284 lines
36 KiB
XML
284 lines
36 KiB
XML
<document id="0F2D2B5635FA2522DAE1283B72692EE9" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2023" docId="03877623620BFFAD2DEFB11DFE625A76" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Bothynoptera) emarginata Shi & Liang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="79" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="78" updateTime="1699506634024" updateUser="plazi">
|
||
<mods:mods id="CBD6133F9228F6D9C7DADCD1C1887676" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="2B6969C74016912CB181222B7EDAB5F0">
|
||
<mods:title id="5CDFDAD2AE239C585955666859F7A75F">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FA1D55469854F4EBBCD8A155F2796452" type="personal">
|
||
<mods:role id="2005FE4C32907E63C4F4833D47D09D7E">
|
||
<mods:roleTerm id="3E95A14DCD183EC53AF8F27AF5153DE7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6A3B8337C3A94EEDCDDB4FF862B9336F">Shi, Hongliang</mods:namePart>
|
||
<mods:nameIdentifier id="852CD5B1E0D17EA2E8BAAEEFDEDCC474" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="EDB540123F5086EC95B1B93A311A49A5" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="3CC9152F5B97C84C2A68EF1A2B283A40" type="personal">
|
||
<mods:role id="065CD66A77A9DDCD06004E4FEB8E384F">
|
||
<mods:roleTerm id="7B39958AAC09C0FF3A3831CE044E31B0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4261A043006FD66993FA687C7B55B9E8">Liang, Hongbin</mods:namePart>
|
||
<mods:nameIdentifier id="DE39A182AF67FCF5199F16C2A6F47816" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="77F50DD8481BA665BFA2F3D53CCDE383" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="FA1F419DB9F311E0C68ED91085EB90AF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F92130AB75C0FD0DEF5966B08EEEFFC4" type="host">
|
||
<mods:titleInfo id="0727E8C2B8978362758DB6D5578D34DE">
|
||
<mods:title id="B6CE70F1C4B16E9988D1B4A578C96192">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="BE69C1AA44CAEE69D1C7B2DA759D8E94">
|
||
<mods:date id="63E1AA4656F188F4685C4A2CEE08837C">2023</mods:date>
|
||
<mods:detail id="97697A27C3C0189383E65D5A38AE21F0" type="pubDate">
|
||
<mods:number id="E98A2CC3B75ED6EA17AF5DC47B7D08D0">2023-05-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D756A93CE9D8C87080E690D4380C9B0F" type="volume">
|
||
<mods:number id="BDAB5F5B84A356CF64F523B364A67567">5286</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="22E52D073B548BCDE1C14FF8972AF513" type="issue">
|
||
<mods:number id="EC690094FDF029A03B57B15C7B42B128">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="83D71D85634ED0E3450331A1C993AF5E" unit="page">
|
||
<mods:start id="E5E342956A757F8BDDDE9D2739E3AC39">1</mods:start>
|
||
<mods:end id="1319312879F7F785FB4107ED765F0601">144</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="61073F7E94A006F7E5BBA7F9EA81B8E9">
|
||
<mods:url id="B895B03A93153F387F71AF5B642F625C">http://dx.doi.org/10.11646/zootaxa.5286.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="E48F8320D21E62FBA4F9FD48E21F704C">journal article</mods:classification>
|
||
<mods:identifier id="B19A54202DCAB749FC2E330B96A678F7" type="CLB-Dataset">263342</mods:identifier>
|
||
<mods:identifier id="C721A70451A36E46CE87D6BE1A62368B" type="DOI">10.11646/zootaxa.5286.1.1</mods:identifier>
|
||
<mods:identifier id="CEF1AF675E9C95D1DEDAB30A0671C98A" type="GBIF-Dataset">67e7a105-1f62-414a-83d9-f758039eb726</mods:identifier>
|
||
<mods:identifier id="B7933DC1A7764F8484B4906DB237B3EA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="9B95FBABCCC19AADB9A1B9B19CC7763E" type="Zenodo-Dep">7958937</mods:identifier>
|
||
<mods:identifier id="1E558AB1D1BCA42C2F037AA11E7FD77C" type="ZooBank">F9834684-24D3-4795-B5EB-77B451DF856D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03877623620BFFAD2DEFB11DFE625A76" ID-DOI="http://doi.org/10.5281/zenodo.7963759" ID-GBIF-Taxon="209629012" ID-Zenodo-Dep="7963759" LSID="urn:lsid:plazi:treatment:03877623620BFFAD2DEFB11DFE625A76" httpUri="http://treatment.plazi.org/id/03877623620BFFAD2DEFB11DFE625A76" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
|
||
<subSubSection id="C33494BE620BFFAE2DEFB11DFD395EDB" box="[151,734,870,897]" pageId="77" pageNumber="78" type="nomenclature">
|
||
<paragraph id="8B91C735620BFFAE2DEFB11DFD395EDB" blockId="77.[151,833,870,933]" box="[151,734,870,897]" pageId="77" pageNumber="78">
|
||
<emphasis id="B95A1B27620BFFAE2DEFB11DFD395EDB" bold="true" box="[151,734,870,897]" pageId="77" pageNumber="78">
|
||
[21]
|
||
<taxonomicName id="4C2EBCB6620BFFAE2DB7B11CFD9E5EDB" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[207,633,870,897]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="emarginata" status="sp. nov." subGenus="Bothynoptera">
|
||
<emphasis id="B95A1B27620BFFAE2DB7B11CFEC05EDB" bold="true" box="[207,295,871,897]" italics="true" pageId="77" pageNumber="78">Parena</emphasis>
|
||
(
|
||
<emphasis id="B95A1B27620BFFAE2C40B11DFE3A5EDB" bold="true" box="[312,477,870,897]" italics="true" pageId="77" pageNumber="78">Bothynoptera</emphasis>
|
||
)
|
||
<emphasis id="B95A1B27620BFFAE2C96B11CFD9E5EDB" bold="true" box="[494,633,871,897]" italics="true" pageId="77" pageNumber="78">emarginata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A269A65C620BFFAE2FF8B11CFD395EDB" box="[640,734,871,897]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE620BFFAE2DEFB1F0FCA65EFF" box="[151,833,907,933]" pageId="77" pageNumber="78" type="description">
|
||
<paragraph id="8B91C735620BFFAE2DEFB1F0FCA65EFF" blockId="77.[151,833,870,933]" box="[151,833,907,933]" pageId="77" pageNumber="78">
|
||
Habitus:
|
||
<figureCitation id="1315DBB0620BFFAE2D85B1F0FE425EFF" box="[253,421,907,933]" captionStart="FIGURE 46" captionStartId="78.[151,250,1595,1619]" captionTargetBox="[159,1436,707,1563]" captionTargetId="figure-274@78.[151,1436,705,1572]" captionTargetPageId="78" captionText="FIGURE 46. Parena (Bothynoptera) emarginata sp. n. A Habitus of holotype. B Habitus of paratype. C Mouthparts of holotype. D Labels of holotype. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959031" httpUri="https://zenodo.org/record/7959031/files/figure.png" pageId="77" pageNumber="78">Figs 46A, 46B</figureCitation>
|
||
. Gonocoxites of ovipositor:
|
||
<figureCitation id="1315DBB0620BFFAE2F9AB1F0FCDA5EFF" box="[738,829,907,933]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="77" pageNumber="78">Fig. 11I</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE620BFFAE2DEFB1A8FD08599F" pageId="77" pageNumber="78" type="materials_examined">
|
||
<paragraph id="8B91C735620BFFAE2DEFB1A8FBD05EB7" blockId="77.[151,1437,978,2013]" box="[151,1079,978,1005]" pageId="77" pageNumber="78">
|
||
<materialsCitation id="3B46CD68620BFFAE2DEFB1A8FBD45EB7" ID-GBIF-Occurrence="4115871479" box="[151,1075,978,1005]" county="Muli County" elevation="3059" latitude="28.11" location="Liziping" longLatPrecision="74" longitude="101.138" pageId="77" pageNumber="78" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
|
||
<emphasis id="B95A1B27620BFFAE2DEFB1A8FED15EB6" bold="true" box="[151,310,978,1005]" pageId="77" pageNumber="78">
|
||
<typeStatus id="54957997620BFFAE2DEFB1A8FF355EB7" box="[151,210,979,1005]" pageId="77" pageNumber="78">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingRegion id="49EA09D7620BFFAE2C44B1A8FE7E5EB7" box="[316,409,979,1005]" country="China" name="Sichuan" pageId="77" pageNumber="78">Sichuan</collectingRegion>
|
||
,
|
||
<collectingCounty id="62F0BFB9620BFFAE2CDDB1A8FDD15EB7" box="[421,566,979,1005]" pageId="77" pageNumber="78">Muli County</collectingCounty>
|
||
,
|
||
<location id="8EF191EE620BFFAE2F39B1A8FD475EB7" LSID="urn:lsid:plazi:treatment:03877623620BFFAD2DEFB11DFE625A76:8EF191EE620BFFAE2F39B1A8FD475EB7" box="[577,672,979,1005]" county="Muli County" latitude="28.11" longLatPrecision="74" longitude="101.138" name="Liziping" pageId="77" pageNumber="78" stateProvince="Sichuan">Liziping</location>
|
||
xiang,
|
||
<geoCoordinate id="EE1AA1F2620BFFAE2F8BB1A8FCB15EB7" box="[755,854,979,1005]" degrees="28.110" direction="north" orientation="latitude" pageId="77" pageNumber="78" precision="55" value="28.11">N28.110</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1AA1F2620BFFAE2E19B1A8FC375EB7" box="[865,976,979,1005]" degrees="101.138" direction="east" orientation="longitude" pageId="77" pageNumber="78" precision="55" value="101.138">E101.138</geoCoordinate>
|
||
,
|
||
<quantity id="4CD66AD0620BFFAE2EA3B1A9FBD45EB7" box="[987,1075,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.0589999999999997" pageId="77" pageNumber="78" unit="m" value="3059.0">
|
||
<elevation id="00032006620BFFAE2EA3B1A9FBD45EB7" box="[987,1075,978,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.0589999999999997" pageId="77" pageNumber="78" unit="m" value="3059.0">3059 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B91C735620BFFAE2DBFB18CFD08599F" blockId="77.[151,1437,978,2013]" pageId="77" pageNumber="78">
|
||
<emphasis id="B95A1B27620BFFAE2DBFB18CFE88594A" bold="true" box="[199,367,1014,1041]" pageId="77" pageNumber="78">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B46CD68620BFFAE2C07B18CFDE25927" ID-GBIF-Occurrence="4115871585" collectorName="Huang Hao" country="China" county="Muli county" elevation="3059" latitude="28.11032" location="Inst. of Zoology" longLatPrecision="1" longitude="101.13768" municipality="Liziping" pageId="77" pageNumber="78" specimenCount="2" stateProvince="Sichuan" typeStatus="holotype">
|
||
<emphasis id="B95A1B27620BFFAE2C07B18CFE0B594B" bold="true" box="[383,492,1015,1041]" pageId="77" pageNumber="78">
|
||
<typeStatus id="54957997620BFFAE2C07B18CFE0B594B" box="[383,492,1015,1041]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(IZAS,
|
||
<figureCitation id="1315DBB0620BFFAE2F33B18CFD53594B" box="[587,692,1015,1041]" captionStart="FIGURE 46" captionStartId="78.[151,250,1595,1619]" captionTargetBox="[159,1436,707,1563]" captionTargetId="figure-274@78.[151,1436,705,1572]" captionTargetPageId="78" captionText="FIGURE 46. Parena (Bothynoptera) emarginata sp. n. A Habitus of holotype. B Habitus of paratype. C Mouthparts of holotype. D Labels of holotype. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959031" httpUri="https://zenodo.org/record/7959031/files/figure.png" pageId="77" pageNumber="78">Fig. 46A</figureCitation>
|
||
): female, body length =
|
||
<quantity id="4CD66AD0620BFFAE2EB4B18CFBC2594B" box="[972,1061,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="77" pageNumber="78" unit="mm" value="6.7">6.7 mm</quantity>
|
||
, board mounted, "
|
||
<collectingCountry id="F33987A5620BFFAE298BB18CFADD594B" box="[1267,1338,1015,1041]" name="China" pageId="77" pageNumber="78">China</collectingCountry>
|
||
<collectingRegion id="49EA09D7620BFFAE283AB18CFA7B594B" box="[1346,1436,1015,1041]" country="China" name="Sichuan" pageId="77" pageNumber="78">Sichuan</collectingRegion>
|
||
<collectingCounty id="62F0BFB9620BFFAE2DEFB660FEC6596F" box="[151,289,1051,1077]" pageId="77" pageNumber="78">Muli county</collectingCounty>
|
||
, /
|
||
<collectingMunicipality id="6BF55D4F620BFFAE2C41B660FE7F596F" box="[313,408,1051,1077]" pageId="77" pageNumber="78">Liziping</collectingMunicipality>
|
||
xiang, river bank with / sparse tree /
|
||
<geoCoordinate id="EE1AA1F2620BFFAE2E48B660FC4B596F" box="[816,940,1051,1077]" degrees="28.11032" direction="north" orientation="latitude" pageId="77" pageNumber="78" precision="1" value="28.11032">N28.11032</geoCoordinate>
|
||
<geoCoordinate id="EE1AA1F2620BFFAE2ECAB660FBA6596F" box="[946,1089,1051,1077]" degrees="101.13768" direction="east" orientation="longitude" pageId="77" pageNumber="78" precision="1" value="101.13768">E101.13768</geoCoordinate>
|
||
", "2012,V,6. D
|
||
<quantity id="4CD66AD0620BFFAE2992B661FAD9596F" box="[1258,1342,1050,1077]" metricMagnitude="3" metricUnit="m" metricValue="3.0589999999999997" pageId="77" pageNumber="78" unit="m" value="3059.0">
|
||
<elevation id="00032006620BFFAE2992B661FAD9596F" box="[1258,1342,1050,1077]" metricMagnitude="3" metricUnit="m" metricValue="3.0589999999999997" pageId="77" pageNumber="78" unit="m" value="3059.0">3059 m</elevation>
|
||
</quantity>
|
||
/
|
||
<collectorName id="26DBA2E3620BFFAE282AB660FF215903" pageId="77" pageNumber="78">Huang Hao</collectorName>
|
||
leg /
|
||
<location id="8EF191EE620BFFAE2C7AB644FE505903" LSID="urn:lsid:plazi:treatment:03877623620BFFAD2DEFB11DFE625A76:8EF191EE620BFFAE2C7AB644FE505903" box="[258,439,1087,1113]" country="China" county="Muli county" latitude="28.11032" longLatPrecision="1" longitude="101.13768" municipality="Liziping" name="Inst. of Zoology" pageId="77" pageNumber="78" stateProvince="Sichuan">Inst. of Zoology</location>
|
||
, CAS", "HOLOTYPE /
|
||
<taxonomicName id="4C2EBCB6620BFFAE2FBCB644FC265903" box="[708,961,1087,1113]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Bothynoptera">Parena (Bothynoptera)</taxonomicName>
|
||
/ emarginata
|
||
<taxonomicNameLabel id="A269A65C620BFFAE2920B644FB495903" box="[1112,1198,1087,1113]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
/ des. Shi H.L. 2022" [red label] <
|
||
<figureCitation id="1315DBB0620BFFAE2C5DB618FE615927" box="[293,390,1123,1149]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="77" pageNumber="78">Figs 11I</figureCitation>
|
||
,
|
||
<figureCitation id="1315DBB0620BFFAE2CEBB618FDE25927" box="[403,517,1123,1149]" captionStart="FIGURE 46" captionStartId="78.[151,250,1595,1619]" captionTargetBox="[159,1436,707,1563]" captionTargetId="figure-274@78.[151,1436,705,1572]" captionTargetPageId="78" captionText="FIGURE 46. Parena (Bothynoptera) emarginata sp. n. A Habitus of holotype. B Habitus of paratype. C Mouthparts of holotype. D Labels of holotype. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959031" httpUri="https://zenodo.org/record/7959031/files/figure.png" pageId="77" pageNumber="78">46C, 46D</figureCitation>
|
||
</materialsCitation>
|
||
>.
|
||
<materialsCitation id="3B46CD68620BFFAE2F5AB618FD3C599F" ID-GBIF-Occurrence="4115871557" collectingDate="2013-06-20" collectorName="Yang Xiaodong & Chinese" country="China" elevation="2500" location="Lai" municipality="Cona county" pageId="77" pageNumber="78" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
|
||
<emphasis id="B95A1B27620BFFAE2F5AB618FD685927" bold="true" box="[546,655,1123,1149]" pageId="77" pageNumber="78">
|
||
<typeStatus id="54957997620BFFAE2F5AB618FD685927" box="[546,655,1123,1149]" pageId="77" pageNumber="78" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
(IZAS):
|
||
<specimenCount id="9D280CBC620BFFAE2F81B618FCB85927" box="[761,863,1123,1149]" pageId="77" pageNumber="78" type="female">1 female</specimenCount>
|
||
, "
|
||
<collectingRegion id="49EA09D7620BFFAE2E0CB618FC2C5927" box="[884,971,1123,1149]" country="China" name="Xizang" pageId="77" pageNumber="78">Xizang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BF55D4F620BFFAE2EA0B618FB8E5927" box="[984,1129,1123,1149]" pageId="77" pageNumber="78">Cona county</collectingMunicipality>
|
||
,
|
||
<location id="8EF191EE620BFFAE290FB618FB7B5927" LSID="urn:lsid:plazi:treatment:03877623620BFFAD2DEFB11DFE625A76:8EF191EE620BFFAE290FB618FB7B5927" box="[1143,1180,1123,1149]" country="China" municipality="Cona county" name="Lai" pageId="77" pageNumber="78" stateProvince="Xizang">Lai</location>
|
||
xiang /
|
||
<quantity id="4CD66AD0620BFFAE2985B619FAB15927" box="[1277,1366,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="77" pageNumber="78" unit="m" value="2500.0">
|
||
<elevation id="00032006620BFFAE2985B619FAB15927" box="[1277,1366,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="77" pageNumber="78" unit="m" value="2500.0">2500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="26DBA2E3620BFFAE281AB618FEE259FB" pageId="77" pageNumber="78">Yang Xiaodong</collectorName>
|
||
leg. /
|
||
<date id="FF90E1F5620BFFAE2C28B6FCFE3459FB" box="[336,467,1159,1185]" pageId="77" pageNumber="78" value="2013-06-20">
|
||
<collectingDate id="EFD4181D620BFFAE2C28B6FCFE3459FB" box="[336,467,1159,1185]" pageId="77" pageNumber="78" value="2013-06-20">2013.VI.20</collectingDate>
|
||
</date>
|
||
" [in Chinese], "IOZ(E)1700238", "PARATYPE /
|
||
<taxonomicName id="4C2EBCB6620BFFAE2970B6FCFAE059FB" box="[1032,1287,1159,1185]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Bothynoptera">Parena (Bothynoptera)</taxonomicName>
|
||
/ emarginata
|
||
<emphasis id="B95A1B27620BFFAE2DEFB6D0FF16599F" bold="true" box="[151,241,1195,1221]" pageId="77" pageNumber="78">
|
||
<taxonomicNameLabel id="A269A65C620BFFAE2DEFB6D0FF16599F" box="[151,241,1195,1221]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
/ des. Shi H.L. 2022" [red label] <
|
||
<figureCitation id="1315DBB0620BFFAE2F0EB6D0FD3C599F" box="[630,731,1195,1221]" captionStart="FIGURE 46" captionStartId="78.[151,250,1595,1619]" captionTargetBox="[159,1436,707,1563]" captionTargetId="figure-274@78.[151,1436,705,1572]" captionTargetPageId="78" captionText="FIGURE 46. Parena (Bothynoptera) emarginata sp. n. A Habitus of holotype. B Habitus of paratype. C Mouthparts of holotype. D Labels of holotype. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959031" httpUri="https://zenodo.org/record/7959031/files/figure.png" pageId="77" pageNumber="78">Fig. 46B</figureCitation>
|
||
</materialsCitation>
|
||
>.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE620BFFAE2DBFB6B5FE7F5B77" pageId="77" pageNumber="78" type="diagnosis">
|
||
<paragraph id="8B91C735620BFFAE2DBFB6B5FB19586B" blockId="77.[151,1437,978,2013]" pageId="77" pageNumber="78">
|
||
<emphasis id="B95A1B27620BFFAE2DBFB6B5FE2B59B3" bold="true" box="[199,460,1230,1257]" pageId="77" pageNumber="78">Diagnostic characters</emphasis>
|
||
. Dorsum pale yellow, pronotum with a pair of dark stripes, elytra with a pair of short basal stripes and a pair of well separate subapical spots; labrum slightly dilated to apex, apex strongly emarginate; antennae extended only slightly beyond elytral base; elytra intervals with very sparse fine punctures.
|
||
</paragraph>
|
||
<paragraph id="8B91C735620BFFAE2DBFB741FE7F5B77" blockId="77.[151,1437,978,2013]" pageId="77" pageNumber="78">
|
||
<emphasis id="B95A1B27620BFFAE2DBFB741FE81580E" bold="true" box="[199,358,1338,1364]" pageId="77" pageNumber="78">Comparisons</emphasis>
|
||
. The new species is different from all other species of the genus
|
||
<taxonomicName id="4C2EBCB6620BFFAE2931B740FB7D580E" box="[1097,1178,1339,1364]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27620BFFAE2931B740FB7D580E" box="[1097,1178,1339,1364]" italics="true" pageId="77" pageNumber="78">Parena</emphasis>
|
||
</taxonomicName>
|
||
by the apex of labrum strongly emarginate (
|
||
<figureCitation id="1315DBB0620BFFAE2CFCB724FE0B5823" box="[388,492,1375,1401]" captionStart="FIGURE 46" captionStartId="78.[151,250,1595,1619]" captionTargetBox="[159,1436,707,1563]" captionTargetId="figure-274@78.[151,1436,705,1572]" captionTargetPageId="78" captionText="FIGURE 46. Parena (Bothynoptera) emarginata sp. n. A Habitus of holotype. B Habitus of paratype. C Mouthparts of holotype. D Labels of holotype. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959031" httpUri="https://zenodo.org/record/7959031/files/figure.png" pageId="77" pageNumber="78">Fig. 46C</figureCitation>
|
||
). Several other species of the
|
||
<taxonomicName id="4C2EBCB6620BFFAE2E4FB724FC295822" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[823,974,1375,1400]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tripunctata">
|
||
<emphasis id="B95A1B27620BFFAE2E4FB724FC295822" box="[823,974,1375,1400]" italics="true" pageId="77" pageNumber="78">P. tripunctata</emphasis>
|
||
</taxonomicName>
|
||
group have the labrum apex very weakly emarginate, but the labrum apex is straight or slightly curved in all other species groups of the genus. The elytral color pattern of the new species is similar to some individuals of
|
||
<taxonomicName id="4C2EBCB6620BFFAE2E14B7DCFC06589A" authorityName="Hua. A" authorityYear="2002" box="[876,993,1447,1472]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="taiwana">
|
||
<emphasis id="B95A1B27620BFFAE2E14B7DCFC06589A" box="[876,993,1447,1472]" italics="true" pageId="77" pageNumber="78">P. taiwana</emphasis>
|
||
</taxonomicName>
|
||
with four spots. However, the new species can be easily distinguished from
|
||
<taxonomicName id="4C2EBCB6620BFFAE2F44B7B0FD5658BE" authorityName="Hua. A" authorityYear="2002" box="[572,689,1483,1508]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="taiwana">
|
||
<emphasis id="B95A1B27620BFFAE2F44B7B0FD5658BE" box="[572,689,1483,1508]" italics="true" pageId="77" pageNumber="78">P. taiwana</emphasis>
|
||
</taxonomicName>
|
||
by: labrum apex emarginate; pronotum with a pair of dark stripes; elytral intervals without dense punctures; and antennae shorter, extended beyond the elytra base by the length of only one antennomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE620BFFAD2DBFB44DFC905F3B" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78" type="description">
|
||
<paragraph id="8B91C735620BFFAE2DBFB44DFA7A5B0B" blockId="77.[151,1437,978,2013]" box="[199,1437,1590,1617]" pageId="77" pageNumber="78">
|
||
<emphasis id="B95A1B27620BFFAE2DBFB44DFEB65B0A" bold="true" box="[199,337,1590,1616]" pageId="77" pageNumber="78">Description</emphasis>
|
||
. Body length
|
||
<quantity id="4CD66AD0620BFFAE2C8AB44CFD9A5B0B" box="[498,637,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="6.7" metricValueMin="6.3" pageId="77" pageNumber="78" unit="mm" value="6.5" valueMax="6.7" valueMin="6.3">6.3–6.7 mm</quantity>
|
||
; body relatively narrow for the genus; dorsum without microsculpture.
|
||
</paragraph>
|
||
<paragraph id="8B91C735620BFFAD2DEFB421FC905F3B" blockId="77.[151,1437,978,2013]" lastBlockId="78.[151,1437,150,681]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
|
||
<emphasis id="B95A1B27620BFFAE2DEFB421FF075B2E" bold="true" box="[151,224,1626,1652]" pageId="77" pageNumber="78">Color.</emphasis>
|
||
Dorsum mostly pale yellow to light brown; head yellowish brown, vertex with indistinct dark brown patch, mouthparts and antennae yellow, apices of mandibles dark. Pronotum disc with a pair of dark stripes aside median line, not extended to anterior or posterior margins. Elytra with four black patches: a pair of short stripes slightly distant from elytra base, about one-fifth as long as elytra, extending from outer half of interval 4 to inner half interval 6; another pair of round spots near apical third of elytra, extending from outer half of interval 3 to inner half of interval 5. Scutellum, epipleura, legs and venter yellow.
|
||
<emphasis id="B95A1B27620BFFAE2FD2B574FD0D5A73" bold="true" box="[682,746,1807,1833]" pageId="77" pageNumber="78">Head</emphasis>
|
||
with sporadic fine punctures on vertex and occiput; eyes large and strongly prominent; tempora short, abruptly narrowed behind eyes, length of tempora plus neck-constriction approximately one-third of diameter of eye; postgenae with a pair of suborbital setae, as long as supraorbital setae. Antennae extended beyond elytral base by length of only one antennomere. Labrum slightly dilated to apex, apex strongly emarginate; mandibles short and wide; mentum with a pair of long median setae, lateral lobes short, apex narrow, inner margins strongly oblique, outer margins rounded, epilobes rather wide.
|
||
<emphasis id="B95A1B27620BFFAE2935B5B8FB215A87" bold="true" box="[1101,1222,1987,2013]" pageId="77" pageNumber="78">Pronotum</emphasis>
|
||
nearly rectangular, PW/PL = 1.35–1.37, about same width as head, PW/HW = 1.00–1.03, widest at anterior third, lateral explanations slightly wide; lateral margins fully rounded at anterior half and then narrowed to base with distinct sinuation before posterior angles; posterior angles weakly pointed outwards, forming rectangular angles with rounded apex; anterior margin nearly straight at middle; posterior margin oblique at sides; disc convex, with very fine transverse wrinkles.
|
||
<emphasis id="B95A1B276208FFAD2DEFB35CFF045C1B" bold="true" box="[151,227,295,321]" pageId="78" pageNumber="79">Elytra</emphasis>
|
||
weakly convex, slightly dilated to apex. Striae very shallowly incised at basal half, almost flat at apical half, with rows of fine punctures on basal five-sixth, punctures absent from apical sixth; intervals weakly convex, with very sparse fine punctures. Disc without depression, lateral sides slightly depressed near anterior third. Elytral basal pore present at base of stria 1; interval 3 with three discal setigerous pores: first one on a level much behind scutellar apex, adjacent to stria 3; second one slightly before middle, adjacent to stria 3; third one on apical eighth, adjacent to stria 2; interval 9 with 23–24 umbilicular pores. Apical truncation indistinct, evenly rounded, sutural angles indistinct.
|
||
<emphasis id="B95A1B276208FFAD2D8FB384FEA05F43" bold="true" box="[247,327,511,537]" pageId="78" pageNumber="79">Venter</emphasis>
|
||
. In females, apex of abdominal sternite VII straight, with two setae on each side.
|
||
<emphasis id="B95A1B276208FFAD29A9B384FA715F42" bold="true" box="[1233,1430,510,537]" pageId="78" pageNumber="79">Female genitalia</emphasis>
|
||
. Gonocoxite II of ovipositor rounded rectangle, basal width much greater than length, apex straight, with five or six very short ensiform setae nearly evenly arranged. Male unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE6208FFAD2DBFB011FEA45FF3" pageId="78" pageNumber="79" type="distribution">
|
||
<paragraph id="8B91C7356208FFAD2DBFB011FEA45FF3" blockId="78.[151,1437,150,681]" pageId="78" pageNumber="79">
|
||
<emphasis id="B95A1B276208FFAD2DBFB011FEBE5FDE" bold="true" box="[199,345,618,644]" pageId="78" pageNumber="79">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1315DBB06208FFAD2C10B010FE545FDF" box="[360,435,619,645]" captionStart="MAP 8" captionStartId="65.[152,211,1703,1727]" captionTargetBox="[180,1408,802,1680]" captionTargetId="figure-242@65.[180,1408,802,1680]" captionText="MAP 8. Distributions for species in the Parena (Bothynoptera) tripunctata group." figureDoi="http://doi.org/10.5281/zenodo.7963675" httpUri="https://zenodo.org/record/7963675/files/figure.png" pageId="78" pageNumber="79">Map 8</figureCitation>
|
||
, dark green).
|
||
<collectingCountry id="F33987A56208FFAD2F36B010FD765FDF" box="[590,657,619,645]" name="China" pageId="78" pageNumber="79">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49EA09D76208FFAD2FE7B010FD195FDF" box="[671,766,619,645]" country="China" name="Sichuan" pageId="78" pageNumber="79">Sichuan</collectingRegion>
|
||
,
|
||
<collectingRegion id="49EA09D76208FFAD2E71B010FCB85FDF" box="[777,863,619,645]" country="China" name="Xizang" pageId="78" pageNumber="79">Xizang</collectingRegion>
|
||
). Only known from
|
||
<specimenCount id="9D280CBC6208FFAD2938B010FB015FDF" box="[1088,1254,619,645]" pageId="78" pageNumber="79" type="generic">two specimens</specimenCount>
|
||
from well separated localities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5197BD6208FFAD2DEFB440FD905B2D" ID-DOI="http://doi.org/10.5281/zenodo.7959031" ID-Zenodo-Dep="7959031" httpUri="https://zenodo.org/record/7959031/files/figure.png" pageId="78" pageNumber="79" startId="78.[151,250,1595,1619]" targetBox="[159,1436,707,1563]" targetPageId="78" targetType="figure">
|
||
<paragraph id="8B91C7356208FFAD2DEFB440FD905B2D" blockId="78.[151,1436,1595,1656]" pageId="78" pageNumber="79">
|
||
<emphasis id="B95A1B276208FFAD2DEFB440FEFD5B09" bold="true" box="[151,282,1595,1619]" pageId="78" pageNumber="79">FIGURE 46</emphasis>
|
||
.
|
||
<taxonomicName id="4C2EBCB66208FFAD2C5FB447FD695B09" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[295,654,1595,1620]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="emarginata" status="sp. nov." subGenus="Bothynoptera">
|
||
<emphasis id="B95A1B276208FFAD2C5FB447FE965B09" box="[295,369,1596,1619]" italics="true" pageId="78" pageNumber="79">Parena</emphasis>
|
||
(
|
||
<emphasis id="B95A1B276208FFAD2CF9B440FDEC5B09" box="[385,523,1595,1619]" italics="true" pageId="78" pageNumber="79">Bothynoptera</emphasis>
|
||
)
|
||
<emphasis id="B95A1B276208FFAD2F62B447FD695B09" box="[538,654,1596,1619]" italics="true" pageId="78" pageNumber="79">emarginata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B276208FFAD2FEDB447FD0F5B09" bold="true" box="[661,744,1595,1620]" pageId="78" pageNumber="79">
|
||
<taxonomicNameLabel id="A269A65C6208FFAD2FEDB447FD285B0E" box="[661,719,1596,1620]" pageId="78" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
Habitus of holotype.
|
||
<emphasis id="B95A1B276208FFAD2EBFB447FC3F5B0E" bold="true" box="[967,984,1596,1620]" pageId="78" pageNumber="79">B</emphasis>
|
||
Habitus of paratype.
|
||
<emphasis id="B95A1B276208FFAD29CDB440FB205B09" bold="true" box="[1205,1223,1595,1619]" pageId="78" pageNumber="79">C</emphasis>
|
||
Mouthparts of holotype.
|
||
<emphasis id="B95A1B276208FFAD2DB7B41BFF065B22" bold="true" box="[207,225,1632,1656]" pageId="78" pageNumber="79">D</emphasis>
|
||
Labels of holotype. Scale bars = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33494BE6208FFAD2DBFB4DDFC525BBE" pageId="78" pageNumber="79" type="etymology">
|
||
<paragraph id="8B91C7356208FFAD2DBFB4DDFC525BBE" blockId="78.[151,1436,1702,1836]" pageId="78" pageNumber="79">
|
||
<emphasis id="B95A1B276208FFAD2DBFB4DDFEAF5B9A" bold="true" box="[199,328,1702,1728]" pageId="78" pageNumber="79">Etymology</emphasis>
|
||
. The scientific name of the new species "
|
||
<taxonomicName id="4C2EBCB66208FFAD2E65B4DCFC7A5B9A" authorityName="Shi & Liang" authorityYear="2023" box="[797,925,1703,1728]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="emarginata">
|
||
<emphasis id="B95A1B276208FFAD2E65B4DCFC7A5B9A" box="[797,925,1703,1728]" italics="true" pageId="78" pageNumber="79">emarginata</emphasis>
|
||
</taxonomicName>
|
||
" is derived from Latin referring to its labrum apex strongly emarginate, which is the unique character of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE6208FFAD2DBFB495FE625A76" pageId="78" pageNumber="79" type="discussion">
|
||
<paragraph id="8B91C7356208FFAD2DBFB495FE625A76" blockId="78.[151,1436,1702,1836]" pageId="78" pageNumber="79">
|
||
<emphasis id="B95A1B276208FFAD2DBFB495FED35A52" bold="true" box="[199,308,1774,1800]" pageId="78" pageNumber="79">Remarks</emphasis>
|
||
. The
|
||
<typeStatus id="549579976208FFAD2C0EB495FE3F5A52" box="[374,472,1774,1800]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
|
||
was collected in a sparse riparian forest, about six meters high in the canopy, using a long-stick sweep net.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |