treatments-xml/data/59/A5/C1/59A5C1EA93DE51C2B10622BC39E70E41.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.96.50821" ID-GBIF-Dataset="fb568d02-c3d7-48b9-9729-2f85ea601db7" ID-GBIF-Taxon="164363160" ID-Pensoft-Pub="1860-0743-1-217" ID-Pensoft-UUID="930A6280AF085CA9B6F8AC6CE0E1C120" ID-ZooBank="0B80FF1B5ED7430BA471A5DE02E8E6D3" LSID="urn:lsid:zoobank.org:act:BE521B67-6769-4EF5-B3C9-56EB43BF2DFE" ModsDocID="1860-0743-1-217" checkinTime="1590646559532" checkinUser="pensoft" docAuthor="Gasiorek, Piotr, Voncina, Katarzyna, Degma, Peter &amp; Michalczyk, Lukasz" docDate="2020" docId="59A5C1EA93DE51C2B10622BC39E70E41" docLanguage="en" docName="ZoosystEvol 96(1): 217-236" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.50821" docTitle="Bryodelphax australasiaticus Gasiorek, Voncina, Degma &amp; Michalczyk 2020, sp. nov." docType="treatment" docUuid="BE521B67-6769-4EF5-B3C9-56EB43BF2DFE" docUuidSource="ZooBank" docVersion="4" id="930A6280AF085CA9B6F8AC6CE0E1C120" lastPageNumber="217" masterDocId="930A6280AF085CA9B6F8AC6CE0E1C120" masterDocTitle="Small is beautiful: the first phylogenetic analysis of Bryodelphax Thulin, 1928 (Heterotardigrada, Echiniscidae)" masterLastPageNumber="236" masterPageNumber="217" pageId="0" pageNumber="217" updateTime="1634708808286" updateUser="ExternalLinkService">
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<mods:title>Small is beautiful: the first phylogenetic analysis of Bryodelphax Thulin, 1928 (Heterotardigrada, Echiniscidae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gasiorek, Piotr</mods:namePart>
<mods:affiliation>Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2814-8117</mods:nameIdentifier>
<mods:nameIdentifier type="email">piotr.lukas.gasiorek@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Voncina, Katarzyna</mods:namePart>
<mods:affiliation>Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Degma, Peter</mods:namePart>
<mods:affiliation>Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6 / B 1, 84215, Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Michalczyk, Lukasz</mods:namePart>
<mods:affiliation>Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2912-4870</mods:nameIdentifier>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>96</mods:number>
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<mods:number>1</mods:number>
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<mods:start>217</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zse.96.50821</mods:url>
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<treatment ID-GBIF-Taxon="164363160" LSID="urn:lsid:zoobank.org:act:BE521B67-6769-4EF5-B3C9-56EB43BF2DFE" httpUri="http://treatment.plazi.org/id/59A5C1EA93DE51C2B10622BC39E70E41" lastPageNumber="217" pageId="0" pageNumber="217">
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<taxonomicName LSID="59A5C1EA-93DE-51C2-B106-22BC39E70E41" authority="Gąsiorek, Vončina, Degma &amp; Michalczyk" authorityName="Gasiorek, Voncina, Degma &amp; Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax australasiaticus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="australasiaticus" status="sp. nov.">
Bryodelphax australasiaticus 
<normalizedToken originalValue="Gąsiorek">Gasiorek</normalizedToken>
, 
<normalizedToken originalValue="Vončina">Voncina</normalizedToken>
, Degma &amp; Michalczyk
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="217">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">, 4</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Schematic arrangement of the ventral plates in all members of the weglarskae group (species are arranged in order of the increasing reduction of the ventral armature). Known cases of ontogenetic variability and sexual dimorphism are depicted. Following species taken from Kaczmarek et al. (2012): B. aaseae, B. iohannis, B. parvuspolaris, B. sinensis and B. weglarskae; Gasiorek et al. (2017 a): B. maculatus; Gasiorek &amp; Degma (2018): B. instabilis." figureDoi="10.3897/zse.96.50821.figure12" httpUri="https://binary.pensoft.net/fig/413959" pageId="0" pageNumber="217">, 12</figureCitation>
, 
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements [in µm] of selected morphological structures of mature females of Bryodelphax australasiaticus sp. nov. mounted in Hoyer's medium (N - number of specimens / structures measured, Range refers to the smallest and the largest structure amongst all measured specimens; SD - standard deviation; sp - the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage)." pageId="0" pageNumber="217">Table 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="reference_group">
<paragraph pageId="0" pageNumber="217">
<taxonomicName family="Echiniscidae" genus="B." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="B. australis" order="Echiniscoidea" pageId="0" pageNumber="217" rank="species" species="australis">B. australis</taxonomicName>
sp. n. in 
<bibRefCitation author="Claxton, SK" journalOrPublisher="PhD Thesis, Macquarie University, Sydney, Australia" pageId="0" pageNumber="217" refId="B9" refString="Claxton, SK, 2004. The taxonomy and distribution of Australian terrestrial tardigrades. PhD Thesis, Macquarie University, Sydney, Australia" title="The taxonomy and distribution of Australian terrestrial tardigrades." year="2004">Claxton (2004)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="type material">
<paragraph pageId="0" pageNumber="217">
<taxonomicName genus="Locus" lsidName="Locus typicus" pageId="0" pageNumber="217" rank="species" species="typicus">
<emphasis italics="true" pageId="0" pageNumber="217">Locus typicus</emphasis>
</taxonomicName>
and type material.
</paragraph>
<paragraph pageId="0" pageNumber="217">
<geoCoordinate degrees="5" direction="north" minutes="27" orientation="latitude" precision="15" seconds="05" value="5.451389">5°27'05&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="100" direction="east" minutes="11" orientation="longitude" precision="15" seconds="00" value="100.183334">100°11'00&quot;E</geoCoordinate>
, 4 m a.s.l.; Pantai Keracut, Pulau Penang, Malaysia. Holotype (adult female; slide MY.240.01) and seven paratypes (5 females, 2 juveniles; slides MY.240.01-04) deposited in the Institute of Zoology and Biomedical Research, Jagiellonian University; two paratypes (2 females; slide MY.240.05) deposited in the Department of Zoology, Comenius University in Bratislava; one paratype (1 female; slide MY.240.06) deposited in the Natural History Museum of Denmark, University of Copenhagen; two paratypes (2 females; slide MY.240.07) deposited in the Department of Animal Biology, University of Catania; three paratypes (2 females, one larva; slides MY.241.02, MY.242.02) deposited in the Raffles Museum of Biodiversity Research, National University of Singapore.
</paragraph>
<caption ID-Table-UUID="B2C2E7D407EDCA696B2AE82A58CD5CEC" httpUri="http://table.plazi.org/id/B2C2E7D407EDCA696B2AE82A58CD5CEC" pageId="0" pageNumber="217" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="217">
<emphasis bold="true" pageId="0" pageNumber="217">Table 3.</emphasis>
Measurements [in 
<normalizedToken originalValue="µm">µm</normalizedToken>
] of selected morphological structures of mature females of 
<taxonomicName authorityName="Gasiorek, Voncina, Degma &amp; Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax australasiaticus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="australasiaticus">
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax australasiaticus</emphasis>
</taxonomicName>
sp. nov. mounted in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium (
<emphasis bold="true" pageId="0" pageNumber="217">N</emphasis>
- number of specimens/structures measured, 
<emphasis bold="true" pageId="0" pageNumber="217">Range</emphasis>
refers to the smallest and the largest structure amongst all measured specimens; 
<emphasis bold="true" pageId="0" pageNumber="217">SD</emphasis>
- standard deviation; 
<emphasis bold="true" italics="true" pageId="0" pageNumber="217">sp</emphasis>
- the ratio of the length of a given structure to the length of the scapular plate expressed as a percentage).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="217">
<table pageId="0" pageNumber="217">
<tr pageId="0" pageNumber="217">
<th colspan="1" pageId="0" pageNumber="217" rowspan="2">Character</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="2">N</th>
<th colspan="6" pageId="0" pageNumber="217" rowspan="1">Range</th>
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">Mean</th>
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">SD</th>
<th colspan="2" pageId="0" pageNumber="217" rowspan="1">Holotype</th>
</tr>
<tr pageId="0" pageNumber="217">
<th colspan="3" pageId="0" pageNumber="217" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="3" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<normalizedToken originalValue="µm">µm</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">sp</emphasis>
</th>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Body length</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">91</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">119</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">584</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">737</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">108</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">667</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">41</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">111</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">666</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Scapular plate length</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15.2</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">17.9</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">16.2</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">16.6</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Head appendages lengths</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
Cirrus 
<emphasis italics="true" pageId="0" pageNumber="217">internus</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.7</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">6.6</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">30.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">42.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.7</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">35.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">4.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">33.9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Cephalic papilla</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">13.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">21.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">18.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">2.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">17.6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
Cirrus 
<emphasis italics="true" pageId="0" pageNumber="217">externus</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">7.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">47.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">58.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">52.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.5</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">3.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">54.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Clava</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">10.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">20.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">14.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">2.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">11.2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
Cirrus 
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">23.2</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">28.7</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">144.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">183.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">25.8</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">160.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">11.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">24.6</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">147.7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
Cirrus 
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
/Body length ratio
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">21%</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">26%</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">24%</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">1%</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">22%</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="14" pageId="0" pageNumber="217" rowspan="1">Claw heights</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw I</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">13</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.5</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.7</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">26.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">33.9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">31.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">2.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">30.4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw II</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.5</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">26.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">33.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.0</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">30.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">2.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.5</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">33.3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw III</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.5</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">26.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">34.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.8</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">29.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.3</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">2.1</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.0</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">30.2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="217">
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">Claw IV</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">13</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">4.9</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">6.2</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">30.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">38.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">33.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">0.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">2.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="217" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="217">32.2</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="etymology">
<paragraph pageId="0" pageNumber="217">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="217">The name refers to the currently identified geographic range of the new species that encompasses Asia and Australia. Adjective in the nominative singular.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="adults">
<paragraph pageId="0" pageNumber="217">Adults.</paragraph>
<paragraph pageId="0" pageNumber="217">
Body pink, pearly opalescent; eyes absent or not visible after mounting in 
<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
medium. Primary and secondary clavae small and conical. Cirri 
<emphasis italics="true" pageId="0" pageNumber="217">interni</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="217">externi</emphasis>
with poorly-developed cirrophores. Cirri 
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
of typical length for 
<taxonomicName authorityName="Thulin" authorityYear="1928" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax</emphasis>
</taxonomicName>
, i.e. reaching around 25% of the total body length. All dorsal plates with barely-discernible intra-cuticular pillars (better visible under a 1000
<normalizedToken originalValue="×">x</normalizedToken>
magnification), the centro-posterior portion of the caudal (terminal) plate has evident, largest pillars (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
). Dark epicuticular granules absent (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3</figureCitation>
; 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">4A, B</figureCitation>
), but lateral margins of all dorsal plates and internal margins of facets constituting the scapular plate distinctly thicker and, consequently, darker (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
). Pores large and easily detectable (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A, B</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">4A, B</figureCitation>
). Pores distributed unevenly, with the largest number present on the antero-central portion of the scapular plate (17-40 pores/100 
<normalizedToken originalValue="μm">μm</normalizedToken>
2, x̅ = 29, N = 16, Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">4A</figureCitation>
) and the central portion of the caudal plate (7-43 pores/100 
<normalizedToken originalValue="μm">μm</normalizedToken>
2, x̅ = 31, N = 16, Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">4B</figureCitation>
); other plates more variable in terms of pore density, which is always lower than in the aforementioned elements of the armour (0-26 pores/100 
<normalizedToken originalValue="μm">μm</normalizedToken>
2, x̅ = 14, N = 16, Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
). Scapular plate gently faceted by transverse cuticular extensions (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">4A</figureCitation>
), with deep sutures separating lateral portions from the central faceted part, extending from the base of cirrophore 
<emphasis italics="true" pageId="0" pageNumber="217">A</emphasis>
to the posterior margin of the plate (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
). Paired plates divided into two roughly equal anterior and posterior parts by a transverse stripe (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A, B</figureCitation>
). Caudal (terminal) plate with poorly developed sutures, not visible under PCM (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
), but present and visible under SEM (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A, B</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Detailed morphology of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - frontal part of the body, faceting of the scapular plate; B - caudal part of the body; C - claws III in close-up. Scale bars: 10 µm." figureDoi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217">4B</figureCitation>
). Median plate 1 subdivided into anterior narrow portion with dark posterior edge and posterior pentagonal portion with transverse suture (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A</figureCitation>
). Median plate 2 is the largest amongst median plates, with well-developed anterior pentagonal portion and poorly sclerotised posterior portion (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A</figureCitation>
). Median plate 3 with only the anterior portion fully developed, the posterior portion triangular and rounded (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3A</figureCitation>
). Supplementary lateral platelets present at the level of median plates (three pairs of platelets on each body side: a pair between the scapular plate and the first pair of the segmental plates, a pair between the paired plates and a pair between the second pair of segmental plates and the caudal plate; Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="217">
<emphasis bold="true" pageId="0" pageNumber="217">Figure 2.</emphasis>
Habitus of adult female of 
<taxonomicName authorityName="Gasiorek, Voncina, Degma &amp; Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax australasiaticus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="australasiaticus">
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax australasiaticus</emphasis>
</taxonomicName>
sp. nov. (holotype, PCM): 
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
- dorsolateral view; 
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
- ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="217">
<emphasis bold="true" pageId="0" pageNumber="217">Figure 3.</emphasis>
Habitus of adult females of 
<taxonomicName authorityName="Gasiorek, Voncina, Degma &amp; Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax australasiaticus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="australasiaticus">
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax australasiaticus</emphasis>
</taxonomicName>
sp. nov. (paratypes, SEM): 
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
- dorsal view; 
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
- lateral view; 
<emphasis bold="true" pageId="0" pageNumber="217">C</emphasis>
- ventrolateral view; 
<emphasis bold="true" pageId="0" pageNumber="217">D</emphasis>
- ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="217">
Venter with seven rows of faint, greyish plates (VII:4-4-2-4-2-2-1), of which two plates of the first, subcephalic row are located in a more ventrolateral position (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2B</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3C, D</figureCitation>
, 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Details of cuticular sculpturing of Bryodelphax nigripunctatus sp. nov.: A - microscope focused on the epicuticular granules (allotype, NCM); B - microscope focused on the intracuticular pillars (allotype, NCM); C - frontal part of the body (allotype, PCM); D - frontal part of the body (holotype, PCM). Scale bars in µm." figureDoi="10.3897/zse.96.50821.figure10" httpUri="https://binary.pensoft.net/fig/413957" pageId="0" pageNumber="217">10</figureCitation>
). Under SEM, only the central subcephalic and genital plates are visible as true cuticular thickenings, whereas other plates are visible only as darker areas on the cuticular surface (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3C, D</figureCitation>
). Leg papillae undetectable under LCM (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2</figureCitation>
), but papillae IV visible under SEM (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3B, C</figureCitation>
). Both pulvini and pedal plates present, the former developed as thin rectangular stripes in the proximal leg portions (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3C</figureCitation>
) and the latter - as large swellings in the central leg portions (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of adult females of Bryodelphax australasiaticus sp. nov. (paratypes, SEM): A - dorsal view; B - lateral view; C - ventrolateral view; D - ventral view. Roman numerals signify ventral plate rows and arrowheads indicate papillae on fourth pair of legs. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure3" httpUri="https://binary.pensoft.net/fig/413950" pageId="0" pageNumber="217">3C</figureCitation>
). Pedal plate IV toothless, but with a distinct dark margin (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2A</figureCitation>
). External claws spurless, but internal ones with minute spurs positioned close to the claw bases (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of adult female of Bryodelphax australasiaticus sp. nov. (holotype, PCM): A - dorsolateral view; B - ventrolateral view (insert with the claws III, arrowhead indicates spur on internal claw). Roman numerals signify ventral plate rows. Scale bars: 50 µm." figureDoi="10.3897/zse.96.50821.figure2" httpUri="https://binary.pensoft.net/fig/413949" pageId="0" pageNumber="217">2B</figureCitation>
[insert], 4C).
</paragraph>
<caption doi="10.3897/zse.96.50821.figure4" httpUri="https://binary.pensoft.net/fig/413951" pageId="0" pageNumber="217" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="217">
<emphasis bold="true" pageId="0" pageNumber="217">Figure 4.</emphasis>
Detailed morphology of 
<taxonomicName authorityName="Gasiorek, Voncina, Degma &amp; Michalczyk" authorityYear="2020" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax australasiaticus" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="australasiaticus">
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax australasiaticus</emphasis>
</taxonomicName>
sp. nov. (paratypes, SEM): 
<emphasis bold="true" pageId="0" pageNumber="217">A</emphasis>
- frontal part of the body, faceting of the scapular plate; 
<emphasis bold="true" pageId="0" pageNumber="217">B</emphasis>
- caudal part of the body; 
<emphasis bold="true" pageId="0" pageNumber="217">C</emphasis>
- claws III in close-up. Scale bars: 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="juveniles">
<paragraph pageId="0" pageNumber="217">Juveniles.</paragraph>
<paragraph pageId="0" pageNumber="217">
Body 73-101 
<normalizedToken originalValue="μm">μm</normalizedToken>
long in the two found juveniles. Dorsal and ventral plates developed similarly to adults. Scapular plate 12-16 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Claws 3.7-4.8 
<normalizedToken originalValue="μm">μm</normalizedToken>
long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="larvae">
<paragraph pageId="0" pageNumber="217">Larvae.</paragraph>
<paragraph pageId="0" pageNumber="217">
Body 80 
<normalizedToken originalValue="μm">μm</normalizedToken>
long in a single found two-clawed specimen. Dorsal and ventral plates developed similarly to adults. Scapular plate 12.7 
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Claws 4.0-4.4 
<normalizedToken originalValue="μm">μm</normalizedToken>
long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="eggs">
<paragraph pageId="0" pageNumber="217">Eggs.</paragraph>
<paragraph pageId="0" pageNumber="217">Up to one egg in exuvia was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="dna sequences">
<paragraph pageId="0" pageNumber="217">DNA sequences.</paragraph>
<paragraph pageId="0" pageNumber="217">Single 18S rRNA haplotype (MT333468), two 28S rRNA haplotypes (MT333460-1) and single ITS-1 haplotype (MT333477).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="phenotypic differential diagnosis">
<paragraph pageId="0" pageNumber="217">Phenotypic differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="217">
Within the 
<taxonomicName baseAuthorityName="Pilato" baseAuthorityYear="1972" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="weglarskae" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis italics="true" pageId="0" pageNumber="217">weglarskae</emphasis>
</taxonomicName>
group, only 
<taxonomicName genus="B." lsidName="B. decoratus" pageId="0" pageNumber="217" rank="species" species="decoratus">
<emphasis italics="true" pageId="0" pageNumber="217">B. decoratus</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName genus="B." lsidName="B. sinensis" pageId="0" pageNumber="217" rank="species" species="sinensis">
<emphasis italics="true" pageId="0" pageNumber="217">B. sinensis</emphasis>
</taxonomicName>
and 
<taxonomicName genus="B." lsidName="B. instabilis" pageId="0" pageNumber="217" rank="species" species="instabilis">
<emphasis italics="true" pageId="0" pageNumber="217">B. instabilis</emphasis>
</taxonomicName>
have seven plate rows, but 
<taxonomicName genus="B." lsidName="B. olszanowskii" pageId="0" pageNumber="217" rank="species" species="olszanowskii">
<emphasis italics="true" pageId="0" pageNumber="217">B. olszanowskii</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.1007/s00300-017-2190-4" author="Kaczmarek, L" journalOrPublisher="Polar Biology" pageId="0" pageNumber="217" pagination="283 - 301" refId="B40" refString="Kaczmarek, L, Parnikoza, I, Gawlak, M, Esefeld, J, Peter, H-U, Kozeretska, I, Roszkowska, M, 2018. Tardigrades from Larus dominicanus Lichtenstein, 1823 nests on the Argentine Islands (maritime Antarctic). Polar Biology 41: 283 - 301, DOI: https://doi.org/10.1007/s00300-017-2190-4" title="Tardigrades from Larus dominicanus Lichtenstein, 1823 nests on the Argentine Islands (maritime Antarctic)." url="https://doi.org/10.1007/s00300-017-2190-4" volume="41" year="2018">Kaczmarek et al., 2018</bibRefCitation>
exhibits peculiar ventrolateral plates in the subcephalic row and that is why this taxon is also taken into consideration in the differential diagnosis. Adult females of 
<taxonomicName genus="B." lsidName="B. australasiaticus" pageId="0" pageNumber="217" rank="species" species="australasiaticus">
<emphasis italics="true" pageId="0" pageNumber="217">B. australasiaticus</emphasis>
</taxonomicName>
sp. nov. are differentiated from:
</paragraph>
<paragraph pageId="0" pageNumber="217">B. decoratus sp. nov., by the ventral plate formula (VII:4-4-2-4-2-2-1 in the new species vs. VII:4-2- 2-4-2-2-1 in B. decoratus sp. nov.) and by dorsal plate sculpturing (merged epicuticular ridges surrounding the borders of all dorsal plates in the new species vs. large, dark epicuticular granules in B. decoratus sp. nov.);</paragraph>
<paragraph pageId="0" pageNumber="217">B. sinensis, known from Caucasus and China (the record from Spitsbergen (Dastych 1985) represents B. parvuspolaris), by the ventral plate formula (VII:4-4-2-4-2-2-1 in the new species vs. VII:2-2-2-2-2-2-1 in B. sinensis) and the caudal (terminal) plate faceting (invisible under LCM in the new species vs. four facets formed by the raised plate areas between two longitudinal and one transversal sutures in B. sinensis);</paragraph>
<paragraph pageId="0" pageNumber="217">B. instabilis, currently considered endemic to the Tatras and northern Slovakia, by the ventral plate formula (VII:4-4-2-4-2-2-1 in the new species vs. VII/IX:(2)-(1)-2/4-2-2/4-2-2-2-1 in B. instabilis), the presence of dentate collar IV (absent in the new species vs. present in B. instabilis), the detectability of papilla IV under LCM (undetectable in the new species vs. detectable in B. instabilis) and by the reproductive mode (parthenogenesis in the new species vs. dioecy in B. instabilis);</paragraph>
<paragraph pageId="0" pageNumber="217">B. olszanowskii, reported from the Antarctic, by the ventral plate formula (VII:4-4-2-4-2-2-1 in the new species vs. VIII:4-1-1-2-2-2-2-2 in B. olszanowskii), the presence of dark epicuticular granules (ab sent in the new species vs. present and accumulated on ventral plates in B. olszanowskii) and the detectability of papilla IV under LM (absent in the new species vs. present in B. olszanowskii).</paragraph>
<paragraph pageId="0" pageNumber="217">
Genotypic differential diagnosis: 
<emphasis italics="true" pageId="0" pageNumber="217">p</emphasis>
-distances between the new species and the remaining 
<taxonomicName authorityName="Thulin" authorityYear="1928" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryodelphax" order="Echiniscoidea" pageId="0" pageNumber="217" phylum="Tardigrada" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="217">Bryodelphax</emphasis>
</taxonomicName>
spp., for which DNA sequences are available, were as follows:
</paragraph>
<paragraph pageId="0" pageNumber="217">18S rRNA: from 0.3% (B. decoratus sp. nov., MT333469-70) to 3.4% (B. cf. parvulus, HM193371);</paragraph>
<paragraph pageId="0" pageNumber="217">28S rRNA from 0.5% (Bryodelphax sp. nov. from Celebes, MT333467) to 9.3% (B. cf. parvulus, MT333466);</paragraph>
<paragraph pageId="0" pageNumber="217">ITS-1: from 2.6% (B. arenosus, MT346599-600) to 3.3% (B. decoratus sp. nov., MT333478).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="217" type="remarks">
<paragraph pageId="0" pageNumber="217">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="217">
Two ventrolateral plates were not drawn in 
<bibRefCitation author="Claxton, SK" journalOrPublisher="PhD Thesis, Macquarie University, Sydney, Australia" pageId="0" pageNumber="217" refId="B9" refString="Claxton, SK, 2004. The taxonomy and distribution of Australian terrestrial tardigrades. PhD Thesis, Macquarie University, Sydney, Australia" title="The taxonomy and distribution of Australian terrestrial tardigrades." year="2004">Claxton (2004)</bibRefCitation>
, which is an unpublished PhD dissertation, thus the species described therein are not valid. However, having ascertained that these structures exist in specimens from Australia, the compared populations from both continents appeared identical in terms of morphology.
</paragraph>
</subSubSection>
</treatment>
</document>