448 lines
54 KiB
XML
448 lines
54 KiB
XML
<document id="9E7D68778633B691F5950A79E58758E0" ID-CLB-Dataset="5815" ID-DOI="10.11646/zootaxa.3779.3.2" ID-GBIF-Dataset="2055f347-d478-4d50-aad0-7e78c4b3b9c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4910403" ID-ZooBank="23584B50-EFAC-4BCE-A20E-9B09C22529B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623118476769" checkinUser="felipe" docAuthor="Deein, Gridsada, Tangjitjaroen, Weerapongse & Page, Lawrence M." docDate="2014" docId="038987E7FFC8DB2AC0EF5FEB5DE7A50D" docLanguage="en" docName="zt03779p352.pdf" docOrigin="Zootaxa 3779 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidocephalus nanensis Deein & Tangjitjaroen & Page 2014, n. sp." docType="treatment" docVersion="8" lastPageNumber="347" masterDocId="FFB0FF9FFFCBDB2CC0785C1C5D08A042" masterDocTitle="A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae)" masterLastPageNumber="352" masterPageNumber="341" pageNumber="344" updateTime="1699003940845" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="EE10449740F08A03D15F6D4A6DA46122" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2E853B12D75BF6BFE3FEA916D7BADFFA">
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<mods:title id="D967C45E2ED90A544C9E5F6ADF2B1BDA">A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9923AB1B88C9D2613D4D1E528C99EF81" type="personal">
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<mods:roleTerm id="C48F731493917233C60B1B52D718980C">Author</mods:roleTerm>
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<mods:namePart id="0E4D30AFF06461594D4301A901B5D13C">Deein, Gridsada</mods:namePart>
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<mods:affiliation id="ADF0BF276140B15A85D49EBD2928A4D9">Inland Fisheries Research and Development Bureau, Department of Fisheries, NIFI Building, Kasetklang, Chatuchak, Bangkok 10900 Thailand. E-mail: dgridsada @ hotmail. com</mods:affiliation>
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</mods:name>
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<mods:name id="C639DBF64816504D9F5019B103518846" type="personal">
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<mods:roleTerm id="79E7BEEF2A435632295DF346B0950DBB">Author</mods:roleTerm>
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<mods:namePart id="A6D3040694B81D36A36AAF829EC498F0">Tangjitjaroen, Weerapongse</mods:namePart>
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<mods:affiliation id="99AF0946BDB14ED9581A7D24F18BD032">Faculty of Veterinary Medicine, Chiang Mai University, Chiang Mai, 50100, Thailand. E-mail: weerapongse @ yahoo. com</mods:affiliation>
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<mods:name id="D0BE4E4AA02725A43FB27E13AB7F3516" type="personal">
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<mods:roleTerm id="6F0D270A01435315CAC44FD16FE28023">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7444D0B2AEDE79DFE2E9A10AE638A297">Page, Lawrence M.</mods:namePart>
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<mods:affiliation id="8DD60BE3449F02D535D1984216521882">Florida Museum of Natural History, University of Florida, Gainesville, FL 32611 - 7800 USA. E-mail: lpage 1 @ ufl. edu</mods:affiliation>
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<mods:typeOfResource id="297139724C3FEA266DFE8A940088D6EB">text</mods:typeOfResource>
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<mods:title id="D2D52DB65CB23A1CC275667964BDD265">Zootaxa</mods:title>
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<mods:part id="D2A3A85AD4C04161295316700C867F53">
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<mods:date id="E8D515555EA35A4B535EFCA792D0F993">2014</mods:date>
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<mods:number id="A68B472E6E122001FC375493629B107E">2014-03-17</mods:number>
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<mods:number id="B0964CD197F70E48D10B4BCEE451D5AA">3779</mods:number>
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<mods:start id="A8A495E3F40FCE37FB2836FF40AA5B73">341</mods:start>
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<mods:identifier id="E0EDAF9CCB3614D42EA0B9E48F90F315" type="Zenodo-Dep">4910403</mods:identifier>
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<treatment id="038987E7FFC8DB2AC0EF5FEB5DE7A50D" ID-DOI="http://doi.org/10.5281/zenodo.4910382" ID-GBIF-Taxon="183237227" ID-Zenodo-Dep="4910382" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D" httpUri="http://treatment.plazi.org/id/038987E7FFC8DB2AC0EF5FEB5DE7A50D" lastPageId="6" lastPageNumber="347" pageId="3" pageNumber="344">
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<subSubSection id="C33A657AFFC8DB2FC0EF5FEB5C54A471" pageId="3" pageNumber="344" type="nomenclature">
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<paragraph id="8B9F36F1FFC8DB2FC0EF5FEB5F11A450" blockId="3.[151,537,1015,1111]" box="[151,537,1015,1042]" pageId="3" pageNumber="344">
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<heading id="D0D7819DFFC8DB2FC0EF5FEB5F11A450" bold="true" box="[151,537,1015,1042]" fontSize="11" level="1" pageId="3" pageNumber="344" reason="1">
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<emphasis id="B954EAE3FFC8DB2FC0EF5FEB5F11A450" bold="true" box="[151,537,1015,1042]" pageId="3" pageNumber="344">
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<taxonomicName id="4C204D72FFC8DB2FC0EF5FEB5CCFA453" ID-CoL="6PB8V" authority="Deein & Tangjitjaroen & Page, 2014" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[151,455,1015,1041]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="nanensis" status="sp. nov.">
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<emphasis id="B954EAE3FFC8DB2FC0EF5FEB5CCFA453" bold="true" box="[151,455,1015,1041]" italics="true" pageId="3" pageNumber="344">Lepidocephalus nanensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2675798FFC8DB2FC1AF5FE45F11A450" box="[471,537,1016,1042]" pageId="3" pageNumber="344" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9F36F1FFC8DB2FC0EF58065C54A471" blockId="3.[151,537,1015,1111]" box="[151,348,1050,1075]" pageId="3" pageNumber="344">Thai Spirit Loach</paragraph>
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</subSubSection>
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<subSubSection id="C33A657AFFC8DB2FC0EF58225DF7A415" box="[151,255,1086,1111]" pageId="3" pageNumber="344" type="description">
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<paragraph id="8B9F36F1FFC8DB2FC0EF58225DF7A415" blockId="3.[151,537,1015,1111]" box="[151,255,1086,1111]" pageId="3" pageNumber="344">
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(
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<figureCitation id="131B2A74FFC8DB2FC0E758225DFEA415" box="[159,246,1086,1111]" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="3" pageNumber="344">Fig. 1A</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A657AFFC8DB2FC0EF589A5837A645" pageId="3" pageNumber="344" type="materials_examined">
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<paragraph id="8B9F36F1FFC8DB2FC0EF589A5CC4A481" blockId="3.[151,1436,1158,2011]" pageId="3" pageNumber="344">
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<materialsCitation id="3B483CACFFC8DB2FC0EF589A5CC4A481" ID-GBIF-Occurrence="3313289305" collectingDate="2008-11" collectionCode="NIFI" collectorName="G. Deein." country="Thailand" county="Khlong Chompu" location="Chao Phraya" municipality="Mae Nam" pageId="3" pageNumber="344" specimenCode="NIFI 4388" specimenCount="1" stateProvince="Phitsanulok" typeStatus="holotype">
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<emphasis id="B954EAE3FFC8DB2FC0EF589A5C03A4DD" bold="true" box="[151,267,1158,1183]" inLineHeading="true" pageId="3" pageNumber="344" reason="1">
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<typeStatus id="549B8853FFC8DB2FC0EF589A5C0EA4DD" box="[151,262,1158,1183]" pageId="3" pageNumber="344" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DB869E8AFFC8DB2FC16C589B5C86A4DD" box="[276,398,1158,1183]" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="3" pageNumber="344">NIFI 4388</specimenCode>
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, 72.0 mm SL;
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<collectingCountry id="F3377661FFC8DB2FC242589A5FA8A4DD" box="[570,672,1158,1183]" name="Thailand" pageId="3" pageNumber="344">Thailand</collectingCountry>
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,
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<collectingRegion id="49E4F813FFC8DB2FC2D6589A5E70A4DC" box="[686,888,1158,1183]" country="Thailand" name="Phitsanulok" pageId="3" pageNumber="344">Phitsanulok Prov.</collectingRegion>
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,
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<collectingCounty id="62FE4E7DFFC8DB2FC3FD589A594DA4DD" box="[901,1093,1158,1183]" pageId="3" pageNumber="344">Khlong Chompu</collectingCounty>
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,
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<collectingMunicipality id="6BFBAC8BFFC8DB2FC42A589B59CBA4DC" box="[1106,1219,1159,1182]" pageId="3" pageNumber="344">Mae Nam</collectingMunicipality>
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<collectingRegion id="49E4F813FFC8DB2FC4B4589B59F6A4DC" box="[1228,1278,1159,1182]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
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,
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<location id="8EFF602AFFC8DB2FC572589A5894A4DD" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC572589A5894A4DD" box="[1290,1436,1158,1183]" country="Thailand" county="Khlong Chompu" municipality="Mae Nam" name="Chao Phraya" pageId="3" pageNumber="344" stateProvince="Phitsanulok">Chao Phraya</location>
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basin,
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<date id="FF9E1031FFC8DB2FC09858B75C5FA481" box="[224,343,1194,1219]" pageId="3" pageNumber="344" value="2008-11">
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<collectingDate id="EFDAE9D9FFC8DB2FC09858B75C5FA481" box="[224,343,1194,1219]" pageId="3" pageNumber="344" value="2008-11">Nov. 2008</collectingDate>
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</date>
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,
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<collectorName id="26D55327FFC8DB2FC11A58B65CC4A481" box="[354,460,1194,1219]" pageId="3" pageNumber="344">G. Deein.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9F36F1FFC8DB2FC0BF58D25837A645" blockId="3.[151,1436,1158,2011]" pageId="3" pageNumber="344">
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<materialsCitation id="3B483CACFFC8DB2FC0BF58D25F6EA549" ID-GBIF-Occurrence="3313289309" collectingDate="2008-11" collectionCode="NIFI" country="Thailand" county="Nern Ma Prang District" location="Chao Phraya" municipality="Mae Nam" pageId="3" pageNumber="344" specimenCode="NIFI 3787" specimenCount="1" stateProvince="Phitsanulok" typeStatus="paratype">
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<emphasis id="B954EAE3FFC8DB2FC0BF58D25C4EA4A5" bold="true" box="[199,326,1230,1255]" inLineHeading="true" pageId="3" pageNumber="344" reason="1">
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<typeStatus id="549B8853FFC8DB2FC0BF58D25C49A4A5" box="[199,321,1230,1255]" pageId="3" pageNumber="344" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCode id="DB869E8AFFC8DB2FC12858D35CC4A4A5" box="[336,460,1230,1255]" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="3" pageNumber="344">NIFI 3787</specimenCode>
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,
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<quantity id="4CD89B14FFC8DB2FC1A258D25F49A4A4" box="[474,577,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="7.93" pageId="3" pageNumber="344" unit="mm" value="79.3">79.3 mm</quantity>
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SL;
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<collectingCountry id="F3377661FFC8DB2FC20658D25FECA4A5" box="[638,740,1230,1255]" name="Thailand" pageId="3" pageNumber="344">Thailand</collectingCountry>
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,
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<collectingRegion id="49E4F813FFC8DB2FC28B58D25EB7A4A4" box="[755,959,1230,1255]" country="Thailand" name="Phitsanulok" pageId="3" pageNumber="344">Phitsanulok Prov.</collectingRegion>
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,
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<collectingCounty id="62FE4E7DFFC8DB2FC3B558D359EAA4A5" box="[973,1250,1230,1255]" pageId="3" pageNumber="344">Nern Ma Prang District</collectingCounty>
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,
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<collectingMunicipality id="6BFBAC8BFFC8DB2FC48858D3586AA4A4" box="[1264,1378,1231,1254]" pageId="3" pageNumber="344">Mae Nam</collectingMunicipality>
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<collectingRegion id="49E4F813FFC8DB2FC51558D35894A4A4" box="[1389,1436,1231,1254]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
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drainage,
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<location id="8EFF602AFFC8DB2FC17F58EE5C90A549" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC17F58EE5C90A549" box="[263,408,1266,1291]" country="Thailand" county="Nern Ma Prang District" municipality="Mae Nam" name="Chao Phraya" pageId="3" pageNumber="344" stateProvince="Phitsanulok">Chao Phraya</location>
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basin,
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<date id="FF9E1031FFC8DB2FC19558EF5F6EA549" box="[493,614,1266,1291]" pageId="3" pageNumber="344" value="2008-11">
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<collectingDate id="EFDAE9D9FFC8DB2FC19558EF5F6EA549" box="[493,614,1266,1291]" pageId="3" pageNumber="344" value="2008-11">Nov. 2008</collectingDate>
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</date>
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</materialsCitation>
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;
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<materialsCitation id="3B483CACFFC8DB2FC20C58EF5FB6A56D" ID-GBIF-Occurrence="3313289301" collectingDate="2011-02-09" collectionCode="NIFI" country="Thailand" latitude="16.76" location="Mae Nam" longLatPrecision="768" longitude="100.11" municipality="Bang Rakam District" pageId="3" pageNumber="344" specimenCode="NIFI 4856" specimenCount="1" stateProvince="Phitsanulok" typeStatus="paratype">
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<specimenCode id="DB869E8AFFC8DB2FC20C58EF5FE3A549" box="[628,747,1266,1291]" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="3" pageNumber="344">NIFI 4856</specimenCode>
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(6, 69.9–76.1),
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<collectingRegion id="49E4F813FFC8DB2FC3DC58EE599FA549" box="[932,1175,1266,1291]" country="Thailand" name="Phitsanulok" pageId="3" pageNumber="344">Phitsanulok Province</collectingRegion>
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,
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<collectingMunicipality id="6BFBAC8BFFC8DB2FC4DD58EF5890A549" box="[1189,1432,1266,1291]" pageId="3" pageNumber="344">Bang Rakam District</collectingMunicipality>
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,
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<location id="8EFF602AFFC8DB2FC0EF590B5C04A56C" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC0EF590B5C04A56C" box="[151,268,1303,1326]" country="Thailand" latitude="16.76" longLatPrecision="768" longitude="100.11" municipality="Bang Rakam District" name="Mae Nam" pageId="3" pageNumber="344" stateProvince="Phitsanulok">Mae Nam</location>
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<collectingRegion id="49E4F813FFC8DB2FC163590B5C45A56C" box="[283,333,1303,1326]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
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,
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<geoCoordinate id="EE145036FFC8DB2FC126590A5CBCA56D" box="[350,436,1302,1327]" degrees="16.76" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="555" value="16.76">16.76N</geoCoordinate>
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,
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,
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<date id="FF9E1031FFC8DB2FC252590A5FB6A56D" box="[554,702,1302,1327]" pageId="3" pageNumber="344" value="2011-02-09">
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<collectingDate id="EFDAE9D9FFC8DB2FC252590A5FB6A56D" box="[554,702,1302,1327]" pageId="3" pageNumber="344" value="2011-02-09">9 Feb. 2011</collectingDate>
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</date>
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</materialsCitation>
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;
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<materialsCitation id="3B483CACFFC8DB2FC2A9590B5CADA535" ID-GBIF-Occurrence="3313289302" collectingDate="2010-08-18" collectionCode="NIFI" country="Thailand" latitude="16.76" location="Mae Nam" longLatPrecision="768" longitude="100.11" municipality="Kongkrairat District" pageId="3" pageNumber="344" specimenCode="NIFI 4857" specimenCount="1" stateProvince="Sukhothai" typeStatus="paratype">
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<specimenCode id="DB869E8AFFC8DB2FC2A9590B5E45A56D" box="[721,845,1302,1327]" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="3" pageNumber="344">NIFI 4857</specimenCode>
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(6, 68.4–80.1), boundary of
|
||
<collectingRegion id="49E4F813FFC8DB2FC4CA590A589FA56D" box="[1202,1431,1302,1327]" country="Thailand" name="Sukhothai" pageId="3" pageNumber="344">Sukhothai Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BFBAC8BFFC8DB2FC0EF59265C74A511" box="[151,380,1338,1363]" pageId="3" pageNumber="344">Kongkrairat District</collectingMunicipality>
|
||
with
|
||
<collectingRegion id="49E4F813FFC8DB2FC1BA59265FBEA511" box="[450,694,1338,1363]" country="Thailand" name="Phitsanulok" pageId="3" pageNumber="344">Phitsanulok Province</collectingRegion>
|
||
, Prompiram and Bang Rakam districts,
|
||
<location id="8EFF602AFFC8DB2FC4FE592759F0A510" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC4FE592759F0A510" box="[1158,1272,1339,1362]" country="Thailand" latitude="16.76" longLatPrecision="768" longitude="100.11" municipality="Kongkrairat District" name="Mae Nam" pageId="3" pageNumber="344" stateProvince="Sukhothai">Mae Nam</location>
|
||
<collectingRegion id="49E4F813FFC8DB2FC57A5927583BA510" box="[1282,1331,1339,1362]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC53A5926589FA511" box="[1346,1431,1338,1363]" degrees="16.76" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="555" value="16.76">16.76N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC0EF59425DFFA535" box="[151,247,1374,1399]" degrees="100.11" direction="east" orientation="longitude" pageId="3" pageNumber="344" precision="555" value="100.11">100.11E</geoCoordinate>
|
||
,
|
||
<date id="FF9E1031FFC8DB2FC17C59425CADA535" box="[260,421,1374,1399]" pageId="3" pageNumber="344" value="2010-08-18">
|
||
<collectingDate id="EFDAE9D9FFC8DB2FC17C59425CADA535" box="[260,421,1374,1399]" pageId="3" pageNumber="344" value="2010-08-18">18 Aug. 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CACFFC8DB2FC1CB59435CFFA5D9" ID-GBIF-Occurrence="3313289303" collectingDate="2010-10-11" collectionCode="NIFI" country="Thailand" latitude="15.71" location="Borraped Lake" longLatPrecision="770" longitude="100.27" municipality="Chum Sang District" pageId="3" pageNumber="344" specimenCode="NIFI 4858" specimenCount="1" stateProvince="Nakon Sawan Province" typeStatus="paratype">
|
||
<specimenCode id="DB869E8AFFC8DB2FC1CB59435F22A535" box="[435,554,1374,1399]" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="3" pageNumber="344">NIFI 4858</specimenCode>
|
||
(4, 56.4–67.6)
|
||
<collectingRegion id="49E4F813FFC8DB2FC2A559425EE3A535" box="[733,1003,1374,1399]" inferredBySuffix="true" pageId="3" pageNumber="344">Nakon Sawan Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BFBAC8BFFC8DB2FC381594259D6A535" box="[1017,1246,1374,1399]" pageId="3" pageNumber="344">Chum Sang District</collectingMunicipality>
|
||
,
|
||
<location id="8EFF602AFFC8DB2FC49459425890A535" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC49459425890A535" box="[1260,1432,1374,1399]" country="Thailand" latitude="15.71" longLatPrecision="770" longitude="100.27" municipality="Chum Sang District" name="Borraped Lake" pageId="3" pageNumber="344" stateProvince="Nakon Sawan Province">Borraped Lake</location>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC0EF599E5DE4A5D9" box="[151,236,1410,1435]" degrees="15.71" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="555" value="15.71">15.71N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC088599E5C5AA5D9" box="[240,338,1410,1435]" degrees="100.27" direction="east" orientation="longitude" pageId="3" pageNumber="344" precision="555" value="100.27">100.27E</geoCoordinate>
|
||
,
|
||
<date id="FF9E1031FFC8DB2FC127599E5CFFA5D9" box="[351,503,1410,1435]" pageId="3" pageNumber="344" value="2010-10-11">
|
||
<collectingDate id="EFDAE9D9FFC8DB2FC127599E5CFFA5D9" box="[351,503,1410,1435]" pageId="3" pageNumber="344" value="2010-10-11">11 Oct. 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CACFFC8DB2FC27C599F5E16A5FD" ID-GBIF-Occurrence="3313289311" accessionNumber="UF 186928" collectingDate="2009-04-23" collectionCode="UF" country="Thailand" county="Prompiram District" latitude="17.04" location="Naraesuan Dam" longLatPrecision="768" longitude="100.2" municipality="Mae Nam" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Phitsanulok" typeStatus="paratype">
|
||
<collectionCode id="ED31AE34FFC8DB2FC27C599F5F20A5D8" box="[516,552,1411,1434]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="344" type="Museum">UF</collectionCode>
|
||
186928 (5, 68.2–75.0),
|
||
<collectingRegion id="49E4F813FFC8DB2FC347599E593AA5D9" box="[831,1074,1410,1435]" country="Thailand" name="Phitsanulok" pageId="3" pageNumber="344">Phitsanulok Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62FE4E7DFFC8DB2FC438599E5816A5D9" box="[1088,1310,1410,1435]" pageId="3" pageNumber="344">Prompiram District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BFBAC8BFFC8DB2FC553599F5893A5D8" box="[1323,1435,1411,1434]" pageId="3" pageNumber="344">Mae Nam</collectingMunicipality>
|
||
<collectingRegion id="49E4F813FFC8DB2FC0EF59BB5DCEA5FC" box="[151,198,1447,1470]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
|
||
near
|
||
<location id="8EFF602AFFC8DB2FC17B59BB5CB0A5FC" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC17B59BB5CB0A5FC" box="[259,440,1447,1470]" country="Thailand" county="Prompiram District" latitude="17.04" longLatPrecision="768" longitude="100.2" municipality="Mae Nam" name="Naraesuan Dam" pageId="3" pageNumber="344" stateProvince="Phitsanulok">Naraesuan Dam</location>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC1BB59BA5F10A5FD" box="[451,536,1446,1471]" degrees="17.04" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="555" value="17.04">17.04N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC26459BA5F76A5FD" box="[540,638,1446,1471]" degrees="100.20" direction="east" orientation="longitude" pageId="3" pageNumber="344" precision="555" value="100.2">100.20E</geoCoordinate>
|
||
,
|
||
<date id="FF9E1031FFC8DB2FC2F159BA5E16A5FD" box="[649,798,1446,1471]" pageId="3" pageNumber="344" value="2009-04-23">
|
||
<collectingDate id="EFDAE9D9FFC8DB2FC2F159BA5E16A5FD" box="[649,798,1446,1471]" pageId="3" pageNumber="344" value="2009-04-23">23 Apr. 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CACFFC8DB2FC35159BB5E0BA5A1" ID-GBIF-Occurrence="3313289306" accessionNumber="UF 186929" collectingDate="2011-10-04" collectionCode="UF" country="Thailand" latitude="16.5" location="Mae Nam" longLatPrecision="769" longitude="100.2" municipality="Sam Ngam District" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Phichit" typeStatus="paratype">
|
||
<collectionCode id="ED31AE34FFC8DB2FC35159BB5E45A5FC" box="[809,845,1447,1470]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="344" type="Museum">UF</collectionCode>
|
||
186929 (2, 65.5–72.7),
|
||
<collectingRegion id="49E4F813FFC8DB2FC42159BA581CA5FD" box="[1113,1300,1446,1471]" country="Thailand" name="Phichit" pageId="3" pageNumber="344">Phichit Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BFBAC8BFFC8DB2FC56759BA5DE6A5A1" pageId="3" pageNumber="344">Sam Ngam District</collectingMunicipality>
|
||
,
|
||
<location id="8EFF602AFFC8DB2FC08259D75C63A5A0" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC08259D75C63A5A0" box="[250,363,1483,1506]" country="Thailand" latitude="16.5" longLatPrecision="769" longitude="100.2" municipality="Sam Ngam District" name="Mae Nam" pageId="3" pageNumber="344" stateProvince="Phichit">Mae Nam</location>
|
||
<collectingRegion id="49E4F813FFC8DB2FC10C59D75CADA5A0" box="[372,421,1483,1506]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC1CA59D65F0FA5A1" box="[434,519,1482,1507]" degrees="16.50" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="555" value="16.5">16.50N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC27359D65F65A5A1" box="[523,621,1482,1507]" degrees="100.20" direction="east" orientation="longitude" pageId="3" pageNumber="344" precision="555" value="100.2">100.20E</geoCoordinate>
|
||
,
|
||
<date id="FF9E1031FFC8DB2FC20259D65E0BA5A1" box="[634,771,1482,1507]" pageId="3" pageNumber="344" value="2011-10-04">
|
||
<collectingDate id="EFDAE9D9FFC8DB2FC20259D65E0BA5A1" box="[634,771,1482,1507]" pageId="3" pageNumber="344" value="2011-10-04">4 Oct. 2011</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B483CACFFC8DB2FC36859D75834A645" ID-GBIF-Occurrence="3313289310" accessionNumber="UF 186930" collectingDate="2009-06-15" collectionCode="UF" country="Thailand" county="Nakon Sawan Province" latitude="15.55" location="Chao Phraya River" longLatPrecision="771" longitude="100.08" municipality="Krok Pra District" pageId="3" pageNumber="344" specimenCount="1" stateProvince="Nan" typeStatus="paratype">
|
||
<collectionCode id="ED31AE34FFC8DB2FC36859D75E3CA5A0" box="[784,820,1483,1506]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="344" type="Museum">UF</collectionCode>
|
||
186930 (2, 65.2–70.0),
|
||
<collectingCounty id="62FE4E7DFFC8DB2FC43159D6585DA5A1" box="[1097,1365,1482,1507]" pageId="3" pageNumber="344">Nakon Sawan Province</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BFBAC8BFFC8DB2FC51A59D65C12A645" pageId="3" pageNumber="344">Krok Pra District</collectingMunicipality>
|
||
,
|
||
<location id="8EFF602AFFC8DB2FC15D59F25CF4A645" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC15D59F25CF4A645" box="[293,508,1518,1543]" country="Thailand" county="Nakon Sawan Province" latitude="15.55" longLatPrecision="771" longitude="100.08" municipality="Krok Pra District" name="Chao Phraya River" pageId="3" pageNumber="344" stateProvince="Nan">Chao Phraya River</location>
|
||
, below confluence of
|
||
<collectingRegion id="49E4F813FFC8DB2FC28A59F35E29A644" box="[754,801,1519,1542]" country="Thailand" name="Nan" pageId="3" pageNumber="344">Nan</collectingRegion>
|
||
and
|
||
<location id="8EFF602AFFC8DB2FC32059F35E85A644" LSID="urn:lsid:plazi:treatment:038987E7FFC8DB2AC0EF5FEB5DE7A50D:8EFF602AFFC8DB2FC32059F35E85A644" box="[856,909,1519,1542]" country="Thailand" county="Nakon Sawan Province" latitude="15.55" longLatPrecision="771" longitude="100.08" municipality="Krok Pra District" name="Yom" pageId="3" pageNumber="344" stateProvince="Nan">Yom</location>
|
||
rivers,
|
||
<geoCoordinate id="EE145036FFC8DB2FC39959F2593EA645" box="[993,1078,1518,1543]" degrees="15.55" direction="north" orientation="latitude" pageId="3" pageNumber="344" precision="555" value="15.55">15.55N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145036FFC8DB2FC44359F25995A645" box="[1083,1181,1518,1543]" degrees="100.08" direction="east" orientation="longitude" pageId="3" pageNumber="344" precision="555" value="100.08">100.08E</geoCoordinate>
|
||
,
|
||
<date id="FF9E1031FFC8DB2FC4D059F25834A645" box="[1192,1340,1518,1543]" pageId="3" pageNumber="344" value="2009-06-15">
|
||
<collectingDate id="EFDAE9D9FFC8DB2FC4D059F25834A645" box="[1192,1340,1518,1543]" pageId="3" pageNumber="344" value="2009-06-15">15 Jun. 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFC8DB2FC0BF5A0E5F21A69D" pageId="3" pageNumber="344" type="diagnosis">
|
||
<paragraph id="8B9F36F1FFC8DB2FC0BF5A0E5F21A69D" blockId="3.[151,1436,1158,2011]" pageId="3" pageNumber="344">
|
||
<emphasis id="B954EAE3FFC8DB2FC0BF5A0E5C49A668" box="[199,321,1554,1579]" inLineHeading="true" pageId="3" pageNumber="344" reason="1">
|
||
<emphasis id="B954EAE3FFC8DB2FC0BF5A0E5C32A669" bold="true" box="[199,314,1554,1579]" pageId="3" pageNumber="344">Diagnosis</emphasis>
|
||
.
|
||
</emphasis>
|
||
A species of
|
||
<taxonomicName id="4C204D72FFC8DB2FC1985A0F5F99A669" authorityName="Bleeker" authorityYear="1858" box="[480,657,1555,1579]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="344" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EAE3FFC8DB2FC1985A0F5F99A669" box="[480,657,1555,1579]" italics="true" pageId="3" pageNumber="344">Lepidocephalus</emphasis>
|
||
</taxonomicName>
|
||
that is readily distinguishable (
|
||
<figureCitation id="131B2A74FFC8DB2FC3805A0E5934A669" box="[1016,1084,1554,1579]" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="3" pageNumber="344">Fig. 1</figureCitation>
|
||
,
|
||
<tableCitation id="C6A2034AFFC8DB2FC4345A0E5997A669" box="[1100,1183,1554,1579]" captionStart="TABLE 1" captionStartId="9.[151,241,476,499]" captionTargetPageId="9" captionText="TABLE 1. Characteristics that vary among species of Lepidocephalus. Measurements are in mm or percentages of standard length (SL). Range is followed by average in parentheses." httpUri="http://table.plazi.org/id/DF5F6679FFC2DB25C0EF5DC05E6AA251" pageId="3" pageNumber="344" tableUuid="DF5F6679FFC2DB25C0EF5DC05E6AA251">Table 1</tableCitation>
|
||
) from
|
||
<taxonomicName id="4C204D72FFC8DB2FC4895A0F587FA668" authorityName="Roberts" authorityYear="1989" box="[1265,1399,1555,1578]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="spectrum">
|
||
<emphasis id="B954EAE3FFC8DB2FC4895A0F587FA668" box="[1265,1399,1555,1578]" italics="true" pageId="3" pageNumber="344">L. spectrum</emphasis>
|
||
</taxonomicName>
|
||
by the presence of eyes and dark pigment on the body, and absence of tubules along the lateral line and large papillae on the lips; from
|
||
<taxonomicName id="4C204D72FFC8DB2FC1235A475F0EA631" authorityName=", de Beaufort" authorityYear="1933" box="[347,518,1627,1651]" family="Cobitidae" genus="Acanthophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="pahangensis">
|
||
<emphasis id="B954EAE3FFC8DB2FC1235A475F0EA631" box="[347,518,1627,1651]" italics="true" pageId="3" pageNumber="344">L. pahangensis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of scales on top of the head and absence of large papillae on the lips (
|
||
<figureCitation id="131B2A74FFC8DB2FC0A95A625C30A6D5" box="[209,312,1662,1687]" captionStart="FIGURE 3" captionStartId="5.[151,249,945,967]" captionTargetBox="[165,1424,354,917]" captionTargetId="figure-182@5.[159,1428,350,920]" captionTargetPageId="5" captionText="FIGURE 3. Mouths of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (C) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos by Z. Randall." figureDoi="http://doi.org/10.5281/zenodo.4910409" httpUri="https://zenodo.org/record/4910409/files/figure.png" pageId="3" pageNumber="344">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A74FFC8DB2FC13E5A625C65A6D5" box="[326,365,1662,1687]" captionStart="FIGURE 4" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[170,1416,1104,1898]" captionTargetId="figure-185@5.[164,1422,1094,1898]" captionTargetPageId="5" captionText="FIGURE 4. Mouths of specimens of Lepidocephalus shown in Figure 3. (A) L. nanensis, (B) L. spectrum. Although the lower lip of L. nanensis is folded more posteriorly than that of L. spectrum, it and the upper lip lack the large papillae shown on the lips of L. spectrum. Drawings by D. Lumbantobing." figureDoi="http://doi.org/10.5281/zenodo.4910411" httpUri="https://zenodo.org/record/4910411/files/figure.png" pageId="3" pageNumber="344">4A</figureCitation>
|
||
); from
|
||
<taxonomicName id="4C204D72FFC8DB2FC1BC5A635F3DA6D4" authorityName="Vaillant" authorityYear="1902" box="[452,565,1662,1686]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="pallens">
|
||
<emphasis id="B954EAE3FFC8DB2FC1BC5A635F3DA6D4" box="[452,565,1662,1686]" italics="true" pageId="3" pageNumber="344">L. pallens</emphasis>
|
||
</taxonomicName>
|
||
by having the dorsal-fin origin well behind (vs. over) the pelvic–fin origin (predorsal length 68.1–71.3 vs. 59.9% SL); and from
|
||
<taxonomicName id="4C204D72FFC8DB2FC2885ABF5E89A6F9" baseAuthorityName="Bleeker" baseAuthorityYear="1854" box="[752,897,1699,1723]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="344" phylum="Chordata" rank="species" species="macrochir">
|
||
<emphasis id="B954EAE3FFC8DB2FC2885ABF5E89A6F9" box="[752,897,1699,1723]" italics="true" pageId="3" pageNumber="344">L. macrochir</emphasis>
|
||
</taxonomicName>
|
||
by having a longer snout (5.6–7.4 vs. 3.9–5.1% SL; 28.6–38.1 vs. 23.3–27.1% HL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFC8DB2AC0BF5AF65CBEA465" lastPageId="6" lastPageNumber="347" pageId="3" pageNumber="344" type="description">
|
||
<paragraph id="8B9F36F1FFC8DB2FC0BF5AF65857A709" blockId="3.[151,1436,1158,2011]" pageId="3" pageNumber="344">
|
||
<emphasis id="B954EAE3FFC8DB2FC0BF5AF65C50A740" box="[199,344,1770,1795]" inLineHeading="true" pageId="3" pageNumber="344" reason="1">
|
||
<emphasis id="B954EAE3FFC8DB2FC0BF5AF65C59A741" bold="true" box="[199,337,1770,1795]" pageId="3" pageNumber="344">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body deep, slab-sided; greatest depth 14.7–21.2% SL. Head narrow, length 16.9–22.2% SL. Eye small, in dorsal half of head in shallow depression above bifid suborbital spine. Origin of dorsal fin behind origin of pelvic fin. No or small axillary lobe on pelvic fin; absent on
|
||
<typeStatus id="549B8853FFC8DB2FC3225B2E5EC8A709" box="[858,960,1842,1867]" pageId="3" pageNumber="344" type="holotype">holotype</typeStatus>
|
||
. Fleshy lobe at origin of pectoral fin.
|
||
</paragraph>
|
||
<paragraph id="8B9F36F1FFC8DB2FC0BF5B4A59D8A7D1" blockId="3.[151,1436,1158,2011]" pageId="3" pageNumber="344">Scales on top of head, cheek and opercle; body completely covered with minute, often deeply embedded scales. Lateral line complete, 185–205 scales; 0–2 pores on caudal fin; no tubules on lateral line.</paragraph>
|
||
<paragraph id="8B9F36F1FFC8DB28C0BF5B825EFFA159" blockId="3.[151,1436,1158,2011]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="345" pageId="3" pageNumber="344">
|
||
Mouth horseshoe-shaped; upper lip smooth on ventral surface, with small papilla on anterior and posterior margins, without median indentation; lower lip plicated, with median indentation on lower edge (
|
||
<figureCitation id="131B2A74FFC8DB2FC48D5BDE5853A799" box="[1269,1371,1986,2011]" captionStart="FIGURE 3" captionStartId="5.[151,249,945,967]" captionTargetBox="[165,1424,354,917]" captionTargetId="figure-182@5.[159,1428,350,920]" captionTargetPageId="5" captionText="FIGURE 3. Mouths of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (C) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos by Z. Randall." figureDoi="http://doi.org/10.5281/zenodo.4910409" httpUri="https://zenodo.org/record/4910409/files/figure.png" pageId="3" pageNumber="344">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A74FFC8DB2FC5115BDE5898A799" box="[1385,1424,1986,2011]" captionStart="FIGURE 4" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[170,1416,1104,1898]" captionTargetId="figure-185@5.[164,1422,1094,1898]" captionTargetPageId="5" captionText="FIGURE 4. Mouths of specimens of Lepidocephalus shown in Figure 3. (A) L. nanensis, (B) L. spectrum. Although the lower lip of L. nanensis is folded more posteriorly than that of L. spectrum, it and the upper lip lack the large papillae shown on the lips of L. spectrum. Drawings by D. Lumbantobing." figureDoi="http://doi.org/10.5281/zenodo.4910411" httpUri="https://zenodo.org/record/4910411/files/figure.png" pageId="3" pageNumber="344">4A</figureCitation>
|
||
). Two pairs of rostral barbels; inner pair reaching to corner of mouth, outer pair slightly longer and reaching slightly past corner of mouth; one pair of maxillary barbels, reaching approximately to vertical at posterior end of groove containing suborbital spine. Large flap on anterior nostril. Dorsal rays iii,8; pectoral rays 9–11; pelvic rays 6–8; anal rays ii,5–6; upper branched caudal rays 7; lower branched caudal rays 7.
|
||
</paragraph>
|
||
<caption id="DF5F6679FFCFDB28C0EF5ABF5E58A6FB" ID-DOI="http://doi.org/10.5281/zenodo.4910407" ID-Zenodo-Dep="4910407" box="[151,848,1699,1721]" httpUri="https://zenodo.org/record/4910407/files/figure.png" pageId="4" pageNumber="345" startId="4.[151,249,1699,1721]" targetBox="[153,1429,349,1673]" targetPageId="4">
|
||
<paragraph id="8B9F36F1FFCFDB28C0EF5ABF5E58A6FB" blockId="4.[151,848,1699,1721]" box="[151,848,1699,1721]" pageId="4" pageNumber="345">
|
||
<emphasis id="B954EAE3FFCFDB28C0EF5ABF5C1AA6FA" bold="true" box="[151,274,1699,1721]" pageId="4" pageNumber="345">FIGURE 2.</emphasis>
|
||
Distribution of specimens of
|
||
<taxonomicName id="4C204D72FFCFDB28C2385ABF5FE9A6FB" authorityName="Bleeker" authorityYear="1858" box="[576,737,1699,1721]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="345" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EAE3FFCFDB28C2385ABF5FE9A6FB" box="[576,737,1699,1721]" italics="true" pageId="4" pageNumber="345">Lepidocephalus</emphasis>
|
||
</taxonomicName>
|
||
examined.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36F1FFCFDB28C0BF5AF95C1CA7DB" blockId="4.[151,1436,1765,2022]" pageId="4" pageNumber="345">Cephalic lateralis system: lateral canal with 1 pore, extends anteriorly from lateral line; suborbital canal extends along lower margin of groove holding suborbital spine with 4 pores, abruptly turns anteroventrally with 4 more pores, ends on side of snout well in front of nostrils; no supraorbital canal at surface. Supratemporal canal branches off lateral canal, then ends or continues under skin without pores at surface. Preoperculomandibular pores not visible.</paragraph>
|
||
<paragraph id="8B9F36F1FFCFDB29C0BF5BBA5F6BA167" blockId="4.[151,1436,1765,2022]" lastBlockId="5.[151,1436,151,293]" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345">
|
||
Pectoral fin falcate in both sexes; second ray longer in both sexes than other pectoral-fin rays. Pectoral fin much longer in some individuals than others (12.2–18.9% SL); extent and significance of inter- and intra-variation, including sexual and ontogenetic variation, is unknown. Second ray of pectoral fin much thicker (forming lamina circularis) in adult male than in female, but not longer. Pectoral fins of male
|
||
<taxonomicName id="4C204D72FFCEDB29C4785CA35989A094" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[1024,1153,191,214]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCEDB29C4785CA35989A094" box="[1024,1153,191,214]" italics="true" pageId="5" pageNumber="346">L. nanensis</emphasis>
|
||
</taxonomicName>
|
||
(N = 4) averaged 15.1% SL; those of females (N = 7) averaged 15.5% SL. The thicked ray presumably has a function related to spawning behavior. Pelvic fins are short, 6–8% SL.
|
||
</paragraph>
|
||
<caption id="DF5F6679FFCEDB29C0EF5FAD5C01A446" ID-DOI="http://doi.org/10.5281/zenodo.4910409" ID-Zenodo-Dep="4910409" httpUri="https://zenodo.org/record/4910409/files/figure.png" pageId="5" pageNumber="346" startId="5.[151,249,945,967]" targetBox="[165,1424,354,917]" targetPageId="5">
|
||
<paragraph id="8B9F36F1FFCEDB29C0EF5FAD5C01A446" blockId="5.[151,1437,945,1028]" pageId="5" pageNumber="346">
|
||
<emphasis id="B954EAE3FFCEDB29C0EF5FAD5C1EA384" bold="true" box="[151,278,945,967]" pageId="5" pageNumber="346">FIGURE 3.</emphasis>
|
||
Mouths of
|
||
<taxonomicName id="4C204D72FFCEDB29C1EE5FAD5F3EA385" authorityName="Bleeker" authorityYear="1858" box="[406,566,945,967]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EAE3FFCEDB29C1EE5FAD5F3EA385" box="[406,566,945,967]" italics="true" pageId="5" pageNumber="346">Lepidocephalus</emphasis>
|
||
</taxonomicName>
|
||
. (A)
|
||
<taxonomicName id="4C204D72FFCEDB29C20B5FAE5FE2A385" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[627,746,946,967]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCEDB29C20B5FAE5FE2A385" box="[627,746,946,967]" italics="true" pageId="5" pageNumber="346">L. nanensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B)
|
||
<taxonomicName id="4C204D72FFCEDB29C5FF5FAE5C1EA3A7" authorityName=", de Beaufort" authorityYear="1933" family="Cobitidae" genus="Acanthophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="pahangensis">
|
||
<emphasis id="B954EAE3FFCEDB29C5FF5FAE5C1EA3A7" italics="true" pageId="5" pageNumber="346">L. pahangensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, ZRC 490, 35.8 mm SL; (C)
|
||
<taxonomicName id="4C204D72FFCEDB29C2D85FCC5E1EA3A7" authorityName="Roberts" authorityYear="1989" box="[672,790,976,997]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="spectrum">
|
||
<emphasis id="B954EAE3FFCEDB29C2D85FCC5E1EA3A7" box="[672,790,976,997]" italics="true" pageId="5" pageNumber="346">L. spectrum</emphasis>
|
||
</taxonomicName>
|
||
, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos by Z. Randall.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6679FFCEDB29C0EF5B805FADA7AD" ID-DOI="http://doi.org/10.5281/zenodo.4910411" ID-Zenodo-Dep="4910411" httpUri="https://zenodo.org/record/4910411/files/figure.png" pageId="5" pageNumber="346" startId="5.[151,249,1948,1970]" targetBox="[170,1416,1104,1898]" targetPageId="5">
|
||
<paragraph id="8B9F36F1FFCEDB29C0EF5B805FADA7AD" blockId="5.[151,1436,1948,2031]" pageId="5" pageNumber="346">
|
||
<emphasis id="B954EAE3FFCEDB29C0EF5B805C1AA7F3" bold="true" box="[151,274,1948,1970]" pageId="5" pageNumber="346">FIGURE 4.</emphasis>
|
||
Mouths of specimens of
|
||
<taxonomicName id="4C204D72FFCEDB29C26C5B805FBCA7F0" authorityName="Bleeker" authorityYear="1858" box="[532,692,1948,1970]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="genus">
|
||
<emphasis id="B954EAE3FFCEDB29C26C5B805FBCA7F0" box="[532,692,1948,1970]" italics="true" pageId="5" pageNumber="346">Lepidocephalus</emphasis>
|
||
</taxonomicName>
|
||
shown in Figure 3. (A)
|
||
<taxonomicName id="4C204D72FFCEDB29C3D25B805915A7F3" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[938,1053,1948,1969]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCEDB29C3D25B805915A7F3" box="[938,1053,1948,1969]" italics="true" pageId="5" pageNumber="346">L. nanensis</emphasis>
|
||
</taxonomicName>
|
||
, (B)
|
||
<taxonomicName id="4C204D72FFCEDB29C42A5B8059C2A7F3" authorityName="Roberts" authorityYear="1989" box="[1106,1226,1948,1969]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="spectrum">
|
||
<emphasis id="B954EAE3FFCEDB29C42A5B8059C2A7F3" box="[1106,1226,1948,1969]" italics="true" pageId="5" pageNumber="346">L. spectrum</emphasis>
|
||
</taxonomicName>
|
||
. Although the lower lip of
|
||
<taxonomicName id="4C204D72FFCEDB29C0AD5BA75C41A792" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[213,329,1979,2000]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCEDB29C0AD5BA75C41A792" box="[213,329,1979,2000]" italics="true" pageId="5" pageNumber="346">L. nanensis</emphasis>
|
||
</taxonomicName>
|
||
is folded more posteriorly than that of
|
||
<taxonomicName id="4C204D72FFCEDB29C2A45BA75E5CA792" authorityName="Roberts" authorityYear="1989" box="[732,852,1979,2000]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="spectrum">
|
||
<emphasis id="B954EAE3FFCEDB29C2A45BA75E5CA792" box="[732,852,1979,2000]" italics="true" pageId="5" pageNumber="346">L. spectrum</emphasis>
|
||
</taxonomicName>
|
||
, it and the upper lip lack the large papillae shown on the lips of
|
||
<taxonomicName id="4C204D72FFCEDB29C0A55BC65C5DA7AD" authorityName="Roberts" authorityYear="1989" box="[221,341,2010,2031]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="346" phylum="Chordata" rank="species" species="spectrum">
|
||
<emphasis id="B954EAE3FFCEDB29C0A55BC65C5DA7AD" box="[221,341,2010,2031]" italics="true" pageId="5" pageNumber="346">L. spectrum</emphasis>
|
||
</taxonomicName>
|
||
. Drawings by D. Lumbantobing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6679FFCDDB2AC0EF5F0F5EE3A305" ID-DOI="http://doi.org/10.5281/zenodo.4910413" ID-Zenodo-Dep="4910413" httpUri="https://zenodo.org/record/4910413/files/figure.png" pageId="6" pageNumber="347" startId="6.[151,249,787,809]" targetBox="[151,1436,195,765]" targetPageId="6">
|
||
<paragraph id="8B9F36F1FFCDDB2AC0EF5F0F5EE3A305" blockId="6.[151,1436,787,839]" pageId="6" pageNumber="347">
|
||
<emphasis id="B954EAE3FFCDDB2AC0EF5F0F5C1CA36A" bold="true" box="[151,276,787,809]" pageId="6" pageNumber="347">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C204D72FFCDDB2AC1655F0F5F17A36A" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[285,543,787,809]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCDDB2AC1655F0F5F17A36A" box="[285,543,787,809]" italics="true" pageId="6" pageNumber="347">Lepidocephalus nanensis</emphasis>
|
||
</taxonomicName>
|
||
in an aquarium. Collected at Naraesuan Dam on Mae Nam Nan, Prompiram District, Phitsanulok Province, Thailand, June 2005 (not vouchered). Photo by N. Panitvong.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36F1FFCDDB2AC0BF5F685CBEA465" blockId="6.[151,1436,884,1359]" pageId="6" pageNumber="347">
|
||
<emphasis id="B954EAE3FFCDDB2AC0BF5F685CA0A3CE" box="[199,424,884,909]" inLineHeading="true" pageId="6" pageNumber="347" reason="1">
|
||
<emphasis id="B954EAE3FFCDDB2AC0BF5F685CA9A3CF" bold="true" box="[199,417,884,909]" pageId="6" pageNumber="347">Coloration in life</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body dusky pink-brown; darker gray-brown along mid-dorsum and along mid-side posteriorly (
|
||
<figureCitation id="131B2A74FFCDDB2AC1675F875C6BA3F6" box="[287,355,923,948]" captionStart="FIGURE 5" captionStartId="6.[151,249,787,809]" captionTargetBox="[151,1436,195,765]" captionTargetId="figure-419@6.[151,1436,195,765]" captionTargetPageId="6" captionText="FIGURE 5. Lepidocephalus nanensis in an aquarium. Collected at Naraesuan Dam on Mae Nam Nan, Prompiram District, Phitsanulok Province, Thailand, June 2005 (not vouchered). Photo by N. Panitvong." figureDoi="http://doi.org/10.5281/zenodo.4910413" httpUri="https://zenodo.org/record/4910413/files/figure.png" pageId="6" pageNumber="347">Fig. 5</figureCitation>
|
||
). Distinctive yellow snout followed by blue-black bar before eye; head dusky yellow behind eye. Cheek and opercle with silver-blue iridescence. Dorsal fin with concentration of melanophores at base, creating dark basal band - deepest anteriorly; melanophores outlining lower halves of fin rays. Other fins clear except for melanophores along rays.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFCDDB2AC0BF58295C0CA437" pageId="6" pageNumber="347" type="etymology">
|
||
<paragraph id="8B9F36F1FFCDDB2AC0BF58295C0CA437" blockId="6.[151,1436,884,1359]" pageId="6" pageNumber="347">
|
||
<emphasis id="B954EAE3FFCDDB2AC0BF58295C45A40F" box="[199,333,1077,1102]" inLineHeading="true" pageId="6" pageNumber="347" reason="1">
|
||
<emphasis id="B954EAE3FFCDDB2AC0BF58295C4EA40C" bold="true" box="[199,326,1077,1102]" pageId="6" pageNumber="347">Etymology</emphasis>
|
||
.
|
||
</emphasis>
|
||
The name
|
||
<taxonomicName id="4C204D72FFCDDB2AC1AE582A5F30A40F" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[470,568,1078,1101]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCDDB2AC1AE582A5F30A40F" box="[470,568,1078,1101]" italics="true" pageId="6" pageNumber="347">nanensis</emphasis>
|
||
</taxonomicName>
|
||
, an adjective, refers to the river system where most specimens have been collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFCDDB2AC0BF589F5DE7A50D" pageId="6" pageNumber="347" type="distribution">
|
||
<paragraph id="8B9F36F1FFCDDB2AC0BF589F5DE7A50D" blockId="6.[151,1436,884,1359]" pageId="6" pageNumber="347">
|
||
<emphasis id="B954EAE3FFCDDB2AC0BF589F5C68A4D8" box="[199,352,1155,1180]" inLineHeading="true" pageId="6" pageNumber="347" reason="1">
|
||
<emphasis id="B954EAE3FFCDDB2AC0BF589F5C51A4DE" bold="true" box="[199,345,1155,1180]" pageId="6" pageNumber="347">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C204D72FFCDDB2AC111589F5F8DA4D9" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[361,645,1155,1179]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCDDB2AC111589F5F8DA4D9" box="[361,645,1155,1179]" italics="true" pageId="6" pageNumber="347">Lepidocephalus nanensis</emphasis>
|
||
</taxonomicName>
|
||
is found in
|
||
<collectingCountry id="F3377661FFCDDB2AC36B589F5E7FA4DE" box="[787,887,1155,1180]" name="Thailand" pageId="6" pageNumber="347">Thailand</collectingCountry>
|
||
in the
|
||
<collectingRegion id="49E4F813FFCDDB2AC3B1589F5EF0A4D8" box="[969,1016,1155,1178]" country="Thailand" name="Nan" pageId="6" pageNumber="347">Nan</collectingRegion>
|
||
River and in the Chao-Phraya River below the confluence of the
|
||
<collectingRegion id="49E4F813FFCDDB2AC1AF58BB5F0EA4FC" box="[471,518,1191,1214]" country="Thailand" name="Nan" pageId="6" pageNumber="347">Nan</collectingRegion>
|
||
and Yom rivers (
|
||
<figureCitation id="131B2A74FFCDDB2AC2B158BA5E06A4FD" box="[713,782,1190,1215]" captionStart="FIGURE 2" captionStartId="4.[151,249,1699,1721]" captionTargetBox="[153,1429,349,1673]" captionTargetId="figure-196@4.[151,1436,339,1677]" captionTargetPageId="4" captionText="FIGURE 2. Distribution of specimens of Lepidocephalus examined." figureDoi="http://doi.org/10.5281/zenodo.4910407" httpUri="https://zenodo.org/record/4910407/files/figure.png" pageId="6" pageNumber="347">Fig. 2</figureCitation>
|
||
). To our knowledge,
|
||
<bibRefCitation id="EFB14B00FFCDDB2AC38658BA59F6A4FD" author="Slechtova, V. & Bohlen, J. & Perdices, A." box="[1022,1278,1190,1215]" pageId="6" pageNumber="347" pagination="812 - 831" refId="ref8173" refString="Slechtova, V., Bohlen, J. & Perdices, A. (2008) Molecular phylogeny of the freshwater fish family Cobitidae (Cypriniformes: Teleostei): Delimination of genera, mitochondrial introgression and evolution of sexual dimorphism. Molecular Phylogenetics and Evolution, 47, 812 - 831. http: // dx. doi. org / 10.1016 / j. ympev. 2007.12.018" type="journal article" year="2008">
|
||
Šlechtová
|
||
<emphasis id="B954EAE3FFCDDB2AC40C58B459ACA4FD" box="[1140,1188,1191,1215]" italics="true" pageId="6" pageNumber="347">et al</emphasis>
|
||
. (2008)
|
||
</bibRefCitation>
|
||
first recorded this species from
|
||
<collectingCountry id="F3377661FFCDDB2AC12258D75CB6A4A6" box="[346,446,1227,1252]" name="Thailand" pageId="6" pageNumber="347">Thailand</collectingCountry>
|
||
(as
|
||
<taxonomicName id="4C204D72FFCDDB2AC19458D05F75A4A1" baseAuthorityName="Bleeker" baseAuthorityYear="1854" box="[492,637,1227,1251]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="347" phylum="Chordata" rank="species" species="macrochir">
|
||
<emphasis id="B954EAE3FFCDDB2AC19458D05F75A4A1" box="[492,637,1227,1251]" italics="true" pageId="6" pageNumber="347">L. macrochir</emphasis>
|
||
</taxonomicName>
|
||
) and included molecular data from specimens collected at a fishpond at Nakom Savan (Nakon Sawan, Nakon Sawan Province),
|
||
<collectingCountry id="F3377661FFCDDB2AC36658F25E8DA545" box="[798,901,1262,1287]" name="Thailand" pageId="6" pageNumber="347">Thailand</collectingCountry>
|
||
. Recent collections suggest that this species is common in the
|
||
<collectingRegion id="49E4F813FFCDDB2AC12A590F5C89A568" box="[338,385,1299,1322]" country="Thailand" name="Nan" pageId="6" pageNumber="347">Nan</collectingRegion>
|
||
and lower Chao Phraya rivers. These large rivers seem to be the preferred habitat of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |