treatments-xml/data/F7/01/87/F70187ECC227FFCC5282FEAEFE00FED0.xml

<document id="F7ABFB68F7EBD90745DC6084B88B2BB6" ID-DOI="10.1649/0010-065X-73.4.1067" ID-ISSN="1938-4394" ID-Zenodo-Dep="10113974" ID-ZooBank="urn:lsid:zoobank.org:pub:50E315DC-95A6-426D-8EE3-EE69A360575E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630340349733" checkinUser="valdenar" docAuthor="Mora-Aguilar, Eder F. &amp; Delgado, Leonardo" docDate="2019" docId="F70187ECC227FFCC5282FEAEFE00FED0" docLanguage="en" docName="0010-065X-73.4.1067.pdf" docOrigin="The Coleopterists Bulletin 73 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-73.4.1067" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Canthidium kohlmanni Mora-Aguilar and Delgado, new species" docType="treatment" docVersion="3" lastPageNumber="1070" masterDocId="0B38FF94C225FFC95249FFFCFF82FFDC" masterDocTitle="A New Species of Canthidium Erichson (Coleoptera: Scarabaeidae: Scarabaeinae) from the Endemic Biota of Los Chimalapas, Oaxaca, Mexico" masterLastPageNumber="1074" masterPageNumber="1067" pageNumber="1068" updateTime="1699723958763" updateUser="ExternalLinkService">
<mods:mods id="559E7BBD63299F9608A7D35A9C7E41D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="007032A0A3656505BF6DADA356BF6711">
<mods:title id="32A0E5C1546A3262826D2CA9ED0FE51C">A New Species of Canthidium Erichson (Coleoptera: Scarabaeidae: Scarabaeinae) from the Endemic Biota of Los Chimalapas, Oaxaca, Mexico</mods:title>
</mods:titleInfo>
<mods:name id="A974FC9B4691A59FBF5506E4A9E09665" type="personal">
<mods:role id="62CC42D61101CEAA2DEC02DDDD4334D8">
<mods:roleTerm id="8756182084CD93D38FCB9271AF1788F4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB562FC8E3053E7DFBD336BCAC36D952">Mora-Aguilar, Eder F.</mods:namePart>
</mods:name>
<mods:name id="072A488721172174214DDB5675920F13" type="personal">
<mods:role id="682D134CD8186280C1D9C9D01D7C5FE6">
<mods:roleTerm id="61AFD608B4D1D0F89CA1507C4CA4F296">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="352235B75FAB5B7E2862B9F63D63DF57">Delgado, Leonardo</mods:namePart>
</mods:name>
<mods:typeOfResource id="FFB0F92CDCD22713C556BD09F97A213E">text</mods:typeOfResource>
<mods:relatedItem id="7AD0846456248244830D1ECE0B558A9E" type="host">
<mods:titleInfo id="04A4425E19D6EF0F31B0DB2B01649230">
<mods:title id="742831BA59D246BD3948CA16B747DB45">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="F4774EA5E13E8AEA9D0C623955ADAF6A">
<mods:date id="81BEA2D186B44F52A3C3EBE15F423C39">2019</mods:date>
<mods:detail id="A9925A7E9B1BC83CD9B93148EE0D4C3E" type="pubDate">
<mods:number id="DFC75BCAF618718E83E87BBA79FD5E2D">2019-12-19</mods:number>
</mods:detail>
<mods:detail id="100B83E18A9DF2DB274077161EAD71D4" type="volume">
<mods:number id="13C5DBB69F095D077A3C67A55B859E7B">73</mods:number>
</mods:detail>
<mods:detail id="5B14CC5EDDCB1E44F226F860B6BF8AAB" type="issue">
<mods:number id="66EC99347497118B20BDCA0681072986">4</mods:number>
</mods:detail>
<mods:extent id="A9982C6B618FDECE4220D5A134AAB623" unit="page">
<mods:start id="023D90A8DEEF7990AE2BA76C909EC1A8">1067</mods:start>
<mods:end id="4F0D48CD971FEB995BB571269CC74D07">1074</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="81511518EC46C1EB474BF5423A43539D">
<mods:url id="357A9B2927AD62465A625695BFB8E01C">http://dx.doi.org/10.1649/0010-065x-73.4.1067</mods:url>
</mods:location>
<mods:classification id="10D8D4B0D69B47C4631DEA5E66C9C176">journal article</mods:classification>
<mods:identifier id="0DB11EB9F375A570C82044D1FD4F39B7" type="DOI">10.1649/0010-065X-73.4.1067</mods:identifier>
<mods:identifier id="6DCF41E9BB4C35D5E704DA7E61EC8D0C" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="42C1D221391F0EED776FBB42D10A2B52" type="Zenodo-Dep">10113974</mods:identifier>
<mods:identifier id="5A68D24FA18029ECDFE51D9CEC7A4AFD" type="ZooBank">urn:lsid:zoobank.org:pub:50E315DC-95A6-426D-8EE3-EE69A360575E</mods:identifier>
</mods:mods>
<treatment id="F70187ECC227FFCC5282FEAEFE00FED0" LSID="urn:lsid:plazi:treatment:F70187ECC227FFCC5282FEAEFE00FED0" httpUri="http://treatment.plazi.org/id/F70187ECC227FFCC5282FEAEFE00FED0" lastPageId="5" lastPageNumber="1070" pageId="2" pageNumber="1068">
<subSubSection id="37B26571C227FFCB5282FEAEFE3DFE0D" pageId="2" pageNumber="1068" type="nomenclature">
<paragraph id="7F1736FAC227FFCB5282FEAEFD8EFE5D" blockId="2.[178,611,338,466]" pageId="2" pageNumber="1068">
<heading id="245F8196C227FFCB5282FEAEFD8EFE5D" centered="true" fontSize="9" level="2" pageId="2" pageNumber="1068" reason="2">
<emphasis id="4DDCEAE8C227FFCB5282FEAEFD8EFE5D" bold="true" pageId="2" pageNumber="1068">
<taxonomicName id="B8A84D79C227FFCB5282FEAEFE09FE5D" ID-CoL="QN9Z" authority="Mora-Aguilar and Delgado" authorityName="Mora-Aguilar and Delgado" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="kohlmanni" status="sp. nov.">
<emphasis id="4DDCEAE8C227FFCB5282FEAEFE37FEBB" bold="true" box="[203,437,338,359]" italics="true" pageId="2" pageNumber="1068">Canthidium kohlmanni</emphasis>
Mora-Aguilar and Delgado
</taxonomicName>
, 
<taxonomicNameLabel id="56EF5793C227FFCB53DEFE90FD8EFE5D" box="[407,524,364,385]" pageId="2" pageNumber="1068" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7F1736FAC227FFCB529DFE7BFE3DFE0D" blockId="2.[178,611,338,466]" pageId="2" pageNumber="1068">
Zoobank.org/ 
<uri id="0B393AF8C227FFCB5313FE7BFDE1FE6B" pageId="2" pageNumber="1068">
urn:lsid:zoobank.org:act: 
<uuid id="0B0E0C2FC227FFCB52FBFE5EFDE1FE6B" box="[178,611,418,439]" pageId="2" pageNumber="1068">0A277144-6EF2-4AE6-9E14-323653931294</uuid>
</uri>
(
<figureCitation id="E7932A7FC227FFCB5314FE40FE3AFE0D" box="[349,440,444,466]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="37B26571C227FFCB52E6FE1BFDC3FD47" pageId="2" pageNumber="1068" type="materials_examined">
<paragraph id="7F1736FAC227FFCB52E6FE1BFDC3FD47" blockId="2.[151,639,486,1730]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB52E6FE1BFEBFFE20" bold="true" box="[175,317,487,508]" pageId="2" pageNumber="1068">Type Material.</emphasis>
<materialsCitation id="CFC03CA7C227FFCB530AFE1BFD98FDBA" collectingDate="2013-07-08" collectorName="L. Delgado" country="Mexico" county="San Miguel Chimalapa" elevation="1250" latitude="16.7159" location="El Puerto" longLatPrecision="7" longitude="-94.1076" municipality="Benito Juarez" pageId="2" pageNumber="1068" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<typeStatus id="A0138858C227FFCB530AFE1BFE14FE20" box="[323,406,487,508]" pageId="2" pageNumber="1068" type="holotype">Holotype</typeStatus>
male labeled “ 
<collectingCountry id="07BF766AC227FFCB5057FE1BFDF9FE20" box="[542,635,487,508]" name="Mexico" pageId="2" pageNumber="1068">MEXICO</collectingCountry>
: 
<collectingRegion id="BD6CF818C227FFCB52DEFDFDFF60FDCA" box="[151,226,513,534]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="1068">Oaxaca</collectingRegion>
, 
<collectingCounty id="96764E76C227FFCB52BBFDFDFE5DFDCA" box="[242,479,513,534]" pageId="2" pageNumber="1068">San Miguel Chimalapa</collectingCounty>
, 
<collectingMunicipality id="9F73AC80C227FFCB53A7FDFDFDF8FDCA" box="[494,634,513,534]" pageId="2" pageNumber="1068">Benito Juárez</collectingMunicipality>
, 
<location id="7A776021C227FFCB52DEFDE0FF75FDED" LSID="urn:lsid:plazi:treatment:F70187ECC227FFCC5282FEAEFE00FED0:7A776021C227FFCB52DEFDE0FF75FDED" box="[151,247,540,561]" country="Mexico" county="San Miguel Chimalapa" latitude="16.7159" longLatPrecision="7" longitude="-94.1076" municipality="Benito Juarez" name="El Puerto" pageId="2" pageNumber="1068" stateProvince="Oaxaca">El Puerto</location>
, 
<date id="0B16103AC227FFCB534AFDE0FEF1FDED" box="[259,371,540,561]" pageId="2" pageNumber="1068" value="2013-07-08">
<collectingDate id="1B52E9D2C227FFCB534AFDE0FEF1FDED" box="[259,371,540,561]" pageId="2" pageNumber="1068" value="2013-07-08">8-VII-2013</collectingDate>
</date>
, 
<geoCoordinate id="1A9C503DC227FFCB53C9FDE0FE74FDED" box="[384,502,540,561]" degrees="16.7159" direction="north" orientation="latitude" pageId="2" pageNumber="1068" precision="5" value="16.7159">16.7159° N</geoCoordinate>
, 
<geoCoordinate id="1A9C503DC227FFCB504BFDE0FDF8FDED" box="[514,634,540,561]" degrees="94.1076" direction="west" orientation="longitude" pageId="2" pageNumber="1068" precision="5" value="-94.1076">94.1076° W</geoCoordinate>
, 
<quantity id="B8509B1FC227FFCB52DEFDCAFF69FD97" box="[151,235,566,587]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="1068" unit="m" value="1250.0">
<elevation id="F485D1C9C227FFCB52DEFDCAFF69FD97" box="[151,235,566,587]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="1068" unit="m" value="1250.0">1,250 m</elevation>
</quantity>
, 
<emphasis id="4DDCEAE8C227FFCB52BEFDCAFD80FD90" box="[247,514,566,588]" italics="true" pageId="2" pageNumber="1068">Pinus-Quercus-Liquidambar</emphasis>
forest, Coprotrampa, E. Mora-A. y 
<collectorName id="D25D532CC227FFCB5332FDADFE67FDBA" box="[379,485,593,614]" pageId="2" pageNumber="1068">L. Delgado</collectorName>
cols”
</materialsCitation>
. 
<materialsCitation id="CFC03CA7C227FFCB506EFDADFDBFFD47" collectorName="Instituto de Ecolog'ia A. C." country="Mexico" location="Coleccion Entomologica del Instituto de Ecolog'ia" pageId="2" pageNumber="1068" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
<typeStatus id="A0138858C227FFCB506EFDADFDFFFDBA" box="[551,637,593,614]" pageId="2" pageNumber="1068" type="holotype">Holotype</typeStatus>
is deposited in the 
<location id="7A776021C227FFCB532CFD97FED9FD47" LSID="urn:lsid:plazi:treatment:F70187ECC227FFCC5282FEAEFE00FED0:7A776021C227FFCB532CFD97FED9FD47" country="Mexico" name="Coleccion Entomologica del Instituto de Ecolog'ia" pageId="2" pageNumber="1068" stateProvince="Mexico">
Colección Entomológica del 
<collectorName id="D25D532CC227FFCB52DEFD7AFED9FD47" box="[151,347,645,668]" pageId="2" pageNumber="1068">Instituto de Ecolog´ıa</collectorName>
</location>
A. C., Xalapa, 
<collectingRegion id="BD6CF818C227FFCB53BAFD7AFDBFFD47" box="[499,573,646,667]" country="Mexico" name="Mexico" pageId="2" pageNumber="1068">Mexico</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="37B26571C227FFCB52E6FD5DFBB5FF2D" pageId="2" pageNumber="1068" type="description">
<paragraph id="7F1736FAC227FFCB52E6FD5DFBB5FF2D" blockId="2.[151,639,486,1730]" lastBlockId="2.[685,1173,194,1191]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB52E6FD5DFEA8FD6A" bold="true" box="[175,298,673,694]" pageId="2" pageNumber="1068">Description.</emphasis>
<typeStatus id="A0138858C227FFCB537FFD5DFE0CFD6A" box="[310,398,673,694]" pageId="2" pageNumber="1068" type="holotype">Holotype</typeStatus>
male. Length 
<quantity id="B8509B1FC227FFCB5060FD5DFDF8FD6A" box="[553,634,673,694]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="1068" unit="mm" value="3.9">3.9 mm</quantity>
, width across humeri 
<quantity id="B8509B1FC227FFCB5311FD47FE20FD0C" box="[344,418,699,720]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="1068" unit="mm" value="2.7">2.7 mm</quantity>
. Body form moderately convex and glabrous dorsally. Head and pronotum dark green and shiny, elytra darker, striae with green reflections, legs reddish brown; clypeus and frons slightly microreticulated (
<figureCitation id="E7932A7FC227FFCB53DDFCD9FE4DFCE6" box="[404,463,805,826]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 1</figureCitation>
). 
<emphasis id="4DDCEAE8C227FFCB53A1FCD9FDA7FCE6" bold="true" box="[488,549,805,826]" pageId="2" pageNumber="1068">Head:</emphasis>
Clypeus slightly concave, apex bidentate, median emargination V-shaped, rugose-punctate anteriorly, transversally reticulated posteriorly; frons almost flat with 2 barely visible swellings, punctures confluent, sparser and shallower toward vertex; vertex slightly convex, almost impunctate on each side of midline. Eyes very narrow, with 2 facets at posterior end of genae; eyes separated by about 9–10 times maximum width of eye (
<figureCitation id="E7932A7FC227FFCB533DFBE9FE36FBF6" box="[372,436,1045,1066]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 3</figureCitation>
). Antennae reddish brown, sericeous. 
<emphasis id="4DDCEAE8C227FFCB5305FBD3FE3EFB98" bold="true" box="[332,444,1071,1092]" pageId="2" pageNumber="1068">Pronotum:</emphasis>
Surface shiny, uniformly punctate; punctures sparse, shallow, minute to small at sides, like those on vertex (
<figureCitation id="E7932A7FC227FFCB5078FB98FDECFBA5" box="[561,622,1124,1145]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 1</figureCitation>
); lateral fovea semicircular and slightly marked; posterior margin lacking elongate punctures or groove. 
<emphasis id="4DDCEAE8C227FFCB52AFFB48FEAEFB15" bold="true" box="[230,300,1204,1225]" pageId="2" pageNumber="1068">Elytra:</emphasis>
Striae moderately impressed, interrupted by annulated punctures separated by about 2–3 diameters of 1 puncture; intervals very finely punctate, surface shiny (
<figureCitation id="E7932A7FC227FFCB53CFFAF8FE43FAC5" box="[390,449,1284,1305]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 1</figureCitation>
). 
<emphasis id="4DDCEAE8C227FFCB5390FAF8FDA2FAC5" bold="true" box="[473,544,1284,1305]" pageId="2" pageNumber="1068">Wings:</emphasis>
Macropterous, wings functional. 
<emphasis id="4DDCEAE8C227FFCB53D3FAE2FE64FAEF" bold="true" box="[410,486,1310,1331]" pageId="2" pageNumber="1068">Venter:</emphasis>
Proepisternum excavate anteriorly, surface microreticulated; sternellum smooth and with a line of setiferous punctures along base; mesosternum microreticulated with minute punctures; metasternum with minute punctures denser than mesosternum (
<figureCitation id="E7932A7FC227FFCB5381FA5FFD81FA64" box="[456,515,1443,1464]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 2</figureCitation>
). 
<emphasis id="4DDCEAE8C227FFCB505CFA5FFDFFFA64" bold="true" box="[533,637,1443,1464]" pageId="2" pageNumber="1068">Abdomen:</emphasis>
Sternites 1–6 microreticulated, with minute punctures; pygidium convex, shiny, with minute, shallow punctures (
<figureCitation id="E7932A7FC227FFCB52B6FA0FFEB8F9D4" box="[255,314,1523,1544]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 2</figureCitation>
). 
<emphasis id="4DDCEAE8C227FFCB5305FA0FFE00F9D4" bold="true" box="[332,386,1523,1544]" pageId="2" pageNumber="1068">Legs:</emphasis>
Protibia with 3 large teeth on external border, apical and middle teeth separated by 2 denticles, inner apical margin of protibia produced into triangular anterior projection and slightly bent downwards; apical spur simple and slightly curved inward (
<figureCitation id="E7932A7FC227FFCB5365F984FEE5F951" box="[300,359,1656,1677]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 4</figureCitation>
); profemora microreticulated; meso- and metafemora densely punctate, punctures minute, shallow, denser apically. 
<emphasis id="4DDCEAE8C227FFCB53F0F951FD9FF91E" bold="true" box="[441,541,1709,1730]" pageId="2" pageNumber="1068">Aedeagus:</emphasis>
Parameres moderately short, apices truncated to slightly rounded, inner margins slightly concave (
<figureCitation id="E7932A7FC227FFCB519AFF20FBA9FF2D" box="[979,1067,220,242]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Figs. 5–7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="37B26571C227FFCB508CFF0BFB16F9BD" pageId="2" pageNumber="1068" type="etymology">
<paragraph id="7F1736FAC227FFCB508CFF0BFB17FE81" blockId="2.[685,1173,194,1191]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB508CFF0BFCBAFED0" bold="true" box="[709,824,247,268]" pageId="2" pageNumber="1068">Etymology.</emphasis>
We dedicate this species to Bert Kohlmann, friend and specialist of the subfamily 
<taxonomicName id="B8A84D79C227FFCB50E4FED1FCAEFE9E" authorityName="Latreille" authorityYear="1802" box="[685,812,301,322]" pageId="2" pageNumber="1068" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
, and author of the works on the species of 
<taxonomicName id="B8A84D79C227FFCB508FFEBBFCB4FE81" box="[710,822,327,349]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="genus">
<emphasis id="4DDCEAE8C227FFCB508FFEBBFCB4FE81" box="[710,822,327,349]" italics="true" pageId="2" pageNumber="1068">Canthidium</emphasis>
</taxonomicName>
from North America and 
<collectingCountry id="07BF766AC227FFCB566EFEB4FB12FE81" box="[1063,1168,328,349]" name="Costa Rica" pageId="2" pageNumber="1068">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="7F1736FAC227FFCB508CFE9EFB86FCF6" blockId="2.[685,1173,194,1191]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB508CFE9EFC26FEAB" bold="true" box="[709,932,354,375]" pageId="2" pageNumber="1068">Taxonomic Remarks.</emphasis>
This species is distinguished by the following combination of characters: body moderately convex; dorsally dark green in color, pronotum and head with a light cast green and reddish; head with rounded swellings, not tuberculate; vertex slightly punctate; eye dorsally with two facets wide at the posterior end of gena, interocular distance separated by about 9–10 times the width of an eye; antennal club reddish brown and sericeous; pronotal surface with shallow and minute punctures on disc, at sides with punctures moderate in size; basal pronotal border not margined; elytra with eight clearly punctate striae; mesosternum with minute punctures; abdominal sternites shagreened, pygidium smooth and with minute, shallow punctures; macropterous; and the shape of the parameres (
<figureCitation id="E7932A7FC227FFCB51D2FCE9FC7AFCF6" box="[923,1016,789,811]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Figs. 5–7</figureCitation>
).
</paragraph>
<paragraph id="7F1736FAC227FFCB508CFCD3FC56FB8A" blockId="2.[685,1173,194,1191]" pageId="2" pageNumber="1068">
<taxonomicName id="B8A84D79C227FFCB508CFCD3FC23FC99" box="[709,929,815,837]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="4DDCEAE8C227FFCB508CFCD3FC23FC99" box="[709,929,815,837]" italics="true" pageId="2" pageNumber="1068">Canthidium kohlmanni</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="B8A84D79C227FFCB566AFCD3FC4DFCBC" authority="Kohlmann" authorityName="Kohlmann" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="pseudoperceptibile">
<emphasis id="4DDCEAE8C227FFCB566AFCD3FCE2FCBD" italics="true" pageId="2" pageNumber="1068">Canthidium pseudoperceptibile</emphasis>
Kohlmann
</taxonomicName>
and Sol´ıs (the new species was compared with 
<typeStatus id="A0138858C227FFCB5188FC9BFB9EFCA0" box="[961,1052,871,892]" pageId="2" pageNumber="1068" type="paratype">paratypes</typeStatus>
of this species), but it is distinguished from 
<taxonomicName id="B8A84D79C227FFCB566DFC7DFC98FC6E" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="pseudoperceptibile">
<emphasis id="4DDCEAE8C227FFCB566DFC7DFC98FC6E" italics="true" pageId="2" pageNumber="1068">C. pseudoperceptibile</emphasis>
</taxonomicName>
by the frons and vertex with denser and shallower punctures (
<figureCitation id="E7932A7FC227FFCB5131FC44FC33FC11" box="[888,945,952,973]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 3</figureCitation>
<emphasis id="4DDCEAE8C227FFCB51F1FC44FC71FC11" box="[952,1011,952,973]" italics="true" pageId="2" pageNumber="1068">versus</emphasis>
<figureCitation id="E7932A7FC227FFCB51B2FC44FBB7FC11" box="[1019,1077,952,973]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Fig. 9</figureCitation>
), the pronotum with finer and shallower punctation (
<figureCitation id="E7932A7FC227FFCB5610FC28FB16FC35" box="[1113,1172,980,1001]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 1</figureCitation>
<emphasis id="4DDCEAE8C227FFCB50E4FC13FD6AFBD8" box="[685,744,1007,1028]" italics="true" pageId="2" pageNumber="1068">versus</emphasis>
<figureCitation id="E7932A7FC227FFCB50A4FC13FCA6FBD8" box="[749,804,1007,1028]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Fig. 8</figureCitation>
), the protibia with the apical and middle teeth separated by denticles (
<figureCitation id="E7932A7FC227FFCB51F6FBF6FC7AFBC3" box="[959,1016,1034,1055]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 4</figureCitation>
<emphasis id="4DDCEAE8C227FFCB5649FBF6FBB9FBC3" box="[1024,1083,1034,1055]" italics="true" pageId="2" pageNumber="1068">versus</emphasis>
<figureCitation id="E7932A7FC227FFCB560BFBF6FB0AFBC3" box="[1090,1160,1034,1055]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Fig. 10</figureCitation>
), and the shape of the parameres of the male genitalia (
<figureCitation id="E7932A7FC227FFCB50FDFBBDFC8CFB8A" box="[692,782,1088,1110]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Figs. 5–7</figureCitation>
versus 
<figureCitation id="E7932A7FC227FFCB5111FBBDFC4BFB8A" box="[856,969,1088,1110]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Figs. 11–12</figureCitation>
).
</paragraph>
<paragraph id="7F1736FAC227FFCB508CFBA7FB17FB7B" blockId="2.[685,1173,194,1191]" pageId="2" pageNumber="1068">
To include 
<taxonomicName id="B8A84D79C227FFCB5170FBA6FC3FFBAC" box="[825,957,1114,1136]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="4DDCEAE8C227FFCB5170FBA6FC3FFBAC" box="[825,957,1114,1136]" italics="true" pageId="2" pageNumber="1068">C. kohlmanni</emphasis>
</taxonomicName>
, we propose the following modifications in Moctezuma 
<emphasis id="4DDCEAE8C227FFCB5647FB8BFBBDFB50" box="[1038,1087,1142,1164]" italics="true" pageId="2" pageNumber="1068">et al.</emphasis>
’s (2019) key to the species of 
<taxonomicName id="B8A84D79C227FFCB5138FB6DFC63FB7B" box="[881,993,1169,1191]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="genus">
<emphasis id="4DDCEAE8C227FFCB5138FB6DFC63FB7B" box="[881,993,1169,1191]" italics="true" pageId="2" pageNumber="1068">Canthidium</emphasis>
</taxonomicName>
of North America.
</paragraph>
<paragraph id="7F1736FAC227FFCB50E2FB41FB16FB0E" blockId="2.[683,1173,1213,1633]" box="[683,1172,1213,1234]" pageId="2" pageNumber="1068">13. Head and pronotum weakly punctate,</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFB24FB16FADB" blockId="2.[683,1173,1213,1633]" pageId="2" pageNumber="1068">
appearing smooth .............................. 13 
<emphasis id="4DDCEAE8C227FFCB56C7FB24FB11FB32" bold="true" box="[1166,1171,1240,1262]" pageId="2" pageNumber="1068">ʹ</emphasis>
- Head and pronotum coarsely punctate,
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFAF1FB11FAE1" blockId="2.[683,1173,1213,1633]" pageId="2" pageNumber="1068">
appearing roughened ........................... 14 13 
<emphasis id="4DDCEAE8C227FFCB508AFADBFD4AFAE1" bold="true" box="[707,712,1319,1341]" pageId="2" pageNumber="1068">ʹ</emphasis>
. Punctures on pronotal disc small, moderately
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFABEFB16FA8B" blockId="2.[683,1173,1213,1633]" box="[739,1172,1346,1367]" pageId="2" pageNumber="1068">
deep (
<figureCitation id="E7932A7FC227FFCB5155FABEFCD6FA8B" box="[796,852,1346,1367]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Fig. 8</figureCitation>
); apical and middle protibial teeth
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFAA1FB11FAAE" blockId="2.[683,1173,1213,1633]" box="[739,1171,1373,1394]" pageId="2" pageNumber="1068">
without denticles between them (
<figureCitation id="E7932A7FC227FFCB5655FAA1FBE1FAAE" box="[1052,1123,1373,1394]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Fig. 10</figureCitation>
); left
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFA8BFB17FA50" blockId="2.[683,1173,1213,1633]" box="[739,1173,1399,1420]" pageId="2" pageNumber="1068">
paramere with projection on inner edge (
<figureCitation id="E7932A7FC227FFCB5624FA8BFB17FA50" box="[1133,1173,1399,1420]" captionStart="Figs" captionStartId="4.[175,214,1310,1329]" captionTargetBox="[157,1166,196,1278]" captionTargetId="figure-195@4.[154,1169,194,1294]" captionTargetPageId="4" captionText="Figs. 8–12. Canthidium pseudoperceptibile, male paratype. 8) Habitus, dorsal view; 9) Head; 10) Protibia; 11–12) Parameres, caudal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="2" pageNumber="1068">Fig.</figureCitation>
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFA6EFB16FA7B" blockId="2.[683,1173,1213,1633]" box="[739,1172,1426,1447]" pageId="2" pageNumber="1068">
11) ........................ 
<taxonomicName id="B8A84D79C227FFCB51F7FA6EFB16FA7B" box="[958,1172,1426,1447]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="pseudoperceptibile">
<emphasis id="4DDCEAE8C227FFCB51F7FA6EFB16FA7B" bold="true" box="[958,1172,1426,1447]" italics="true" pageId="2" pageNumber="1068">C. pseudoperceptibile</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="7F1736FAC227FFCB518AFA50FB16FA00" blockId="2.[683,1173,1213,1633]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB518AFA50FB11FA1D" bold="true" box="[963,1171,1452,1474]" pageId="2" pageNumber="1068">Kohlmann and Sol´ıs</emphasis>
- Punctures on pronotal disc minute, shallow
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFA1EFB16FA2B" blockId="2.[683,1173,1213,1633]" box="[739,1172,1506,1527]" pageId="2" pageNumber="1068">
(
<figureCitation id="E7932A7FC227FFCB50A3FA1EFCA0FA2B" box="[746,802,1506,1527]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 1</figureCitation>
); apical and middle protibial teeth with
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAFA00FB16F9CD" blockId="2.[683,1173,1213,1633]" box="[739,1172,1532,1553]" pageId="2" pageNumber="1068">
denticles between them (
<figureCitation id="E7932A7FC227FFCB5190FA00FB9AF9CD" box="[985,1048,1532,1553]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 4</figureCitation>
); parameres
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAF9EBFB16F9F0" blockId="2.[683,1173,1213,1633]" box="[739,1172,1559,1580]" pageId="2" pageNumber="1068">
without projection on inner edge (
<figureCitation id="E7932A7FC227FFCB567FF9EBFBF3F9F0" box="[1078,1137,1559,1580]" captionStart="Figs" captionStartId="3.[197,236,1439,1458]" captionTargetBox="[185,1187,197,1421]" captionTargetId="figure-99@3.[175,1190,194,1423]" captionTargetPageId="3" captionText="Figs. 1–7. Canthidium kohlmanni, holotype. 1–2) Habitus, dorsal and ventral views, respectively; 3) Head;" figureDoi="http://doi.org/10.5281/zenodo.10113976" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="2" pageNumber="1068">Fig. 5</figureCitation>
) ...
</paragraph>
<paragraph id="7F1736FAC227FFCB50AAF9CDFB16F99A" blockId="2.[683,1173,1213,1633]" box="[739,1172,1585,1606]" pageId="2" pageNumber="1068">
........... 
<taxonomicName id="B8A84D79C227FFCB5109F9CDFB16F99A" authority="Mora-Aguilar and" box="[832,1172,1585,1606]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="4DDCEAE8C227FFCB5109F9CDFB16F99A" bold="true" box="[832,1172,1585,1606]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB5109F9CDFC4EF99A" bold="true" box="[832,972,1585,1606]" italics="true" pageId="2" pageNumber="1068">C. kohlmanni</emphasis>
Mora-Aguilar and
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="7F1736FAC227FFCB51F1F9B0FB16F9BD" blockId="2.[683,1173,1213,1633]" box="[952,1172,1612,1633]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB51F1F9B0FB16F9BD" bold="true" box="[952,1172,1612,1633]" pageId="2" pageNumber="1068">
Delgado, 
<taxonomicNameLabel id="56EF5793C227FFCB5652F9B0FB16F9BD" box="[1051,1172,1612,1633]" pageId="2" pageNumber="1068" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="37B26571C227FFCC50E4F984FE00FED0" lastPageId="5" lastPageNumber="1071" pageId="2" pageNumber="1068" type="distribution">
<paragraph id="7F1736FAC227FFCB50E4F984FB16F91E" blockId="2.[685,1172,1655,1730]" pageId="2" pageNumber="1068">
<emphasis id="4DDCEAE8C227FFCB50E4F984FCADF951" bold="true" box="[685,815,1656,1677]" pageId="2" pageNumber="1068">Distribution.</emphasis>
<taxonomicName id="B8A84D79C227FFCB517AF98BFB8EF951" box="[819,1036,1655,1677]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1068" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="4DDCEAE8C227FFCB517AF98BFB8EF951" box="[819,1036,1655,1677]" italics="true" pageId="2" pageNumber="1068">Canthidium kohlmanni</emphasis>
</taxonomicName>
is only known from the 
<typeStatus id="A0138858C227FFCB5140F96EFCB6F97B" box="[777,820,1682,1703]" pageId="2" pageNumber="1068">type</typeStatus>
locality, in the undisturbed forests of the mountainous areas of Benito Juárez in the
</paragraph>
<caption id="2BD76672C226FFCA528CFA63FB28FA6E" ID-DOI="http://doi.org/10.5281/zenodo.10113976" ID-Zenodo-Dep="10113976" box="[197,1194,1438,1458]" httpUri="https://zenodo.org/record/10113976/files/figure.png" pageId="3" pageNumber="1069" startId="3.[197,236,1439,1458]" targetBox="[185,1187,197,1421]" targetPageId="3">
<paragraph id="7F1736FAC226FFCA528CFA63FB28FA6E" blockId="3.[173,1194,1438,1482]" box="[197,1194,1438,1458]" pageId="3" pageNumber="1069">
<emphasis id="4DDCEAE8C226FFCA528CFA63FEA2FA6E" bold="true" box="[197,288,1438,1458]" pageId="3" pageNumber="1069">Figs. 1–7.</emphasis>
<taxonomicName id="B8A84D79C226FFCA537FFA62FE7CFA6D" box="[310,510,1438,1457]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1069" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="4DDCEAE8C226FFCA537FFA62FE7CFA6D" box="[310,510,1438,1457]" italics="true" pageId="3" pageNumber="1069">Canthidium kohlmanni</emphasis>
</taxonomicName>
, holotype. 
<emphasis id="4DDCEAE8C226FFCA5022FA63FD13FA6E" bold="true" box="[619,657,1438,1458]" pageId="3" pageNumber="1069">1–2)</emphasis>
Habitus, dorsal and ventral views, respectively; 
<emphasis id="4DDCEAE8C226FFCA5610FA63FBE8FA6E" bold="true" box="[1113,1130,1439,1458]" pageId="3" pageNumber="1069">3)</emphasis>
Head;
</paragraph>
</caption>
<paragraph id="7F1736FAC226FFCA52E4FA4BFCACFA16" blockId="3.[173,1194,1438,1482]" box="[173,814,1462,1482]" pageId="3" pageNumber="1069">
<emphasis id="4DDCEAE8C226FFCA52E4FA4BFF3CFA16" bold="true" box="[173,190,1463,1482]" pageId="3" pageNumber="1069">4)</emphasis>
Protibia; 
<emphasis id="4DDCEAE8C226FFCA535DFA4BFEB8FA16" bold="true" box="[276,314,1462,1482]" pageId="3" pageNumber="1069">5–7)</emphasis>
Parameres, caudal, dorsal, and lateral views, respectively.
</paragraph>
<paragraph id="7F1736FAC226FFCA52E4F9F2FC88F9AE" blockId="3.[173,660,1550,1730]" lastBlockId="3.[707,1193,1549,1730]" pageId="3" pageNumber="1069">
Los Chimalapas region of 
<collectingRegion id="BD6CF818C226FFCA53F0F9F2FD85F9FF" box="[441,519,1550,1571]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="1069">Oaxaca</collectingRegion>
, 
<collectingCountry id="07BF766AC226FFCA505BF9F2FDE0F9FF" box="[530,610,1550,1571]" name="Mexico" pageId="3" pageNumber="1069">Mexico</collectingCountry>
. The locality and surrounding areas have a rugged topography with ravines and hills with mixed pine-oak forest and tropical deciduous forests at 
<quantity id="B8509B1FC226FFCA5285F984FEA6F951" box="[204,292,1656,1677]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="1069" unit="m" value="1250.0">1,250 m</quantity>
elevation. The distribution of 
<taxonomicName id="B8A84D79C226FFCA5035F98BFEECF974" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1069" phylum="Arthropoda" rank="species" species="pseudoperceptibile">
<emphasis id="4DDCEAE8C226FFCA5035F98BFEECF974" italics="true" pageId="3" pageNumber="1069">C. pseudoperceptibile</emphasis>
</taxonomicName>
is allopatric with 
<taxonomicName id="B8A84D79C226FFCA5075F96EFF6EF91E" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1069" phylum="Arthropoda" rank="species" species="kohlmanni">
<emphasis id="4DDCEAE8C226FFCA5075F96EFF6EF91E" italics="true" pageId="3" pageNumber="1069">C. kohlmanni</emphasis>
</taxonomicName>
, since the former species inhabits tropical forests in the lowlands of the states of 
<collectingRegion id="BD6CF818C226FFCA5606F9F1FB2BF9FE" box="[1103,1193,1549,1570]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="1069">Veracruz</collectingRegion>
and 
<collectingRegion id="BD6CF818C226FFCA50BFF9D4FCC5F9E1" box="[758,839,1576,1597]" country="Mexico" name="Chiapas" pageId="3" pageNumber="1069">Chiapas</collectingRegion>
in 
<collectingCountry id="07BF766AC226FFCA5130F9D4FC47F9E1" box="[889,965,1576,1597]" name="Mexico" pageId="3" pageNumber="1069">Mexico</collectingCountry>
and 
<collectingCountry id="07BF766AC226FFCA5640F9D4FBF4F9E1" box="[1033,1142,1576,1597]" name="Guatemala" pageId="3" pageNumber="1069">Guatemala</collectingCountry>
and 
<collectingCountry id="07BF766AC226FFCA508AF9BFFC81F984" box="[707,771,1603,1624]" name="Belize" pageId="3" pageNumber="1069">Belize</collectingCountry>
but on the Gulf of 
<collectingCountry id="07BF766AC226FFCA519DF9BFFBA2F984" box="[980,1056,1603,1624]" name="Mexico" pageId="3" pageNumber="1069">Mexico</collectingCountry>
and Atlantic slopes.
</paragraph>
<paragraph id="7F1736FAC226FFCD5092F984FCE1FA0F" blockId="3.[707,1193,1549,1730]" lastBlockId="4.[685,1172,1417,1730]" lastPageId="4" lastPageNumber="1070" pageId="3" pageNumber="1069">
Los Chimalapas has remained as a biologically unexplored region in 
<collectingCountry id="07BF766AC226FFCA51D7F96EFC6BF97B" box="[926,1001,1682,1703]" name="Mexico" pageId="3" pageNumber="1069">Mexico</collectingCountry>
, principally at elevations above 
<quantity id="B8509B1FC226FFCA511EF951FC2FF91E" box="[855,941,1709,1730]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="1069" unit="m" value="1000.0">1,000 m</quantity>
. However, since the last decade of the last century to present, many species from this region have been described. Currently, one genus and 23 species of animals and 20 species of plants are endemic in the region (Appendix 1). Two groups stand out by the greater number of endemic species in this region, plants and beetles. 
<taxonomicName id="B8A84D79C221FFCD505FF9F1FDFCF9FE" authorityName="Linnaeus" authorityYear="1758" box="[534,638,1549,1570]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="1070" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
is the most diverse group yet less studied. In Los Chimalapas, there are currently known one genus and one species of rove beetles, one species of zopherid beetle, 11 species of scarab beetles, and two species of passalid beetles that are endemic in this region (Appendix 1). Most of the species endemic species to this region are restricted to cloud and mixed forests with conifers located above 
<quantity id="B8509B1FC221FFCD50E4FA42FD7FFA0F" box="[685,765,1470,1491]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="1070" unit="m" value="1000.0">1,000 m</quantity>
elevation.
</paragraph>
<caption id="2BD76672C221FFCD52E6FAE2FDC2FA95" ID-DOI="http://doi.org/10.5281/zenodo.10113978" ID-Zenodo-Dep="10113978" httpUri="https://zenodo.org/record/10113978/files/figure.png" pageId="4" pageNumber="1070" startId="4.[175,214,1310,1329]" targetBox="[157,1166,196,1278]" targetPageId="4">
<paragraph id="7F1736FAC221FFCD52E6FAE2FB11FAED" blockId="4.[151,1171,1309,1353]" box="[175,1171,1309,1329]" pageId="4" pageNumber="1070">
<emphasis id="4DDCEAE8C221FFCD52E6FAE2FE93FAED" bold="true" box="[175,273,1309,1329]" pageId="4" pageNumber="1070">Figs. 8–12.</emphasis>
<taxonomicName id="B8A84D79C221FFCD536FFAE1FDADFAEC" box="[294,559,1309,1328]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1070" phylum="Arthropoda" rank="species" species="pseudoperceptibile">
<emphasis id="4DDCEAE8C221FFCD536FFAE1FDADFAEC" box="[294,559,1309,1328]" italics="true" pageId="4" pageNumber="1070">Canthidium pseudoperceptibile</emphasis>
</taxonomicName>
, male paratype. 
<emphasis id="4DDCEAE8C221FFCD5089FAE2FD53FAED" bold="true" box="[704,721,1310,1329]" pageId="4" pageNumber="1070">8)</emphasis>
Habitus, dorsal view; 
<emphasis id="4DDCEAE8C221FFCD51DFFAE2FC25FAED" bold="true" box="[918,935,1310,1329]" pageId="4" pageNumber="1070">9)</emphasis>
Head; 
<emphasis id="4DDCEAE8C221FFCD51A1FAE2FB86FAED" bold="true" box="[1000,1028,1310,1329]" pageId="4" pageNumber="1070">10)</emphasis>
Protibia; 
<emphasis id="4DDCEAE8C221FFCD5613FAE2FB11FAED" bold="true" box="[1114,1171,1309,1329]" pageId="4" pageNumber="1070">11–12)</emphasis>
</paragraph>
<paragraph id="7F1736FAC221FFCD52DEFACAFDC2FA95" blockId="4.[151,1171,1309,1353]" box="[151,576,1334,1353]" pageId="4" pageNumber="1070">Parameres, caudal and lateral views, respectively.</paragraph>
</caption>
<paragraph id="7F1736FAC221FFCC508CFA24FE00FED0" blockId="4.[685,1172,1417,1730]" lastBlockId="5.[173,658,194,268]" lastPageId="5" lastPageNumber="1071" pageId="4" pageNumber="1070">
It is remarkable that this region conserves endemic species of large size, such as sturdy trees, 
<emphasis id="4DDCEAE8C221FFCD5622FA0FFB0AF9D4" box="[1131,1160,1523,1544]" italics="true" pageId="4" pageNumber="1070">e.g</emphasis>
., 
<taxonomicName id="B8A84D79C221FFCD50E4F9F2FBBEF9FE" authority="Lorea-Hern." authorityName="Lorea-Hern. Lorea-Hernandez" authorityYear="2008" box="[685,1084,1549,1571]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="4" pageNumber="1070" phylum="Tracheophyta" rank="species" species="obscura">
<emphasis id="4DDCEAE8C221FFCD50E4F9F2FCC7F9FF" box="[685,837,1549,1571]" italics="true" pageId="4" pageNumber="1070">Persea obscura</emphasis>
Lorea-Hern. (Lauraceae)
</taxonomicName>
, or a robust beetle species, 
<emphasis id="4DDCEAE8C221FFCD5136F9D4FC1EF9E1" box="[895,924,1576,1597]" italics="true" pageId="4" pageNumber="1070">e.g</emphasis>
., 
<taxonomicName id="B8A84D79C221FFCD51FCF9DBFBA7F984" authority="Delgado and Mora-Aguilar" authorityName="Delgado and Mora-Aguilar" class="Insecta" family="Passalidae" genus="Proculus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1070" phylum="Arthropoda" rank="species" species="reyescastilloi">
<emphasis id="4DDCEAE8C221FFCD51FCF9DBFB16F9E1" box="[949,1172,1575,1597]" italics="true" pageId="4" pageNumber="1070">Proculus reyescastilloi</emphasis>
Delgado and Mora-Aguilar (Passalidae)
</taxonomicName>
. The area of Los Chimalapas has remained as a little-disturbed region due to its mountainous and steep topography, which make accessibility difficult. Priority efforts to conserve this region must be given to maintain not only particular species, but its huge composite biodiversity of species and landscapes (Navarro- Sigüenza 
<emphasis id="4DDCEAE8C220FFCC5342FF0BFEBFFED0" box="[267,317,246,268]" italics="true" pageId="5" pageNumber="1071">et al.</emphasis>
2008).
</paragraph>
</subSubSection>
</treatment>
</document>