594 lines
71 KiB
XML
594 lines
71 KiB
XML
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<mods:title id="4AADBDF4F9E1DE63309349B36C5E373C">New insights in the taxonomy of Lepismatidae (Insecta, Zygentoma) with an updated key to genera and future challenges</mods:title>
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<mods:namePart id="9D8B746B9BC3987F27F8BF8541A3EFCD">Molero-Baltanás, Rafael</mods:namePart>
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<mods:affiliation id="D302D44961C7F43598ACCF336E4EB6B0">Department of Zoology, University of Córdoba, C- 1 Campus de Rabanales, 14071 - Córdoba, Spain.</mods:affiliation>
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<mods:title id="6B4EE495BEBD0B253265A5260CC78CD4">European Journal of Taxonomy</mods:title>
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<treatment id="03D11A22FFC0F758FEB16DB12085AD86" ID-DOI="http://doi.org/10.5281/zenodo.12667746" ID-GBIF-Taxon="234495138" ID-Zenodo-Dep="12667746" LSID="urn:lsid:plazi:treatment:03D11A22FFC0F758FEB16DB12085AD86" httpUri="http://treatment.plazi.org/id/03D11A22FFC0F758FEB16DB12085AD86" lastPageId="31" lastPageNumber="111" pageId="28" pageNumber="108">
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<paragraph id="8BC7AB34FFC0F75BFEB16DB126DBAFF0" blockId="28.[380,1208,945,973]" box="[380,1208,945,973]" pageId="28" pageNumber="108">
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<heading id="D08F1C58FFC0F75BFEB16DB126DBAFF0" box="[380,1208,945,973]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="108" reason="2">
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Genus
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<taxonomicName id="4C78D0B7FFC0F75BFE036DB22621AFF1" authority="Wygodzinsky, 1955 sensu Irish, 1987" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[462,1090,945,973]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="genus" status="stat. nov.">
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<emphasis id="B90C7726FFC0F75BFE036DB220E2AFF1" bold="true" box="[462,641,945,972]" italics="true" pageId="28" pageNumber="108">Sceletolepisma</emphasis>
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<bibRefCitation id="EFE9D6C5FFC0F75BFD446DB12111AFF0" author="Wygodzinsky P." box="[649,882,946,973]" pageId="28" pageNumber="108" pagination="83 - 190" refId="ref22565" refString="Wygodzinsky P. 1955. Thysanura. In: Hanstrom B., Brink B. & Rudebeck G. (eds) South African Animal Life 2: 83 - 190. AlmQvist & Wiksells Boktryckeri AB, Uppsala." type="book chapter" year="1955">Wygodzinsky, 1955</bibRefCitation>
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sensu
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<bibRefCitation id="EFE9D6C5FFC0F75BFC0C6DB12621AFF1" author="Irish J." box="[961,1090,946,972]" pageId="28" pageNumber="108" pagination="147 - 207" refId="ref20284" refString="Irish J. 1987. Revision of the genus Ctenolepisma Escherich (Thysanura: Lepismatidae) in southern Africa. Cimbebasia (A) 7 (11): 147 - 207." type="journal article" year="1987">Irish, 1987</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23FCA5DFFC0F75BFB826DB026DBAFF0" box="[1103,1208,947,973]" pageId="28" pageNumber="108">stat. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C362F8BFFFC0F75BFF706DF0213EA833" box="[189,861,1011,1038]" pageId="28" pageNumber="108" type="reference_group">
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<paragraph id="8BC7AB34FFC0F75BFF706DF0213EA833" blockId="28.[189,1399,1011,1144]" box="[189,861,1011,1038]" pageId="28" pageNumber="108">
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<treatmentCitation id="0AD98D25FFC0F75BFF706DF0213BA833" author="Wygodzinsky P." box="[189,856,1011,1038]" page="154" pageId="28" pageNumber="108" year="1955">
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<taxonomicName id="4C78D0B7FFC0F75BFF706DF0213BA833" authority="Wygodzinsky, 1955: 154" authorityName="Wygodzinsky" authorityPageNumber="154" authorityYear="1955" box="[189,856,1011,1038]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Sceletolepisma">
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<emphasis id="B90C7726FFC0F75BFF706DF0233DA830" box="[189,350,1011,1037]" italics="true" pageId="28" pageNumber="108">Ctenolepisma</emphasis>
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(
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<emphasis id="B90C7726FFC0F75BFEBD6DF02043A830" box="[368,544,1011,1037]" italics="true" pageId="28" pageNumber="108">Sceletolepisma</emphasis>
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)
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<bibRefCitation id="EFE9D6C5FFC0F75BFDFC6DF0213BA833" author="Wygodzinsky P." box="[561,856,1011,1038]" pageId="28" pageNumber="108" pagination="83 - 190" refId="ref22565" refString="Wygodzinsky P. 1955. Thysanura. In: Hanstrom B., Brink B. & Rudebeck G. (eds) South African Animal Life 2: 83 - 190. AlmQvist & Wiksells Boktryckeri AB, Uppsala." type="book chapter" year="1955">Wygodzinsky, 1955: 154</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C362F8BFFFC0F75BFCA96DF7231DA845" pageId="28" pageNumber="108" type="type_taxon">
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<paragraph id="8BC7AB34FFC0F75BFCA96DF7231DA845" blockId="28.[189,1399,1011,1144]" pageId="28" pageNumber="108">
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<typeStatus id="54C31596FFC0F75BFCA96DF721FDA833" box="[868,926,1012,1038]" pageId="28" pageNumber="108">Type</typeStatus>
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species:
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<taxonomicName id="4C78D0B7FFC0F75BFBC76DF02001A80C" authority="Wygodzinsky, 1955" authorityName="Wygodzinsky" authorityYear="1955" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="species" species="arenicola" subGenus="Sceletolepisma">
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<emphasis id="B90C7726FFC0F75BFBC76DF026C8A830" box="[1034,1195,1011,1037]" italics="true" pageId="28" pageNumber="108">Ctenolepisma</emphasis>
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(
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<emphasis id="B90C7726FFC0F75BFB706DF0270EA830" box="[1213,1389,1011,1037]" italics="true" pageId="28" pageNumber="108">Sceletolepisma</emphasis>
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)
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<emphasis id="B90C7726FFC0F75BFF276A14233AA80C" box="[234,345,1047,1073]" italics="true" pageId="28" pageNumber="108">arenicola</emphasis>
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||
<bibRefCitation id="EFE9D6C5FFC0F75BFEA06A142001A80C" author="Wygodzinsky P." box="[365,610,1046,1073]" pageId="28" pageNumber="108" pagination="83 - 190" refId="ref22565" refString="Wygodzinsky P. 1955. Thysanura. In: Hanstrom B., Brink B. & Rudebeck G. (eds) South African Animal Life 2: 83 - 190. AlmQvist & Wiksells Boktryckeri AB, Uppsala." type="book chapter" year="1955">Wygodzinsky, 1955</bibRefCitation>
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</taxonomicName>
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by original designation [created as monotypic subgenus of
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<taxonomicName id="4C78D0B7FFC0F75BFF276A392064A869" ID-CoL="3WZW" authority="Escherich" authorityName="Escherich" authorityYear="1905" box="[234,519,1082,1108]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="B90C7726FFC0F75BFF276A3923E8A869" box="[234,395,1082,1108]" italics="true" pageId="28" pageNumber="108">Ctenolepisma</emphasis>
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Escherich
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</taxonomicName>
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for
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<taxonomicName id="4C78D0B7FFC0F75BFDF56A392188A868" authority="Wygodzinsky, 1955" authorityName="Wygodzinsky" authorityYear="1955" box="[568,1003,1082,1109]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="species" species="arenicola" subGenus="Sceletolepisma">
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<emphasis id="B90C7726FFC0F75BFDF56A392030A869" box="[568,595,1082,1108]" italics="true" pageId="28" pageNumber="108">C.</emphasis>
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(
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<emphasis id="B90C7726FFC0F75BFDA96A392019A869" box="[612,634,1082,1108]" italics="true" pageId="28" pageNumber="108">S.</emphasis>
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)
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<emphasis id="B90C7726FFC0F75BFD466A392099A869" box="[651,762,1082,1108]" italics="true" pageId="28" pageNumber="108">arenicola</emphasis>
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||
<bibRefCitation id="EFE9D6C5FFC0F75BFCCF6A392188A868" author="Wygodzinsky P." box="[770,1003,1082,1109]" pageId="28" pageNumber="108" pagination="83 - 190" refId="ref22565" refString="Wygodzinsky P. 1955. Thysanura. In: Hanstrom B., Brink B. & Rudebeck G. (eds) South African Animal Life 2: 83 - 190. AlmQvist & Wiksells Boktryckeri AB, Uppsala." type="book chapter" year="1955">Wygodzinsky, 1955</bibRefCitation>
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</taxonomicName>
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on the basis of the 2+2 combs on urotergite I].
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C362F8BFFFC0F75BFF706A9D269FA8E1" pageId="28" pageNumber="108" type="reference_group">
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<paragraph id="8BC7AB34FFC0F75BFF706A9D269FA8E1" blockId="28.[189,1399,1182,1244]" pageId="28" pageNumber="108">
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<treatmentCitationGroup id="AB688C1AFFC0F75BFF706A9D269FA8E1" pageId="28" pageNumber="108">
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||
<taxonomicName id="4C78D0B7FFC0F75BFF706A9D204AA884" authorityName="Wygodzinsky" authorityYear="1955" box="[189,553,1182,1209]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Sceletolepisma">
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||
<emphasis id="B90C7726FFC0F75BFF706A9D233DA885" box="[189,350,1182,1208]" italics="true" pageId="28" pageNumber="108">Ctenolepisma</emphasis>
|
||
(
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<emphasis id="B90C7726FFC0F75BFEA26A9D207CA885" box="[367,543,1182,1208]" italics="true" pageId="28" pageNumber="108">Sceletolepisma</emphasis>
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||
)
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</taxonomicName>
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–
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<treatmentCitation id="0AD98D25FFC0F75BFD886A9D2094A884" author="Irish J." box="[581,759,1182,1209]" page="149" pageId="28" pageNumber="108" year="1987">
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||
<bibRefCitation id="EFE9D6C5FFC0F75BFD886A9D2094A884" author="Irish J." box="[581,759,1182,1209]" pageId="28" pageNumber="108" pagination="147 - 207" refId="ref20284" refString="Irish J. 1987. Revision of the genus Ctenolepisma Escherich (Thysanura: Lepismatidae) in southern Africa. Cimbebasia (A) 7 (11): 147 - 207." type="journal article" year="1987">Irish 1987: 149</bibRefCitation>
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</treatmentCitation>
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[redefined subgenus to include all species with at least one pair of medial urosternal combs irrespective of the arrangement of urotergal combs].
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C362F8BFFFC0F75BFF706B002714AB05" pageId="28" pageNumber="108" type="diagnosis">
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<paragraph id="8BC7AB34FFC0F75BFF706B002355A920" blockId="28.[189,310,1283,1309]" box="[189,310,1283,1309]" pageId="28" pageNumber="108">
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<heading id="D08F1C58FFC0F75BFF706B002355A920" bold="true" box="[189,310,1283,1309]" fontSize="11" level="3" pageId="28" pageNumber="108" reason="3">
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<emphasis id="B90C7726FFC0F75BFF706B002355A920" bold="true" box="[189,310,1283,1309]" pageId="28" pageNumber="108">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="8BC7AB34FFC0F75BFF706B312714AB05" blockId="28.[189,1401,1330,1849]" pageId="28" pageNumber="108">
|
||
Silverfish with fusiform or subcylindrical shape, bearing feathered macrochaetae and lacking pronged sensilla on apical article of maxillary palp. Frontal area with a bare gap between two big lateral tufts of macrochaetae; clypeus and labrum usually with 1+1 tufts. Apical article of labial palp with (2)3–5 papillae arranged in one row. Pronotum with setal collar; all thoracic nota with several lateral combs and usually with 1+1 posterolateral combs. Anterior trichobothrial areas of pronotum and mesonotum associated with antepenultimate lateral combs (N−2); sometimes those of pronotum in anterior to antepenultimate (N−3) lateral combs. Thoracic sternites variable in shape, usually cordiform with convex hind margin, prosternum usually smaller than meso- and metasternum. Coxae and femora covered with rounded orbicular scales similar to those of body; tibiae and tarsi without scales, only covered by setae; remaining appendages without scales. Several urotergites (usually II–V, II–VI, II–VII or II–VIII) with 3+3 combs of macrochaetae. Urotergite I with 1+1 or 2+2 combs. Tenth urotergite trapezoidal, with straight hind margin, sometimes slightly concave or convex. Several urosternites (at least four) with one median comb, usually present on abdominal sternites I–VI, I–VII, II–VI or II–VII); 1+1 lateral combs on some urosternites (usually on II–VIII or III–VIII). Coxites IX with or without a transverse comb. One to three pairs of abdominal styli. Males without paramera. Ovipositor of primary
|
||
<typeStatus id="54C31596FFC0F75BFBF6691C260EAB04" box="[1083,1133,1823,1849]" pageId="28" pageNumber="108">type</typeStatus>
|
||
or apically sclerotized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362F8BFFFC0F759FF70696321C9A8C5" lastPageId="30" lastPageNumber="110" pageId="28" pageNumber="108" type="discussion">
|
||
<paragraph id="8BC7AB34FFC0F75BFF706963234CAB47" blockId="28.[189,303,1888,1914]" box="[189,303,1888,1914]" pageId="28" pageNumber="108">
|
||
<heading id="D08F1C58FFC0F75BFF706963234CAB47" bold="true" box="[189,303,1888,1914]" fontSize="11" level="3" pageId="28" pageNumber="108" reason="3">
|
||
<emphasis id="B90C7726FFC0F75BFF706963234CAB47" bold="true" box="[189,303,1888,1914]" pageId="28" pageNumber="108">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC0F75AFF70698D2677AA97" blockId="28.[189,1399,1934,1996]" lastBlockId="29.[186,1399,1257,1706]" lastPageId="29" lastPageNumber="109" pageId="28" pageNumber="108">
|
||
This taxon was erected to include only one species,
|
||
<taxonomicName id="4C78D0B7FFC0F75BFC98698D2712AB94" authority="Wygodzinsky, 1955" authorityName="Wygodzinsky" authorityYear="1955" box="[853,1393,1934,1961]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="108" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="B90C7726FFC0F75BFC98698D2610AB95" box="[853,1139,1934,1960]" italics="true" pageId="28" pageNumber="108">Ctenolepisma arenicola</emphasis>
|
||
<bibRefCitation id="EFE9D6C5FFC0F75BFB4D698D2712AB94" author="Wygodzinsky P." box="[1152,1393,1934,1961]" pageId="28" pageNumber="108" pagination="83 - 190" refId="ref22565" refString="Wygodzinsky P. 1955. Thysanura. In: Hanstrom B., Brink B. & Rudebeck G. (eds) South African Animal Life 2: 83 - 190. AlmQvist & Wiksells Boktryckeri AB, Uppsala." type="book chapter" year="1955">Wygodzinsky, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has 2+2 combs on urotergite I; this character was exclusive to this species at that moment within
|
||
<taxonomicName id="4C78D0B7FFC1F75AFF706AEA233DA93E" authorityName="Escherich" authorityYear="1905" box="[189,350,1257,1283]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFF706AEA233DA93E" box="[189,350,1257,1283]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
, but several additional species sharing this character were discovered afterwards.
|
||
<bibRefCitation id="EFE9D6C5FFC1F75AFA8C6AEA236FA91B" author="Irish J." pageId="29" pageNumber="109" pagination="147 - 207" refId="ref20284" refString="Irish J. 1987. Revision of the genus Ctenolepisma Escherich (Thysanura: Lepismatidae) in southern Africa. Cimbebasia (A) 7 (11): 147 - 207." type="journal article" year="1987">Irish (1987)</bibRefCitation>
|
||
redefined
|
||
<taxonomicName id="4C78D0B7FFC1F75AFE406B0F205EA91B" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[397,573,1292,1318]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFE406B0F205EA91B" box="[397,573,1292,1318]" italics="true" pageId="29" pageNumber="109">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
using a different criterion for distinguishing this subgenus from the remaining species of
|
||
<taxonomicName id="4C78D0B7FFC1F75AFE716B2C203FA974" authorityName="Escherich" authorityYear="1905" box="[444,604,1327,1353]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFE716B2C203FA974" box="[444,604,1327,1353]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
, a group that includes the
|
||
<typeStatus id="54C31596FFC1F75AFC566B3321ADA977" box="[923,974,1328,1354]" pageId="29" pageNumber="109">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="4C78D0B7FFC1F75AFB026B2C270CA974" authorityName="Escherich" authorityYear="1905" box="[1231,1391,1327,1353]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFB026B2C270CA974" box="[1231,1391,1327,1353]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C78D0B7FFC1F75AFF706B502326A951" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[189,325,1362,1389]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="species" species="lineatum">
|
||
<emphasis id="B90C7726FFC1F75AFF706B502326A951" box="[189,325,1362,1389]" italics="true" pageId="29" pageNumber="109">C. lineatum</emphasis>
|
||
</taxonomicName>
|
||
. The new criterion was the possession of at least one median comb of macrochaetae on some urosternites, a character also present in
|
||
<taxonomicName id="4C78D0B7FFC1F75AFDB96B752161A9B2" authorityName="Wygodzinsky" authorityYear="1955" box="[628,770,1397,1424]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="B90C7726FFC1F75AFDB96B752161A9B2" box="[628,770,1397,1424]" italics="true" pageId="29" pageNumber="109">C. arenicola</emphasis>
|
||
</taxonomicName>
|
||
.The subgenus
|
||
<taxonomicName id="4C78D0B7FFC1F75AFC626B762633A9B2" authorityName="Escherich" authorityYear="1905" box="[943,1104,1397,1423]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC1F75AFC626B762633A9B2" box="[943,1104,1397,1423]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC1F75AFB956B7526F1A9AD" box="[1112,1170,1398,1424]" pageId="29" pageNumber="109" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
includes, according to Irish, all species without median bristle-combs, only having 1+1 lateral combs of macrochaetae on most urosternites (usually III–VIII), but he suggested that this group could be heterogeneous. At the moment, the group with median urosternal bristle combs includes about 80 species. We consider that
|
||
<taxonomicName id="4C78D0B7FFC1F75AFF726801230EAA21" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[191,365,1538,1564]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFF726801230EAA21" box="[191,365,1538,1564]" italics="true" pageId="29" pageNumber="109">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
, in the sense established by
|
||
<bibRefCitation id="EFE9D6C5FFC1F75AFD7168012128AA21" author="Irish J." box="[700,843,1538,1564]" pageId="29" pageNumber="109" pagination="147 - 207" refId="ref20284" refString="Irish J. 1987. Revision of the genus Ctenolepisma Escherich (Thysanura: Lepismatidae) in southern Africa. Cimbebasia (A) 7 (11): 147 - 207." type="journal article" year="1987">Irish (1987)</bibRefCitation>
|
||
, should be considered as an independent genus because all species examined share several characters apart from the presence of the median combs of macrochaetae; these characters are constant in
|
||
<taxonomicName id="4C78D0B7FFC1F75AFCD6684A21A8AA5E" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[795,971,1609,1635]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFCD6684A21A8AA5E" box="[795,971,1609,1635]" italics="true" pageId="29" pageNumber="109">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
, while in
|
||
<taxonomicName id="4C78D0B7FFC1F75AFB81684A268EAA5E" authorityName="Escherich" authorityYear="1905" box="[1100,1261,1609,1635]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC1F75AFB81684A268EAA5E" box="[1100,1261,1609,1635]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC1F75AFB35684A275AAA5E" box="[1272,1337,1609,1635]" pageId="29" pageNumber="109" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
they are variable. Although the presence or absence of median combs is diagnostic in most cases, some exceptions have occasionally been observed in some
|
||
<taxonomicName id="4C78D0B7FFC1F75AFCE3688C21ACAA94" authorityName="Escherich" authorityYear="1905" box="[814,975,1679,1705]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC1F75AFCE3688C21ACAA94" box="[814,975,1679,1705]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC1F75AFC1B68932677AA97" box="[982,1044,1680,1706]" pageId="29" pageNumber="109" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="DF07FBBCFFC1F75AFF706F0A2197ADB1" ID-Table-UUID="DF07FBBCFFC1F75AFF706F0A2197ADB1" httpUri="http://table.plazi.org/id/DF07FBBCFFC1F75AFF706F0A2197ADB1" pageId="29" pageNumber="109" startId="29.[189,259,265,291]" targetBox="[213,1382,455,1157]" targetIsTable="true" targetPageId="29">
|
||
<paragraph id="8BC7AB34FFC1F75AFF706F0A2197ADB1" blockId="29.[189,1400,265,397]" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFF706F0A2341AD1E" bold="true" box="[189,290,265,291]" pageId="29" pageNumber="109">Table 1.</emphasis>
|
||
Abdominal chaetotaxy of
|
||
<taxonomicName id="4C78D0B7FFC1F75AFDA86F0A265EAD19" authority="(Escherich, 1905)" authorityName="Molero-Baltanás & Gaju-Ricart & Smith" authorityYear="2024" baseAuthorityName="Escherich" baseAuthorityYear="1905" box="[613,1085,265,292]" class="Insecta" family="Lepismatidae" genus="Caribesella" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="species" species="impudica" status="comb. nov.">
|
||
<emphasis id="B90C7726FFC1F75AFDA86F0A2107AD1E" box="[613,868,265,291]" italics="true" pageId="29" pageNumber="109">Caribesella impudica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE9D6C5FFC1F75AFCBE6F0A2656AD19" author="Escherich K." box="[883,1077,265,292]" pageId="29" pageNumber="109" pagination="1 - 164" refId="ref20193" refString="Escherich K. 1905. Das System der Lepismatiden. Zoologica (Stuttgart) 43: 1 - 164. https: // doi. org / 10.5962 / bhl. title. 7909" type="journal article" year="1905">Escherich, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC1F75AFB8A6F0A26AEAD19" box="[1095,1229,265,292]" pageId="29" pageNumber="109" rank="species">comb. nov.</taxonomicNameLabel>
|
||
, from Uverito (Venezuela). The number of macrochaetae per comb is indicated in each position (minimum and maximum number observed). The variability indicated by
|
||
<bibRefCitation id="EFE9D6C5FFC1F75AFCB06F4C26E0AD57" author="Wygodzinsky P." box="[893,1155,335,362]" pageId="29" pageNumber="109" pagination="28 - 49" refId="ref22606" refString="Wygodzinsky P. 1959 a. Thysanura and Machilida of the Lesser Antilles and northern South America. Studies on the Fauna of Curacao and other Caribbean Islands 36: 28 - 49." type="journal article" year="1959">Wygodzinsky (1959a)</bibRefCitation>
|
||
for specimens from different localities fits inside the range of variation shown in the table.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC7AB34FFC1F75AFF186FC42779A802" pageId="29" pageNumber="109">
|
||
<table id="F9785994FFC108B8FF186FC42705A8B8" box="[213,1382,455,1157]" gridcols="6" gridrows="11" pageId="29" pageNumber="109">
|
||
<tr id="3548A976FFC108B8FF186FC42705AE1A" box="[213,1382,455,551]" gridrow="0" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186FC42302AE1A" box="[213,353,455,551]" gridcol="0" gridrow="0" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFF186FC4232CAE39" bold="true" box="[213,353,455,516]" pageId="29" pageNumber="109">Abdominal segment</emphasis>
|
||
</th>
|
||
<th id="7699C00AFFC108B8FE426FC42051AE1A" box="[399,562,455,551]" gridcol="1" gridrow="0" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFE696FC42051AE1A" bold="true" pageId="29" pageNumber="109">Urotergal comb A (infralateral)</emphasis>
|
||
</th>
|
||
<th id="7699C00AFFC108B8FDB96FC4208DAE1A" box="[628,750,455,551]" gridcol="2" gridrow="0" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFDB96FC42080AE1A" bold="true" pageId="29" pageNumber="109">Urotergal comb B (lateral)</emphasis>
|
||
</th>
|
||
<th id="7699C00AFFC108B8FCF36FC421ACAE1A" box="[830,975,455,551]" gridcol="3" gridrow="0" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFC876FC421ACAE1A" bold="true" pageId="29" pageNumber="109">Urotergal comb C (sublateral)</emphasis>
|
||
</th>
|
||
<th id="7699C00AFFC108B8FBCF6FC426FCAE1A" box="[1026,1183,455,551]" gridcol="4" gridrow="0" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFBC06FC42607AE1A" bold="true" pageId="29" pageNumber="109">Urosternal lateral comb (L)</emphasis>
|
||
</th>
|
||
<th id="7699C00AFFC108B8FB736FC42705AE1A" box="[1214,1382,455,551]" gridcol="5" gridrow="0" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFB026FC42749AE1A" bold="true" pageId="29" pageNumber="109">Urosternal median comb (M)</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186C4B2705AE5E" box="[213,1382,584,611]" gridrow="1" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186C4B2302AE5E" box="[213,353,584,611]" gridcol="0" gridrow="1" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEDB6C4B2342AE5F" bold="true" box="[278,289,584,610]" pageId="29" pageNumber="109">I</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426C4B2051AE5E" box="[399,562,584,611]" gridcol="1" gridrow="1" pageId="29" pageNumber="109">7–9</td>
|
||
<td id="7699C00AFFC108B8FDB96C4B208DAE5E" box="[628,750,584,611]" gridcol="2" gridrow="1" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FCF36C4B21ACAE5E" box="[830,975,584,611]" gridcol="3" gridrow="1" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FBCF6C4B26FCAE5E" box="[1026,1183,584,611]" gridcol="4" gridrow="1" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FB736C4B2705AE5E" box="[1214,1382,584,611]" gridcol="5" gridrow="1" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186C7E2705AEAA" box="[213,1382,637,663]" gridrow="2" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186C7E2302AEAA" box="[213,353,637,663]" gridcol="0" gridrow="2" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEDD6C7E2344AEAA" bold="true" box="[272,295,637,663]" pageId="29" pageNumber="109">II</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426C7E2051AEAA" box="[399,562,637,663]" gridcol="1" gridrow="2" pageId="29" pageNumber="109">6–10</td>
|
||
<td id="7699C00AFFC108B8FDB96C7E208DAEAA" box="[628,750,637,663]" gridcol="2" gridrow="2" pageId="29" pageNumber="109">4–7</td>
|
||
<td id="7699C00AFFC108B8FCF36C7E21ACAEAA" box="[830,975,637,663]" gridcol="3" gridrow="2" pageId="29" pageNumber="109">4–7</td>
|
||
<td id="7699C00AFFC108B8FBCF6C7E26FCAEAA" box="[1026,1183,637,663]" gridcol="4" gridrow="2" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FB736C7E2705AEAA" box="[1214,1382,637,663]" gridcol="5" gridrow="2" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186CB22705AEF1" box="[213,1382,689,716]" gridrow="3" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186CB22302AEF1" box="[213,353,689,716]" gridcol="0" gridrow="3" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEC76CB2234FAEF6" bold="true" box="[266,300,689,715]" pageId="29" pageNumber="109">III</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426CB22051AEF1" box="[399,562,689,716]" gridcol="1" gridrow="3" pageId="29" pageNumber="109">6–9</td>
|
||
<td id="7699C00AFFC108B8FDB96CB2208DAEF1" box="[628,750,689,716]" gridcol="2" gridrow="3" pageId="29" pageNumber="109">4–7</td>
|
||
<td id="7699C00AFFC108B8FCF36CB221ACAEF1" box="[830,975,689,716]" gridcol="3" gridrow="3" pageId="29" pageNumber="109">4–7</td>
|
||
<td id="7699C00AFFC108B8FBCF6CB226FCAEF1" box="[1026,1183,689,716]" gridcol="4" gridrow="3" pageId="29" pageNumber="109">9–11</td>
|
||
<td id="7699C00AFFC108B8FB736CB22705AEF1" box="[1214,1382,689,716]" gridcol="5" gridrow="3" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186CE52705AF3F" box="[213,1382,742,770]" gridrow="4" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186CE52302AF3F" box="[213,353,742,770]" gridcol="0" gridrow="4" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEC66CE5234FAF3D" bold="true" box="[267,300,742,768]" pageId="29" pageNumber="109">IV</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426CE52051AF3F" box="[399,562,742,770]" gridcol="1" gridrow="4" pageId="29" pageNumber="109">7–9</td>
|
||
<td id="7699C00AFFC108B8FDB96CE5208DAF3F" box="[628,750,742,770]" gridcol="2" gridrow="4" pageId="29" pageNumber="109">5–8</td>
|
||
<td id="7699C00AFFC108B8FCF36CE521ACAF3F" box="[830,975,742,770]" gridcol="3" gridrow="4" pageId="29" pageNumber="109">5–7</td>
|
||
<td id="7699C00AFFC108B8FBCF6CE526FCAF3F" box="[1026,1183,742,770]" gridcol="4" gridrow="4" pageId="29" pageNumber="109">10–12</td>
|
||
<td id="7699C00AFFC108B8FB736CE52705AF3F" box="[1214,1382,742,770]" gridcol="5" gridrow="4" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186D182705AF0B" box="[213,1382,795,822]" gridrow="5" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186D182302AF0B" box="[213,353,795,822]" gridcol="0" gridrow="5" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEDC6D182345AF08" bold="true" box="[273,294,795,821]" pageId="29" pageNumber="109">V</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426D182051AF0B" box="[399,562,795,822]" gridcol="1" gridrow="5" pageId="29" pageNumber="109">9–11</td>
|
||
<td id="7699C00AFFC108B8FDB96D18208DAF0B" box="[628,750,795,822]" gridcol="2" gridrow="5" pageId="29" pageNumber="109">5–8</td>
|
||
<td id="7699C00AFFC108B8FCF36D1821ACAF0B" box="[830,975,795,822]" gridcol="3" gridrow="5" pageId="29" pageNumber="109">5–7</td>
|
||
<td id="7699C00AFFC108B8FBCF6D1826FCAF0B" box="[1026,1183,795,822]" gridcol="4" gridrow="5" pageId="29" pageNumber="109">9–12</td>
|
||
<td id="7699C00AFFC108B8FB736D182705AF0B" box="[1214,1382,795,822]" gridcol="5" gridrow="5" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186D532705AF56" box="[213,1382,848,875]" gridrow="6" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186D532302AF56" box="[213,353,848,875]" gridcol="0" gridrow="6" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEC66D53234FAF57" bold="true" box="[267,300,848,874]" pageId="29" pageNumber="109">VI</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426D532051AF56" box="[399,562,848,875]" gridcol="1" gridrow="6" pageId="29" pageNumber="109">9–13</td>
|
||
<td id="7699C00AFFC108B8FDB96D53208DAF56" box="[628,750,848,875]" gridcol="2" gridrow="6" pageId="29" pageNumber="109">5–8</td>
|
||
<td id="7699C00AFFC108B8FCF36D5321ACAF56" box="[830,975,848,875]" gridcol="3" gridrow="6" pageId="29" pageNumber="109">5–8</td>
|
||
<td id="7699C00AFFC108B8FBCF6D5326FCAF56" box="[1026,1183,848,875]" gridcol="4" gridrow="6" pageId="29" pageNumber="109">10–14</td>
|
||
<td id="7699C00AFFC108B8FB736D532705AF56" box="[1214,1382,848,875]" gridcol="5" gridrow="6" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186D862705AF9D" box="[213,1382,901,928]" gridrow="7" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186D862302AF9D" box="[213,353,901,928]" gridcol="0" gridrow="7" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEC86D862352AFA2" bold="true" box="[261,305,901,927]" pageId="29" pageNumber="109">VII</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426D862051AF9D" box="[399,562,901,928]" gridcol="1" gridrow="7" pageId="29" pageNumber="109">8–12</td>
|
||
<td id="7699C00AFFC108B8FDB96D86208DAF9D" box="[628,750,901,928]" gridcol="2" gridrow="7" pageId="29" pageNumber="109">6–11</td>
|
||
<td id="7699C00AFFC108B8FCF36D8621ACAF9D" box="[830,975,901,928]" gridcol="3" gridrow="7" pageId="29" pageNumber="109">6–9</td>
|
||
<td id="7699C00AFFC108B8FBCF6D8626FCAF9D" box="[1026,1183,901,928]" gridcol="4" gridrow="7" pageId="29" pageNumber="109">10–14</td>
|
||
<td id="7699C00AFFC108B8FB736D862705AF9D" box="[1214,1382,901,928]" gridcol="5" gridrow="7" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186DB92705AFE8" box="[213,1382,954,981]" gridrow="8" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186DB92302AFE8" box="[213,353,954,981]" gridcol="0" gridrow="8" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFECD6DB92354AFE9" bold="true" box="[256,311,954,980]" pageId="29" pageNumber="109">VIII</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426DB92051AFE8" box="[399,562,954,981]" gridcol="1" gridrow="8" pageId="29" pageNumber="109">9–10</td>
|
||
<td id="7699C00AFFC108B8FDB96DB9208DAFE8" box="[628,750,954,981]" gridcol="2" gridrow="8" pageId="29" pageNumber="109">6–8</td>
|
||
<td id="7699C00AFFC108B8FCF36DB921ACAFE8" box="[830,975,954,981]" gridcol="3" gridrow="8" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FBCF6DB926FCAFE8" box="[1026,1183,954,981]" gridcol="4" gridrow="8" pageId="29" pageNumber="109">10–13</td>
|
||
<td id="7699C00AFFC108B8FB736DB92705AFE8" box="[1214,1382,954,981]" gridcol="5" gridrow="8" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186DEC2705A837" box="[213,1382,1007,1034]" gridrow="9" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186DEC2302A837" box="[213,353,1007,1034]" gridcol="0" gridrow="9" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEC66DEC234FA834" bold="true" box="[267,300,1007,1033]" pageId="29" pageNumber="109">IX</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426DEC2051A837" box="[399,562,1007,1034]" gridcol="1" gridrow="9" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FDB96DEC208DA837" box="[628,750,1007,1034]" gridcol="2" gridrow="9" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FCF36DEC21ACA837" box="[830,975,1007,1034]" gridcol="3" gridrow="9" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FBCF6DEC26FCA837" box="[1026,1183,1007,1034]" gridcol="4" gridrow="9" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FB736DEC2705A837" box="[1214,1382,1007,1034]" gridcol="5" gridrow="9" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
<tr id="3548A976FFC108B8FF186A272705A8B8" box="[213,1382,1060,1157]" gridrow="10" pageId="29" pageNumber="109">
|
||
<th id="7699C00AFFC108B8FF186A272302A8B8" box="[213,353,1060,1157]" gridcol="0" gridrow="10" pageId="29" pageNumber="109">
|
||
<emphasis id="B90C7726FFC1F75AFEDC6A272345A803" bold="true" box="[273,294,1060,1086]" pageId="29" pageNumber="109">X</emphasis>
|
||
</th>
|
||
<td id="7699C00AFFC108B8FE426A272051A8B8" box="[399,562,1060,1157]" gridcol="1" gridrow="10" pageId="29" pageNumber="109">6–8 5–8 3–6</td>
|
||
<td id="7699C00AFFC108B8FDB96A27208DA8B8" box="[628,750,1060,1157]" gridcol="2" gridrow="10" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FCF36A2721ACA8B8" box="[830,975,1060,1157]" gridcol="3" gridrow="10" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FBCF6A2726FCA8B8" box="[1026,1183,1060,1157]" gridcol="4" gridrow="10" pageId="29" pageNumber="109">–</td>
|
||
<td id="7699C00AFFC108B8FB736A272705A8B8" box="[1214,1382,1060,1157]" gridcol="5" gridrow="10" pageId="29" pageNumber="109">–</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC1F75AFF7068D32649AAD7" blockId="29.[189,1066,1744,1771]" box="[189,1066,1744,1771]" pageId="29" pageNumber="109">The additional characters that need to be considered include the following:</paragraph>
|
||
<paragraph id="8BC7AB34FFC1F759FF70691120B8AD7B" blockId="29.[189,1399,1810,2013]" lastBlockId="30.[189,1399,265,749]" lastPageId="30" lastPageNumber="110" pageId="29" pageNumber="109">
|
||
a) The scales of the appendages of all species of
|
||
<taxonomicName id="4C78D0B7FFC1F75AFCCB691121D5AB11" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[774,950,1810,1836]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFCCB691121D5AB11" box="[774,950,1810,1836]" italics="true" pageId="29" pageNumber="109">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
have a similar shape and distribution, where all appendages are covered with setae (not scales) with the exception of the scapus of the antennae and coxae of legs, which are covered with rounded scales similar to those covering the body. This distribution is, however, not exclusive to
|
||
<taxonomicName id="4C78D0B7FFC1F75AFCAF697F2671ABAB" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[866,1042,1916,1942]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC1F75AFCAF697F2671ABAB" box="[866,1042,1916,1942]" italics="true" pageId="29" pageNumber="109">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
since it can also be found in
|
||
<taxonomicName id="4C78D0B7FFC1F75AFF2E699C2086AB84" authority="(Dufour, 1831)" baseAuthorityName="Dufour" baseAuthorityYear="1831" box="[227,741,1951,1977]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="species" species="ciliatum" subGenus="C.">
|
||
<emphasis id="B90C7726FFC1F75AFF2E699C23E7AB84" box="[227,388,1951,1977]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
(
|
||
<emphasis id="B90C7726FFC1F75AFE55699C23D0AB84" box="[408,435,1951,1977]" italics="true" pageId="29" pageNumber="109">C.</emphasis>
|
||
)
|
||
<emphasis id="B90C7726FFC1F75AFE0A699C2044AB84" box="[455,551,1951,1977]" italics="true" pageId="29" pageNumber="109">ciliatum</emphasis>
|
||
(Dufour, 1831)
|
||
</taxonomicName>
|
||
and related Palaearctic species of
|
||
<taxonomicName id="4C78D0B7FFC1F75AFB46699C274FAB84" authorityName="Escherich" authorityYear="1905" box="[1163,1324,1951,1977]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="109" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC1F75AFB46699C274FAB84" box="[1163,1324,1951,1977]" italics="true" pageId="29" pageNumber="109">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC1F75AFAFB699C2714AB84" box="[1334,1399,1951,1977]" pageId="29" pageNumber="109" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
with a trapezoidal tenth urotergite (Molero-Baltanás
|
||
<emphasis id="B90C7726FFC1F75AFC8669C0211DABE1" box="[843,894,1986,2012]" italics="true" pageId="29" pageNumber="109">et al</emphasis>
|
||
. 2010), but different to other species of the genus, such as
|
||
<taxonomicName id="4C78D0B7FFC2F759FE5C6F0A2033AD1E" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[401,592,265,291]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="lineatum" subGenus="C.">
|
||
<emphasis id="B90C7726FFC2F759FE5C6F0A23CFAD1E" box="[401,428,265,291]" italics="true" pageId="30" pageNumber="110">C.</emphasis>
|
||
(
|
||
<emphasis id="B90C7726FFC2F759FE706F0A23BBAD1E" box="[445,472,265,291]" italics="true" pageId="30" pageNumber="110">C.</emphasis>
|
||
)
|
||
<emphasis id="B90C7726FFC2F759FE246F0A2033AD1E" box="[489,592,265,291]" italics="true" pageId="30" pageNumber="110">lineatum</emphasis>
|
||
</taxonomicName>
|
||
. In this species, for example, scales are also present on the femora as described in
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FEB56F2F20B4AD7B" author="Molero Baltanas R. & Gaju Ricart M. & Bach de Roca C." box="[376,727,300,327]" pageId="30" pageNumber="110" pagination="66 - 80" refId="ref21744" refString="Molero Baltanas R., Gaju Ricart M. & Bach de Roca C. 2012. New data for a revision of the genus Ctenolepisma (Zygentoma: Lepismatidae): redescription of Ctenolepisma lineata and new status for Ctenolepisma nicoletii. Annales de la Societe entomologique de France 48 (1 - 2): 66 - 80. https: // doi. org / 10.1080 / 00379271.2012.10697753" type="journal article" year="2012">
|
||
Molero Baltanás
|
||
<emphasis id="B90C7726FFC2F759FD8E6F2E2015AD7B" box="[579,630,300,326]" italics="true" pageId="30" pageNumber="110">et al</emphasis>
|
||
. (2012)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC2F759FF706F4C2746AE00" blockId="30.[189,1399,265,749]" pageId="30" pageNumber="110">
|
||
b) The arrangement of their trichobothrial areas is similar (contiguous on the metanotum, i.e., associated with the two last lateral combs; separated by one lateral comb on the mesonotum, i.e., associated with the last (N) and antepenultimate lateral comb (N−2) and separated by one or two lateral combs on the pronotum (associated with N and N−2 or N−3 combs)). This arrangement is also not exclusive to
|
||
<taxonomicName id="4C78D0B7FFC2F759FECA6FDF23D4ADCB" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[263,439,476,502]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FECA6FDF23D4ADCB" box="[263,439,476,502]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
, but present in several genera of
|
||
<taxonomicName id="4C78D0B7FFC2F759FCA66FDF2626ADCB" authorityName="Mendes" authorityYear="1991" box="[875,1093,476,502]" class="Insecta" family="Lepismatidae" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="subFamily" subFamily="Ctenolepismatinae">Ctenolepismatinae</taxonomicName>
|
||
and in some species of
|
||
<taxonomicName id="4C78D0B7FFC2F759FF2E6FFC23E7AE24" authorityName="Escherich" authorityYear="1905" box="[227,388,511,537]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC2F759FF2E6FFC23E7AE24" box="[227,388,511,537]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC2F759FE466C0323A9AE27" box="[395,458,512,538]" pageId="30" pageNumber="110" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
, but in this latter genus several
|
||
<typeStatus id="54C31596FFC2F759FCF46C032114AE27" box="[825,887,512,538]" pageId="30" pageNumber="110">types</typeStatus>
|
||
of arrangement of trichobothrial areas have been detected, suggesting that it represents a more heterogeneous group than
|
||
<taxonomicName id="4C78D0B7FFC2F759FBA36C21277DAE01" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[1134,1310,546,572]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FBA36C21277DAE01" box="[1134,1310,546,572]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC2F759FF706C45210DAED0" blockId="30.[189,1399,265,749]" pageId="30" pageNumber="110">
|
||
c) Within
|
||
<taxonomicName id="4C78D0B7FFC2F759FEF26C452383AE5D" authorityName="Escherich" authorityYear="1905" box="[319,480,582,608]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B90C7726FFC2F759FEF26C452383AE5D" box="[319,480,582,608]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC2F759FE216C452052AE5D" box="[492,561,582,608]" pageId="30" pageNumber="110" sensu="lato">s. lat.</taxonomicNameLabel>
|
||
, species having transverse bristle-combs on the inner process of the coxite IX belong only to
|
||
<taxonomicName id="4C78D0B7FFC2F759FDDA6C6A20A4AEBE" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[535,711,617,643]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FDDA6C6A20A4AEBE" box="[535,711,617,643]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
, suggesting that only the lineage with median urosternal combs has developed this character (perhaps more than once). Probably the same happens with the occurrence of 2+2 pairs of combs on urotergite I and other apomorphies that are shared only by several species from
|
||
<collectingCountry id="F36FEBA4FFC2F759FE176CD12011AED1" box="[474,626,722,748]" name="South Africa" pageId="30" pageNumber="110">South Africa</collectingCountry>
|
||
, such as
|
||
<taxonomicName id="4C78D0B7FFC2F759FD176CD02104AED1" authorityName="Wygodzinsky" authorityYear="1955" box="[730,871,722,749]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="B90C7726FFC2F759FD176CD02104AED1" box="[730,871,722,749]" italics="true" pageId="30" pageNumber="110">S. arenicola</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC2F759FF706D1721C9A8C5" blockId="30.[187,1400,788,1272]" pageId="30" pageNumber="110">
|
||
According to
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FEA96D172396AF13" author="Irish J." box="[356,501,788,814]" pageId="30" pageNumber="110" pagination="147 - 207" refId="ref20284" refString="Irish J. 1987. Revision of the genus Ctenolepisma Escherich (Thysanura: Lepismatidae) in southern Africa. Cimbebasia (A) 7 (11): 147 - 207." type="journal article" year="1987">Irish (1987)</bibRefCitation>
|
||
, the species
|
||
<taxonomicName id="4C78D0B7FFC2F759FD5D6D1726F4AF13" authority="Mendes, 1982" authorityName="Mendes" authorityYear="1982" box="[656,1175,788,814]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="unipectinatum">
|
||
<emphasis id="B90C7726FFC2F759FD5D6D172187AF13" box="[656,996,788,814]" italics="true" pageId="30" pageNumber="110">Ctenolepisma unipectinatum</emphasis>
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FC236D1726F4AF13" author="Mendes L. F." box="[1006,1175,788,814]" pageId="30" pageNumber="110" pagination="591 - 661" refId="ref20909" refString="Mendes L. F. 1982. Notas e descricoes de Lepismatidae afrotropicais (Zygentoma: Apterygota) II. Generos Ctenolepisma e Namunukulina. Revue zoologique africaine 93 (3): 591 - 661." type="journal article" year="1982">Mendes, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C78D0B7FFC2F759FB186D1723DEAF6C" authority="Escherich, 1910" authorityName="Escherich" authorityYear="1910" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="howa">
|
||
<emphasis id="B90C7726FFC2F759FB186D17229EAF6C" italics="true" pageId="30" pageNumber="110">Ctenolepisma howa</emphasis>
|
||
Escherich, 1910
|
||
</taxonomicName>
|
||
could be included inside
|
||
<taxonomicName id="4C78D0B7FFC2F759FD116D3421EFAF6C" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[732,908,823,849]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FD116D3421EFAF6C" box="[732,908,823,849]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
, as bearing one and two median urosternal combs respectively. Nevertheless, types of both species have been re-examined (loaned by MUHNAC and MFN), concluding that the first one should be excluded from
|
||
<taxonomicName id="4C78D0B7FFC2F759FC146D7D26EAAFA5" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[985,1161,894,920]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FC146D7D26EAAFA5" box="[985,1161,894,920]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
and the second has several additional median combs and not only two, fitting into the present diagnosis of the genus. A slide mounted
|
||
<typeStatus id="54C31596FFC2F759FEE76DC623ECAFE2" box="[298,399,965,991]" pageId="30" pageNumber="110" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C78D0B7FFC2F759FE796DC72167AFE3" authorityName="Mendes" authorityYear="1982" box="[436,772,964,990]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="unipectinatum">
|
||
<emphasis id="B90C7726FFC2F759FE796DC72167AFE3" box="[436,772,964,990]" italics="true" pageId="30" pageNumber="110">Ctenolepisma unipectinatum</emphasis>
|
||
</taxonomicName>
|
||
, loaned by Luis F. Mendes, described as bearing only one small median comb on urosternite II, was checked and the occurrence of scales on the tibiae and narrow truncate or emarginate scales on the femora has been confirmed, so it should be excluded from
|
||
<taxonomicName id="4C78D0B7FFC2F759FF706A2D230EA875" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[189,365,1070,1096]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FF706A2D230EA875" box="[189,365,1070,1096]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
. It is probably related to African species of
|
||
<taxonomicName id="4C78D0B7FFC2F759FCB66A2D267FA875" authorityName="Escherich" authorityYear="1905" box="[891,1052,1070,1096]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC2F759FCB66A2D267FA875" box="[891,1052,1070,1096]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC2F759FBE96A2D2601A875" box="[1060,1122,1070,1096]" pageId="30" pageNumber="110" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
showing similar scales on the legs. We have also examined a specimen collected in the Atlas Mountains in
|
||
<collectingCountry id="F36FEBA4FFC2F759FB606A522779A856" box="[1197,1306,1105,1131]" name="Morocco" pageId="30" pageNumber="110">Morocco</collectingCountry>
|
||
, related to
|
||
<taxonomicName id="4C78D0B7FFC2F759FF136A772332A8B3" authorityName="Wygodzinsky" authorityYear="1941" box="[222,337,1140,1166]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="brauni">
|
||
<emphasis id="B90C7726FFC2F759FF136A772332A8B3" box="[222,337,1140,1166]" italics="true" pageId="30" pageNumber="110">C. brauni</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C78D0B7FFC2F759FEB26A772068A8B3" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[383,523,1140,1166]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="lineatum">
|
||
<emphasis id="B90C7726FFC2F759FEB26A772068A8B3" box="[383,523,1140,1166]" italics="true" pageId="30" pageNumber="110">C. lineatum</emphasis>
|
||
</taxonomicName>
|
||
(probably belonging to an undescribed species), that also has one small median comb on one abdominal sternite, so the occurrence of urosternal combs, at least if they are small and occurring on only one urosternite, is not a diagnostic character to separate
|
||
<taxonomicName id="4C78D0B7FFC2F759FB4E6AB92750A8E9" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[1155,1331,1210,1236]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FB4E6AB92750A8E9" box="[1155,1331,1210,1236]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C78D0B7FFC2F759FF706ADD233DA8C5" authorityName="Escherich" authorityYear="1905" box="[189,350,1246,1272]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FF706ADD233DA8C5" box="[189,350,1246,1272]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
, but a combination of characteristics is necessary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362F8BFFFC2F759FF706B1C2724AB93" pageId="30" pageNumber="110" type="reference_group">
|
||
<paragraph id="8BC7AB34FFC2F759FF706B1C229CA99E" blockId="30.[189,1399,1311,1443]" pageId="30" pageNumber="110">
|
||
Although
|
||
<taxonomicName id="4C78D0B7FFC2F759FEF96B1C215EA907" authority="Wygodzinsky, 1955" authorityName="Wygodzinsky" authorityYear="1955" box="[308,829,1311,1338]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="arenicola">
|
||
<emphasis id="B90C7726FFC2F759FEF96B1C202FA904" box="[308,588,1311,1337]" italics="true" pageId="30" pageNumber="110">Ctenolepisma arenicola</emphasis>
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FD9E6B1C215EA907" author="Wygodzinsky P." box="[595,829,1311,1338]" pageId="30" pageNumber="110" pagination="83 - 190" refId="ref22565" refString="Wygodzinsky P. 1955. Thysanura. In: Hanstrom B., Brink B. & Rudebeck G. (eds) South African Animal Life 2: 83 - 190. AlmQvist & Wiksells Boktryckeri AB, Uppsala." type="book chapter" year="1955">Wygodzinsky, 1955</bibRefCitation>
|
||
</taxonomicName>
|
||
is the
|
||
<typeStatus id="54C31596FFC2F759FC466B2321DDA907" box="[907,958,1312,1338]" pageId="30" pageNumber="110">type</typeStatus>
|
||
species of the genus, since it was the first to be included in the genus
|
||
<taxonomicName id="4C78D0B7FFC2F759FD8B6B412095A961" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[582,758,1346,1372]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FD8B6B412095A961" box="[582,758,1346,1372]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
, it is not the first species to be described that can be assigned to this genus as defined here (sensu
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FD1D6B662130A9BD" author="Irish J." box="[720,851,1381,1408]" pageId="30" pageNumber="110" pagination="147 - 207" refId="ref20284" refString="Irish J. 1987. Revision of the genus Ctenolepisma Escherich (Thysanura: Lepismatidae) in southern Africa. Cimbebasia (A) 7 (11): 147 - 207." type="journal article" year="1987">Irish, 1987</bibRefCitation>
|
||
), which would be
|
||
<taxonomicName id="4C78D0B7FFC2F759FBE16B652298A99E" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Lepismatidae" genus="Lepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="species" species="villosum">
|
||
<emphasis id="B90C7726FFC2F759FBE16B652699A942" box="[1068,1274,1381,1407]" italics="true" pageId="30" pageNumber="110">Lepisma villosum</emphasis>
|
||
Fabricius, 1775
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC2F759FF706BC92724AB93" blockId="30.[188,1401,1482,1966]" pageId="30" pageNumber="110">
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FF706BC9230AA9D9" author="Kaplin V. G." box="[189,361,1482,1508]" pageId="30" pageNumber="110" pagination="34 - 51" refId="ref20666" refString="Kaplin V. G. 1993. On the systematics and phylogeny of the genera Ctenolepisma and Sceletolepisma stat. n. (Thysanura, Lepismatidae). Zoologicheskii Zhurnal 72 (7): 34 - 51. [In Russian. English translation in Entomological Review 73 (8): 124 - 140.]" type="journal article" year="1993">Kaplin (1993)</bibRefCitation>
|
||
, considering urotergal chaetotaxy as the key character for subdividing
|
||
<taxonomicName id="4C78D0B7FFC2F759FB036BC9270CA9D9" authorityName="Escherich" authorityYear="1905" box="[1230,1391,1482,1508]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FB036BC9270CA9D9" box="[1230,1391,1482,1508]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
, raised the subgenus
|
||
<taxonomicName id="4C78D0B7FFC2F759FE7C6BEE2002AA3A" authorityName="Wygodzinsky, 1955 sensu Irish" authorityYear="1987" box="[433,609,1517,1543]" kingdom="Animalia" pageId="30" pageNumber="110" rank="subGenus" subGenus="Sceletolepisma">
|
||
<emphasis id="B90C7726FFC2F759FE7C6BEE2002AA3A" box="[433,609,1517,1543]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
to the generic level but maintaining the criterion of Wygodzinsky (op. cit.); he also created three new subgenera within
|
||
<taxonomicName id="4C78D0B7FFC2F759FC8968132186AA17" authorityName="Escherich" authorityYear="1905" box="[836,997,1552,1578]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FC8968132186AA17" box="[836,997,1552,1578]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C78D0B7FFC2F759FC34681326E6AA17" authorityName="Kaplan" authorityYear="1993" box="[1017,1157,1552,1578]" class="Insecta" family="Lepismatidae" genus="Allolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FC34681326E6AA17" box="[1017,1157,1552,1578]" italics="true" pageId="30" pageNumber="110">Allolepisma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C78D0B7FFC2F759FB5B68132720AA17" authorityName="Kaplan" authorityYear="1993" box="[1174,1347,1552,1578]" class="Insecta" family="Lepismatidae" genus="Escherichisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FB5B68132720AA17" box="[1174,1347,1552,1578]" italics="true" pageId="30" pageNumber="110">Escherichisma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C78D0B7FFC2F759FF706837230AAA73" authorityName="Kaplan" authorityYear="1993" box="[189,361,1588,1614]" class="Insecta" family="Lepismatidae" genus="Silvestrellisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FF706837230AAA73" box="[189,361,1588,1614]" italics="true" pageId="30" pageNumber="110">Silvestrellisma</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EFE9D6C5FFC2F759FE4B68372075AA73" author="Irish J." box="[390,534,1588,1614]" pageId="30" pageNumber="110" pagination="453 - 496" refId="ref20469" refString="Irish J. 1994. Biogeography and systematics of South African Lepismatidae (Thysanura, Insecta). I. The Orange Free State. Navorsinge van die Nasionale Museum Bloemfontein 10 (11): 453 - 496." type="journal article" year="1994">Irish (1994)</bibRefCitation>
|
||
rejected Kaplin’s arrangement as probably not reflecting phylogenetic history, retaining his earlier definitions when describing new species from southern Africa. Here the criterion of Irish is followed. Although a more extensive revision is required, Kaplin’s criteria make neither phylogenetic nor biogeographic sense. The character separating
|
||
<taxonomicName id="4C78D0B7FFC2F759FBB4689E233DAAE7" authority="sensu Wygodzinsky" authorityName="sensu Wygodzinsky" authorityYear="1955" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FBB4689E274AAA8A" box="[1145,1321,1693,1719]" italics="true" pageId="30" pageNumber="110">Sceletolepisma</emphasis>
|
||
sensu Wygodzinsky
|
||
</taxonomicName>
|
||
from the remaining
|
||
<taxonomicName id="4C78D0B7FFC2F759FDAD68C32162AAE7" authorityName="Escherich" authorityYear="1905" box="[608,769,1728,1754]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FDAD68C32162AAE7" box="[608,769,1728,1754]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
taxa, i.e., those bearing 2+2 combs on urotergite I instead of 1+1 combs, appears in several South African species that are closely related with those that do not have this character. However, species of
|
||
<taxonomicName id="4C78D0B7FFC2F759FD37690421F8AB1C" authorityName="Escherich" authorityYear="1905" box="[762,923,1799,1825]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B90C7726FFC2F759FD37690421F8AB1C" box="[762,923,1799,1825]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC2F759FC6E69042180AB1C" box="[931,995,1799,1825]" pageId="30" pageNumber="110" sensu="lato">s. lat.</taxonomicNameLabel>
|
||
bearing median urosternal combs can be considered as a more uniform group sharing the characters indicated in the present diagnosis, and species lacking several median combs on abdominal sternites (in the present arrangement, the only representatives of the genus
|
||
<taxonomicName id="4C78D0B7FFC2F759FDC8697320C5ABB7" authorityName="Escherich" authorityYear="1905" box="[517,678,1904,1930]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="30" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC2F759FDC8697320C5ABB7" box="[517,678,1904,1930]" italics="true" pageId="30" pageNumber="110">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
) form a more heterogeneous group which requires a thorough revision, but not using the urotergal chaetotaxy as the unique character for splitting into subgroups.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362F8BFFFC3F758FF706F0B2085AD86" pageId="31" pageNumber="111" type="distribution">
|
||
<paragraph id="8BC7AB34FFC3F758FF706F0B2335AD1F" blockId="31.[189,342,264,290]" box="[189,342,264,290]" pageId="31" pageNumber="111">
|
||
<heading id="D08F1C58FFC3F758FF706F0B2335AD1F" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="31" pageNumber="111" reason="3">
|
||
<emphasis id="B90C7726FFC3F758FF706F0B2335AD1F" bold="true" box="[189,342,264,290]" pageId="31" pageNumber="111">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7AB34FFC3F758FF706F342085AD86" blockId="31.[189,1400,311,443]" pageId="31" pageNumber="111">
|
||
Representatives of
|
||
<taxonomicName id="4C78D0B7FFC3F758FE6F6F342031AD6C" authorityName="sensu Wygodzinsky" authorityYear="1955" box="[418,594,311,337]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="31" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC3F758FE6F6F342031AD6C" box="[418,594,311,337]" italics="true" pageId="31" pageNumber="111">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
are absent from America, while species belonging to
|
||
<taxonomicName id="4C78D0B7FFC3F758FB1A6F34271BAD6C" authorityName="Escherich" authorityYear="1905" box="[1239,1400,311,337]" class="Insecta" family="Lepismatidae" genus="Ctenolepisma" kingdom="Animalia" order="Zygentoma" pageId="31" pageNumber="111" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B90C7726FFC3F758FB1A6F34271BAD6C" box="[1239,1400,311,337]" italics="true" pageId="31" pageNumber="111">Ctenolepisma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23FCA5DFFC3F758FF706F58229AAD48" box="[189,249,347,373]" pageId="31" pageNumber="111" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
are widespread on both sides of the Atlantic Ocean. This suggests that
|
||
<taxonomicName id="4C78D0B7FFC3F758FBEF6F5926B1AD49" authorityName="sensu Wygodzinsky" authorityYear="1955" box="[1058,1234,346,372]" class="Insecta" family="Lepismatidae" genus="Sceletolepisma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="31" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90C7726FFC3F758FBEF6F5926B1AD49" box="[1058,1234,346,372]" italics="true" pageId="31" pageNumber="111">Sceletolepisma</emphasis>
|
||
</taxonomicName>
|
||
first appeared after the opening of this ocean, but we cannot discard a previous origin and a later extinction event in South America after its separation from Africa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |