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<document id="F441487545CD3005CFA6F1927ECF4C5A" ID-CLB-Dataset="29103" ID-DOI="10.11646/zootaxa.4455.2.5" ID-GBIF-Dataset="568a5364-52f2-4feb-b08b-87eb85d4cf77" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457283" ID-ZooBank="8CB311F1-431C-4861-956E-3200329E4F0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539261916473" checkinUser="plazi" docAuthor="Nada, B. &amp; Ballantyne, L. A." docDate="2018" docId="039887E6FFF27A0D9ECB2040A29B65EB" docLanguage="en" docName="zootaxa.4455.2.5.pdf" docOrigin="Zootaxa 4455 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygoluciola" docType="key" docVersion="7" lastPageNumber="348" masterDocId="FFA1FF9EFFF67A089E5C2657A7076769" masterDocTitle="A new species of Pygoluciola Wittmer with unusual abdominal configuration, from lowland dipterocarp forest in Peninsular Malaysia (Coleoptera: Lampyridae: Luciolinae)" masterLastPageNumber="362" masterPageNumber="343" pageNumber="347" updateTime="1698575593942" updateUser="ExternalLinkService">
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<mods:title id="1BD9F0512C2E705B1ADC8BEFA8D493F2">A new species of Pygoluciola Wittmer with unusual abdominal configuration, from lowland dipterocarp forest in Peninsular Malaysia (Coleoptera: Lampyridae: Luciolinae)</mods:title>
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<mods:namePart id="22DEC64496B3238B9EFB77899EECC490">Nada, B.</mods:namePart>
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<mods:namePart id="D95A584D58434C8038F98E54A2D22FC7">Ballantyne, L. A.</mods:namePart>
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<heading id="D0C6819CFFF27A0C9ECB2040A5926158" bold="true" box="[151,661,1559,1585]" fontSize="11" level="1" pageId="4" pageNumber="347" reason="1">
<emphasis id="B945EAE2FFF27A0C9ECB2040A5926158" bold="true" box="[151,661,1559,1585]" pageId="4" pageNumber="347">
Key to species of
<taxonomicName id="4C314D73FFF27A0C9F312040A6FB6158" ID-CoL="63QVY" box="[365,508,1559,1585]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAE2FFF27A0C9F312040A6FB6158" bold="true" box="[365,508,1559,1585]" italics="true" pageId="4" pageNumber="347">Pygoluciola</emphasis>
</taxonomicName>
using males
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36F0FFF27A0C9ECB206EA5B7613B" blockId="4.[151,688,1559,1618]" box="[151,688,1593,1618]" pageId="4" pageNumber="347">Modified and expanded from Ballantyne (2008)</paragraph>
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<keyLead id="30C095C5FFF27A0C9ECB20D6A29261DD" pageId="4" pageNumber="347">
1. Median posterior margin of
<date id="FF8F1030FFF27A0C9FB220D6A50C61FF" box="[494,523,1665,1686]" pageId="4" pageNumber="347">V7</date>
and T8 prolonged and often curving,
<date id="FF8F1030FFF27A0C9D2A20D6A49461FF" box="[886,915,1665,1686]" pageId="4" pageNumber="347">V7</date>
dorsally and T8 ventrally and partially engulfing each other............................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF27A0C9ECB20EBA2926187" blockId="4.[151,1436,1665,2009]" pageId="4" pageNumber="347">
<keyLead id="30C095C5FFF27A0C9ECB20EBA2926187" pageId="4" pageNumber="347">
- Median posterior margin of
<date id="FF8F1030FFF27A0C9FAC20EBA50A61B8" box="[496,525,1724,1745]" pageId="4" pageNumber="347">V7</date>
and T8 not prolonged; if
<date id="FF8F1030FFF27A0C9D5920EBA42561B8" box="[773,802,1724,1745]" pageId="4" pageNumber="347">V7</date>
prolonged then not curving strongly dorsally and not engulfed by T8; posterior margin of T
<date id="FF8F1030FFF27A0C9F8A208EA5176187" box="[470,528,1753,1774]" pageId="4" pageNumber="347">8 may</date>
be slightly downturned but not narrowed........................................ 7
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</paragraph>
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<keyLead id="30C095C5FFF27A0C9ECB20A0A29B6065" box="[151,1436,1783,1804]" pageId="4" pageNumber="347">
2. All tibiae curved; lateral margins of elytra tapering posteriorly (
<bibRefCitation id="EFA04B01FFF27A0C9D1E20A0A39C6065" box="[834,1179,1783,1804]" pageId="4" pageNumber="347" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin, 2006, fig. 6</bibRefCitation>
)................... 3
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF27A0C9ECB2143A29C6040" blockId="4.[151,1436,1665,2009]" box="[151,1435,1812,1833]" pageId="4" pageNumber="347">
<keyLead id="30C095C5FFF27A0C9ECB2143A29C6040" box="[151,1435,1812,1833]" pageId="4" pageNumber="347">- No tibiae curved; lateral margins of elytra usually sub-parallel-sided............................................. 4</keyLead>
</paragraph>
</keyStep>
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<keyLead id="30C095C5FFF27A0C9ECB2166A29B600D" pageId="4" pageNumber="347">
3. Apex of median posterior projection of tergite 8 no wider than rest and rounded, not emarginate; lateral margins of pronotum slightly sinuate (
<bibRefCitation id="EFA04B01FFF27A0C9F232118A419600D" box="[383,798,1871,1892]" pageId="4" pageNumber="347" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin, 2006 figs 1, 19, 21</bibRefCitation>
)..................................
<taxonomicName id="4C314D73FFF27A0C9A932118A29B600D" ID-CoL="4QP8S" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[1231,1436,1871,1892]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="guigliae">
<emphasis id="B945EAE2FFF27A0C9A932118A21A600D" box="[1231,1309,1871,1892]" italics="true" pageId="4" pageNumber="347">guigliae</emphasis>
(Ballantyne)
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</keyLead>
</paragraph>
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<keyLead id="30C095C5FFF27A0C9ECB213BA29B60F7" pageId="4" pageNumber="347">
- Apex of median posterior projection of tergite 8 wider than rest; median emarginate; lateral margins of pronotum not slightly sinuate (
<bibRefCitation id="EFA04B01FFF27A0C9F6D21DEA5AB60F7" box="[305,684,1929,1950]" pageId="4" pageNumber="347" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin, 2006 figs 4, 25</bibRefCitation>
).................................................
<taxonomicName id="4C314D73FFF27A0C9B5F21DEA29B60F7" ID-CoL="4QP8X" authority="Wittmer" authorityName="Wittmer" box="[1283,1436,1929,1950]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="stylifer">
<emphasis id="B945EAE2FFF27A0C9B5F21DEA24160F7" box="[1283,1350,1929,1950]" italics="true" pageId="4" pageNumber="347">stylifer</emphasis>
Wittmer
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
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<keyLead id="30C095C5FFF27A0C9ECB21F0A29B60D5" box="[151,1436,1959,1980]" pageId="4" pageNumber="347">
4. Median posterior projection of abdominal
<date id="FF8F1030FFF27A0C9C2B21F0A59C60D5" box="[631,667,1959,1980]" pageId="4" pageNumber="347">V 7</date>
bifurcate at apex (
<bibRefCitation id="EFA04B01FFF27A0C9D1121F0A3D360D5" box="[845,1236,1959,1980]" pageId="4" pageNumber="347" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin, 2006 figs 18, 26</bibRefCitation>
).............. 5
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF27A0C9ECB2193A29C60B0" blockId="4.[151,1436,1665,2009]" box="[151,1435,1988,2009]" pageId="4" pageNumber="347">
<keyLead id="30C095C5FFF27A0C9ECB2193A29C60B0" box="[151,1435,1988,2009]" pageId="4" pageNumber="347">
- Median posterior projection of abdominal
<date id="FF8F1030FFF27A0C9C2B2193A59C60B0" box="[631,667,1988,2009]" pageId="4" pageNumber="347">V 7</date>
not bifurcate at apex.............................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB26C0A29B666D" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB26C0A29B67A3" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB26C0A29B67A3" pageId="5" pageNumber="348">
5 Apex of median posterior projection of abdominal
<date id="FF8F1030FFF37A0D9C9A26C0A5ED67C5" box="[710,746,151,172]" pageId="5" pageNumber="348">V 7</date>
deeply emarginate, laterally ensheathing the downturned apex of tergite 8 and projecting laterally beside it (
<bibRefCitation id="EFA04B01FFF37A0D9C6826E2A44967A3" box="[564,846,181,202]" pageId="5" pageNumber="348" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin, 2006</bibRefCitation>
figs 18, 26;
<bibRefCitation id="EFA04B01FFF37A0D9D9B26E2A32867A3" box="[967,1071,181,202]" pageId="5" pageNumber="348" refId="ref9031" refString="Chey, V. K. (2008) Fireflies of Sepilok. Sepilok Bulletin, 9, 3 ‾ 11." type="journal article">Chey 2008</bibRefCitation>
fig 2)........
<emphasis id="B945EAE2FFF37A0D9A9526E2A21A67A3" box="[1225,1309,181,202]" italics="true" pageId="5" pageNumber="348">wittmeri</emphasis>
(Ballantyne)
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB2685A29B666D" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2685A29B666D" pageId="5" pageNumber="348">
- Apex of median posterior projection of abdominal
<date id="FF8F1030FFF37A0D9C9B2685A5ED678E" box="[711,746,210,231]" pageId="5" pageNumber="348">V 7</date>
shallowly emarginate, not laterally ensheathing the downturned apex of tergite 8 and not projecting beside it (
<bibRefCitation id="EFA04B01FFF37A0D9C1B26B8A4C5666D" box="[583,962,239,260]" pageId="5" pageNumber="348" refId="ref8435" refString="Ballantyne, L. A. &amp; Lambkin, C. (2001) A new firefly, Luciola (Pygoluciola) kinabalua sp. nov. (Coleoptera: Lampyridae), from Malaysia, with observations on a possible copulation clamp. Raffles Bulletin of Zoology, 49 (2), 363 - 377." type="journal article">Ballantyne &amp; Lambkin, 2001 figs 5, 10</bibRefCitation>
)...................
<taxonomicName id="4C314D73FFF37A0D9AE226B8A29B666D" ID-CoL="3WCSR" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne &amp; Lambkin" baseAuthorityYear="2001" box="[1214,1436,239,260]" class="Insecta" family="Lampyridae" genus="Luciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="kinabalua">
<emphasis id="B945EAE2FFF37A0D9AE226B8A219666D" box="[1214,1310,239,260]" italics="true" pageId="5" pageNumber="348">kinabalua</emphasis>
(Ballantyne)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB275AA29B66FE" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB275AA29B6656" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB275AA29B6656" pageId="5" pageNumber="348">
6 MPP of
<date id="FF8F1030FFF37A0D9F73275AA64B664B" box="[303,332,269,290]" pageId="5" pageNumber="348">V7</date>
elongate slender, ventral surface shallowly depressed along its length, and bearing on its median dorsal surface two slender teeth; posterior apex of MPP not differentiated; lateral margins of T8 downturned................
<taxonomicName id="4C314D73FFF37A0D9AA3277DA29B6656" ID-CoL="4QP8W" authority="Ballantyne" authorityName="Ballantyne" authorityYear="2008" box="[1279,1436,298,319]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="B945EAE2FFF37A0D9AA3277DA22A6656" box="[1279,1325,298,319]" italics="true" pageId="5" pageNumber="348">satoi</emphasis>
Ballantyne
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB2710A29B66FE" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2710A29B66FE" pageId="5" pageNumber="348">
- MPP of
<date id="FF8F1030FFF37A0D9F692710A6556635" box="[309,338,327,348]" pageId="5" pageNumber="348">V7</date>
shorter, about as wide as long, ventral surface not shallowly depressed along its length, and not bearing dorsal teeth; posterior apex (face) of MPP differentiated and shallowly depressed; lateral margins T 8 not downturned.................................................................................................
<emphasis id="B945EAE2FFF37A0D9ABB27D5A24566FE" box="[1255,1346,386,407]" italics="true" pageId="5" pageNumber="348">hamulata</emphasis>
(Olivier)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB27C8A29C6565" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB27C8A29B6699" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB27C8A29B6699" pageId="5" pageNumber="348">
7. MPP of
<date id="FF8F1030FFF37A0D9F6927C8A65566DD" box="[309,338,415,436]" pageId="5" pageNumber="348">V7</date>
defined, longer than wide and either horizontal or inclined slightly upwards along its length and grooved along its ventral surface (Figs 3, 6, 10, 12, 14); T8 much narrower than preceding tergite and apically pointed (
<figureCitation id="130A2A75FFF37A0D9AF227EAA3F366BB" box="[1198,1268,445,466]" captionStart="FIGURES 1016" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 1016. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral wings arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="5" pageNumber="348">Fig. 15</figureCitation>
)............................................ …………………………………………………………………..
<taxonomicName id="4C314D73FFF37A0D9AE0278DA24E6686" ID-CoL="8N8PY" authority="Nada" authorityName="Nada &amp; Ballantyne" authorityYear="2018" box="[1212,1353,474,495]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="dunguna" status="sp. nov.">
<emphasis id="B945EAE2FFF37A0D9AE0278DA2176686" box="[1212,1296,474,495]" italics="true" pageId="5" pageNumber="348">dunguna</emphasis>
Nada
</taxonomicName>
<taxonomicNameLabel id="A2765799FFF37A0D9B13278DA29B6699" box="[1359,1436,474,496]" pageId="5" pageNumber="348" rank="species">
<emphasis id="B945EAE2FFF37A0D9B13278DA29B6699" bold="true" box="[1359,1436,474,496]" pageId="5" pageNumber="348">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB27A0A29C6565" blockId="5.[151,1436,151,642]" box="[151,1435,503,524]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB27A0A29C6565" box="[151,1435,503,524]" pageId="5" pageNumber="348">
- MPP not defined, posterior margin of
<date id="FF8F1030FFF37A0D9C1227A0A56C6565" box="[590,619,503,524]" pageId="5" pageNumber="348">V7</date>
rounded; T8 not much narrower than preceding tergite and not apically pointed.. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E55FFF37A0D9ECB2442A29B65EB" pageId="5" pageNumber="348">
<paragraph id="8B8E36F0FFF37A0D9ECB2442A29C652E" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2442A29C652E" pageId="5" pageNumber="348">
8. Australian species; pronotum with wide median black band reaching to anterior and posterior margins; eyes large, subcontiguous ventrally, with well defined posterolateral eye emargination; elytra brown with pale lateral margins..
<taxonomicName id="4C314D73FFF37A0D9A882465A29C652E" ID-CoL="4QP8R" authority="(Blackburn)" authorityName="Blackburn" baseAuthorityName="Blackburn" baseAuthorityYear="1897" box="[1236,1435,562,583]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="cowleyi">
<emphasis id="B945EAE2FFF37A0D9A882465A226652E" box="[1236,1313,562,583]" italics="true" pageId="5" pageNumber="348">cowleyi</emphasis>
(Blackburn)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB2418A29B65EB" blockId="5.[151,1436,151,642]" pageId="5" pageNumber="348">
<keyLead id="30C095C5FFF37A0D9ECB2418A29B65EB" pageId="5" pageNumber="348">
- Chinese species; pronotum pale whitish cream with restricted median brown spots; eyes not subcontiguous ventrally nor with well defined eye emargination; elytra black...............................................
<taxonomicName id="4C314D73FFF37A0D9AEE243AA29B65EB" ID-CoL="4QP8V" authority="Fu et Ballantyne" authorityName="Fu et Ballantyne" box="[1202,1436,621,642]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="qingyu">
<emphasis id="B945EAE2FFF37A0D9AEE243AA3F165EB" box="[1202,1270,621,642]" italics="true" pageId="5" pageNumber="348">qingyu</emphasis>
Fu
<emphasis id="B945EAE2FFF37A0D9B40243AA22A65EB" box="[1308,1325,621,642]" italics="true" pageId="5" pageNumber="348">et</emphasis>
Ballantyne
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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