 1 short crenula in juvenile pattern at facet center in distal mesistele (
<figureCitation id="136D00B6FFCDFFEE5561FE62AAE6FE75" box="[373,458,439,464]" captionStart="FIGURE 16" captionStartId="22.[151,250,1830,1852]" captionTargetBox="[151,1436,913,1808]" captionTargetId="figure@22.[151,1436,913,1809]" captionTargetPageId="22" captionText="FIGURE 16. Columnal articulations in mid and distal stalk of Ptilocrinus brucei paratype. A: symplexy in proximal mesistele, B, D: transition between symplexy and syzygy; B: in distal mesistele; D: in proximal dististele; C, E: in distalmost stalk at a distance of 36 mm from D." httpUri="https://zenodo.org/record/243065/files/figure.png" pageId="23" pageNumber="404">Fig. 16</figureCitation>
B). Transition between mesistele and dististele with multiradiate crenularium of up to about 20 wide crenulae (
<figureCitation id="136D00B6FFCDFFEE55A0FE09A92BFE50" box="[436,519,476,501]" captionStart="FIGURE 16" captionStartId="22.[151,250,1830,1852]" captionTargetBox="[151,1436,913,1808]" captionTargetId="figure@22.[151,1436,913,1809]" captionTargetPageId="22" captionText="FIGURE 16. Columnal articulations in mid and distal stalk of Ptilocrinus brucei paratype. A: symplexy in proximal mesistele, B, D: transition between symplexy and syzygy; B: in distal mesistele; D: in proximal dististele; C, E: in distalmost stalk at a distance of 36 mm from D." httpUri="https://zenodo.org/record/243065/files/figure.png" pageId="23" pageNumber="404">Fig. 16</figureCitation>
B and D); distalmost syzygies examined with numerous thinner radial crenulae becoming irregular in outer facet (
<figureCitation id="136D00B6FFCDFFEE5603FE2AA94AFDBD" box="[535,614,511,536]" captionStart="FIGURE 16" captionStartId="22.[151,250,1830,1852]" captionTargetBox="[151,1436,913,1808]" captionTargetId="figure@22.[151,1436,913,1809]" captionTargetPageId="22" captionText="FIGURE 16. Columnal articulations in mid and distal stalk of Ptilocrinus brucei paratype. A: symplexy in proximal mesistele, B, D: transition between symplexy and syzygy; B: in distal mesistele; D: in proximal dististele; C, E: in distalmost stalk at a distance of 36 mm from D." httpUri="https://zenodo.org/record/243065/files/figure.png" pageId="23" pageNumber="404">Fig. 16</figureCitation>
C and E).
</paragraph>
</subSubSection>
<subSubSection id="C34C4FB8FFCDFFEE54D3FDF1AA5DF807" pageId="23" pageNumber="404" type="discussion">
<paragraph id="8BE91C33FFCDFFEE54D3FDF1AA32FC48" blockId="23.[151,1437,151,1005]" pageId="23" pageNumber="404">
<emphasis id="B922C021FFCDFFEE54D3FDF1AA17FD98" bold="true" box="[199,315,548,573]" pageId="23" pageNumber="404">Remarks.</emphasis>
<bibRefCitation id="EFC761C2FFCDFFEE5552FDF1A9BDFD99" author="Mironov" box="[326,657,548,573]" pageId="23" pageNumber="404" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina (1998)</bibRefCitation>
described two other poorly preserved specimens collected off the 
<collectingCountry id="F3415CA3FFCDFFEE5483FD92AA6FFDC5" box="[151,323,583,608]" name="United Kingdom" pageId="23" pageNumber="404">Orkney Islands</collectingCountry>
by Russian cruises. The specimen attributed to 
<taxonomicName id="4C5667B0FFCDFFEE577EFD9CA8E3FDC5" box="[874,975,583,608]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCDFFEE577EFD9CA8E3FDC5" box="[874,975,583,608]" italics="true" pageId="23" pageNumber="404">P. brucei</emphasis>
</taxonomicName>
was collected at R/V 
<emphasis id="B922C021FFCDFFEE50DBFD9CAEB0FDC5" box="[1231,1436,583,608]" italics="true" pageId="23" pageNumber="404">Dmitri Mendeleev</emphasis>
sta. 4094, 60°42.8’–43.1’S to 41°03.0–01.9’W, depth 
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. Proximal arm patterns included two variants: 1+2 3 4 5+6 with first pinnule on Br4 (3 cases) and with one of these arms with the third brachial pair at 8+9 followed by 6 successive muscular articulations, and 1+2 3 4+5 with first pinnule on Br5 (2 cases) and with one of these arms with the third brachial pair at 7+8, number of successive muscular articulations 
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mid arm fragment and 
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distal arm. Anal cone height about 
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. Three stalk fragments with total length 
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(
<tableCitation id="C6D42988FFCDFFEE548AFCCAAA28FC9D" box="[158,260,799,824]" captionStart="TABLE 14" captionStartId="23.[151,239,1052,1075]" captionTargetBox="[159,1320,1134,1322]" captionTargetPageId="23" captionText="TABLE 14. Ptilocrinus (Chambersaecrinus) brucei, R / V Dmitri Mendeleev sta. 4094. Stalk measurements. D: maximum columnal diameter, H: columnal height. Values in mm, except H / D." httpUri="http://table.plazi.org/id/DF294CBBFFCDFFEE5483FBC9A8F5FBF6" pageId="23" pageNumber="404" tableUuid="DF294CBBFFCDFFEE5483FBC9A8F5FBF6">Table 14</tableCitation>
); estimated proximalmost diameter about 
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; symplexies with 9–10 crenular units of 1–2 crenulae; ratio of lumen to columnal diameter 
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mid stalk. The second specimen, attributed to a new species 
<taxonomicName id="4C5667B0FFCDFFEE5134FC90AAFAFC25" authority="Mironov &amp; Sorokina, 1998" authorityName="Mironov &amp; Sorokina" authorityYear="1998" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="flexibilis">
<emphasis id="B922C021FFCDFFEE5134FC90AEB7FCF8" box="[1312,1435,836,861]" italics="true" pageId="23" pageNumber="404">P. flexibilis</emphasis>
<bibRefCitation id="EFC761C2FFCDFFEE5483FCB2AAFAFC25" author="Mironov" box="[151,470,871,896]" pageId="23" pageNumber="404" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina, 1998</bibRefCitation>
</taxonomicName>
, was collected at R/V 
<emphasis id="B922C021FFCDFFEE56F7FCB2A8FDFC25" box="[739,977,871,896]" italics="true" pageId="23" pageNumber="404">Akademic Kurchatov</emphasis>
sta. 908, 60°13.5’–12.9’S to 44°10.6’– 12.2’W, depth 
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(Moscow State Univ. Zoological Museum cat. no. C-20). Proximal arm patterns 1+2 3 4 5+6 7 8 (1 case) and 1+2 3 4 5+6 7+8 (1). 
<bibRefCitation id="EFC761C2FFCDFFEE56AFFC65A8FEFC6D" author="Roux" box="[699,978,943,968]" pageId="23" pageNumber="404" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert (2011)</bibRefCitation>
considered 
<taxonomicName id="4C5667B0FFCDFFEE5049FC64AFEEFC6D" box="[1117,1218,943,968]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFCDFFEE5049FC64AFEEFC6D" box="[1117,1218,943,968]" italics="true" pageId="23" pageNumber="404">P. brucei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5667B0FFCDFFEE50E9FC64AE50FC6D" box="[1277,1404,943,968]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="flexibilis">
<emphasis id="B922C021FFCDFFEE50E9FC64AE50FC6D" box="[1277,1404,943,968]" italics="true" pageId="23" pageNumber="404">P. flexibilis</emphasis>
</taxonomicName>
as conspecific.
</paragraph>
<caption id="DF294CBBFFCDFFEE5483FBC9A8F5FBF6" ID-Table-UUID="DF294CBBFFCDFFEE5483FBC9A8F5FBF6" httpUri="http://table.plazi.org/id/DF294CBBFFCDFFEE5483FBC9A8F5FBF6" pageId="23" pageNumber="404" targetBox="[159,1320,1134,1322]" targetIsTable="true" targetPageId="23">
<paragraph id="8BE91C33FFCDFFEE5483FBC9A8F5FBF6" blockId="23.[151,1436,1052,1107]" pageId="23" pageNumber="404">
<emphasis id="B922C021FFCDFFEE5483FBC9AA0CFB96" bold="true" box="[151,288,1052,1075]" pageId="23" pageNumber="404">TABLE 14.</emphasis>
<emphasis id="B922C021FFCDFFEE553BFBC9A9C6FB96" box="[303,746,1052,1075]" italics="true" pageId="23" pageNumber="404">
<taxonomicName id="4C5667B0FFCDFFEE553BFBC9A9C9FB96" box="[303,741,1052,1075]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">Ptilocrinus (Chambersaecrinus) brucei</taxonomicName>
,
</emphasis>
R/V 
<emphasis id="B922C021FFCDFFEE5721FBC9A8D2FB96" box="[821,1022,1052,1075]" italics="true" pageId="23" pageNumber="404">Dmitri Mendeleev</emphasis>
sta. 4094. Stalk measurements. D: maximum columnal diameter, H: columnal height. Values in mm, except H/D.
</paragraph>
</caption>
<paragraph id="8BE91C33FFCDFFEE548BFBBBAE04FA8F" pageId="23" pageNumber="404">
<table id="F956EE93FFCD0006548BFBBBAE04FA8F" box="[159,1320,1134,1322]" gridcols="4" gridrows="5" pageId="23" pageNumber="404">
<tr id="35661E71FFCD0006548BFBBBAE04FB21" box="[159,1320,1134,1156]" gridrow="0" pageId="23" pageNumber="404">
<th id="76B7770DFFCD0006548BFBBBAA14FB21" box="[159,312,1134,1156]" gridcol="0" gridrow="0" pageId="23" pageNumber="404">Stalk fragment</th>
<th id="76B7770DFFCD000655ADFBBBA9A0FB21" box="[441,652,1134,1156]" gridcol="1" gridrow="0" pageId="23" pageNumber="404">Mid-distal mesistele</th>
<th id="76B7770DFFCD00065713FBBBA8F6FB21" box="[775,986,1134,1156]" gridcol="2" gridrow="0" pageId="23" pageNumber="404">Proximal dististele</th>
<th id="76B7770DFFCD00065041FBBBAE04FB21" box="[1109,1320,1134,1156]" gridcol="3" gridrow="0" pageId="23" pageNumber="404">Distal dististele</th>
</tr>
<tr id="35661E71FFCD0006548BFB4DAE04FB0B" box="[159,1320,1176,1198]" gridrow="1" pageId="23" pageNumber="404">
<th id="76B7770DFFCD0006548BFB4DAA14FB0B" box="[159,312,1176,1198]" gridcol="0" gridrow="1" pageId="23" pageNumber="404">From proximal</th>
<td id="76B7770DFFCD000655ADFB4DA9A0FB0B" box="[441,652,1176,1198]" gridcol="1" gridrow="1" pageId="23" pageNumber="404">0 83.2</td>
<td id="76B7770DFFCD00065713FB4DA8F6FB0B" box="[775,986,1176,1198]" gridcol="2" gridrow="1" pageId="23" pageNumber="404">0 31</td>
<td id="76B7770DFFCD00065041FB4DAE04FB0B" box="[1109,1320,1176,1198]" gridcol="3" gridrow="1" pageId="23" pageNumber="404">0 49</td>
</tr>
<tr id="35661E71FFCD0006548BFB14AE04FB72" box="[159,1320,1217,1239]" gridrow="2" pageId="23" pageNumber="404">
<th id="76B7770DFFCD0006548BFB14AA14FB72" box="[159,312,1217,1239]" gridcol="0" gridrow="2" pageId="23" pageNumber="404">Diameter (D)</th>
<td id="76B7770DFFCD000655ADFB14A9A0FB72" box="[441,652,1217,1239]" gridcol="1" gridrow="2" pageId="23" pageNumber="404">2.2 2.3</td>
<td id="76B7770DFFCD00065713FB14A8F6FB72" box="[775,986,1217,1239]" gridcol="2" gridrow="2" pageId="23" pageNumber="404">2.6 2.7</td>
<td id="76B7770DFFCD00065041FB14AE04FB72" box="[1109,1320,1217,1239]" gridcol="3" gridrow="2" pageId="23" pageNumber="404">2.8 3.2</td>
</tr>
<tr id="35661E71FFCD0006548BFB3FAE04FAA5" box="[159,1320,1258,1280]" gridrow="3" pageId="23" pageNumber="404">
<th id="76B7770DFFCD0006548BFB3FAA14FAA5" box="[159,312,1258,1280]" gridcol="0" gridrow="3" pageId="23" pageNumber="404">Height (H)</th>
<td id="76B7770DFFCD000655ADFB3FA9A0FAA5" box="[441,652,1258,1280]" gridcol="1" gridrow="3" pageId="23" pageNumber="404">1.1 1.1</td>
<td id="76B7770DFFCD00065713FB3FA8F6FAA5" box="[775,986,1258,1280]" gridcol="2" gridrow="3" pageId="23" pageNumber="404">1.1 1.2</td>
<td id="76B7770DFFCD00065041FB3FAE04FAA5" box="[1109,1320,1258,1280]" gridcol="3" gridrow="3" pageId="23" pageNumber="404">1.1 0.7</td>
</tr>
<tr id="35661E71FFCD0006548BFAC1AE04FA8F" box="[159,1320,1300,1322]" gridrow="4" pageId="23" pageNumber="404">
<th id="76B7770DFFCD0006548BFAC1AA14FA8F" box="[159,312,1300,1322]" gridcol="0" gridrow="4" pageId="23" pageNumber="404">H/D</th>
<td id="76B7770DFFCD000655ADFAC1A9A0FA8F" box="[441,652,1300,1322]" gridcol="1" gridrow="4" pageId="23" pageNumber="404">0.5 0.48</td>
<td id="76B7770DFFCD00065713FAC1A8F6FA8F" box="[775,986,1300,1322]" gridcol="2" gridrow="4" pageId="23" pageNumber="404">0.42 0.44</td>
<td id="76B7770DFFCD00065041FAC1AE04FA8F" box="[1109,1320,1300,1322]" gridcol="3" gridrow="4" pageId="23" pageNumber="404">0.39 0.22</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE91C33FFCDFFEE54D3FABBAA5DF807" blockId="23.[151,1436,1390,2027]" pageId="23" pageNumber="404">
The current study documents the wide range of intraspecific variation in 
<taxonomicName id="4C5667B0FFCDFFEE5019FABAAF8BFA22" box="[1037,1191,1390,1415]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFCDFFEE5019FABAAF8BFA22" box="[1037,1191,1390,1415]" italics="true" pageId="23" pageNumber="404">P. (C.) brucei</emphasis>
</taxonomicName>
. 
<figureCitation id="136D00B6FFCDFFEE50A3FABBAFD7FA22" box="[1207,1275,1390,1415]" captionStart="FIGURE 9" captionStartId="14.[151,250,1495,1517]" captionTargetBox="[295,1291,921,1473]" captionTargetId="figure@14.[295,1291,921,1474]" captionTargetPageId="14" captionText="FIGURE 9. Theca and proximal arms of Ptilocrinus brucei. A – B: specimen 1005439; C: specimen 1005433 - A; A: view of anal interray; B: view of A – B interray; C: anterior view; as: anal sac; o: oral." httpUri="https://zenodo.org/record/243058/files/figure.png" pageId="23" pageNumber="404">Fig. 9</figureCitation>
illustrates the range in variation in external morphology. Arm pattern varies from chiefly alternating free brachials and brachial pairs to long series of successive muscular articulations beyond the third brachial pair (
<tableCitation id="C6D42988FFCDFFEE5095FA63AFC5FA6A" box="[1153,1257,1462,1487]" captionStart="TABLE 15" captionStartId="24.[151,239,440,463]" captionText="TABLE 15. Ptilocrinus (Chambersaecrinus) brucei. Variations in proximal arm pattern. * holotype, ** paratype." pageId="23" pageNumber="404">Table 15</tableCitation>
); proximalmost arm pattern 1+2 3 4 5+6 7 (75.9%), 1+2 3 4 5+6 7+8 (13.8%); first pinnule on Br4 (94.7%); brachial pair at 8+9 (44.0%), 9+10 (13.0%), 10+11 (22.7%), 13+14 (15.0%), 15+16 (18.7%), 18+19 (27.3%), and other brachial pairs &lt;13%. The adoral process in brachial muscular synarthries has never been described in the other species of 
<taxonomicName id="4C5667B0FFCDFFEE5483F993AA3FF9FA" box="[151,275,1606,1631]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="genus">
<emphasis id="B922C021FFCDFFEE5483F993AA3FF9FA" box="[151,275,1606,1631]" italics="true" pageId="23" pageNumber="404">Ptilocrinus</emphasis>
</taxonomicName>
. Genital pinnules display an asymmetrical architecture. Exceptionally preserved submature oocytes&gt;100 µm across (
<figureCitation id="136D00B6FFCDFFEE5575F9BFAA9CF926" box="[353,432,1642,1667]" captionStart="FIGURE 11" captionStartId="17.[151,250,1593,1615]" captionTargetBox="[215,1372,193,1568]" captionTargetId="figure@17.[215,1372,193,1569]" captionTargetPageId="17" captionText="FIGURE 11. Architecture and articulations in Ptilocrinus brucei pinnules. A – B: genital pinnule inflation in specimen USNM 1005439; A: proximal part, B: distal part; C – D: specimen USNM 1005438; C: proximal part with multiplated adoral skeleton covering female gonad (g); D: detail of gonad showing submature oocytes; E – I: specimen USNM 1005439; E: proximal muscular synarthry of Pn 2; F: aboral distal view of Pn 2; G: proximal synostose of Pn 3; H: pinnule broken proximally to Pn 4 (cv: cover plates, Pn: pinnular, arrow indicating convex side of genital inflation); I: proximal synostose of asymetrical Pn 4 (adoral plates removed, orientation like in H)." httpUri="https://zenodo.org/record/243060/files/figure.png" pageId="23" pageNumber="404">Fig. 11</figureCitation>
C–D) suggest that larvae could be lecithotrophic. The number of crenular units in stalk symplexies increases throughout ontogeny from 6 to 12 (possibly 14), but the rate of this change varies from an individual to another; usually each crenular unit has 1 crenula of various lengths, but sometime up to 3 crenulae or with subsidiary crenulae between units in the largest specimens. The enlargement of the axial canal in the proximal mesistele through ontogeny is also variable. Small specimens have relatively tall columnals (maximum ratio of columnal height to diameter 1.43) compared to the largest specimens (0.41). The anal cone is always significantly higher than the oral cone. The arrangement of interadial tegminal plates into a median row is irregular and frequently absent. 
<collectingCountry id="F3415CA3FFCDFFEE557EF8B2AAA5F8DA" box="[362,393,1895,1919]" name="American Samoa" pageId="23" pageNumber="404">As</collectingCountry>
pointed out by 
<bibRefCitation id="EFC761C2FFCDFFEE562BF8B2A87AF8DA" author="Roux" box="[575,854,1894,1919]" pageId="23" pageNumber="404" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert (2011)</bibRefCitation>
, this character cannot be used to identify subgenus 
<taxonomicName id="4C5667B0FFCDFFEE5483F85FAA47F806" box="[151,363,1930,1955]" class="Crinoidea" family="Hyocrinidae" genus="Chambersaecrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Millericrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="genus">
<emphasis id="B922C021FFCDFFEE5483F85FAA47F806" box="[151,363,1930,1955]" italics="true" pageId="23" pageNumber="404">Chambersaecrinus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C4FB8FFCDFFE154D3F87BAA60FBBF" lastPageId="24" lastPageNumber="405" pageId="23" pageNumber="404" type="reference_group">
<paragraph id="8BE91C33FFCDFFE154D3F87BAE67FEE5" blockId="23.[151,1436,1390,2027]" lastBlockId="24.[151,1436,151,392]" lastPageId="24" lastPageNumber="405" pageId="23" pageNumber="404">
<taxonomicName id="4C5667B0FFCDFFEE54D3F87BAAE8F862" ID-CoL="8GHQM" box="[199,452,1966,1991]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFCDFFEE54D3F87BAA6FF862" box="[199,323,1966,1991]" italics="true" pageId="23" pageNumber="404">Ptilocrinus</emphasis>
(
<emphasis id="B922C021FFCDFFEE5547F87AAA41F863" box="[339,365,1967,1990]" italics="true" pageId="23" pageNumber="404">C.</emphasis>
) 
<emphasis id="B922C021FFCDFFEE5569F87BAAE8F862" box="[381,452,1966,1991]" italics="true" pageId="23" pageNumber="404">brucei</emphasis>
</taxonomicName>
shares with 
<taxonomicName id="4C5667B0FFCDFFEE5644F87AAF62F862" ID-CoL="8GHQT" authority="Roux &amp; Lambert, 2011" authorityName="Roux &amp; Lambert" authorityYear="2011" box="[592,1102,1966,1991]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="clarki" subGenus="Ptilocrinus">
<emphasis id="B922C021FFCDFFEE5644F87AA94EF863" box="[592,610,1967,1990]" italics="true" pageId="23" pageNumber="404">P.</emphasis>
(
<emphasis id="B922C021FFCDFFEE5660F87BA9DCF862" box="[628,752,1966,1991]" italics="true" pageId="23" pageNumber="404">Ptilocrinus</emphasis>
) 
<emphasis id="B922C021FFCDFFEE5715F87BA86EF862" box="[769,834,1966,1991]" italics="true" pageId="23" pageNumber="404">clarki</emphasis>
<bibRefCitation id="EFC761C2FFCDFFEE575DF87AAF62F862" author="Roux" box="[841,1102,1966,1991]" pageId="23" pageNumber="404" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert, 2011</bibRefCitation>
</taxonomicName>
, an asymmetrical architecture in the proximal part of the genital pinnules, and, with 
<taxonomicName id="4C5667B0FFCDFFEE56E9F801AFEDF84E" ID-CoL="8GHQR" authority="Eleaume et al., 2011" authorityName="Eleaume et al." authorityYear="2011" box="[765,1217,2002,2027]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="amezianeae" subGenus="Ptilocrinus">
<emphasis id="B922C021FFCDFFEE56E9F801A823F84E" box="[765,783,2004,2027]" italics="true" pageId="23" pageNumber="404">P.</emphasis>
(
<emphasis id="B922C021FFCDFFEE5737F801A819F84E" box="[803,821,2004,2027]" italics="true" pageId="23" pageNumber="404">P.</emphasis>
) 
<emphasis id="B922C021FFCDFFEE575CF801A8E1F84E" box="[840,973,2004,2027]" italics="true" pageId="23" pageNumber="404">amezianeae</emphasis>
<bibRefCitation id="EFC761C2FFCDFFEE57C1F807AFEDF84E" author="Eleaume" box="[981,1217,2002,2027]" pageId="23" pageNumber="404" refString="Eleaume, M., Hemery, L. G., Bowden, D. A. &amp; Roux, M. (2011) A large new species of the genus Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae) from Antarctic seamounts, Polar Biology, 34, 1385 - 1397." type="journal article" year="2011">
Eléaume 
<emphasis id="B922C021FFCDFFEE502BF801AF54F84E" box="[1087,1144,2002,2027]" italics="true" pageId="23" pageNumber="404">et al.</emphasis>
, 2011
</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C5667B0FFCDFFE150E9F801AA82FF15" ID-CoL="8G7Z6" authority="Bather, 1908" authorityName="Bather" authorityYear="1908" class="Crinoidea" family="Hyocrinidae" genus="Dumetocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="24" lastPageNumber="405" order="Hyocrinida" pageId="23" pageNumber="404" phylum="Echinodermata" rank="species" species="antarcticus">
<emphasis id="B922C021FFCDFFE150E9F801AA38FF15" italics="true" lastPageId="24" lastPageNumber="405" pageId="23" pageNumber="404">Dumetocrinus antarcticus</emphasis>
<bibRefCitation id="EFC761C2FFC2FFE1550FFF42AA82FF15" author="Bather" box="[283,430,151,176]" pageId="24" pageNumber="405" refString="Bather, F. A. (1908) Ptilocrinus antarcticus n. sp. a crinoid dredged by the Belgian Antarctic expedition. Bulletin de l'Academie royale de Belgique, 3, 296 - 299." type="journal article" year="1908">Bather, 1908</bibRefCitation>
</taxonomicName>
, significant enlargement of the axial canal through ontogeny, more than 10 crenular units in symplexies of large specimens, and frequently long series of free brachials in mid arm. 
<bibRefCitation id="EFC761C2FFC2FFE1509FFF68AEB0FF71" author="Roux" box="[1163,1436,188,213]" pageId="24" pageNumber="405" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert (2011)</bibRefCitation>
placed 
<taxonomicName id="4C5667B0FFC2FFE154F3FF34AA65FF5D" ID-CoL="8GHQM" box="[231,329,223,248]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="species" species="brucei">
<emphasis id="B922C021FFC2FFE154F3FF34AA65FF5D" box="[231,329,223,248]" italics="true" pageId="24" pageNumber="405">P. brucei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5667B0FFC2FFE15594FF34A923FF5D" ID-CoL="8GHQP" box="[384,527,223,248]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="species" species="stukalinae">
<emphasis id="B922C021FFC2FFE15594FF34A923FF5D" box="[384,527,223,248]" italics="true" pageId="24" pageNumber="405">P. stukalinae</emphasis>
</taxonomicName>
within subgenus 
<taxonomicName id="4C5667B0FFC2FFE156C7FF0AA88AFF5D" box="[723,934,223,248]" class="Crinoidea" family="Hyocrinidae" genus="Chambersaecrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Millericrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="genus">
<emphasis id="B922C021FFC2FFE156C7FF0AA88AFF5D" box="[723,934,223,248]" italics="true" pageId="24" pageNumber="405">Chambersaecrinus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C5667B0FFC2FFE157A1FF34AF6FFF5D" ID-CoL="8GHQP" box="[949,1091,223,248]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="species" species="stukalinae">
<emphasis id="B922C021FFC2FFE157A1FF34AF6FFF5D" box="[949,1091,223,248]" italics="true" pageId="24" pageNumber="405">P. stukalinae</emphasis>
</taxonomicName>
mainly differs in having a less inflated tegmen, cover plates with a long terminal projection, symplexies with 6–8 crenular units of 1–3 crenulae, ligamentary pits between crenular units, and without enlargement of the stalk axial canal through ontogeny.
</paragraph>
<paragraph id="8BE91C33FFC2FFE154D3FE99AE72FE2D" blockId="24.[151,1436,151,392]" pageId="24" pageNumber="405">
<emphasis id="B922C021FFC2FFE154D3FE99AA75FEC0" bold="true" box="[199,345,332,357]" pageId="24" pageNumber="405">Occurrence.</emphasis>
Southern Ocean from 33° to 128°W and from 54° to 63°S (NW Weddell Sea; 
<collectingCountry id="F3415CA3FFC2FFE150E4FE99AEB4FEC0" box="[1264,1432,332,357]" name="South Georgia and the South Sandwich Islands" pageId="24" pageNumber="405">South Georgia</collectingCountry>
, Orcade, 
<collectingCountry id="F3415CA3FFC2FFE154E2FEBAAA66FE2D" box="[246,330,367,392]" name="United Kingdom" pageId="24" pageNumber="405">Orkney</collectingCountry>
and 
<collectingCountry id="F3415CA3FFC2FFE15594FEBAA910FE2D" box="[384,572,367,392]" name="United Kingdom" pageId="24" pageNumber="405">Shetland Islands</collectingCountry>
; SE Pacific basin), depth 
<quantity id="4CAEB1D6FFC2FFE1574FFEA5AF36FE22" box="[859,1050,367,392]" metricMagnitude="3" metricUnit="m" metricValue="4.0915" metricValueMax="5.465" metricValueMin="2.718" pageId="24" pageNumber="405" unit="m" value="4091.5" valueMax="5465.0" valueMin="2718.0">2,718 to 5,465 m</quantity>
(possibly 
<quantity id="4CAEB1D6FFC2FFE1509BFEBAAE7DFE2D" box="[1167,1361,367,392]" metricMagnitude="3" metricUnit="m" metricValue="4.0685" metricValueMax="5.473999999999999" metricValueMin="2.663" pageId="24" pageNumber="405" unit="m" value="4068.5" valueMax="5474.0" valueMin="2663.0">2,663 to 5,474 m</quantity>
).
</paragraph>
<caption id="DF294CBBFFC2FFE15483FE6DAE6CFE6A" box="[151,1344,440,463]" pageId="24" pageNumber="405">
<paragraph id="8BE91C33FFC2FFE15483FE6DAE6CFE6A" blockId="24.[151,1344,440,463]" box="[151,1344,440,463]" pageId="24" pageNumber="405">
<emphasis id="B922C021FFC2FFE15483FE6DAA3BFE6A" bold="true" box="[151,279,440,463]" pageId="24" pageNumber="405">TABLE 15.</emphasis>
<taxonomicName id="4C5667B0FFC2FFE1550AFE6DA9EDFE6A" ID-CoL="8GHQM" box="[286,705,440,463]" class="Crinoidea" family="Hyocrinidae" genus="Ptilocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hyocrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="species" species="brucei" subGenus="Chambersaecrinus">
<emphasis id="B922C021FFC2FFE1550AFE6DA9EDFE6A" box="[286,705,440,463]" italics="true" pageId="24" pageNumber="405">Ptilocrinus (Chambersaecrinus) brucei</emphasis>
</taxonomicName>
. Variations in proximal arm pattern. *holotype, **paratype.
</paragraph>
</caption>
<paragraph id="8BE91C33FFC2FFE154B6FE3FA9FDFD8F" blockId="24.[162,1182,490,1050]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FE3FABDCFE5A" bold="true" box="[227,240,490,511]" pageId="24" pageNumber="405">4</emphasis>
5 6 7 8+ 
<date id="FFE83AF3FFC2FFE1555DFE3FAABFFDA5" box="[329,403,490,512]" pageId="24" pageNumber="405" value="2011-10-09">9 10 11</date>
+ 
<date id="FFE83AF3FFC2FFE155B6FE3FAAD0FDA5" box="[418,508,490,512]" pageId="24" pageNumber="405" value="2014-12-13">12 13 14</date>
15+16 17 18+19 20 21+22 23 24+25 26 27+28 29 30+31 ** 1+2 3 
<emphasis id="B922C021FFC2FFE154F7FDC1ABDCFD8C" bold="true" box="[227,240,532,553]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 
<date id="FFE83AF3FFC2FFE15522FDC1AA59FD8F" box="[310,373,532,554]" pageId="24" pageNumber="405" value="2010-09-08">8 9 10</date>
<date id="FFE83AF3FFC2FFE15568FDC1AAF8FD8F" box="[380,468,532,554]" pageId="24" pageNumber="405" value="2013-12-11">11 12 13</date>
14 15 16 17 18 19 20 21
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FDE8A917FD6A" blockId="24.[162,1182,490,1050]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FDE8ABDCFDF7" bold="true" box="[227,240,573,594]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 
<date id="FFE83AF3FFC2FFE15522FDE8AA5AFDF6" box="[310,374,573,595]" pageId="24" pageNumber="405" value="2010-09-08">8 9 10</date>
+ 
<date id="FFE83AF3FFC2FFE15591FDE8AAF0FDF6" box="[389,476,573,595]" pageId="24" pageNumber="405" value="2013-12-11">11 12 13</date>
14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 
<date id="FFE83AF3FFC2FFE157C8FDE8AB87FDD9" pageId="24" pageNumber="405" value="1930-01-31">30 31 1</date>
+2 3 
<emphasis id="B922C021FFC2FFE154F7FDB2ABDCFDD9" bold="true" box="[227,240,615,636]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8 9+ 
<date id="FFE83AF3FFC2FFE15570FDB3AA90FDD9" box="[356,444,614,636]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 1+2 3 
<emphasis id="B922C021FFC2FFE154F7FD45ABDCFD00" bold="true" box="[227,240,656,677]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8 9+ 
<date id="FFE83AF3FFC2FFE15571FD45AA90FD03" box="[357,444,656,678]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
13 14 15 16+17 18 19 20 21 22 23 24+25 26 27 28 29 30 31 32 33+34 * 1+2 3 
<emphasis id="B922C021FFC2FFE154F7FD6CABDCFD6B" bold="true" box="[227,240,697,718]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8+9 
<date id="FFE83AF3FFC2FFE15570FD6CAA90FD6A" box="[356,444,697,719]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
13 14 15 16
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FD37A98EFD5D" blockId="24.[162,1182,490,1050]" box="[162,674,738,760]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FD37ABDCFD52" bold="true" box="[227,240,738,759]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8+9 
<date id="FFE83AF3FFC2FFE15570FD37AA90FD5D" box="[356,444,738,760]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
13 14 15+16 17 18 19
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FCD9A9F7FC87" blockId="24.[162,1182,490,1050]" box="[162,731,780,802]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FCD9ABDCFC84" bold="true" box="[227,240,780,801]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8+9 
<date id="FFE83AF3FFC2FFE15570FCD9AA90FC87" box="[356,444,780,802]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
13 14+15 16 17 18+19 20 *
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FCE0A990FCEE" blockId="24.[162,1182,490,1050]" box="[162,700,821,843]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FCE0ABDCFCEF" bold="true" box="[227,240,821,842]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8+9 
<date id="FFE83AF3FFC2FFE15570FCE0AA90FCEE" box="[356,444,821,843]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
13+14 15 16 17+18 19 *
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FC8BA96EFC3B" blockId="24.[162,1182,490,1050]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FC8AABDCFCD1" bold="true" box="[227,240,863,884]" pageId="24" pageNumber="405">4</emphasis>
5+6 7 8+9 
<date id="FFE83AF3FFC2FFE15570FC8BAA91FCD1" box="[356,445,862,884]" pageId="24" pageNumber="405" value="2012-11-10">10 11 12</date>
+13 14 15+16 17 18+19 20 21+22 23 24+25 26 27+28 29 ** 1+2 3 4 5+6 7 8+ 
<date id="FFE83AF3FFC2FFE15545FC5DAAB7FC3B" box="[337,411,904,926]" pageId="24" pageNumber="405" value="2011-10-09">9 10 11</date>
+ 
<date id="FFE83AF3FFC2FFE155BEFC5DA92FFC3B" box="[426,515,904,926]" pageId="24" pageNumber="405" value="2014-12-13">12 13 14</date>
15 16
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FC64A946FC62" blockId="24.[162,1182,490,1050]" box="[162,618,945,967]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FC64ABDCFC63" bold="true" box="[227,240,945,966]" pageId="24" pageNumber="405">4</emphasis>
5+6 7+ 
<date id="FFE83AF3FFC2FFE15554FC64AA52FC62" box="[320,382,945,967]" pageId="24" pageNumber="405" value="2010-09-08">8 9 10</date>
+ 
<date id="FFE83AF3FFC2FFE15599FC64AAC8FC62" box="[397,484,945,967]" pageId="24" pageNumber="405" value="2013-12-11">11 12 13</date>
14+15 16 **
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FC0FA834FC55" blockId="24.[162,1182,490,1050]" box="[162,792,986,1008]" pageId="24" pageNumber="405">
1+2 3 
<emphasis id="B922C021FFC2FFE154F7FC0FABDCFC4A" bold="true" box="[227,240,986,1007]" pageId="24" pageNumber="405">4</emphasis>
5+6 7+ 
<date id="FFE83AF3FFC2FFE15554FC0FAA52FC55" box="[320,382,986,1008]" pageId="24" pageNumber="405" value="2010-09-08">8 9 10</date>
+ 
<date id="FFE83AF3FFC2FFE15599FC0FAACEFC55" box="[397,482,986,1008]" pageId="24" pageNumber="405" value="2013-12-11">11 12 13</date>
+14 15 16+17 18 19+20 21 **
</paragraph>
<paragraph id="8BE91C33FFC2FFE154B6FBD1AA60FBBF" blockId="24.[162,1182,490,1050]" box="[162,332,1028,1050]" pageId="24" pageNumber="405">
1+2 3 4+ 
<emphasis id="B922C021FFC2FFE154E9FBD1AA26FBBC" bold="true" box="[253,266,1028,1049]" pageId="24" pageNumber="405">5</emphasis>
6 7+8
</paragraph>
</subSubSection>
</treatment>
</document>