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<document id="F1E2CB3FAF8BF193DC1A6393997B661E" ID-DOI="10.11646/zootaxa.3620.2.2" ID-GBIF-Dataset="4b142fea-210e-4145-b876-38b099dfa650" ID-ISSN="1175-5326" ID-Zenodo-Dep="215672" ID-ZooBank="611E561A-E74D-4145-81C7-AD87E143DDC3" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460579426272" checkinUser="plazi" docAuthor="Zhang, Haomiao &amp; Tong, Xiaoli" docDate="2013" docId="F70187F9FE219578FF270A4430822FB4" docLanguage="en" docName="zt03620p244.pdf" docOrigin="Zootaxa 3620 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlorogomphus usudai Ishida 1996" docType="treatment" docVersion="8" lastPageNumber="236" masterDocId="0B38FF81FE2B9575FFB00C6832292C74" masterDocTitle="Descriptions of the final instar larvae of seven Chinese Chlorogomphidae species, with taxonomic notes on adults (Odonata: Anisoptera)" masterLastPageNumber="244" masterPageNumber="223" pageNumber="233" updateTime="1698323969956" updateUser="plazi">
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<mods:title id="255188FCF649F1DA00753BAF2DB811F8">Descriptions of the final instar larvae of seven Chinese Chlorogomphidae species, with taxonomic notes on adults (Odonata: Anisoptera)</mods:title>
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<mods:namePart id="361131F1D241E80F268F13902BDB5798">Zhang, Haomiao</mods:namePart>
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<treatment id="F70187F9FE219578FF270A4430822FB4" ID-DOI="http://doi.org/10.5281/zenodo.6153033" ID-GBIF-Taxon="119560841" ID-Zenodo-Dep="6153033" LSID="urn:lsid:plazi:treatment:F70187F9FE219578FF270A4430822FB4" httpUri="http://treatment.plazi.org/id/F70187F9FE219578FF270A4430822FB4" lastPageId="13" lastPageNumber="236" pageId="10" pageNumber="233">
<subSubSection id="37B26564FE21957FFF270A44301A2ABF" pageId="10" pageNumber="233" type="nomenclature">
<paragraph id="7F1736EFFE21957FFF270A44307B2A31" blockId="10.[151,594,1579,1638]" box="[151,594,1579,1606]" pageId="10" pageNumber="233">
<heading id="245F8183FE21957FFF270A44307B2A31" bold="true" box="[151,594,1579,1606]" fontSize="11" level="1" pageId="10" pageNumber="233" reason="1">
<taxonomicName id="B8A84D6CFE21957FFF270A44307B2A31" ID-CoL="5XY7S" authority="Ishida, 1996" authorityName="Ishida" authorityYear="1996" box="[151,594,1579,1606]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE21957FFF270A44307B2A31" bold="true" box="[151,594,1579,1606]" pageId="10" pageNumber="233">
<emphasis id="4DDCEAFDFE21957FFF270A44339A2A32" bold="true" box="[151,435,1580,1606]" italics="true" pageId="10" pageNumber="233">Chlorogomphus usudai</emphasis>
Ishida, 1996
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7F1736EFFE21957FFF270A2633402A12" blockId="10.[151,594,1579,1638]" box="[151,361,1614,1638]" pageId="10" pageNumber="233">
<figureCitation id="E7932A6AFE21957FFF270A2633132A12" box="[151,314,1614,1638]" captionStart="FIGURES 31 35" captionStartId="12.[151,264,1977,2000]" captionTargetBox="[203,1391,198,1929]" captionTargetId="figure@12.[182,1406,193,1954]" captionTargetPageId="12" captionText="FIGURES 31 35. Larva of Chlorogomphus usudai: (31) right mandible; (32) left mandible; (33) prementum, dorsal view; (34) bifid branches of frontal margin of prementum; (35) frontal shelf, dorsal view." httpUri="https://zenodo.org/record/215679/files/figure.png" pageId="10" pageNumber="233">Figures 3135</figureCitation>
,
<figureCitation id="E7932A6AFE21957FFEF60A26334C2A12" box="[326,357,1614,1638]" captionStart="FIGURES 54 59" captionStartId="19.[151,264,1925,1948]" captionTargetBox="[301,1259,204,1879]" captionTargetId="figure@19.[295,1291,193,1904]" captionTargetPageId="19" captionText="FIGURES 54 59. Larvae of Chlorogomphidae: (54) Chlorogomphus usudai; (55 58) Chlorogomphus yokoii; (59) Chloropetalia soarer." httpUri="https://zenodo.org/record/215684/files/figure.png" pageId="10" pageNumber="233">54</figureCitation>
.
</paragraph>
<paragraph id="7F1736EFFE21957FFF270AFF301A2ABF" blockId="10.[151,957,1686,1800]" pageId="10" pageNumber="233">
<taxonomicName id="B8A84D6CFE21957FFF270AFF30052AD8" authority="Ishida, 1996: 51" authorityName="Ishida" authorityPageNumber="51" authorityYear="1996" box="[151,556,1686,1709]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE21957FFF270AFF33AB2AD9" box="[151,386,1687,1709]" italics="true" pageId="10" pageNumber="233">Chlorogomphus usudai</emphasis>
Ishida, 1996: 51
</taxonomicName>
54,
<typeStatus id="A013884DFE21957FFDD00AFF30BB2AD8" box="[608,658,1687,1708]" pageId="10" pageNumber="233">Type</typeStatus>
Locality: Tongza, Hainan.
<emphasis id="4DDCEAFDFE21957FFF270ADD33022ABF" box="[151,299,1717,1739]" italics="true" pageId="10" pageNumber="233">Aurorachlorus</emphasis>
<taxonomicName id="B8A84D6CFE21957FFE810ADD303A2ABF" authority="Chao, 1999: 4" authorityName="Chao" authorityPageNumber="4" authorityYear="1999" box="[305,531,1717,1739]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE21957FFE810ADD33572ABF" box="[305,382,1717,1739]" italics="true" pageId="10" pageNumber="233">usudai:</emphasis>
Chao, 1999: 4
</taxonomicName>
5.
</paragraph>
</subSubSection>
<subSubSection id="37B26564FE21957FFF270ABC312E2B7C" pageId="10" pageNumber="233" type="description">
<paragraph id="7F1736EFFE21957FFF270ABC312E2B7C" blockId="10.[151,957,1686,1800]" pageId="10" pageNumber="233">
<taxonomicName id="B8A84D6CFE21957FFF270ABC30892A9D" authority="Wilson &amp; Reels, 2001: 172" authorityName="Wilson &amp; Reels" authorityPageNumber="172" authorityYear="2001" box="[151,672,1747,1770]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="icarus">
<emphasis id="4DDCEAFDFE21957FFF270ABC33572A9E" box="[151,382,1748,1770]" italics="true" pageId="10" pageNumber="233">Chlorogomphus icarus</emphasis>
: Wilson &amp; Reels, 2001: 172
</taxonomicName>
173, figs. 6775. Syn. nov.
<taxonomicName id="B8A84D6CFE21957FFF270A9A30F32B7C" authority="Wilson, 2002: 69" authorityName="Wilson" authorityPageNumber="69" authorityYear="2002" box="[151,730,1778,1800]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="usudai" subGenus="Orogomphus">
<emphasis id="4DDCEAFDFE21957FFF270A9A300F2B7C" box="[151,550,1778,1800]" italics="true" pageId="10" pageNumber="233">Chlorogomphus (Orogomphus) usudai:</emphasis>
Wilson, 2002: 69
</taxonomicName>
70.
</paragraph>
</subSubSection>
<subSubSection id="37B26564FE219578FF270B5D37552E2C" lastPageId="13" lastPageNumber="236" pageId="10" pageNumber="233" type="materials_examined">
<paragraph id="7F1736EFFE21957FFF270B5D306F2BE1" blockId="10.[151,1436,1844,1977]" pageId="10" pageNumber="233">
<emphasis id="4DDCEAFDFE21957FFF270B5D33A92B3A" bold="true" box="[151,384,1845,1870]" pageId="10" pageNumber="233">Material examined.</emphasis>
final stage larvae: 13,
<date id="0B16102FFE21957FFD360B5D31322B39" box="[646,795,1845,1869]" pageId="10" pageNumber="233" value="2005-08-30">30.VIII.2005</date>
, Yinggeling Nature Reserve (18°58ʹ76ʺN, 109°24ʹ27ʺE), Hainan Province, Liusheng Chen leg.; 13,
<date id="0B16102FFE21957FFD340B31312B2B05" box="[644,770,1881,1905]" pageId="10" pageNumber="233" value="2009-04-23">23.IV.2009</date>
, Diaoluoshan Nature Reserve (18°43ʹ37ʺN, 109°52ʹ20ʺE), Hainan Province, Haomiao Zhang leg.
</paragraph>
<paragraph id="7F1736EFFE21957FFF790BC936472BCD" blockId="10.[151,1436,1844,1977]" box="[201,1134,1953,1977]" pageId="10" pageNumber="233">Small chlorogomphid larva, ground color brownish yellow with brown maculation.</paragraph>
<caption id="2BD76667FE20957EFF270BDB31C92B9C" httpUri="https://zenodo.org/record/215678/files/figure.png" pageId="11" pageNumber="234" targetBox="[180,1410,203,1922]" targetPageId="11">
<paragraph id="7F1736EFFE20957EFF270BDB31C92B9C" blockId="11.[151,1436,1971,2025]" pageId="11" pageNumber="234">
<emphasis id="4DDCEAFDFE20957EFF270BDB33012BBD" bold="true" box="[151,296,1971,1994]" pageId="11" pageNumber="234">FIGURES 26</emphasis>
<emphasis id="4DDCEAFDFE20957EFE850BDC337C2BBD" bold="true" box="[309,341,1972,1993]" pageId="11" pageNumber="234">30.</emphasis>
Larva of
<taxonomicName id="B8A84D6CFE20957EFE0C0BDC30972BBE" box="[444,702,1972,1994]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="234" phylum="Arthropoda" rank="species" species="shanicus">
<emphasis id="4DDCEAFDFE20957EFE0C0BDC30972BBE" box="[444,702,1972,1994]" italics="true" pageId="11" pageNumber="234">Chlorogomphus shanicus</emphasis>
</taxonomicName>
: (26) right mandible; (27) left mandible; (28) prementum, dorsal view; (29) bifid branches of frontal margin of prementum; (30) frontal shelf, dorsal view.
</paragraph>
</caption>
<caption id="2BD76667FE279579FF270BD131862B9A" httpUri="https://zenodo.org/record/215679/files/figure.png" pageId="12" pageNumber="235" targetBox="[203,1391,198,1929]" targetPageId="12">
<paragraph id="7F1736EFFE279579FF270BD131862B9A" blockId="12.[151,1435,1977,2030]" pageId="12" pageNumber="235">
<emphasis id="4DDCEAFDFE279579FF270BD1330F2BBB" bold="true" box="[151,294,1977,2000]" pageId="12" pageNumber="235">FIGURES 31</emphasis>
<emphasis id="4DDCEAFDFE279579FE830BD2337A2BBB" bold="true" box="[307,339,1978,1999]" pageId="12" pageNumber="235">35.</emphasis>
Larva of
<taxonomicName id="B8A84D6CFE279579FE040BD130892BBB" box="[436,672,1977,1999]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="235" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE279579FE040BD130892BBB" box="[436,672,1977,1999]" italics="true" pageId="12" pageNumber="235">Chlorogomphus usudai</emphasis>
</taxonomicName>
: (31) right mandible; (32) left mandible; (33) prementum, dorsal view; (34) bifid branches of frontal margin of prementum; (35) frontal shelf, dorsal view.
</paragraph>
</caption>
<paragraph id="7F1736EFFE269578FF790CF036532CA0" blockId="13.[151,1436,151,212]" pageId="13" pageNumber="236">
<emphasis id="4DDCEAFDFE269578FF790CF033202CC4" bold="true" box="[201,265,152,176]" pageId="13" pageNumber="236">Head</emphasis>
Labrum and clypeus pale yellow. Labrum and postclypeus with brown central spots. Frontal shelf similar to that of
<taxonomicName id="B8A84D6CFE269578FEE80CD533E42CA0" box="[344,461,189,212]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="4DDCEAFDFE269578FEE80CD533E42CA0" box="[344,461,189,212]" italics="true" pageId="13" pageNumber="236">C. nasutus</emphasis>
</taxonomicName>
(
<figureCitation id="E7932A6AFE269578FE6D0CD430182CA0" box="[477,561,188,212]" captionStart="FIGURES 31 35" captionStartId="12.[151,264,1977,2000]" captionTargetBox="[203,1391,198,1929]" captionTargetId="figure@12.[182,1406,193,1954]" captionTargetPageId="12" captionText="FIGURES 31 35. Larva of Chlorogomphus usudai: (31) right mandible; (32) left mandible; (33) prementum, dorsal view; (34) bifid branches of frontal margin of prementum; (35) frontal shelf, dorsal view." httpUri="https://zenodo.org/record/215679/files/figure.png" pageId="13" pageNumber="236">Fig. 35</figureCitation>
). Mandibles as in
<figureCitation id="E7932A6AFE269578FD4D0CD431AC2CA0" box="[765,901,188,212]" captionStart="FIGURES 31 35" captionStartId="12.[151,264,1977,2000]" captionTargetBox="[203,1391,198,1929]" captionTargetId="figure@12.[182,1406,193,1954]" captionTargetPageId="12" captionText="FIGURES 31 35. Larva of Chlorogomphus usudai: (31) right mandible; (32) left mandible; (33) prementum, dorsal view; (34) bifid branches of frontal margin of prementum; (35) frontal shelf, dorsal view." httpUri="https://zenodo.org/record/215679/files/figure.png" pageId="13" pageNumber="236">Figs. 3132</figureCitation>
. Mandibular formula:
</paragraph>
<paragraph id="7F1736EFFE269578FF790DE5370B2E64" blockId="13.[151,1436,397,961]" pageId="13" pageNumber="236">
Left mandible with 4 rounded molar crest,
<emphasis id="4DDCEAFDFE269578FD030DE630E82DD1" box="[691,705,398,421]" italics="true" pageId="13" pageNumber="236">a</emphasis>
complanate,
<emphasis id="4DDCEAFDFE269578FCEA0DE531412DD1" box="[858,872,397,421]" italics="true" pageId="13" pageNumber="236">b</emphasis>
obtuse,
<emphasis id="4DDCEAFDFE269578FC790DE631FE2DD1" box="[969,983,398,421]" italics="true" pageId="13" pageNumber="236">a</emphasis>
&gt;
<emphasis id="4DDCEAFDFE269578FC570DE531DC2DD1" box="[999,1013,397,421]" italics="true" pageId="13" pageNumber="236">b</emphasis>
; incisors pointed, 4&gt;3&gt;2&gt;1&gt;2; right mandible without molar crest,
<emphasis id="4DDCEAFDFE269578FE4B0DDA30202DBD" box="[507,521,434,457]" italics="true" pageId="13" pageNumber="236">a</emphasis>
and
<emphasis id="4DDCEAFDFE269578FDF60DD9307D2DBD" box="[582,596,433,457]" italics="true" pageId="13" pageNumber="236">b</emphasis>
pointed,
<emphasis id="4DDCEAFDFE269578FD740DDA30FB2DBD" box="[708,722,434,457]" italics="true" pageId="13" pageNumber="236">a</emphasis>
&gt;
<emphasis id="4DDCEAFDFE269578FD520DD930D92DBD" box="[738,752,433,457]" italics="true" pageId="13" pageNumber="236">b</emphasis>
; teeth of the incisors pointed, 4&gt;1&gt;3&gt;2, additional tooth
<emphasis id="4DDCEAFDFE269578FA3F0DDA37B22DBD" box="[1423,1435,434,457]" italics="true" pageId="13" pageNumber="236">y</emphasis>
present on the right mandible. Labial mask as in
<figureCitation id="E7932A6AFE269578FD040DBD312C2D99" box="[692,773,469,493]" captionStart="FIGURES 31 35" captionStartId="12.[151,264,1977,2000]" captionTargetBox="[203,1391,198,1929]" captionTargetId="figure@12.[182,1406,193,1954]" captionTargetPageId="12" captionText="FIGURES 31 35. Larva of Chlorogomphus usudai: (31) right mandible; (32) left mandible; (33) prementum, dorsal view; (34) bifid branches of frontal margin of prementum; (35) frontal shelf, dorsal view." httpUri="https://zenodo.org/record/215679/files/figure.png" pageId="13" pageNumber="236">Fig. 33</figureCitation>
. Prementum with row of 5 long outer setae and 35 smaller inner setae on each side. Distal process as in
<figureCitation id="E7932A6AFE269578FD3F0D9030C82E64" box="[655,737,504,528]" captionStart="FIGURES 31 35" captionStartId="12.[151,264,1977,2000]" captionTargetBox="[203,1391,198,1929]" captionTargetId="figure@12.[182,1406,193,1954]" captionTargetPageId="12" captionText="FIGURES 31 35. Larva of Chlorogomphus usudai: (31) right mandible; (32) left mandible; (33) prementum, dorsal view; (34) bifid branches of frontal margin of prementum; (35) frontal shelf, dorsal view." httpUri="https://zenodo.org/record/215679/files/figure.png" pageId="13" pageNumber="236">Fig. 34</figureCitation>
; lobes angled and each lobe bent slightly outwards.
</paragraph>
<paragraph id="7F1736EFFE269578FF790E7537552E2C" blockId="13.[151,1436,397,961]" pageId="13" pageNumber="236">
<emphasis id="4DDCEAFDFE269578FF790E75330B2E41" bold="true" box="[201,290,541,565]" pageId="13" pageNumber="236">Thorax</emphasis>
Brownish yellow with long hair-like setae; pronotum with pair of black spots close to anterior margin; wing pads brownish yellow and broad, reaching middle of S6. femora and tibiae with black stripes submedially.
</paragraph>
</subSubSection>
<subSubSection id="37B26564FE269578FF790E0C30AE2F79" pageId="13" pageNumber="236" type="description">
<paragraph id="7F1736EFFE269578FF790E0C36642EB1" blockId="13.[151,1436,397,961]" pageId="13" pageNumber="236">
<emphasis id="4DDCEAFDFE269578FF790E0C33172E09" bold="true" box="[201,318,612,637]" pageId="13" pageNumber="236">Abdomen</emphasis>
Brownish yellow with brown spots. S14 with two pairs of inconspicuous spots; S56 largely black centrally, with pair of scaphoid spots laterally; S79 with pair of oblique spots centrally and pair of scaphoid spots laterally, less developed than those on S56; S10 and anal appendages without spots.
</paragraph>
<paragraph id="7F1736EFFE269578FF790EB930AE2F79" blockId="13.[151,1436,397,961]" pageId="13" pageNumber="236">
<emphasis id="4DDCEAFDFE269578FF790EB933E52E9D" bold="true" box="[201,460,720,745]" pageId="13" pageNumber="236">Measurements (mm).</emphasis>
Total length 33.033.5, head width 8.08.2, hind wing length 8.5, width 4.0, hind femur 6.0, prementum length 7.0, width 7.5.
</paragraph>
</subSubSection>
<subSubSection id="37B26564FE269578FF770F7030352F44" box="[199,540,792,817]" pageId="13" pageNumber="236" type="distribution">
<paragraph id="7F1736EFFE269578FF770F7030352F44" blockId="13.[151,1436,397,961]" box="[199,540,792,817]" pageId="13" pageNumber="236">
<emphasis id="4DDCEAFDFE269578FF770F7033492F45" bold="true" box="[199,352,792,817]" pageId="13" pageNumber="236">Distribution.</emphasis>
<collectingCountry id="07BF767FFE269578FED70F7033832F44" box="[359,426,792,816]" name="China" pageId="13" pageNumber="236">China</collectingCountry>
(Hainan).
</paragraph>
</subSubSection>
<subSubSection id="37B26564FE269578FF770F5530822FB4" pageId="13" pageNumber="236" type="discussion">
<paragraph id="7F1736EFFE269578FF770F5530822FB4" blockId="13.[151,1436,397,961]" pageId="13" pageNumber="236">
<emphasis id="4DDCEAFDFE269578FF770F5533122F21" bold="true" box="[199,315,829,853]" pageId="13" pageNumber="236">Remarks.</emphasis>
This is a relatively small larva with very broad wing pads. It is only known from Hainan island and is widespread in the montane area. Two
<taxonomicName id="B8A84D6CFE269578FDE80F0931222F0D" box="[600,779,865,889]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="4DDCEAFDFE269578FDE80F0931222F0D" box="[600,779,865,889]" italics="true" pageId="13" pageNumber="236">Chlorogomphus</emphasis>
</taxonomicName>
species were recorded from Hainan island,
<taxonomicName id="B8A84D6CFE269578FB460F0937452F0D" box="[1270,1388,865,889]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="4DDCEAFDFE269578FB460F0937452F0D" box="[1270,1388,865,889]" italics="true" pageId="13" pageNumber="236">C. gracilis</emphasis>
</taxonomicName>
and
<taxonomicName id="B8A84D6CFE269578FF270FEE332F2FE9" box="[151,262,901,925]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE269578FF270FEE332F2FE9" box="[151,262,901,925]" italics="true" pageId="13" pageNumber="236">C. usudai</emphasis>
</taxonomicName>
, and larva of
<taxonomicName id="B8A84D6CFE269578FE1D0FEE30352FE9" box="[429,540,901,925]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE269578FE1D0FEE30352FE9" box="[429,540,901,925]" italics="true" pageId="13" pageNumber="236">C. usudai</emphasis>
</taxonomicName>
was confirmed from the streams in Yinggeling Reserve, where
<taxonomicName id="B8A84D6CFE269578FAB50FEE37562FE9" box="[1285,1407,901,925]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="4DDCEAFDFE269578FAB50FEE37562FE9" box="[1285,1407,901,925]" italics="true" pageId="13" pageNumber="236">C. gracilis</emphasis>
</taxonomicName>
is absent and the adults of
<taxonomicName id="B8A84D6CFE269578FE160FC1303B2FB5" box="[422,530,937,961]" class="Insecta" family="Cordulegastridae" genus="Chlorogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="236" phylum="Arthropoda" rank="species" species="usudai">
<emphasis id="4DDCEAFDFE269578FE160FC1303B2FB5" box="[422,530,937,961]" italics="true" pageId="13" pageNumber="236">C. usudai</emphasis>
</taxonomicName>
are common.
</paragraph>
</subSubSection>
</treatment>
</document>