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<document id="6A482800F6D0782E35E4F1F7DFA775E6" ID-DOI="10.11646/zootaxa.3904.4.1" ID-GBIF-Dataset="a1c49726-9afa-4be5-85ef-9d0caabac1fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="239899" ID-ZooBank="38457A27-A15B-4E87-BE72-9A3AB007CAA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460890290884" checkinUser="plazi" docAuthor="Yoshitomi, Hiroyuki" docDate="2015" docId="03AC6D78C810215149B2EE6C3651F926" docLanguage="en" docName="zt03904p500.pdf" docOrigin="Zootaxa 3904 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ypsiloncyphon philippinensis Yoshitomi, 2015, sp. n." docType="treatment" docUuid="A5E803AD-270D-4945-AD73-F76939724489" docUuidSource="ZooBank" docVersion="9" lastPageNumber="469" masterDocId="FF951500C80121434925E96E356EFFBD" masterDocTitle="New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae)" masterLastPageNumber="500" masterPageNumber="451" pageNumber="468" updateTime="1698359547105" updateUser="plazi">
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<mods:title id="764F402A167DDB439C1EAF48C245AFFC">New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae)</mods:title>
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<mods:name id="EA78D6F96D56089BCD6EE2935F926896" type="personal">
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<mods:roleTerm id="86751CA573F081B41921BAD1F635B248">Author</mods:roleTerm>
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<mods:namePart id="4FA8C50BF0EC631CA61F2E36E9A05F53">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:title id="E5CB47F73BBD494AFA53E083153C2F9C">Zootaxa</mods:title>
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<mods:part id="2A87CA922851E3043D8D6ADD144B3EBF">
<mods:date id="64BD5A26C73752F024A257947F65EA8A">2015</mods:date>
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<mods:number id="05EA5B5EFFF861D16E4B3AEADE7B55CF">3904</mods:number>
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<treatment id="03AC6D78C810215149B2EE6C3651F926" ID-DOI="http://doi.org/10.5281/zenodo.6111873" ID-GBIF-Taxon="119586910" ID-Zenodo-Dep="6111873" LSID="urn:lsid:zoobank.org:act:A5E803AD-270D-4945-AD73-F76939724489" httpUri="http://treatment.plazi.org/id/03AC6D78C810215149B2EE6C3651F926" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468">
<subSubSection id="C31F8FE5C810215249B2EE6C342EF883" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="8BBADC6EC810215249B2EE6C372CF8A1" blockId="17.[151,578,1794,1854]" box="[151,578,1794,1820]" pageId="17" pageNumber="468">
<heading id="D0F26B02C810215249B2EE6C372CF8A1" bold="true" box="[151,578,1794,1820]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<emphasis id="B971007CC810215249B2EE6C372CF8A1" bold="true" box="[151,578,1794,1820]" pageId="17" pageNumber="468">
<taxonomicName id="4C05A7EDC810215249B2EE6C3496F8A1" box="[151,504,1794,1820]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="B971007CC810215249B2EE6C3496F8A1" bold="true" box="[151,504,1794,1820]" italics="true" pageId="17" pageNumber="468">Ypsiloncyphon philippinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C81021524B25EE6C372CF8A1" box="[512,578,1794,1820]" pageId="17" pageNumber="468" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBADC6EC810215249B2EE4B342EF883" blockId="17.[151,578,1794,1854]" box="[151,320,1829,1854]" pageId="17" pageNumber="468">
(
<figureCitation id="133EC0EBC810215249BAEE4B3595F883" box="[159,251,1829,1854]" captionStart="FIGURE 13" captionStartId="17.[151,250,1610,1632]" captionTargetBox="[155,1420,202,1578]" captionTargetId="figure@17.[151,1430,193,1583]" captionTargetPageId="17" captionText="FIGURE 13. Habitus of Ypsiloncyphon spp., holotype, male (except for F). A) Y. guamensis sp. n., B) Y. philippinensis sp. n., C) Y. panayensis sp. n., D) Y. chlorizanoides sp. n., E) Y. bicolor sp. n., F) Y. bicolor sp. n., paratype, female, G) Y. sumatranus sp. n., H) Y. acer sp. n., I) Y. nokrekensis sp. n., J) Y. boukali sp. n., K) Y. javanicus sp. n., L) Y. nepalensis sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239912/files/figure.png" pageId="17" pageNumber="468">Figs. 13</figureCitation>
B, 15)
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C810215249B2EE033779F873" pageId="17" pageNumber="468" type="materials_examined">
<paragraph id="8BBADC6EC810215249B2EE033779F873" blockId="17.[151,1436,1901,1998]" pageId="17" pageNumber="468">
<emphasis id="B971007CC810215249B2EE033428F83B" bold="true" box="[151,326,1901,1926]" pageId="17" pageNumber="468">
<typeStatus id="54BE62CCC810215249B2EE0335BCF83B" box="[151,210,1901,1926]" pageId="17" pageNumber="468">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BE62CCC8102152486AEE0334D9F83B" box="[335,439,1901,1926]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED1444ABC810215248EDEE003776F83B" box="[456,536,1902,1926]" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="17" pageNumber="468">EUMJ</collectionCode>
): male, “
<collectingCountry id="F3129CFEC81021524B59EE00364AF83B" box="[636,804,1902,1926]" name="Philippines" pageId="17" pageNumber="468">PHILIPPINES</collectingCountry>
Mt. Polis,
<quantity id="4CFD718BC81021524A81EE00369CF83B" box="[932,1010,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="17" pageNumber="468" unit="m" value="1900.0">1900m</quantity>
Ifugao Prov., Luzon June, 4, 5, 1977 M. Sato leg.”, genit. s. no. HY 803.
<typeStatus id="54BE62CCC81021524B18EEFC37C4F817" box="[573,682,1938,1962]" pageId="17" pageNumber="468" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="ED1444ABC81021524B9EEEFF3665F814" box="[699,779,1937,1961]" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="17" pageNumber="468">EUMJ</collectionCode>
):
<specimenCount id="9D0317E7C81021524A05EEFF3610F817" box="[800,894,1937,1962]" pageId="17" pageNumber="468" type="male">2 Males</specimenCount>
, “
<collectingCountry id="F3129CFEC81021524AB3EEFF3153F814" box="[918,1085,1937,1961]" name="Philippines" pageId="17" pageNumber="468">PHILIPPINES</collectingCountry>
Kinakin,
<quantity id="4CFD718BC81021524D94EEFF306EF817" box="[1201,1280,1937,1962]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="17" pageNumber="468" unit="m" value="1050.0">1050m</quantity>
Ifugao Prov., Luzon June, 3, 1977 M. Sato leg.”
</paragraph>
</subSubSection>
<caption id="DF7A8CE6C813215149B2EB8B3474FCA7" httpUri="https://zenodo.org/record/239913/files/figure.png" pageId="18" pageNumber="469" targetBox="[279,1307,203,715]" targetPageId="18">
<paragraph id="8BBADC6EC813215149B2EB8B3474FCA7" blockId="18.[151,1436,741,794]" pageId="18" pageNumber="469">
<emphasis id="B971007CC813215149B2EB8B3477FD47" bold="true" box="[151,281,741,763]" pageId="18" pageNumber="469">FIGURE 14</emphasis>
.
<taxonomicName id="4C05A7EDC81321514803EB8B3745FD46" box="[294,555,741,763]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="guamensis" status="sp. nov.">
<emphasis id="B971007CC81321514803EB8B3745FD46" box="[294,555,741,763]" italics="true" pageId="18" pageNumber="469">Ypsiloncyphon guamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C81321514B14EB883705FD46" box="[561,619,742,763]" pageId="18" pageNumber="469" rank="species">
<emphasis id="B971007CC81321514B14EB883705FD46" bold="true" box="[561,619,742,763]" pageId="18" pageNumber="469">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, male. A) sternites VVII, B) sternite VIII, C) sternite IX, D) penis in ventral view.
</paragraph>
</caption>
<subSubSection id="C31F8FE5C813215149E2EA28305CF9B7" pageId="18" pageNumber="469" type="description">
<paragraph id="8BBADC6EC813215149E2EA283406FC1A" blockId="18.[151,1437,838,1691]" pageId="18" pageNumber="469">
<emphasis id="B971007CC813215149E2EA283437FCE2" bold="true" box="[199,345,838,863]" pageId="18" pageNumber="469">Description.</emphasis>
Male. Body (
<figureCitation id="133EC0EBC81321514B25EA28373BFCE2" box="[512,597,838,863]" captionStart="FIGURE 13" captionStartId="17.[151,250,1610,1632]" captionTargetBox="[155,1420,202,1578]" captionTargetId="figure@17.[151,1430,193,1583]" captionTargetPageId="17" captionText="FIGURE 13. Habitus of Ypsiloncyphon spp., holotype, male (except for F). A) Y. guamensis sp. n., B) Y. philippinensis sp. n., C) Y. panayensis sp. n., D) Y. chlorizanoides sp. n., E) Y. bicolor sp. n., F) Y. bicolor sp. n., paratype, female, G) Y. sumatranus sp. n., H) Y. acer sp. n., I) Y. nokrekensis sp. n., J) Y. boukali sp. n., K) Y. javanicus sp. n., L) Y. nepalensis sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/239912/files/figure.png" pageId="18" pageNumber="469">Fig. 13</figureCitation>
B) oval, convex dorsally, shining, closely covered with yellowish-white setae throughout. Coloration of body blackish-brown; antennae, mouthparts, and legs yellowish-brown, except for infuscated femora.
</paragraph>
<paragraph id="8BBADC6EC813215149E2EADC377EFBC2" blockId="18.[151,1437,838,1691]" pageId="18" pageNumber="469">Head moderate in size, dorsal surface almost flat, granulated, emarginate in front margin. Eyes moderate in size, prominent; the distance between eyes about 2.0 times as long as the maximum diameter of an eye. Antennae relatively short. Pronotum granulated as on head; front margin almost straight in dorsal aspect; lateral margin gently arcuate; antero- and postero-lateral corners obtuse; PW/PL 2.34. Scutellar shield subtriangular, finely punctate. Elytra oval, widest at the middle, finely and densely punctate; apices rectangular; EL/EW 1.25; EL/PL 4.69; EW/PW 1.60; TL/EW 1.52.</paragraph>
<paragraph id="8BBADC6EC813215149E2EDE431CDFA7F" blockId="18.[151,1437,838,1691]" pageId="18" pageNumber="469">
Caudal margin of sternite VII (
<figureCitation id="133EC0EBC81321514B1CEDE437E0FB1E" box="[569,654,1162,1187]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
A) arcuate. Tergite VIII (
<figureCitation id="133EC0EBC81321514A9FEDE4317EFB1E" box="[954,1040,1162,1187]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
B) lightly sclerotized, trapezoidal, bearing short setae and spines along posterior margin, with a pair of long and slender apodemes protruding from near antero-lateral corners; sternite VIII (
<figureCitation id="133EC0EBC81321514B44EDBC37DEFB56" box="[609,688,1234,1259]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
C) well sclerotized, Y-shaped, relatively long; tergite IX (
<figureCitation id="133EC0EBC81321514C0AEDBC3013FB56" box="[1327,1405,1234,1259]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
D) lightly sclerotized in lateral part, consisting of a pair of hemitergites; sternite IX (
<figureCitation id="133EC0EBC81321514D61ED9831F8FAB2" box="[1092,1174,1270,1295]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
E) well sclerotized, Ushaped, slender, pointed at apices, bearing minute serrations in apical 1/5. Tegmen (
<figureCitation id="133EC0EBC81321514D12EC7431E8FA8E" box="[1079,1158,1306,1331]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
F) long, well sclerotized, Y-shaped, bifid in apical 1/3; proximal part slightly expanded antero-laterally, anterior margin concave; parameres covered with short spines in apical part, pointed at apex. Penis (
<figureCitation id="133EC0EBC81321514A46EC0C36DFFAC6" box="[867,945,1378,1403]" captionStart="FIGURE 15" captionStartId="19.[151,250,1632,1654]" captionTargetBox="[305,1282,199,1605]" captionTargetId="figure@19.[297,1289,193,1611]" captionTargetPageId="19" captionText="FIGURE 15. Ypsiloncyphon philippinensis sp. n., holotype, male. A) sternites V VII, B) tergite VIII, C) sternite VIII, D) tergite IX, E) sternite IX, F) tegmen, G) penis in ventral view, H) penis in lateral view." httpUri="https://zenodo.org/record/239914/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
G, H) well sclerotized, shorter than tegmen; parameroids obtuse at apex, finely punctate, strongly expanded laterally in proximal 1/3; pala short, subparallelsided, arcuate in basal margin; trigonium gently curved ventrally, somewhat pointed at apex.
</paragraph>
<paragraph id="8BBADC6EC813215149E2ECA034F3FA5A" blockId="18.[151,1437,838,1691]" box="[199,413,1486,1511]" pageId="18" pageNumber="469">Female. Unknown.</paragraph>
<paragraph id="8BBADC6EC813215149E2EC9C305CF9B7" blockId="18.[151,1437,838,1691]" box="[199,1330,1522,1547]" pageId="18" pageNumber="469">
<emphasis id="B971007CC813215149E2EC9C3413F9B6" bold="true" box="[199,381,1522,1547]" pageId="18" pageNumber="469">Measurements.</emphasis>
Male (n = 1): TL
<quantity id="4CFD718BC81321514B6CEC9D37DEF9B7" box="[585,688,1523,1547]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="18" pageNumber="469" unit="mm" value="1.82">1.82 mm</quantity>
; PW
<quantity id="4CFD718BC81321514BC9EC9C363DF9B7" box="[748,851,1522,1547]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="469" unit="mm" value="0.75">0.75 mm</quantity>
; PL
<quantity id="4CFD718BC81321514AA3EC9D3683F9B7" box="[902,1005,1523,1547]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="18" pageNumber="469" unit="mm" value="0.32">0.32 mm</quantity>
; EW
<quantity id="4CFD718BC81321514D0EEC9D31FCF9B7" box="[1067,1170,1523,1547]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="469" unit="mm" value="1.2">1.20 mm</quantity>
; EL
<quantity id="4CFD718BC81321514DE2EC9C3040F9B7" box="[1223,1326,1522,1547]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="469" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C813215149E2EF783719F992" box="[199,631,1558,1583]" pageId="18" pageNumber="469" type="distribution">
<paragraph id="8BBADC6EC813215149E2EF783719F992" blockId="18.[151,1437,838,1691]" box="[199,631,1558,1583]" pageId="18" pageNumber="469">
<emphasis id="B971007CC813215149E2EF78340EF992" bold="true" box="[199,352,1558,1583]" pageId="18" pageNumber="469">Distribution.</emphasis>
<collectingCountry id="F3129CFEC81321514842EF78348BF992" box="[359,485,1558,1583]" name="Philippines" pageId="18" pageNumber="469">Philippines</collectingCountry>
(Luzon Isl.).
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C813215149E2EF54318DF9CA" pageId="18" pageNumber="469" type="discussion">
<paragraph id="8BBADC6EC813215149E2EF54318DF9CA" blockId="18.[151,1437,838,1691]" pageId="18" pageNumber="469">
<emphasis id="B971007CC813215149E2EF543455F9EE" bold="true" box="[199,315,1594,1619]" pageId="18" pageNumber="469">Remarks.</emphasis>
This species is related to
<taxonomicName id="4C05A7EDC81321514B47EF5536ECF9EF" box="[610,898,1595,1619]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="guamensis" status="sp. nov.">
<emphasis id="B971007CC81321514B47EF5536ECF9EF" box="[610,898,1595,1619]" italics="true" pageId="18" pageNumber="469">Ypsiloncyphon guamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C81321514AAFEF5436A5F9EE" box="[906,971,1594,1619]" pageId="18" pageNumber="469" rank="species">
<emphasis id="B971007CC81321514AAFEF5436A5F9EE" bold="true" box="[906,971,1594,1619]" pageId="18" pageNumber="469">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C05A7EDC81321514D26EF5531F4F9EF" box="[1027,1178,1595,1618]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="panayensis" status="sp. nov.">
<emphasis id="B971007CC81321514D26EF553161F9EF" box="[1027,1039,1595,1618]" italics="true" pageId="18" pageNumber="469">Y</emphasis>
.
<emphasis id="B971007CC81321514D38EF5531F4F9EF" box="[1053,1178,1595,1618]" italics="true" pageId="18" pageNumber="469">panayensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A242BD07C81321514D87EF54318CF9EE" box="[1186,1250,1594,1619]" pageId="18" pageNumber="469" rank="species">
<emphasis id="B971007CC81321514D87EF54318CF9EE" bold="true" box="[1186,1250,1594,1619]" pageId="18" pageNumber="469">sp. n.</emphasis>
</taxonomicNameLabel>
in the shapes of tergite XI, tegmen and penis. It differs from them by lacking keels in the apical part of sternite IX.
</paragraph>
</subSubSection>
<subSubSection id="C31F8FE5C813215149E2EFEC3651F926" box="[199,831,1666,1691]" pageId="18" pageNumber="469" type="etymology">
<paragraph id="8BBADC6EC813215149E2EFEC3651F926" blockId="18.[151,1437,838,1691]" box="[199,831,1666,1691]" pageId="18" pageNumber="469">
<emphasis id="B971007CC813215149E2EFEC3420F926" bold="true" box="[199,334,1666,1691]" pageId="18" pageNumber="469">Etymology.</emphasis>
The species is named after the
<typeStatus id="54BE62CCC81321514B8BEFED37B0F926" box="[686,734,1667,1691]" pageId="18" pageNumber="469">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>