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<document id="D69460BF6438BF7D0D0D6BD301330067" ID-DOI="10.24823/EJB.2022.407" ID-ISSN="1747-0036" ID-Zenodo-Dep="10553332" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1704825216459" checkinUser="felipe" docAuthor="Moonlight, Peter &amp; Fuentes, Alfredo F." docDate="2022" docId="03E53311FF90FFE9B62CFECFFD21FBA5" docLanguage="en" docName="EdinbJBot.79.407.1-66.pdf" docOrigin="Edinburgh Journal of Botany 79 (407)" docSource="http://dx.doi.org/10.24823/ejb.2022.407" docStyle="DocumentStyle:73DE8CF451BB5BCBD336E07A8276E690.4:EdinbJBot.2021-.journal_article" docStyleId="73DE8CF451BB5BCBD336E07A8276E690" docStyleName="EdinbJBot.2021-.journal_article" docStyleVersion="4" docTitle="Begonia andina Rusby, Bull." docType="treatment" docVersion="4" lastPageNumber="31" masterDocId="FFDC4B69FF8EFFF7B655FFBDFFD6FF96" masterDocTitle="AN UPDATED CHECKLIST AND KEY TO THE SPECIES OF BOLIVIAN BEGONIA, INCLUDING ONE NEW SPECIES" masterLastPageNumber="66" masterPageNumber="1" pageNumber="31" updateTime="1706008907290" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9097CF6B00520C8A21893B601F67C358">AN UPDATED CHECKLIST AND KEY TO THE SPECIES OF BOLIVIAN BEGONIA, INCLUDING ONE NEW SPECIES</mods:title>
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<mods:namePart id="8D49E543DBAC77F7A0AA257C8FD6CB9D">Moonlight, Peter</mods:namePart>
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<mods:namePart id="1F3D61B49E3F1CAF68D3521F770C8F4E">Fuentes, Alfredo F.</mods:namePart>
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<mods:title id="A73759C56BAC5B899B8E53DD71814DA4">Edinburgh Journal of Botany</mods:title>
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<mods:date id="70C683026A08AFBB3F616AAF2351999D">2022</mods:date>
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<mods:identifier id="6BC2113EFB4B4B0DE98E837B1AA8B739" type="DOI">10.24823/EJB.2022.407</mods:identifier>
<mods:identifier id="C856E96773FD94730D6E3EFCE16F4AE7" type="ISSN">1747-0036</mods:identifier>
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<treatment id="03E53311FF90FFE9B62CFECFFD21FBA5" LSID="urn:lsid:plazi:treatment:03E53311FF90FFE9B62CFECFFD21FBA5" httpUri="http://treatment.plazi.org/id/03E53311FF90FFE9B62CFECFFD21FBA5" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="C356D18CFF90FFE9B62CFECFFCB1FE1D" box="[121,871,370,395]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BF38207FF90FFE9B62CFECFFCB1FE1D" blockId="30.[121,1128,370,395]" box="[121,871,370,395]" pageId="30" pageNumber="31">
<heading id="D0BB356BFF90FFE9B62CFECFFCB1FE1D" box="[121,871,370,395]" fontSize="36" level="2" pageId="30" pageNumber="31" reason="2">
<emphasis id="B9385E15FF90FFE9B62CFECFFF67FE1D" bold="true" box="[121,177,370,395]" pageId="30" pageNumber="31">5.22.</emphasis>
<taxonomicName id="4C4CF984FF90FFE9B6E2FECFFE33FE1D" ID-CoL="87Y6D" authority="Rusby, Bull." authorityName="Rusby, Bull." box="[183,485,370,395]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="andina">
<treatmentCitation id="0AEDA416FF90FFE9B6E2FECFFEB4FE1D" author="Rusby" box="[183,354,370,395]" journal="Bull. New York Bot. Gard." page="108" pageId="30" pageNumber="31" score="366.1451280949614" volume="8" year="1912">
<emphasis id="B9385E15FF90FFE9B6E2FECFFEB4FE1D" bold="true" box="[183,354,370,395]" pageId="30" pageNumber="31">Begonia andina</emphasis>
</treatmentCitation>
<materialsCitation id="3B24885AFF90FFE9B73DFECFFE33FE1D" box="[360,485,370,395]" collectorName="La Paz" country="Bolivia" location="Bolivia" pageId="30" pageNumber="31" specimenCount="1">Rusby, Bull.</materialsCitation>
</taxonomicName>
New York Bot. Gard. 8: 108 (1912).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFF90FFE9B539FECFFB5EFE6D" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BF38207FF90FFE9B539FECFFB5EFE6D" blockId="30.[121,1128,370,395]" lastBlockId="30.[151,1163,408,619]" pageId="30" pageNumber="31">
<heading id="D0BB356BFF90FFE9B539FECFFBBEFE1D" box="[876,1128,370,395]" fontSize="36" level="2" pageId="30" pageNumber="31" reason="2">
<materialsCitation id="3B24885AFF90FFE9B5D7FECFFBBEFE1D" box="[898,1128,370,395]" collectingDate="1902-08-30" collectionCode="NY" collectorName="R. S. Williams" country="Bolivia" county="Schubert" elevation="1524" latitude="-14.733334" location="Santa Barbara" longLatPrecision="1287" longitude="-68.61667" municipality="Smith" pageId="30" pageNumber="31" specimenCode="NY00112290" specimenCount="1" stateProvince="La Paz Department" typeStatus="lectotype">
Type:
<collectingCountry id="F35BC297FF90FFE9B596FECFFBD8FE1D" box="[963,1038,370,395]" name="Bolivia" pageId="30" pageNumber="31">Bolivia</collectingCountry>
, [
<collectingRegion id="49884CE5FF90FFE9B249FECFFBBEFE1D" box="[1052,1128,370,395]" pageId="30" pageNumber="31">La Paz</collectingRegion>
</materialsCitation>
</heading>
Department],
<location id="8E93D4DCFF90FFE9B77EFE25FE1EFE27" LSID="urn:lsid:plazi:treatment:03E53311FF90FFE9B62CFECFFD21FBA5:8E93D4DCFF90FFE9B77EFE25FE1EFE27" box="[299,456,408,433]" country="Bolivia" county="Schubert" latitude="-14.733334" longLatPrecision="1287" longitude="-68.61667" municipality="Smith" name="Santa Barbara" pageId="30" pageNumber="31" stateProvince="La Paz Department">Santa Barbara</location>
, [
<geoCoordinate id="EE78E4C0FF90FFE9B783FE25FDFBFE27" box="[470,557,408,433]" degrees="14" direction="south" minutes="44" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="-14.733334">14°44S</geoCoordinate>
,
<geoCoordinate id="EE78E4C0FF90FFE9B463FE25FD43FE27" box="[566,661,408,433]" degrees="68" direction="west" minutes="37" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="-68.61667">68°37W</geoCoordinate>
],
<quantity id="4CB42FE2FF90FFE9B4F6FE25FD24FE27" box="[675,754,408,433]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="30" pageNumber="31" unit="ft" value="5000.0">
<elevation id="00616534FF90FFE9B4F6FE25FD24FE27" box="[675,754,408,433]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="30" pageNumber="31" unit="ft" value="5000.0">5000 ft</elevation>
</quantity>
,
<date id="FFF2A4C7FF90FFE9B4A9FE25FCADFE27" box="[764,891,408,433]" pageId="30" pageNumber="31" value="1902-08-30">
<collectingDate id="EFB65D2FFF90FFE9B4A9FE25FCADFE27" box="[764,891,408,433]" pageId="30" pageNumber="31" value="1902-08-30">30 viii 1902</collectingDate>
</date>
,
<emphasis id="B9385E15FF90FFE9B5D0FE25FBC7FE27" box="[901,1041,408,433]" italics="true" pageId="30" pageNumber="31">
<collectorName id="26B9E7D1FF90FFE9B5D0FE25FBC7FE27" box="[901,1041,408,433]" pageId="30" pageNumber="31">R.S. Williams</collectorName>
</emphasis>
1566 (
<typeStatus id="54F73CA5FF90FFE9B6C8FE00FED2FE40" box="[157,260,445,470]" pageId="30" pageNumber="31" type="lectotype">lectotype</typeStatus>
<collectionCode id="ED5D1AC2FF90FFE9B75FFE00FEFCFE40" box="[266,298,445,470]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="30" pageNumber="31" type="Herbarium">NY</collectionCode>
[
<specimenCode id="DBEA2A7CFF90FFE9B762FE00FE10FE40" box="[311,454,445,470]" collectionCode="NY" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="30" pageNumber="31" type="Herbarium">NY00112290</specimenCode>
] designated in:
<bibCitation id="5E668A6BFF90FFE9B426FE00FBC9FE40" box="[627,1055,445,470]" issue="87" journal="Revista Univ. (Cuzco)" page="81" pageId="30" pageNumber="31" score="0.150130185271274" volume="33" year="1944">Revista Univ. (Cuzco) 33(87): 81 (1944)</bibCitation>
by Smith, L.B. &amp; Schubert, B.G.;
<materialsCitation id="3B24885AFF90FFE9B7D6FE5FFBADFE6D" box="[387,1147,482,507]" collectionCode="BM, K, US" country="United States of America" location="US" pageId="30" pageNumber="31" specimenCode="BM001191441, K000322980, US00115238" specimenCount="1" typeStatus="isolectotype">
<typeStatus id="54F73CA5FF90FFE9B7D6FE5FFDC3FE6D" box="[387,533,482,507]" pageId="30" pageNumber="31" type="isolectotype">isolectotypes</typeStatus>
<collectionCode id="ED5D1AC2FF90FFE9B44EFE5FFD96FE6D" box="[539,576,482,507]" country="United Kingdom" name="Bristol Museum" pageId="30" pageNumber="31">BM</collectionCode>
[
<specimenCode id="DBEA2A7CFF90FFE9B418FE5FFD39FE6D" box="[589,751,482,507]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="30" pageNumber="31">BM001191441</specimenCode>
],
<collectionCode id="ED5D1AC2FF90FFE9B555FE5FFCD9FE6D" box="[768,783,482,507]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="30" pageNumber="31" type="Herbarium">K</collectionCode>
[
<specimenCode id="DBEA2A7CFF90FFE9B548FE5FFC7EFE6D" box="[797,936,482,507]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="30" pageNumber="31" type="Herbarium">K000322980</specimenCode>
],
<collectionCode id="ED5D1AC2FF90FFE9B5EFFE5FFC0FFE6D" box="[954,985,482,507]" pageId="30" pageNumber="31">US</collectionCode>
[
<specimenCode id="DBEA2A7CFF90FFE9B5B3FE5FFBA5FE6D" box="[998,1139,482,507]" pageId="30" pageNumber="31">US00115238</specimenCode>
]
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFF90FFE9B6C2FDB5FCDFFDFD" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8BF38207FF90FFE9B6C2FDB5FCDFFDFD" blockId="30.[151,1163,408,619]" pageId="30" pageNumber="31">
<bibCitation id="5E668A6BFF90FFE9B6C2FDB5FC5DFDB7" author="L. B. Smith &amp; B. G. Schubert" box="[151,907,520,545]" issue="87" journal="Revista Univ. (Cuzco)" page="81" pageId="30" pageNumber="31" score="0.150130185271274" volume="33" year="1944">
L.B. Smith &amp; B.G.
<location id="8E93D4DCFF90FFE9B708FDB5FE12FDB7" LSID="urn:lsid:plazi:treatment:03E53311FF90FFE9B62CFECFFD21FBA5:8E93D4DCFF90FFE9B708FDB5FE12FDB7" box="[349,452,520,545]" country="United States of America" county="Cuzco" municipality="Revista Univ." name="Schubert" pageId="30" pageNumber="31">Schubert</location>
,
<bibCitation id="5E668A6BFF90FFE9B79BFDB5FC5DFDB7" box="[462,907,520,545]" issue="87" journal="Revista Univ. (Cuzco)" page="81" pageId="30" pageNumber="31" score="0.150130185271274" volume="33" year="1944">
<collectingMunicipality id="6B97187DFF90FFE9B79BFDB5FD89FDB7" box="[462,607,520,545]" pageId="30" pageNumber="31">Revista Univ.</collectingMunicipality>
(
<collectingCounty id="6292FA8BFF90FFE9B43EFDB5FD61FDB7" box="[619,695,520,545]" pageId="30" pageNumber="31">Cuzco</collectingCounty>
) 33(87): 81 (1944)
</bibCitation>
</bibCitation>
;
<bibCitation id="5E668A6BFF90FFE9B5C0FDB5FE89FDD0" author="L. J. Dorr" issue="4" journal="Brittonia" page="223" pageId="30" pageNumber="31" score="9.729594737742866" volume="43" year="1991">
L.J.
<location id="8E93D4DCFF90FFE9B590FDB5FC20FDB7" LSID="urn:lsid:plazi:treatment:03E53311FF90FFE9B62CFECFFD21FBA5:8E93D4DCFF90FFE9B590FDB5FC20FDB7" box="[965,1014,520,545]" country="United States of America" municipality="Brittonia" name="Dorr" pageId="30" pageNumber="31">Dorr</location>
,
<bibCitation id="5E668A6BFF90FFE9B5AAFDB5FE89FDD0" issue="4" journal="Brittonia" page="223" pageId="30" pageNumber="31" score="9.729594737742866" volume="43" year="1991">
<collectingMunicipality id="6B97187DFF90FFE9B5AAFDB5FB88FDB7" box="[1023,1118,520,545]" pageId="30" pageNumber="31">Brittonia</collectingMunicipality>
43(4): 223 (1991)
</bibCitation>
</bibCitation>
; D.C. Wasshausen
<emphasis id="B9385E15FF90FFE9B461FD90FDBDFDD0" box="[564,619,557,582]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
in P.M. JØrgensen
<emphasis id="B9385E15FF90FFE9B515FD90FCA1FDD0" box="[832,887,557,582]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
(eds),
<location id="8E93D4DCFF90FFE9B591FD90FC24FDD0" LSID="urn:lsid:plazi:treatment:03E53311FF90FFE9B62CFECFFD21FBA5:8E93D4DCFF90FFE9B591FD90FC24FDD0" box="[964,1010,557,582]" country="Bolivia" municipality="Monogr. Syst. Bot. Missouri Bot." name="Cat." pageId="30" pageNumber="31">Cat.</location>
<bibCitation id="5E668A6BFF90FFE9B5ADFD90FCD3FDFD" author="Bolivia" journal="Monogr. Syst. Bot. Missouri Bot. Gard." page="383" pageId="30" pageNumber="31" score="1124.4052770756598" volume="127" year="2013">
<collectingCountry id="F35BC297FF90FFE9B5ADFD90FB93FDD0" box="[1016,1093,557,582]" name="Bolivia" pageId="30" pageNumber="31">Bolivia</collectingCountry>
,
<bibCitation id="5E668A6BFF90FFE9B6CDFDEFFCD3FDFD" box="[152,773,594,619]" journal="Monogr. Syst. Bot. Missouri Bot. Gard." page="383" pageId="30" pageNumber="31" score="1222.4916941493243" volume="127" year="2013">
<collectingMunicipality id="6B97187DFF90FFE9B6CDFDEFFDD2FDFD" box="[152,516,594,619]" pageId="30" pageNumber="31">Monogr. Syst. Bot. Missouri Bot.</collectingMunicipality>
Gard. 127: 383 (2013)
</bibCitation>
</bibCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFF90FFE9B62CFD3AFD39FD36" box="[121,751,647,672]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BF38207FF90FFE9B62CFD3AFD39FD36" blockId="30.[121,751,647,672]" box="[121,751,647,672]" pageId="30" pageNumber="31">
<emphasis id="B9385E15FF90FFE9B62CFD3AFF2EFD36" box="[121,248,647,672]" italics="true" pageId="30" pageNumber="31">Distribution</emphasis>
. Endemic to
<collectingCountry id="F35BC297FF90FFE9B7D9FD3AFE01FD36" box="[396,471,647,672]" name="Bolivia" pageId="30" pageNumber="31">Bolivia</collectingCountry>
and La Paz Department.
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFF90FFE9B62CFD06FD21FBA5" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BF38207FF90FFE9B62CFD06FD48FC22" blockId="30.[121,1151,699,948]" pageId="30" pageNumber="31">
<emphasis id="B9385E15FF90FFE9B62CFD06FEB6FD42" box="[121,352,699,724]" italics="true" pageId="30" pageNumber="31">Nomenclatural notes</emphasis>
. The protologue of
<taxonomicName id="4C4CF984FF90FFE9B46BFD06FCE2FD42" authority="Rusby" authorityName="Rusby" box="[574,820,699,724]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B9385E15FF90FFE9B46BFD06FD3EFD42" box="[574,744,699,724]" italics="true" pageId="30" pageNumber="31">Begonia andina</emphasis>
Rusby
</taxonomicName>
cited the collection
<emphasis id="B9385E15FF90FFE9B24CFD06FF01FD6C" italics="true" pageId="30" pageNumber="31">R.S. Williams</emphasis>
1566 but did not mention a herbarium (
<bibRefCitation id="EFDDFFF6FF90FFE9B4C3FD5CFCF7FD6C" author="Rusby HH" box="[662,801,737,762]" pageId="30" pageNumber="31" pagination="89 - 135" refId="ref41033" refString="Rusby HH. 1912. New species from Bolivia, collected by R. S. Williams. Bulletin of the New York Botanic Garden. 8 (28): 89 - 135." type="journal article" year="1912">Rusby, 1912</bibRefCitation>
).
<bibRefCitation id="EFDDFFF6FF90FFE9B562FD5CFB8DFD6C" author="Smith LB &amp; Schubert BG" box="[823,1115,737,762]" pageId="30" pageNumber="31" pagination="71 - 87" refId="ref41249" refString="Smith LB, Schubert BG. 1944 a. Revision de las especies Bolivianas de genero Begonia. Revista Universitat [Universidad Nacional de Cusco]. 87: 71 - 87." type="journal article" year="1944">Smith &amp; Schubert (1944a)</bibRefCitation>
cited the
<typeStatus id="54F73CA5FF90FFE9B6B7FCBBFEC4FC89" box="[226,274,774,799]" pageId="30" pageNumber="31">type</typeStatus>
as in New York herbarium, which is an effective lectotypification of the name. The protologue also cited a collection made by Richard Spruce in
<collectingRegion id="49884CE5FF90FFE9B508FC96FB86FCD2" box="[861,1104,811,836]" country="Ecuador" name="Chimborazo" pageId="30" pageNumber="31">Chimborazo Province</collectingRegion>
,
<collectingCountry id="F35BC297FF90FFE9B62FFCECFF01FCFC" box="[122,215,849,874]" name="Ecuador" pageId="30" pageNumber="31">Ecuador</collectingCountry>
, in
<date id="FFF2A4C7FF90FFE9B6AEFCECFEA5FCFC" box="[251,371,849,874]" pageId="30" pageNumber="31" value="1860-06">June 1860</date>
as the same species. The only sheet we know of that matches this description is a collection of
<taxonomicName id="4C4CF984FF90FFE9B795FCCBFD17FC19" authority="A. DC." authorityName="A. DC." box="[448,705,886,911]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="holtonis">
<emphasis id="B9385E15FF90FFE9B795FCCBFDAEFC19" box="[448,632,886,911]" italics="true" pageId="30" pageNumber="31">Begonia holtonis</emphasis>
A.DC.
</taxonomicName>
, which is a superficially similar but distantly related species (
<bibRefCitation id="EFDDFFF6FF90FFE9B7CDFC26FD58FC22" author="Moonlight PW &amp; Ardi WH &amp; Padilla LA &amp; Chung K-F &amp; Fuller D &amp; Girmansyah D &amp; Hollands R &amp; Jara-Munoz A &amp; Kiew R &amp; Leong W-C &amp; Liu Y &amp; Mahardika A &amp; Marasinghe LDK &amp; O'Connor M &amp; Peng C-I &amp; Perez AJ &amp; Phutthai T &amp; Pullan M &amp; Rajbhandary S &amp; Reynel C &amp; Rubite RR &amp; Sang J &amp; Scherberich D &amp; Shui Y-M &amp; Tebbitt MC &amp; Thomas DC &amp; Wilson HP &amp; Zaini NH &amp; Hughes M." box="[408,654,923,948]" pageId="30" pageNumber="31" pagination="267 - 323" refId="ref40678" refString="Moonlight PW, Ardi WH, Padilla LA, Chung K-F, Fuller D, Girmansyah D, Hollands R, Jara-Munoz A, Kiew R, Leong W-C, Liu Y, Mahardika A, Marasinghe LDK, O'Connor M, Peng C-I, Perez AJ, Phutthai T, Pullan M, Rajbhandary S, Reynel C, Rubite RR, Sang J, Scherberich D, Shui Y-M, Tebbitt MC, Thomas DC, Wilson HP, Zaini NH, Hughes M. 2018. Dividing and conquering the fastest-growing genus: towards a natural sectional classification of the mega-diverse genus Begonia (Begoniaceae). Taxon. 67 (2): 267 - 323. https: // doi. org / 10.12705 / 672.3" type="journal article" year="2018">
Moonlight
<emphasis id="B9385E15FF90FFE9B441FC26FD9CFC22" box="[532,586,923,948]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
, 2018
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF38207FF90FFE9B62CFC6DFD21FBA5" blockId="30.[121,1102,976,1075]" pageId="30" pageNumber="31">
<emphasis id="B9385E15FF90FFE9B62CFC6DFE98FC7F" box="[121,334,976,1001]" italics="true" pageId="30" pageNumber="31">Identification notes</emphasis>
.
<taxonomicName id="4C4CF984FF90FFE9B70EFC6DFDD0FC7F" box="[347,518,976,1001]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B9385E15FF90FFE9B70EFC6DFDD0FC7F" box="[347,518,976,1001]" italics="true" pageId="30" pageNumber="31">Begonia andina</emphasis>
</taxonomicName>
is an easy species to determine on account of the densely tomentose-stellate indumentum on its stems, petioles, and the underside of its leaves, which is unique among Bolivian
<taxonomicName id="4C4CF984FF90FFE9B46DFBA7FD21FBA5" box="[568,759,1050,1075]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9385E15FF90FFE9B46DFBA7FD44FBA5" box="[568,658,1050,1075]" italics="true" pageId="30" pageNumber="31">Begonia</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>