treatments-xml/data/B3/1B/08/B31B0803242EFF97FF4FC56DABE5FE4A.xml
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<document id="14693143597F726C92BF749260326632" ID-CLB-Dataset="26820" ID-DOI="10.11646/zootaxa.4604.3.2" ID-GBIF-Dataset="9db4515c-7f05-4cd4-b281-7b45861fe49c" ID-ISSN="1175-5326" ID-Zenodo-Dep="2839265" ID-ZooBank="8EF9BFFE-BC10-41C0-AEF1-178984F955F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557917591468" checkinUser="plazi" docAuthor="López-García, Margarita M., Giraldo-Mendoza, Alfredo E. &amp; Deloya, Cuauhtémoc" docDate="2019" docId="B31B0803242EFF97FF4FC56DABE5FE4A" docLanguage="en" docName="zootaxa.4604.3.2.pdf.imf" docOrigin="Zootaxa 4604 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomarus villosus" docType="treatment" docVersion="13" lastPageNumber="438" masterDocId="4F22707B2426FF9DFFD8C251AA63FF99" masterDocTitle="Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru" masterLastPageNumber="440" masterPageNumber="428" pageNumber="436" updateTime="1698728627633" updateUser="ExternalLinkService">
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<mods:title id="6FC862D91E28C2267B7F3BD9DCABEFB5">Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru</mods:title>
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<mods:namePart id="262DB47534AA7EE5F92EB23FF38CDEFE">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="00C56CF7E5A7FC86D39ACD35BBEC1A1A">Giraldo-Mendoza, Alfredo E.</mods:namePart>
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<mods:namePart id="2000422120341BE246812161597EDB74">Deloya, Cuauhtémoc</mods:namePart>
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<treatment id="B31B0803242EFF97FF4FC56DABE5FE4A" ID-DOI="http://doi.org/10.5281/zenodo.5412554" ID-GBIF-Taxon="157141509" ID-Zenodo-Dep="5412554" LSID="urn:lsid:plazi:treatment:B31B0803242EFF97FF4FC56DABE5FE4A" httpUri="http://treatment.plazi.org/id/B31B0803242EFF97FF4FC56DABE5FE4A" lastPageId="10" lastPageNumber="438" pageId="8" pageNumber="436">
<subSubSection id="73A8EA9E242EFF95FF4FC56DA830F8CC" box="[151,595,1851,1878]" pageId="8" pageNumber="436" type="nomenclature">
<paragraph id="3B0DB915242EFF95FF4FC56DA830F8CC" blockId="8.[151,595,1851,1913]" box="[151,595,1851,1878]" pageId="8" pageNumber="436">
<heading id="60450E79242EFF95FF4FC56DA830F8CC" bold="true" box="[151,595,1851,1878]" fontSize="11" level="1" pageId="8" pageNumber="436" reason="1">
<taxonomicName id="FCB2C296242EFF95FF4FC56DA830F8CC" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[151,595,1851,1878]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="09C66507242EFF95FF4FC56DA830F8CC" bold="true" box="[151,595,1851,1878]" pageId="8" pageNumber="436">
<emphasis id="09C66507242EFF95FF4FC56DAB07F8CF" bold="true" box="[151,356,1851,1878]" italics="true" pageId="8" pageNumber="436">Tomarus villosus</emphasis>
(
<bibRefCitation id="5F23C4E4242EFF95FEABC56AA828F8CC" author="Burmeister, H." box="[371,587,1851,1877]" pageId="8" pageNumber="436" refId="ref6200" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="73A8EA9E242EFF95FF4FC50EAB4DF8E0" box="[151,302,1886,1913]" pageId="8" pageNumber="436" type="description">
<paragraph id="3B0DB915242EFF95FF4FC50EAB4DF8E0" blockId="8.[151,595,1851,1913]" box="[151,302,1886,1913]" pageId="8" pageNumber="436">
(
<figureCitation id="A389A590242EFF95FF47C50EAB45F8E0" box="[159,294,1886,1913]" captionStart="FIGURES 3538" captionStartId="9.[151,264,1809,1834]" captionTargetBox="[163,1433,1010,1743]" captionTargetId="figure@9.[153,1437,1031,1751]" captionTargetPageId="9" captionText="FIGURES 3538. Tomarus villosus: 35, habitus; 36, parameres in frontal view; 37, parameres in lateral view; 38, locality records in Peru. Habitus photograph by Yony Callohauri." httpUri="https://zenodo.org/record/2839311/files/figure.png" pageId="8" pageNumber="436">Figs. 3538</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73A8EA9E242EFF97FF4FC5F2A814FF41" lastPageId="10" lastPageNumber="438" pageId="8" pageNumber="436" type="reference_group">
<paragraph id="3B0DB915242EFF95FF4FC5F2A850F822" blockId="8.[151,563,1955,2041]" box="[151,563,1955,1980]" pageId="8" pageNumber="436">
<treatmentCitationGroup id="1BA29E3B242EFF95FF4FC5F2A850F822" box="[151,563,1955,1980]" pageId="8" pageNumber="436">
<taxonomicName id="FCB2C296242EFF95FF4FC5F2A850F822" authority="Burmeister, 1847: 120" authorityName="Burmeister" authorityPageNumber="120" authorityYear="1847" box="[151,563,1955,1980]" class="Insecta" family="Dynastidae" genus="Podalgus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="09C66507242EFF95FF4FC5F2AB29F822" box="[151,330,1955,1979]" italics="true" pageId="8" pageNumber="436">Podalgus villosus</emphasis>
<treatmentCitation id="BA139F04242EFF95FE89C5F2A850F822" author="Burmeister, H." box="[337,563,1955,1980]" page="120" pageId="8" pageNumber="436" year="1847">
<bibRefCitation id="5F23C4E4242EFF95FE89C5F2A860F822" author="Burmeister, H." box="[337,515,1955,1980]" pageId="8" pageNumber="436" refId="ref6200" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
: 120
</treatmentCitation>
</taxonomicName>
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</paragraph>
<paragraph id="3B0DB915242EFF95FF4FC593A863F843" blockId="8.[151,563,1955,2041]" box="[151,512,1985,2010]" pageId="8" pageNumber="436">
<treatmentCitationGroup id="1BA29E3B242EFF95FF4FC593A863F843" box="[151,512,1985,2010]" pageId="8" pageNumber="436">
<taxonomicName id="FCB2C296242EFF95FF4FC593A863F843" authority="Solier, 1851: 79" authorityName="Solier" authorityPageNumber="79" authorityYear="1851" box="[151,512,1985,2010]" class="Insecta" family="Dynastidae" genus="Oryctes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="nitidicollis">
<emphasis id="09C66507242EFF95FF4FC593AB3BF843" box="[151,344,1986,2010]" italics="true" pageId="8" pageNumber="436">Oryctes nitidicollis</emphasis>
<treatmentCitation id="BA139F04242EFF95FE87C590A863F843" author="Solier, A. J. J." box="[351,512,1985,2010]" page="79" pageId="8" pageNumber="436" year="1851">
<bibRefCitation id="5F23C4E4242EFF95FE87C590ABBEF843" author="Solier, A. J. J." box="[351,477,1985,2010]" pageId="8" pageNumber="436" refId="ref7526" refString="Solier, A. J. J. (1851) Coleopteros. In: Gay, C. (Ed.), Historia fisica y politica de Chile. Vol. 5. C. Gay, Paris, 574 pp." type="book" year="1851">Solier, 1851</bibRefCitation>
: 79
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</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="3B0DB915242EFF94FF4FC5B0AFFFF847" blockId="8.[151,563,1955,2041]" lastBlockId="9.[151,1437,1915,2015]" lastPageId="9" lastPageNumber="437" pageId="8" pageNumber="436">
<treatmentCitationGroup id="1BA29E3B242EFF94FF4FC5B0AFFFF847" infered="true" lastPageId="9" lastPageNumber="437" pageId="8" pageNumber="436">
<taxonomicName id="FCB2C296242EFF95FF4FC5B0A84BF860" ID-CoL="7WYJV" authority="Steinheil, 1872: 560" authorityName="Steinheil" authorityPageNumber="560" authorityYear="1872" box="[151,552,2016,2041]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="patagonus">
<emphasis id="09C66507242EFF95FF4FC5B0AB36F861" box="[151,341,2017,2040]" italics="true" pageId="8" pageNumber="436">Ligyrus patagonus</emphasis>
<treatmentCitation id="BA139F04242EFF95FE83C5B1A84BF860" author="Steinheil, E." box="[347,552,2016,2041]" page="560" pageId="8" pageNumber="436" year="1872">
<bibRefCitation id="5F23C4E4242EFF95FE83C5B1AB9BF860" author="Steinheil, E." box="[347,504,2016,2041]" pageId="8" pageNumber="436" pagination="554 - 578" refId="ref7570" refString="Steinheil, E. (1872) Symbolae ad historiam Coleopterorum Argentinae meridionalis, ossia enumerazione dei coleotteri raccolti dal Prof. Strobel, durante il suo soggiorno in Buenos Aires e nei viaggi de la intrapresi a Mendoza e nal Chili, indi a Bahia Blanca ed al Carmen de los Patagones; e descrizione delle specie nuove. Atti della Societa Italiana di Scienze Naturali, 15, 554 - 578." type="journal article" year="1872">Steinheil, 1872</bibRefCitation>
: 560
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<emphasis id="09C66507242FFF94FF4FC52AAB71F80F" bold="true" box="[151,274,1915,1942]" pageId="9" pageNumber="437">Diagnosis.</emphasis>
<taxonomicName id="FCB2C296242FFF94FEC2C52CABB9F80F" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[282,474,1916,1942]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="09C66507242FFF94FEC2C52CABB9F80F" box="[282,474,1916,1942]" italics="true" pageId="9" pageNumber="437">Tomarus villosus</emphasis>
</taxonomicName>
can be recognized by the antennal club longer than antennomeres 27; clypeus narrowed towards apex (base 2.5 times wider than apex); frontoclypeal tubercles cariniform; pronotal punctation minute and sparse; pronotum without fovea; pronotal tubercle small; males with the inner claw of the protarsi simple;
</treatmentCitationGroup>
</paragraph>
<caption id="6FCDE99D242FFF94FF4FC1E5A8EBFC68" httpUri="https://zenodo.org/record/2839309/files/figure.png" pageId="9" pageNumber="437" startId="9.[151,264,948,973]" targetBox="[165,1435,180,865]" targetPageId="9">
<paragraph id="3B0DB915242FFF94FF4FC1E5A8EBFC68" blockId="9.[151,1437,948,1010]" pageId="9" pageNumber="437">
<emphasis id="09C66507242FFF94FF4FC1E5AB30FC54" bold="true" box="[151,339,948,973]" pageId="9" pageNumber="437">FIGURES 3134.</emphasis>
<taxonomicName id="FCB2C296242FFF94FE80C1E7AB94FC54" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[344,503,949,973]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="09C66507242FFF94FE80C1E7AB94FC54" box="[344,503,949,973]" italics="true" pageId="9" pageNumber="437">Tomarus similis</emphasis>
</taxonomicName>
: 31, habitus; 32, parameres in frontal view; 33, parameres in lateral view; 34, locality records in Peru. Habitus photograph by Yony Callohauri.
</paragraph>
</caption>
<caption id="6FCDE99D242FFF94FF4FC540A8BAF8D7" httpUri="https://zenodo.org/record/2839311/files/figure.png" pageId="9" pageNumber="437" startId="9.[151,264,1809,1834]" targetBox="[163,1433,1010,1743]" targetPageId="9">
<paragraph id="3B0DB915242FFF94FF4FC540A8BAF8D7" blockId="9.[151,1436,1809,1870]" pageId="9" pageNumber="437">
<emphasis id="09C66507242FFF94FF4FC540AB34F8B3" bold="true" box="[151,343,1809,1834]" pageId="9" pageNumber="437">FIGURES 3538.</emphasis>
<taxonomicName id="FCB2C296242FFF94FEB8C542A86DF8B3" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[352,526,1810,1834]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="09C66507242FFF94FEB8C542A86DF8B3" box="[352,526,1810,1834]" italics="true" pageId="9" pageNumber="437">Tomarus villosus</emphasis>
</taxonomicName>
: 35, habitus; 36, parameres in frontal view; 37, parameres in lateral view; 38, locality records in Peru. Habitus photograph by Yony Callohauri.
</paragraph>
</caption>
<paragraph id="3B0DB915242CFF97FF4FC2C8A814FF41" blockId="10.[151,1437,152,467]" pageId="10" pageNumber="438">
protibia tridentate, without additional basal denticle; apex of metatibia with 1825 spinules; parameres with 2 acute ventral teeth, apices straight (
<figureCitation id="A389A590242CFF97FE39C2ECA808FF41" box="[481,619,189,216]" captionStart="FIGURES 3538" captionStartId="9.[151,264,1809,1834]" captionTargetBox="[163,1433,1010,1743]" captionTargetId="figure@9.[153,1437,1031,1751]" captionTargetPageId="9" captionText="FIGURES 3538. Tomarus villosus: 35, habitus; 36, parameres in frontal view; 37, parameres in lateral view; 38, locality records in Peru. Habitus photograph by Yony Callohauri." httpUri="https://zenodo.org/record/2839311/files/figure.png" pageId="10" pageNumber="438">Figs. 3637</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="73A8EA9E242CFF97FF1FC2B1A8B9FEDA" pageId="10" pageNumber="438" type="description">
<paragraph id="3B0DB915242CFF97FF1FC2B1AA93FE86" blockId="10.[151,1437,152,467]" pageId="10" pageNumber="438">
<emphasis id="09C66507242CFF97FF1FC2B1AB99FF62" bold="true" box="[199,506,224,251]" pageId="10" pageNumber="438">
Locality records (
<figureCitation id="A389A590242CFF97FE40C2B1AB8DFF62" box="[408,494,224,251]" captionStart="FIGURES 3538" captionStartId="9.[151,264,1809,1834]" captionTargetBox="[163,1433,1010,1743]" captionTargetId="figure@9.[153,1437,1031,1751]" captionTargetPageId="9" captionText="FIGURES 3538. Tomarus villosus: 35, habitus; 36, parameres in frontal view; 37, parameres in lateral view; 38, locality records in Peru. Habitus photograph by Yony Callohauri." httpUri="https://zenodo.org/record/2839311/files/figure.png" pageId="10" pageNumber="438">Fig. 38</figureCitation>
).
</emphasis>
<materialsCitation id="8BDAB348242CFF97FDDAC2B0AA8FFE86" ID-GBIF-Occurrence="2242235146" collectionCode="MEKRB" location="La" municipality="Socabaya" pageId="10" pageNumber="438" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Arequipa">
<specimenCount id="2DB4729C242CFF97FDDAC2B0A83BFF62" box="[514,600,225,251]" pageId="10" pageNumber="438" type="male">2 males</specimenCount>
and
<specimenCount id="2DB4729C242CFF97FD56C2B0A88CFF62" box="[654,751,225,251]" pageId="10" pageNumber="438" type="female">1 female</specimenCount>
from
<collectionCode id="5DA321D0242CFF97FCEBC2B0A9F9FF62" box="[819,922,225,251]" pageId="10" pageNumber="438">MEKRB</collectionCode>
.
<emphasis id="09C66507242CFF97FC7CC2B1AE25FF62" bold="true" box="[932,1094,224,251]" pageId="10" pageNumber="438">
<collectingRegion id="F97677F7242CFF97FC7CC2B1AE76FF62" box="[932,1045,224,251]" country="Peru" name="Arequipa" pageId="10" pageNumber="438">Arequipa</collectingRegion>
(2):
</emphasis>
<collectingMunicipality id="DB69236F242CFF97FB95C2B0AED8FF62" box="[1101,1211,225,251]" pageId="10" pageNumber="438">Socabaya</collectingMunicipality>
,
<collectingRegion id="F97677F7242CFF97FB1EC2B0AE9EFF62" box="[1222,1277,225,251]" country="Venezuela" name="Lara" pageId="10" pageNumber="438">Lara</collectingRegion>
.
<emphasis id="09C66507242CFF97FADFC2B1AF1BFF62" bold="true" box="[1287,1400,224,251]" pageId="10" pageNumber="438">
<collectingRegion id="F97677F7242CFF97FADFC2B1AF24FF62" box="[1287,1351,224,251]" country="Peru" name="Lima" pageId="10" pageNumber="438">Lima</collectingRegion>
(1):
</emphasis>
<location id="3E6DEFCE242CFF97FAA7C2B0AFFEFF62" LSID="urn:lsid:plazi:treatment:B31B0803242EFF97FF4FC56DABE5FE4A:3E6DEFCE242CFF97FAA7C2B0AFFEFF62" box="[1407,1437,225,251]" municipality="Socabaya" name="La" pageId="10" pageNumber="438" stateProvince="Arequipa">La</location>
<taxonomicName id="FCB2C296242CFF97FF4FC354AA8FFE86" box="[151,236,261,287]" class="Magnoliopsida" family="Asteraceae" genus="Molina" higherTaxonomySource="CoL" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="428" phylum="Tracheophyta" rank="genus">Molina</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="3B0DB915242CFF97FF1FC378A8B9FEDA" blockId="10.[151,1437,152,467]" box="[199,730,296,323]" pageId="10" pageNumber="438">
<emphasis id="09C66507242CFF97FF1FC378ABB4FEDA" bold="true" box="[199,471,296,323]" pageId="10" pageNumber="438">Temporal distribution.</emphasis>
February (2), June (1).
</paragraph>
</subSubSection>
<subSubSection id="73A8EA9E242CFF97FF1FC31DABE5FE4A" pageId="10" pageNumber="438" type="distribution">
<paragraph id="3B0DB915242CFF97FF1FC31DAEDEFEFE" blockId="10.[151,1437,152,467]" box="[199,1213,332,359]" pageId="10" pageNumber="438">
<emphasis id="09C66507242CFF97FF1FC31DAB02FEFE" bold="true" box="[199,353,332,359]" pageId="10" pageNumber="438">Distribution.</emphasis>
<collectingCountry id="43A5F985242CFF97FEBFC31CABFEFEFE" box="[359,413,333,359]" name="Peru" pageId="10" pageNumber="438">Peru</collectingCountry>
,
<collectingCountry id="43A5F985242CFF97FE71C31CAB8BFEFE" box="[425,488,333,359]" name="Chile" pageId="10" pageNumber="438">Chile</collectingCountry>
, and
<collectingCountry id="43A5F985242CFF97FDF9C31CA8F3FEFE" box="[545,656,333,359]" name="Argentina" pageId="10" pageNumber="438">Argentina</collectingCountry>
(Endr̂di 1985;
<bibRefCitation id="5F23C4E4242CFF97FC91C31CAED2FEFE" author="Neita-Moreno, J. C. &amp; Ratcliffe, B. C." box="[841,1201,333,359]" pageId="10" pageNumber="438" pagination="1 - 36" refId="ref7007" refString="Neita-Moreno, J. C. &amp; Ratcliffe, B. C. (2017) The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay. Insecta Mundi, 547, 1 - 36." type="journal article" year="2017">Neita-Moreno &amp; Ratcliffe 2017</bibRefCitation>
).
</paragraph>
<paragraph id="3B0DB915242CFF97FF1FC320ABE5FE4A" blockId="10.[151,1437,152,467]" pageId="10" pageNumber="438">
<emphasis id="09C66507242CFF97FF1FC320ABEBFE12" bold="true" box="[199,392,368,395]" pageId="10" pageNumber="438">Natural history.</emphasis>
<bibRefCitation id="5F23C4E4242CFF97FE49C320A82CFE12" author="Gutierrez, R." box="[401,591,368,395]" pageId="10" pageNumber="438" pagination="112 - 118" refId="ref6754" refString="Gutierrez, R. (1945). Contribuciones al conocimiento de los Scarabaeidae chilenos. Dynastinae Oryctini. Revista Chilena de Historia Natural, 48, 112 - 118." type="journal article" year="1945">Gutiérrez (1945)</bibRefCitation>
observed the mating behavior of some specimens of
<taxonomicName id="FCB2C296242CFF97FB64C323AF53FE12" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1212,1328,369,395]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="09C66507242CFF97FB64C323AF53FE12" box="[1212,1328,369,395]" italics="true" pageId="10" pageNumber="438">T. villosus</emphasis>
</taxonomicName>
in
<collectingCountry id="43A5F985242CFF97FA81C320AFFBFE12" box="[1369,1432,369,395]" name="Chile" pageId="10" pageNumber="438">Chile</collectingCountry>
, where the males flew searching for females and the copulation occurred below soil, in the same hole from where the females emerged.
</paragraph>
</subSubSection>
</treatment>
</document>