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<document id="422FCFF537555ECDA4FD73C4AB2B490E" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="0387762362C7FF612DEFB059FB0C594B" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Parena) valeriae Facchini. E 2011" docType="treatment" docVersion="8" lastPageNumber="131" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="130" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="441184E2D962BE286CC32D03BAAAE1B1">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<treatment id="0387762362C7FF612DEFB059FB0C594B" ID-DOI="http://doi.org/10.5281/zenodo.7963815" ID-GBIF-Taxon="209628985" ID-Zenodo-Dep="7963815" LSID="urn:lsid:plazi:treatment:0387762362C7FF612DEFB059FB0C594B" httpUri="http://treatment.plazi.org/id/0387762362C7FF612DEFB059FB0C594B" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130">
<subSubSection id="C33494BE62C7FF622DEFB059FD5B5F66" box="[151,700,546,573]" pageId="129" pageNumber="130" type="nomenclature">
<paragraph id="8B91C73562C7FF622DEFB059FD5B5F66" blockId="129.[151,700,546,609]" box="[151,700,546,573]" pageId="129" pageNumber="130">
<heading id="D0D9705962C7FF622DEFB059FD5B5F66" bold="true" box="[151,700,546,573]" fontSize="11" level="2" pageId="129" pageNumber="130" reason="3">
<emphasis id="B95A1B2762C7FF622DEFB059FD5B5F66" bold="true" box="[151,700,546,573]" pageId="129" pageNumber="130">
[41]
<taxonomicName id="4C2EBCB662C7FF622DB7B058FD5B5F66" ID-CoL="75RGX" authority="Facchini, 2011" authorityName="Facchini. E" authorityYear="2011" box="[207,700,546,573]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="valeriae" subGenus="Parena">
<emphasis id="B95A1B2762C7FF622DB7B058FEC05F67" bold="true" box="[207,295,547,573]" italics="true" pageId="129" pageNumber="130">Parena</emphasis>
(
<emphasis id="B95A1B2762C7FF622C40B058FE775F67" bold="true" box="[312,400,547,573]" italics="true" pageId="129" pageNumber="130">Parena</emphasis>
)
<emphasis id="B95A1B2762C7FF622CDAB059FDE55F67" bold="true" box="[418,514,546,573]" italics="true" pageId="129" pageNumber="130">valeriae</emphasis>
<bibRefCitation id="EFBFBAC462C7FF622F71B059FD5B5F66" author="Facchini, S." box="[521,700,546,572]" pageId="129" pageNumber="130" pagination="275 - 279" refId="ref110543" refString="Facchini, S. (2011) A new species of Parena Motschulsky, 1859 from Zambia (Coleoptera, Carabidae, Lebiini). Giornale Italiano di Entomologia, 12 (56), 275 - 279." type="journal article" year="2011">Facchini, 2011</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE62C7FF622DEFB03CFD925F3B" box="[151,629,583,609]" pageId="129" pageNumber="130" type="description">
<paragraph id="8B91C73562C7FF622DEFB03CFD925F3B" blockId="129.[151,700,546,609]" box="[151,629,583,609]" pageId="129" pageNumber="130">
Habitus:
<figureCitation id="1315DBB062C7FF622D85B03CFE835F3B" box="[253,356,583,609]" captionStart="FIGURE 76" captionStartId="126.[151,250,1902,1926]" captionTargetBox="[175,1421,510,1873]" captionTargetId="figure-211@126.[158,1430,510,1879]" captionTargetPageId="126" captionText="FIGURE 76. Habitus of Parena (Parena) scutata species group. A Lectotype of Parena madagascariensis (Alluaud). B Holotype of Parena alluaudi Jeannel [= Parena madagascariensis (Alluaud)]. C P. madagascariensis (Alluaud), female from Madagascar. D Holotype of Parena valeriae Facchini. E Lectotype of Parena scutata (Alluaud). F Holotype of Parena ruficornis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959096" httpUri="https://zenodo.org/record/7959096/files/figure.png" pageId="129" pageNumber="130">Fig. 76D</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB062C7FF622F66B03CFD965F3B" box="[542,625,583,609]" captionStart="FIGURE 79" captionStartId="129.[152,255,1974,1998]" captionTargetBox="[162,1426,1057,1943]" captionTargetId="figure-337@129.[152,1436,1044,1950]" captionTargetPageId="129" captionText="FIGURE 79. Median lobe of male genitalia for Parena (Parena) valeriae Facchini, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959102" httpUri="https://zenodo.org/record/7959102/files/figure.png" pageId="129" pageNumber="130">Fig. 79</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE62C7FF622DEFB0F0FC815FF9" box="[151,870,651,675]" pageId="129" pageNumber="130" type="reference_group">
<paragraph id="8B91C73562C7FF622DEFB0F0FC815FF9" blockId="129.[151,870,651,675]" box="[151,870,651,675]" pageId="129" pageNumber="130">
<materialsCitation id="3B46CD6862C7FF622DEFB0F0FC855FF9" ID-GBIF-Occurrence="4115871353" box="[151,866,651,675]" country="Zambia" location="Chisasa" pageId="129" pageNumber="130" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFBFBAC462C7FF622DEFB0F0FE875FF9" author="Facchini, S." box="[151,352,651,675]" pageId="129" pageNumber="130" pagination="275 - 279" refId="ref110543" refString="Facchini, S. (2011) A new species of Parena Motschulsky, 1859 from Zambia (Coleoptera, Carabidae, Lebiini). Giornale Italiano di Entomologia, 12 (56), 275 - 279." type="journal article" year="2011">Facchini, 2011: 276</bibRefCitation>
(type locality:
<collectingCountry id="F33987A562C7FF622C82B0F0FDAD5FF9" box="[506,586,651,675]" name="Zambia" pageId="129" pageNumber="130">Zambia</collectingCountry>
,
<location id="8EF191EE62C7FF622F2CB0F0FD415FF9" LSID="urn:lsid:plazi:treatment:0387762362C7FF612DEFB059FB0C594B:8EF191EE62C7FF622F2CB0F0FD415FF9" box="[596,678,651,675]" country="Zambia" name="Chisasa" pageId="129" pageNumber="130">Chisasa</location>
;
<typeStatus id="5495799762C7FF622FC9B0F0FCED5FF9" box="[689,778,651,675]" pageId="129" pageNumber="130" type="holotype">holotype</typeStatus>
in CSF)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE62C7FF622DEFB0AAFD465E69" pageId="129" pageNumber="130" type="materials_examined">
<paragraph id="8B91C73562C7FF622DEFB0AAFD465E69" blockId="129.[151,1436,721,1036]" pageId="129" pageNumber="130">
<emphasis id="B95A1B2762C7FF622DEFB0AAFE5C5FB1" bold="true" box="[151,443,721,747]" pageId="129" pageNumber="130">Type material examined</emphasis>
.
<emphasis id="B95A1B2762C7FF622CB3B0A9FC955FB1" bold="true" box="[459,882,721,748]" pageId="129" pageNumber="130">
<taxonomicName id="4C2EBCB662C7FF622CB3B0A9FD105FB1" authority="Facchini" authorityName="Facchini. E" authorityYear="2011" box="[459,759,721,748]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="valeriae">
<emphasis id="B95A1B2762C7FF622CB3B0A9FD635FB6" bold="true" box="[459,644,721,748]" italics="true" pageId="129" pageNumber="130">Parena valeriae</emphasis>
Facchini
</taxonomicName>
:
<typeStatus id="5495799762C7FF622E7DB0AAFC955FB1" box="[773,882,721,747]" pageId="129" pageNumber="130" type="holotype">Holotype</typeStatus>
</emphasis>
(CSF,
<figureCitation id="1315DBB062C7FF622EBBB0AAFBCA5FB1" box="[963,1069,721,747]" captionStart="FIGURE 76" captionStartId="126.[151,250,1902,1926]" captionTargetBox="[175,1421,510,1873]" captionTargetId="figure-211@126.[158,1430,510,1879]" captionTargetPageId="126" captionText="FIGURE 76. Habitus of Parena (Parena) scutata species group. A Lectotype of Parena madagascariensis (Alluaud). B Holotype of Parena alluaudi Jeannel [= Parena madagascariensis (Alluaud)]. C P. madagascariensis (Alluaud), female from Madagascar. D Holotype of Parena valeriae Facchini. E Lectotype of Parena scutata (Alluaud). F Holotype of Parena ruficornis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959096" httpUri="https://zenodo.org/record/7959096/files/figure.png" pageId="129" pageNumber="130">Fig. 76D</figureCitation>
): male, body length = 10.0 mm, board mounted, &quot;
<collectingCountry id="F33987A562C7FF622C2DB08EFE495E55" box="[341,430,757,783]" name="Zambia" pageId="129" pageNumber="130">Zambia</collectingCountry>
NW / E of Chisasa /
<date id="FF90E1F562C7FF622FE4B08EFCFB5E4A" box="[668,796,757,784]" pageId="129" pageNumber="130" value="2005-11-18">18.XI.2005</date>
leg. Snížek&quot;, &quot;
<typeStatus id="5495799762C7FF622EB8B08EFB8D5E55" box="[960,1130,757,783]" pageId="129" pageNumber="130" type="holotype">HOLOTYPUS</typeStatus>
/
<emphasis id="B95A1B2762C7FF6229F8B08DFA7B5E4A" box="[1152,1436,757,784]" italics="true" pageId="129" pageNumber="130">
<taxonomicName id="4C2EBCB662C7FF6229F8B08DFB365E55" box="[1152,1233,758,783]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/
<taxonomicName id="4C2EBCB662C7FF62299FB08EFAA45E55" authorityName="Facchini. E" authorityYear="2011" box="[1255,1347,757,783]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="valeriae">valeriae</taxonomicName>
/ n.sp. /
</emphasis>
Det. Facchini 20
<emphasis id="B95A1B2762C7FF622C29B162FE8C5E69" box="[337,363,793,819]" italics="true" pageId="129" pageNumber="130">11</emphasis>
&quot; [red label] &lt;
<figureCitation id="1315DBB062C7FF622F7FB162FD855E6E" box="[519,610,793,820]" captionStart="FIGURE 5" captionStartId="12.[151,250,1858,1882]" captionTargetBox="[185,1402,181,1834]" captionTargetId="figure-28@12.[185,1402,181,1834]" captionTargetPageId="12" captionText="FIGURE 5. Elytral apex (left) of Parena spp. A P. (Parena) africana (Alluaud), Cameroon. B P. (Parena) ruficornis sp. n., Central Africa, paratype. C P. (Parena) valeriae Facchini, holotype. D P. (Parena) madagascariensis (Alluaud), Madagascar. E P. (Bothynoptera) tesari (Jedlička), Taiwan. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Bothynoptera) kurosai Habu, Japan. H P. (Bothynoptera) dorsigera (Schaum), Yunnan. I P. (Bothynoptera) dorsigera (Schaum), Beijing. J P. (Bothynoptera) dorsigera (Schaum), Japan. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.7958947" httpUri="https://zenodo.org/record/7958947/files/figure.png" pageId="129" pageNumber="130">Figs 5C</figureCitation>
,
<figureCitation id="1315DBB062C7FF622F16B162FD6A5E69" box="[622,653,793,819]" captionStart="FIGURE 79" captionStartId="129.[152,255,1974,1998]" captionTargetBox="[162,1426,1057,1943]" captionTargetId="figure-337@129.[152,1436,1044,1950]" captionTargetPageId="129" captionText="FIGURE 79. Median lobe of male genitalia for Parena (Parena) valeriae Facchini, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959102" httpUri="https://zenodo.org/record/7959102/files/figure.png" pageId="129" pageNumber="130">79</figureCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE62C7FF612DBFB146FC015C1B" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130" type="diagnosis">
<paragraph id="8B91C73562C7FF622DBFB146FBB75956" blockId="129.[151,1436,721,1036]" pageId="129" pageNumber="130">
<emphasis id="B95A1B2762C7FF622DBFB146FE815E0D" bold="true" box="[199,358,829,855]" pageId="129" pageNumber="130">Comparisons</emphasis>
. This species differs from other African species of
<taxonomicName id="4C2EBCB662C7FF622ED7B145FBE75E0D" box="[943,1024,830,855]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B2762C7FF622ED7B145FBE75E0D" box="[943,1024,830,855]" italics="true" pageId="129" pageNumber="130">Parena</emphasis>
</taxonomicName>
by the combination of: dorsum reddish brown, elytra with a pair of dark stripes gradually fading toward apices; postgenae without suborbital setae; antennae and legs uniformly reddish brown. As indicated in the original description (
<bibRefCitation id="EFBFBAC462C7FF622922B1FEFAE15EC5" author="Facchini, S." box="[1114,1286,901,927]" pageId="129" pageNumber="130" pagination="275 - 279" refId="ref110543" refString="Facchini, S. (2011) A new species of Parena Motschulsky, 1859 from Zambia (Coleoptera, Carabidae, Lebiini). Giornale Italiano di Entomologia, 12 (56), 275 - 279." type="journal article" year="2011">Facchini, 2011</bibRefCitation>
), this species is similar to
<taxonomicName id="4C2EBCB662C7FF622C59B1D1FE555E99" box="[289,434,937,963]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B95A1B2762C7FF622C59B1D1FE555E99" box="[289,434,937,963]" italics="true" pageId="129" pageNumber="130">P. ferruginea</emphasis>
</taxonomicName>
, but differs in its unique color: elytra with dark stripes, antennae unicolor, and tarsomeres not darkened. Furthermore, we found one more difference between these two species: the elytra sutural angles are slightly pointed, forming short denticles in
<taxonomicName id="4C2EBCB662C7FF622F04B189FD145951" authorityName="Facchini. E" authorityYear="2011" box="[636,755,1009,1035]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="valeriae">
<emphasis id="B95A1B2762C7FF622F04B189FD145951" box="[636,755,1009,1035]" italics="true" pageId="129" pageNumber="130">P. valeriae</emphasis>
</taxonomicName>
, but indistinct in
<taxonomicName id="4C2EBCB662C7FF622EC0B189FBAE5951" box="[952,1097,1009,1035]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B95A1B2762C7FF622EC0B189FBAE5951" box="[952,1097,1009,1035]" italics="true" pageId="129" pageNumber="130">P. ferruginea</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF5197BD62C7FF622DE0B5CDFD425AA8" ID-DOI="http://doi.org/10.5281/zenodo.7959102" ID-Zenodo-Dep="7959102" httpUri="https://zenodo.org/record/7959102/files/figure.png" pageId="129" pageNumber="130" startId="129.[152,255,1974,1998]" targetBox="[162,1426,1057,1943]" targetPageId="129" targetType="figure">
<paragraph id="8B91C73562C7FF622DE0B5CDFD425AA8" blockId="129.[151,1436,1974,2034]" pageId="129" pageNumber="130">
<emphasis id="B95A1B2762C7FF622DE0B5CDFEF95A94" bold="true" box="[152,286,1974,1998]" pageId="129" pageNumber="130">FIGURE 79</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB662C7FF622FFDB5CCFC0D5A94" authority="Facchini" authorityName="Facchini. E" authorityYear="2011" box="[645,1002,1974,1998]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="valeriae" subGenus="Parena">
<emphasis id="B95A1B2762C7FF622FFDB5CCFD285A94" box="[645,719,1975,1998]" italics="true" pageId="129" pageNumber="130">Parena</emphasis>
(
<emphasis id="B95A1B2762C7FF622FA6B5CCFCCF5A94" box="[734,808,1975,1998]" italics="true" pageId="129" pageNumber="130">Parena</emphasis>
)
<emphasis id="B95A1B2762C7FF622E4EB5CDFC6E5A94" box="[822,905,1974,1998]" italics="true" pageId="129" pageNumber="130">valeriae</emphasis>
Facchini
</taxonomicName>
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B91C73562C4FF612DBFB2ECFC015C1B" blockId="130.[151,1437,151,1041]" pageId="130" pageNumber="131">
We considered this species to be closest to
<taxonomicName id="4C2EBCB662C4FF612FD6B2ECFC735DEB" box="[686,916,151,177]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B95A1B2762C4FF612FD6B2ECFC735DEB" box="[686,916,151,177]" italics="true" pageId="130" pageNumber="131">P. madagascariensis</emphasis>
</taxonomicName>
based on having the elytral apex truncate, sutural angles pointed; antennae completely brown, and apical lamella of aedeagus elongate. Compared to the latter species,
<taxonomicName id="4C2EBCB662C4FF612D8FB2A4FE975DA3" authorityName="Facchini. E" authorityYear="2011" box="[247,368,223,249]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="valeriae">
<emphasis id="B95A1B2762C4FF612D8FB2A4FE975DA3" box="[247,368,223,249]" italics="true" pageId="130" pageNumber="131">P. valeriae</emphasis>
</taxonomicName>
differs in having the elytra with black lateral stripes, elytral outer apical angles rounded (
<figureCitation id="1315DBB062C4FF612817B2A4FF5D5C47" captionStart="FIGURE 5" captionStartId="12.[151,250,1858,1882]" captionTargetBox="[185,1402,181,1834]" captionTargetId="figure-28@12.[185,1402,181,1834]" captionTargetPageId="12" captionText="FIGURE 5. Elytral apex (left) of Parena spp. A P. (Parena) africana (Alluaud), Cameroon. B P. (Parena) ruficornis sp. n., Central Africa, paratype. C P. (Parena) valeriae Facchini, holotype. D P. (Parena) madagascariensis (Alluaud), Madagascar. E P. (Bothynoptera) tesari (Jedlička), Taiwan. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Bothynoptera) kurosai Habu, Japan. H P. (Bothynoptera) dorsigera (Schaum), Yunnan. I P. (Bothynoptera) dorsigera (Schaum), Beijing. J P. (Bothynoptera) dorsigera (Schaum), Japan. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.7958947" httpUri="https://zenodo.org/record/7958947/files/figure.png" pageId="130" pageNumber="131">Fig. 5C</figureCitation>
), elytral striae shallower, elytra without microsculpture, apical lamella of the aedeagus shorter, and the endophallus of male genitalia with the primary sclerite present only in the basal half.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE62C4FF612DBFB331FAFB5EB7" pageId="130" pageNumber="131" type="description">
<paragraph id="8B91C73562C4FF612DBFB331FAFB5EB7" blockId="130.[151,1437,151,1041]" pageId="130" pageNumber="131">
<emphasis id="B95A1B2762C4FF612DBFB331FEB65C3E" bold="true" box="[199,337,330,356]" pageId="130" pageNumber="131">Description</emphasis>
. Body length 10.0 mm. Dorsum reddish brown, moderately polished, elytra with a pair of dark stripes, occupying intervals 6 to 8, dark stripes gradually fading toward apices; antennae uniformly reddish brown; venter reddish brown; legs uniformly reddish brown. Vertex with fine punctures; postgenae without suborbital setae; mentum with a pair of median setae, which are slightly longer than terminal labial palpomeres; antennae extended only to elytral base. Pronotum sub-cordate, PW/PL = 1.45, slightly wider than head, PW/HW = 1.06; widest at anterior third, lateral margins evenly rounded before middle, shallowly sinuate before posterior angles; posterior angles rounded-obtuse, not prominent; lateral explanations slightly wide; disc with fine punctures aside median line. Elytra slightly dilated to apex; without microsculpture; striae shallowly incised, with fine puncture rows; intervals shallowly convex, with sparse fine punctures; discal depressions shallow, sub-triangular, near basal third of intervals 3 to 6; apical truncation distinct, completely rounded at outer apical angles, and gradually turn to straight to sutural angle; sutural angles pointed, forming short denticles. Males with biseriate adhesive setae on apical half of mesotarsomere 1 and almost full length of mesotarsomeres 2–3. Apex of abdominal sternite VII straight, with two or three setae on each side in males (the
<typeStatus id="5495799762C4FF612F79B080FD845E4F" box="[513,611,763,789]" pageId="130" pageNumber="131" type="holotype">holotype</typeStatus>
with three setae on left side, two on right side). Median lobe of aedeagus slender (AL/AW = 4.6), ventral margin slightly expanded near middle; apical lamella thin and narrow, oblong in dorsal view, LL much greater than LW, apex rounded; in lateral view, apical lamella slightly bent upward. Endophallus with wide flared basal expansion of primary sclerite; flagellum very short, ending before middle of median lobe; apical sclerite widely V-shaped, moderately defined, basal core indistinct; basal sheath sparsely and coarsely scaled, apical sheath densely and finely scaled, larger than basal sheath; squamate sac not divided, large and triangular (in lateral view), not chitinized, near middle of median lobe, ventral to squamate sheath. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE62C4FF612DBFB18DFB0C594B" box="[199,1259,1014,1041]" pageId="130" pageNumber="131" type="distribution">
<paragraph id="8B91C73562C4FF612DBFB18DFB0C594B" blockId="130.[151,1437,151,1041]" box="[199,1259,1014,1041]" pageId="130" pageNumber="131">
<emphasis id="B95A1B2762C4FF612DBFB18DFEBE594A" bold="true" box="[199,345,1014,1040]" pageId="130" pageNumber="131">Distribution</emphasis>
(
<figureCitation id="1315DBB062C4FF612C10B18CFE25594B" box="[360,450,1015,1041]" captionStart="MAP 13" captionStartId="125.[152,211,1494,1518]" captionTargetBox="[151,1436,633,1470]" captionTargetId="figure-216@125.[151,1436,633,1470]" captionText="MAP 13. Distributions for species in the Parena (Parena) scutata group" figureDoi="http://doi.org/10.5281/zenodo.7963701" httpUri="https://zenodo.org/record/7963701/files/figure.png" pageId="130" pageNumber="131">Map 13</figureCitation>
, green). Only known from the
<typeStatus id="5495799762C4FF612E64B18CFCAB594B" box="[796,844,1015,1041]" pageId="130" pageNumber="131">type</typeStatus>
locality in the northwestern
<collectingCountry id="F33987A562C4FF6129F6B18CFB00594B" box="[1166,1255,1015,1041]" name="Zambia" pageId="130" pageNumber="131">Zambia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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