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<document id="E554F25F330F65DA3B1089E7110B429A" ID-DOI="10.5281/zenodo.206036" ID-GBIF-Dataset="28370cf1-101b-4a8a-a3c9-16793ec82652" ID-ISSN="1175-5326" ID-Zenodo-Dep="206036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219556155" checkinUser="plazi" docAuthor="Kong, Fanqing &amp; Wang, Xinhua" docDate="2011" docId="039B879DFFEEFF93628CFCBD08E2B331" docLanguage="en" docName="zt02733p068.pdf" docOrigin="Zootaxa 2733" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterotrissocladius quartus Kong &amp; Wang, 2011, sp. n." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FFA2FFE5FFECFF90621BFFA30972B758" masterDocTitle="Heterotrissocladius Spärck from China (Diptera: Chironomidae)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="65" updateTime="1698237436967" updateUser="plazi">
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<mods:title id="503888B87501D6EB1BF3B69073EBC046">Heterotrissocladius Spärck from China (Diptera: Chironomidae)</mods:title>
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<mods:namePart id="459DFFBF4B4217CB761AA285142DC8B0">Kong, Fanqing</mods:namePart>
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<mods:namePart id="227141EE1EAD6C2628C3DC2C6F40595D">Wang, Xinhua</mods:namePart>
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<mods:date id="FAB7B09AC9974BEB18FFE448D2A5BACC">2011</mods:date>
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<treatment id="039B879DFFEEFF93628CFCBD08E2B331" ID-DOI="http://doi.org/10.5281/zenodo.6185365" ID-GBIF-Taxon="119385018" ID-Zenodo-Dep="6185365" LSID="urn:lsid:plazi:treatment:039B879DFFEEFF93628CFCBD08E2B331" httpUri="http://treatment.plazi.org/id/039B879DFFEEFF93628CFCBD08E2B331" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
<subSubSection id="C3286500FFEEFF92628CFCBD0840B401" pageId="2" pageNumber="65" type="nomenclature">
<paragraph id="8B8D368BFFEEFF92628CFCBD0B5DB460" blockId="2.[151,559,798,857]" box="[151,559,798,824]" pageId="2" pageNumber="65">
<heading id="D0C581E7FFEEFF92628CFCBD0B5DB460" bold="true" box="[151,559,798,824]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
<emphasis id="B946EA99FFEEFF92628CFCBD0B5DB460" bold="true" box="[151,559,798,824]" pageId="2" pageNumber="65">
<taxonomicName id="4C324D08FFEEFF92628CFCBD0894B460" ID-CoL="8CC3R" box="[151,486,798,824]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="quartus" status="sp. nov.">
<emphasis id="B946EA99FFEEFF92628CFCBD0894B460" bold="true" box="[151,486,798,824]" italics="true" pageId="2" pageNumber="65">Heterotrissocladius quartus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557E2FFEEFF9263F7FCBD0B5DB460" box="[492,559,798,824]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D368BFFEEFF92628CFCE20840B401" blockId="2.[151,559,798,857]" box="[151,306,833,857]" pageId="2" pageNumber="65">
(
<figureCitation id="13092A0EFFEEFF926284FCE2099EB401" box="[159,236,833,857]" captionStart="FIGURE 2" captionStartId="3.[151,250,750,773]" captionTargetBox="[179,1404,202,719]" captionTargetId="figure@3.[170,1417,181,729]" captionTargetPageId="3" captionText="FIGURE 2. Heterotrissocladius quartus sp. n.: A. Palp; B. Wing; C. Hypopygium." httpUri="https://zenodo.org/record/206038/files/figure.png" pageId="2" pageNumber="65">Figs. 2</figureCitation>
AC)
</paragraph>
</subSubSection>
<subSubSection id="C3286500FFEEFF92628CFC290D71B355" pageId="2" pageNumber="65" type="materials_examined">
<paragraph id="8B8D368BFFEEFF92628CFC290A79B49E" blockId="2.[151,1436,905,1585]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF92628CFC29083AB4FA" bold="true" box="[151,328,905,930]" pageId="2" pageNumber="65">
<typeStatus id="54898829FFEEFF92628CFC2909A2B4FA" box="[151,208,906,930]" pageId="2" pageNumber="65">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54898829FFEEFF92634DFC2A08CCB4F9" box="[342,446,905,929]" pageId="2" pageNumber="65" type="holotype">Holotype</typeStatus>
male (BDN No.1740),
<collectingCountry id="F325761BFFEEFF9260C5FC2A0A49B4F9" box="[734,827,905,929]" name="China" pageId="2" pageNumber="65">CHINA</collectingCountry>
: Guangxi Autonomous Region, Jinxiu Autonomous County, Luoxiang Town,
<date id="FF8C104BFFEEFF9263AEFC0E0B53B49D" box="[437,545,941,965]" pageId="2" pageNumber="65" value="1990-06-09">9.vi.1990</date>
, light trap, X. Wang.
</paragraph>
<paragraph id="8B8D368BFFEEFF9262DCFC720D71B355" blockId="2.[151,1436,905,1585]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DCFC7208A2B4B2" bold="true" box="[199,464,977,1002]" pageId="2" pageNumber="65">Diagnostic characters.</emphasis>
The new species can be separated from other members of the genus by having a 4-segmented palp, short Cu1, low AR (0.56), and a short triangular acute anal point.
</paragraph>
</subSubSection>
<subSubSection id="C3286500FFEEFF9262DCFBBF0DE8B36C" box="[199,1178,1052,1076]" pageId="2" pageNumber="65" type="etymology">
<paragraph id="8B8D368BFFEEFF9262DCFBBF0DE8B36C" blockId="2.[151,1436,905,1585]" box="[199,1178,1052,1076]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DCFBBF083CB36C" bold="true" box="[199,334,1052,1076]" pageId="2" pageNumber="65">Etymology.</emphasis>
From Latin,
<emphasis id="B946EA99FFEEFF9263FAFBBE0B6CB36C" box="[481,542,1053,1076]" italics="true" pageId="2" pageNumber="65">quart</emphasis>
means four, referring to the palp with four palpomeres.
</paragraph>
</subSubSection>
<subSubSection id="C3286500FFEEFF9362DCFB9C0A02B4EE" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65" type="description">
<paragraph id="8B8D368BFFEEFF9262DCFB9C0B4BB300" blockId="2.[151,1436,905,1585]" box="[199,569,1087,1112]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DCFB9C082BB300" bold="true" box="[199,345,1087,1112]" pageId="2" pageNumber="65">Description.</emphasis>
Male imago (n =1).
</paragraph>
<paragraph id="8B8D368BFFEEFF9262DCFBC709BDB3F8" blockId="2.[151,1436,905,1585]" pageId="2" pageNumber="65">
Total length
<quantity id="4CCA9B6EFFEEFF926348FBC708C8B324" box="[339,442,1124,1148]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="65" unit="mm" value="2.1">2.10 mm</quantity>
. Wing length
<quantity id="4CCA9B6EFFEEFF92604EFBC70BC9B324" box="[597,699,1124,1148]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="65" unit="mm" value="1.45">1.45 mm</quantity>
. Total length / wing length 1.48. Wing length / length of profemur 2.64.
</paragraph>
<paragraph id="8B8D368BFFEEFF9262DCFB0809C9B268" blockId="2.[151,1436,905,1585]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DCFB08083DB39C" bold="true" box="[199,335,1195,1220]" pageId="2" pageNumber="65">Coloration.</emphasis>
Head brown. Thorax dark brown. Abdomen pale yellow. Legs yellow. Wing nearly transparent.
<emphasis id="B946EA99FFEEFF92628CFB7309A6B3B0" box="[151,212,1232,1256]" italics="true" pageId="2" pageNumber="65">Head</emphasis>
. AR 0.56. Temporal setae 10, including 4 inner verticals, 3 outer verticals and 3 postorbitals. Clypeus with 13 setae. Tentorium 79 Μm long, 18 Μm wide. Palp 4-segmented; palpomeres lengths (in Μm): 18; 26; 62; 66. (
<figureCitation id="13092A0EFFEEFF926742FB570CEEB254" box="[1369,1436,1268,1292]" captionStart="FIGURE 2" captionStartId="3.[151,250,750,773]" captionTargetBox="[179,1404,202,719]" captionTargetId="figure@3.[170,1417,181,729]" captionTargetPageId="3" captionText="FIGURE 2. Heterotrissocladius quartus sp. n.: A. Palp; B. Wing; C. Hypopygium." httpUri="https://zenodo.org/record/206038/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="8B8D368BFFEEFF9262DEFA9E08ABB2F9" blockId="2.[151,1436,905,1585]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DEFA9E098FB20C" box="[197,253,1341,1364]" italics="true" pageId="2" pageNumber="65">Wing</emphasis>
. (
<figureCitation id="13092A0EFFEEFF92630CFA9F0829B20C" box="[279,347,1340,1364]" captionStart="FIGURE 2" captionStartId="3.[151,250,750,773]" captionTargetBox="[179,1404,202,719]" captionTargetId="figure@3.[170,1417,181,729]" captionTargetPageId="3" captionText="FIGURE 2. Heterotrissocladius quartus sp. n.: A. Palp; B. Wing; C. Hypopygium." httpUri="https://zenodo.org/record/206038/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
B). Anal lobe well developed. Cu1 short, 252 Μm, without setae. VR 1.15. R with 13 setae, R1 with 16, R4+5 with 27, Cu with 19, M3+4 with 4 setae; r2+3 with 9 setae, r4+5 with 30, m1+2 with 13 and cell an with 10 setae. Squama with 12 setae.
</paragraph>
<paragraph id="8B8D368BFFEEFF9262DEFA0D0AC5B29D" blockId="2.[151,1436,905,1585]" box="[197,951,1453,1478]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DEFA0D086CB29E" box="[197,286,1454,1478]" italics="true" pageId="2" pageNumber="65">Thorax.</emphasis>
Dorsocentrals 10; acrostichals 14; prealars 4; scutellars 12.
</paragraph>
<paragraph id="8B8D368BFFEEFF9262DEFA700B65B169" blockId="2.[151,1436,905,1585]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF9262DEFA700873B2B2" box="[197,257,1491,1514]" italics="true" pageId="2" pageNumber="65">Legs.</emphasis>
Spur of front tibia 22 Μm long; of mid tibia 18 Μm and 12 Μm long; of hind tibia 35 Μm and 9 Μm long; with 26 Μm long comb. Width at apex of front tibia 9 Μm, of mid tibia
<quantity id="4CCA9B6EFFEEFF9261AEFA560A82B155" box="[949,1008,1525,1549]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="65" unit="m" value="13.0">13 m</quantity>
, of hind tibia 20 Μm. Lengths (in Μm) and proportions of legs in
<tableCitation id="C6B00330FFEEFF9263A6F9BA0B61B169" box="[445,531,1561,1585]" captionStart="TABLE 2" captionStartId="2.[151,235,1628,1649]" captionText="TABLE 2. Lengths (in µm) and proportions of legs segments of male H. quartus sp. n." pageId="2" pageNumber="65">Table 2</tableCitation>
.
</paragraph>
<caption id="DF4D6603FFEEFF92628CF9FF0D7BB12A" box="[151,1033,1628,1650]" pageId="2" pageNumber="65">
<paragraph id="8B8D368BFFEEFF92628CF9FF0D7BB12A" blockId="2.[151,1289,1628,2001]" box="[151,1033,1628,1650]" pageId="2" pageNumber="65">
<emphasis id="B946EA99FFEEFF92628CF9FF098CB129" bold="true" box="[151,254,1628,1649]" pageId="2" pageNumber="65">TABLE 2</emphasis>
. Lengths (in µm) and proportions of legs segments of male
<taxonomicName id="4C324D08FFEEFF926146F9FE0ABBB12A" box="[861,969,1629,1650]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="quartus" status="sp. nov.">
<emphasis id="B946EA99FFEEFF926146F9FE0ABBB12A" box="[861,969,1629,1650]" italics="true" pageId="2" pageNumber="65">H. quartus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557E2FFEEFF9261CBF9FE0D7BB12A" box="[976,1033,1629,1650]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8D368BFFEEFF9262F1F92E0C7BB1FF" blockId="2.[151,1289,1628,2001]" box="[234,1289,1677,1703]" pageId="2" pageNumber="65">fe ti ta1 ta2 ta3 ta4</paragraph>
<paragraph id="8B8D368BFFEEFF926284F9190C7BB197" blockId="2.[151,1289,1628,2001]" box="[159,1289,1721,1744]" pageId="2" pageNumber="65">p1 550 625 400 200 125 75</paragraph>
<paragraph id="8B8D368BFFEEFF926284F9460C7BB1A2" blockId="2.[151,1289,1628,2001]" box="[159,1289,1764,1787]" pageId="2" pageNumber="65">p2 575 500 225 125 100 50</paragraph>
<paragraph id="8B8D368BFFEEFF926284F8AC0D3FB017" blockId="2.[151,1289,1628,2001]" pageId="2" pageNumber="65">p3 550 600 375 200 125 75 ta5 LR BV SV BR</paragraph>
<paragraph id="8B8D368BFFEEFF926284F8C50D2AB023" blockId="2.[151,1289,1628,2001]" box="[159,1112,1893,1916]" pageId="2" pageNumber="65">p1 50 0.64 1.85 2.94 3.60</paragraph>
<paragraph id="8B8D368BFFEEFF926284F8320D2AB0FD" blockId="2.[151,1289,1628,2001]" box="[159,1112,1936,1958]" pageId="2" pageNumber="65">p2 50 0.45 2.89 4.28 3.00</paragraph>
<paragraph id="8B8D368BFFEEFF926284F8180D2AB088" blockId="2.[151,1289,1628,2001]" box="[159,1112,1978,2001]" pageId="2" pageNumber="65">p3 50 0.63 3.39 3.07 3.30</paragraph>
<caption id="DF4D6603FFEFFF93628CFD4D0A94B45D" box="[151,998,750,773]" httpUri="https://zenodo.org/record/206038/files/figure.png" pageId="3" pageNumber="66" targetBox="[179,1404,202,719]" targetPageId="3">
<paragraph id="8B8D368BFFEFFF93628CFD4D0A94B45D" blockId="3.[151,998,750,773]" box="[151,998,750,773]" pageId="3" pageNumber="66">
<emphasis id="B946EA99FFEFFF93628CFD4D0861B45C" bold="true" box="[151,275,750,773]" pageId="3" pageNumber="66">FIGURE 2.</emphasis>
<taxonomicName id="4C324D08FFEFFF936302FD4C0B41B45D" box="[281,563,751,773]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="quartus" status="sp. nov.">
<emphasis id="B946EA99FFEFFF936302FD4C0B41B45D" box="[281,563,751,773]" italics="true" pageId="3" pageNumber="66">Heterotrissocladius quartus</emphasis>
</taxonomicName>
<emphasis id="B946EA99FFEFFF936021FD4C0B01B45C" bold="true" box="[570,627,751,772]" pageId="3" pageNumber="66">
<taxonomicNameLabel id="A27557E2FFEFFF936021FD4C0B01B45C" box="[570,627,751,772]" pageId="3" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: A. Palp; B. Wing; C. Hypopygium.
</paragraph>
</caption>
<paragraph id="8B8D368BFFEFFF9362DEFC900A02B4EE" blockId="3.[151,1436,817,1130]" pageId="3" pageNumber="66">
<emphasis id="B946EA99FFEFFF9362DEFC900826B412" box="[197,340,819,842]" italics="true" pageId="3" pageNumber="66">Hypopygium</emphasis>
(
<figureCitation id="13092A0EFFEFFF936378FC9208D4B411" box="[355,422,817,841]" captionStart="FIGURE 2" captionStartId="3.[151,250,750,773]" captionTargetBox="[179,1404,202,719]" captionTargetId="figure@3.[170,1417,181,729]" captionTargetPageId="3" captionText="FIGURE 2. Heterotrissocladius quartus sp. n.: A. Palp; B. Wing; C. Hypopygium." httpUri="https://zenodo.org/record/206038/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
C). Anal point roughly triangular with pointed apex, 26 Μm long. Tergite IX with 12 setae on the anal point. Laterosternite IX with 6 setae. Phallapodeme 26 Μm long; transverse sternapodeme 88 Μm long. Gonocoxite 132 Μm long. Inferior volsella 62 Μm long, with 11 long setae. Gonostylus 57 Μm long, with small triangular crista dorsalis; megaseta 10 Μm long. HR 2.27; HV 3.65.
</paragraph>
</subSubSection>
<subSubSection id="C3286500FFEFFF9362DEFC610D2FB379" pageId="3" pageNumber="66" type="discussion">
<paragraph id="8B8D368BFFEFFF9362DEFC610D2FB379" blockId="3.[151,1436,817,1130]" pageId="3" pageNumber="66">
<emphasis id="B946EA99FFEFFF9362DEFC61084BB482" bold="true" box="[197,313,962,986]" pageId="3" pageNumber="66">Remarks.</emphasis>
According to
<bibRefCitation id="EFA34B7AFFEFFF9363C2FC620B08B481" author="Saether" box="[473,634,961,985]" pageId="3" pageNumber="68" refString="Saether, O. A. (1975) Nearctic and Palaearctic Heterotrissocladius (Diptera: Chironomidae). Bulletin of the Fisheries Research Board of Canada, 193: 1 - 67." type="journal article" year="1975">Saether (1975)</bibRefCitation>
,
<taxonomicName id="4C324D08FFEFFF93609DFC600B8FB482" box="[646,765,963,986]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="quartus">
<emphasis id="B946EA99FFEFFF93609DFC600B8FB482" box="[646,765,963,986]" italics="true" pageId="3" pageNumber="66">H. quartus</emphasis>
</taxonomicName>
can be included in the
<taxonomicName id="4C324D08FFEFFF93661BFC610D1AB482" box="[1024,1128,962,986]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="marcidus">
<emphasis id="B946EA99FFEFFF93661BFC610D1AB482" box="[1024,1128,962,986]" italics="true" pageId="3" pageNumber="66">marcidus</emphasis>
</taxonomicName>
species group. It is close to
<taxonomicName id="4C324D08FFEFFF93628CFC4408ABB4A6" authority="Stur &amp; Wiedenbrug" authorityName="Stur &amp; Wiedenbrug" box="[151,473,998,1022]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="zierli">
<emphasis id="B946EA99FFEFFF93628CFC440981B4A6" box="[151,243,998,1022]" italics="true" pageId="3" pageNumber="66">H. zierli</emphasis>
Stur &amp; Wiedenbrug
</taxonomicName>
having nearly the same low AR and hairy wings, but it differs in the number of palpomeres (
<taxonomicName id="4C324D08FFEFFF93631DFBA80813B37A" box="[262,353,1034,1058]" class="Insecta" family="Chironomidae" genus="Heterotrissocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="zierli">
<emphasis id="B946EA99FFEFFF93631DFBA80813B37A" box="[262,353,1034,1058]" italics="true" pageId="3" pageNumber="66">H. zierli</emphasis>
</taxonomicName>
with 5 palpomeres), and in having the much wider inferior volsella.
</paragraph>
</subSubSection>
<subSubSection id="C3286500FFEFFF9362DEFB8E08E2B331" pageId="3" pageNumber="66" type="distribution">
<paragraph id="8B8D368BFFEFFF9362DEFB8E08E2B331" blockId="3.[151,1436,817,1130]" pageId="3" pageNumber="66">
<emphasis id="B946EA99FFEFFF9362DEFB8E082CB31E" bold="true" box="[197,350,1069,1094]" pageId="3" pageNumber="66">Distribution.</emphasis>
The present species was collected in subtropical mountain area in Oriental
<collectingCountry id="F325761BFFEFFF9366FFFB8D0C55B31E" box="[1252,1319,1070,1094]" name="China" pageId="3" pageNumber="66">China</collectingCountry>
(Guangxi Autonomous Region).
</paragraph>
</subSubSection>
</treatment>
</document>