301 lines
33 KiB
XML
301 lines
33 KiB
XML
<document id="76D99926041E667B6994C38B2CFC825F" ID-CLB-Dataset="26820" ID-DOI="10.11646/zootaxa.4604.3.2" ID-GBIF-Dataset="9db4515c-7f05-4cd4-b281-7b45861fe49c" ID-ISSN="1175-5326" ID-Zenodo-Dep="2839265" ID-ZooBank="8EF9BFFE-BC10-41C0-AEF1-178984F955F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557917591468" checkinUser="plazi" docAuthor="López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc" docDate="2019" docId="B31B0803242CFF97FF4FC074AFFFF905" docLanguage="en" docName="zootaxa.4604.3.2.pdf.imf" docOrigin="Zootaxa 4604 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomarus Erichson 1847" docType="key" docVersion="13" lastPageNumber="438" masterDocId="4F22707B2426FF9DFFD8C251AA63FF99" masterDocTitle="Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru" masterLastPageNumber="440" masterPageNumber="428" pageNumber="438" updateTime="1698728627633" updateUser="ExternalLinkService">
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<mods:title id="5404337414E884BCBE1547F39F057369">Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru</mods:title>
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<mods:namePart id="BDAE35B82E39FA43883B8D6E73070FC3">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="94265A1505BD04570057AD8518CC6561">Giraldo-Mendoza, Alfredo E.</mods:namePart>
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<mods:namePart id="A56E24274C710CACD7FEAAD133871123">Deloya, Cuauhtémoc</mods:namePart>
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<mods:title id="0C75BEC1E038D4038615369BFE24B4F7">Zootaxa</mods:title>
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<mods:date id="3721CEAA15C57388B8163CC1FED52283">2019</mods:date>
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<treatment id="B31B0803242CFF97FF4FC074AFFFF905" ID-DOI="http://doi.org/10.5281/zenodo.5412556" ID-GBIF-Taxon="157141508" ID-Zenodo-Dep="5412556" LSID="urn:lsid:plazi:treatment:B31B0803242CFF97FF4FC074AFFFF905" httpUri="http://treatment.plazi.org/id/B31B0803242CFF97FF4FC074AFFFF905" lastPageNumber="438" pageId="10" pageNumber="438">
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<subSubSection id="73A8EA9E242CFF97FF4FC074AA95FD56" pageId="10" pageNumber="438" type="nomenclature">
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<paragraph id="3B0DB915242CFF97FF4FC074A809FDA6" blockId="10.[151,618,549,575]" box="[151,618,549,575]" pageId="10" pageNumber="438">
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<heading id="60450E79242CFF97FF4FC074A809FDA6" bold="true" box="[151,618,549,575]" fontSize="11" level="1" pageId="10" pageNumber="438" reason="1">
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<emphasis id="09C66507242CFF97FF4FC074A809FDA6" bold="true" box="[151,618,549,575]" pageId="10" pageNumber="438">
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Key to the species of
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<taxonomicName id="FCB2C296242CFF97FE45C074A86BFDA6" authorityName="Erichson" authorityYear="1847" box="[413,520,549,575]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="genus">
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<emphasis id="09C66507242CFF97FE45C074A86BFDA6" bold="true" box="[413,520,549,575]" italics="true" pageId="10" pageNumber="438">Tomarus</emphasis>
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</taxonomicName>
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of
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<collectingCountry id="43A5F985242CFF97FDF6C074A809FDA6" box="[558,618,549,575]" name="Peru" pageId="10" pageNumber="438">Peru</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3B0DB915242CFF97FF4FC03CAA95FD56" blockId="10.[151,1437,621,719]" pageId="10" pageNumber="438">
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Males and females can be differentiated by the shape of the apical margin of sternite VIII that is widely emarginate in males and entire to slightly sinuate in females, except in
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<taxonomicName id="FCB2C296242CFF97FCB3C0C3A98FFD32" authorityName="Dupuis" authorityYear="2014" box="[875,1004,658,683]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="rostratus">
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<emphasis id="09C66507242CFF97FCB3C0C3A98FFD32" box="[875,1004,658,683]" italics="true" pageId="10" pageNumber="438">T. rostratus</emphasis>
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</taxonomicName>
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in which it is narrowly emarginate in females.
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</paragraph>
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</subSubSection>
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<subSubSection id="73A8EA9E242CFF97FF4FC0ABAFFFF905" pageId="10" pageNumber="438" type="key">
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<keyStep id="8046A1B0242CFF97FF4FC0ABAFFFFCB5" pageId="10" pageNumber="438">
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<paragraph id="3B0DB915242CFF97FF4FC0ABAFFFFC96" blockId="10.[151,1437,761,1693]" box="[151,1436,761,784]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC0ABAFFFFC96" box="[151,1436,761,784]" pageId="10" pageNumber="438">
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1. Pronotum without tubercle (
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<figureCitation id="A389A590242CFF97FE28C0A8A839FC89" box="[496,602,761,784]" captionStart="FIGURES 19–22" captionStartId="6.[151,264,1816,1841]" captionTargetBox="[158,1435,1038,1747]" captionTargetId="figure@6.[153,1438,1059,1757]" captionTargetPageId="6" captionText="FIGURES 19–22. Tomarus peruvianus: 19, habitus; 20, head and pronotum in dorsal view; 21, parameres in frontal view; 22, locality records in Peru. Habitus photograph by Yony Callohuari." httpUri="https://zenodo.org/record/2839291/files/figure.png" pageId="10" pageNumber="438">Fig. 19–20</figureCitation>
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)................................................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="3B0DB915242CFF97FF4FC146AFFFFCB5" blockId="10.[151,1437,761,1693]" box="[151,1436,791,813]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC146AFFFFCB5" box="[151,1436,791,813]" pageId="10" pageNumber="438">
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- Pronotum with tubercle (
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<figureCitation id="A389A590242CFF97FE0AC146A869FCB4" box="[466,522,791,813]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1946,1971]" captionTargetBox="[159,1429,1129,1853]" captionTargetId="figure@1.[153,1437,1151,1859]" captionTargetPageId="1" captionText="FIGURES 1–4. Tomarus bituberculatus: 1, habitus; 2, parameres in frontal view; 3, parameres in lateral view; 4, locality records in Peru. Habitus photograph by Yony Callohuari." httpUri="https://zenodo.org/record/2839267/files/figure.png" pageId="10" pageNumber="438">Fig. 1</figureCitation>
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)...........................................................................3
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="8046A1B0242CFF97FF4FC165AFFEFC3B" pageId="10" pageNumber="438">
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||
<paragraph id="3B0DB915242CFF97FF4FC165AFFEFCFE" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC165AFFEFCFE" pageId="10" pageNumber="438">
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2. Protibia tridentate, with an additional basal denticle. Males with inner claw of protarsi enlarged and with an acute projection...........................................................................
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<taxonomicName id="FCB2C296242CFF97FBB6C103AFFEFCFE" authority="(Endrodi, 1970)" baseAuthorityName="Endrodi" baseAuthorityYear="1970" box="[1134,1437,849,871]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="peruvianus">
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<emphasis id="09C66507242CFF97FBB6C103AFFEFCFE" bold="true" box="[1134,1437,849,871]" pageId="10" pageNumber="438">
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<emphasis id="09C66507242CFF97FBB6C103AE96FCFE" bold="true" box="[1134,1269,850,871]" italics="true" pageId="10" pageNumber="438">T. peruvianus</emphasis>
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(Endrödi, 1970)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="3B0DB915242CFF97FF4FC13EAFFEFC3B" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC13EAFFEFC3B" pageId="10" pageNumber="438">
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- Protibia tridentate, without additional denticles. Males with inner claw of protarsi entire, equal in size and shape to the outer claw.......................................................................
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<taxonomicName id="FCB2C296242CFF97FBBFC1DCAFFEFC3B" authority="(Steinheil, 1872)" baseAuthorityName="Steinheil" baseAuthorityYear="1872" box="[1127,1437,908,930]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="burmeisteri">
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<emphasis id="09C66507242CFF97FBBFC1DCAFFEFC3B" bold="true" box="[1127,1437,908,930]" pageId="10" pageNumber="438">
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<emphasis id="09C66507242CFF97FBBFC1DCAE92FC3B" bold="true" box="[1127,1265,908,930]" italics="true" pageId="10" pageNumber="438">T. burmeisteri</emphasis>
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(
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<bibRefCitation id="5F23C4E4242CFF97FB25C1DDAFF6FC3B" author="Steinheil, E." box="[1277,1429,908,930]" pageId="10" pageNumber="438" pagination="554 - 578" refId="ref7570" refString="Steinheil, E. (1872) Symbolae ad historiam Coleopterorum Argentinae meridionalis, ossia enumerazione dei coleotteri raccolti dal Prof. Strobel, durante il suo soggiorno in Buenos Aires e nei viaggi de la intrapresi a Mendoza e nal Chili, indi a Bahia Blanca ed al Carmen de los Patagones; e descrizione delle specie nuove. Atti della Societa Italiana di Scienze Naturali, 15, 554 - 578." type="journal article" year="1872">Steinheil, 1872</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="8046A1B0242CFF97FF4FC1FBAFFFFC60" pageId="10" pageNumber="438">
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<paragraph id="3B0DB915242CFF97FF4FC1FBAFFEFC44" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC1FBAFFEFC44" pageId="10" pageNumber="438">
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3. Pronotum without fovea or rugose area behind tubercle. Antennal club longer than antennomeres 2–7...............................................................................................
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<taxonomicName id="FCB2C296242CFF97FBADC196AFFEFC44" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1141,1437,967,989]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="villosus">
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<emphasis id="09C66507242CFF97FBADC196AFFEFC44" bold="true" box="[1141,1437,967,989]" pageId="10" pageNumber="438">
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<emphasis id="09C66507242CFF97FBADC196AEBAFC44" bold="true" box="[1141,1241,967,989]" italics="true" pageId="10" pageNumber="438">T. villosus</emphasis>
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(
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<bibRefCitation id="5F23C4E4242CFF97FB3DC196AFF6FC44" author="Burmeister, H." box="[1253,1429,967,989]" pageId="10" pageNumber="438" refId="ref6200" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="3B0DB915242CFF97FF4FC1B4AFFFFC60" blockId="10.[151,1437,761,1693]" box="[151,1436,996,1018]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC1B4AFFFFC60" box="[151,1436,996,1018]" pageId="10" pageNumber="438">- Pronotum with deep fovea or rugose area behind tubercle. Antennal club subequal to antennomeres 2–7.................4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="8046A1B0242CFF97FF4FC653AFFFFBAD" pageId="10" pageNumber="438">
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<paragraph id="3B0DB915242CFF97FF4FC653AFFFFB8E" blockId="10.[151,1437,761,1693]" box="[151,1436,1025,1048]" pageId="10" pageNumber="438">
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||
<keyLead id="80431A20242CFF97FF4FC653AFFFFB8E" box="[151,1436,1025,1048]" pageId="10" pageNumber="438">
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4. Clypeus trapezoidal (base 2.0 times wider than apex) (
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<figureCitation id="A389A590242CFF97FD34C650A951FB81" box="[748,818,1025,1048]" captionStart="FIGURES 31–34" captionStartId="9.[151,264,948,973]" captionTargetBox="[165,1435,180,865]" captionTargetId="figure@9.[153,1438,179,871]" captionTargetPageId="9" captionText="FIGURES 31–34. Tomarus similis: 31, habitus; 32, parameres in frontal view; 33, parameres in lateral view; 34, locality records in Peru. Habitus photograph by Yony Callohauri." httpUri="https://zenodo.org/record/2839309/files/figure.png" pageId="10" pageNumber="438">Fig. 31</figureCitation>
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)................................................. 5
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</keyLead>
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</paragraph>
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<paragraph id="3B0DB915242CFF97FF4FC64EAFFFFBAD" blockId="10.[151,1437,761,1693]" box="[151,1436,1055,1077]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC64EAFFFFBAD" box="[151,1436,1055,1077]" pageId="10" pageNumber="438">
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- Clypeus strongly constricted towards apex (base 2.5–3.0 times wider than apex) (
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<figureCitation id="A389A590242CFF97FC37C64EAE57FBAC" box="[1007,1076,1055,1077]" captionStart="FIGURES 13–15" captionStartId="4.[151,264,1854,1879]" captionTargetBox="[166,1432,1063,1755]" captionTargetId="figure@4.[153,1438,1082,1764]" captionTargetPageId="4" captionText="FIGURES 13–15. Tomarus maimon: 13, habitus; 14, parameres in frontal view; 15, locality records in Peru. Habitus photograph by Yony Callohuari." httpUri="https://zenodo.org/record/2839279/files/figure.png" pageId="10" pageNumber="438">Fig. 13</figureCitation>
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).............................7
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="8046A1B0242CFF97FF4FC66DAFFFFB33" pageId="10" pageNumber="438">
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||
<paragraph id="3B0DB915242CFF97FF4FC66DAFFFFBF7" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
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<keyLead id="80431A20242CFF97FF4FC66DAFFFFBF7" pageId="10" pageNumber="438">
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5. Protibia with 3 large teeth and an additional basal denticle. Apex of metatibia slightly crenulate. Males with inner claw of pro- tarsi simple. Parameres strongly elongate, without lateral teeth, apices acute (
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<figureCitation id="A389A590242CFF97FC1FC608AE59FBE9" box="[967,1082,1113,1136]" captionStart="FIGURES 27–30" captionStartId="8.[151,264,1076,1101]" captionTargetBox="[153,1433,314,1031]" captionTargetId="figure@8.[153,1438,309,1037]" captionTargetPageId="8" captionText="FIGURES 27–30. Tomarus rostratus: 27, habitus; 28, parameres in frontal view; 29, parameres in lateral view; 30, locality records in Peru. Habitus photograph by Yony Callohauri." httpUri="https://zenodo.org/record/2839303/files/figure.png" pageId="10" pageNumber="438">Figs. 28–29</figureCitation>
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).......
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<taxonomicName id="FCB2C296242CFF97FB78C60BAFFFFBF7" authority="Dupuis, 2014" authorityName="Dupuis" authorityYear="2014" box="[1184,1436,1113,1135]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="rostratus">
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<emphasis id="09C66507242CFF97FB78C60BAFFFFBF7" bold="true" box="[1184,1436,1113,1135]" pageId="10" pageNumber="438">
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<emphasis id="09C66507242CFF97FB78C60BAF72FBF6" bold="true" box="[1184,1297,1114,1135]" italics="true" pageId="10" pageNumber="438">T. rostratus</emphasis>
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<bibRefCitation id="5F23C4E4242CFF97FACFC608AFFFFBF7" author="Dupuis, F." box="[1303,1436,1113,1135]" pageId="10" pageNumber="438" pagination="1 - 4" refId="ref6382" refString="Dupuis, F. (2014) Tomarus rostratus nouvelle espece du Perou (Coleoptera, Dynastidae). Coleopteres, 20, 1 - 4." type="journal article" year="2014">Dupuis, 2014</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</keyLead>
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</paragraph>
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<paragraph id="3B0DB915242CFF97FF4FC626AFFFFB33" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC626AFFFFB33" pageId="10" pageNumber="438">- Protibia with 3 large teeth, without additional basal denticle. Apex of metatibia dentate. Males with inner claw of protarsi en- larged and bifid. Parameres shorter and wider, with a small ventral tooth each side, apices rounded to truncate.............6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8046A1B0242CFF97FF4FC6E0AFFEFB7C" pageId="10" pageNumber="438">
|
||
<paragraph id="3B0DB915242CFF97FF4FC6E0AFFEFB5E" blockId="10.[151,1437,761,1693]" box="[151,1437,1201,1223]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC6E0AFFEFB5E" box="[151,1437,1201,1223]" pageId="10" pageNumber="438">
|
||
6. Parameres wide at base, strongly narrowed at apical third and dilated at apex (
|
||
<figureCitation id="A389A590242CFF97FC17C6E0AE48FB5E" box="[975,1067,1201,1223]" captionStart="FIGURES 5–8" captionStartId="2.[151,264,1872,1897]" captionTargetBox="[153,1435,1126,1812]" captionTargetId="figure@2.[153,1437,1121,1812]" captionTargetPageId="2" captionText="FIGURES 5–8. Tomarus ebenus: 5, habitus; 6, parameres in frontal view; 7, parameres in lateral view; 8, locality records in Peru." httpUri="https://zenodo.org/record/2839273/files/figure.png" pageId="10" pageNumber="438">Figs. 6–7</figureCitation>
|
||
).........
|
||
<taxonomicName id="FCB2C296242CFF97FB42C6E3AFFEFB5E" authority="(De Geer, 1774)" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[1178,1437,1201,1223]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="ebenus">
|
||
<emphasis id="09C66507242CFF97FB42C6E3AFFEFB5E" bold="true" box="[1178,1437,1201,1223]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FB42C6E3AE94FB5E" bold="true" box="[1178,1271,1201,1223]" italics="true" pageId="10" pageNumber="438">T. ebenus</emphasis>
|
||
(
|
||
<bibRefCitation id="5F23C4E4242CFF97FADDC6E0AFF6FB5E" author="De Geer, C." box="[1285,1429,1201,1223]" pageId="10" pageNumber="438" refId="ref6351" refString="De Geer, C. (1774) Memories pour Servir a l'Historie des Insectes. Vol. 4. L. L. Grefing, Stockholm, 456 pp." type="book" year="1774">De Geer, 1774</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="3B0DB915242CFF97FF4FC69EAFFEFB7C" blockId="10.[151,1437,761,1693]" box="[151,1437,1231,1253]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC69EAFFEFB7C" box="[151,1437,1231,1253]" pageId="10" pageNumber="438">
|
||
- Parameres slender, sides nearly parallel (
|
||
<figureCitation id="A389A590242CFF97FDB1C69EA8BFFB7C" box="[617,732,1231,1253]" captionStart="FIGURES 31–34" captionStartId="9.[151,264,948,973]" captionTargetBox="[165,1435,180,865]" captionTargetId="figure@9.[153,1438,179,871]" captionTargetPageId="9" captionText="FIGURES 31–34. Tomarus similis: 31, habitus; 32, parameres in frontal view; 33, parameres in lateral view; 34, locality records in Peru. Habitus photograph by Yony Callohauri." httpUri="https://zenodo.org/record/2839309/files/figure.png" pageId="10" pageNumber="438">Figs. 32–33</figureCitation>
|
||
)....................................
|
||
<taxonomicName id="FCB2C296242CFF97FB44C69EAFFEFB7C" authority="(Endrodi, 1968)" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[1180,1437,1231,1253]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="09C66507242CFF97FB44C69EAFFEFB7C" bold="true" box="[1180,1437,1231,1253]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FB44C69EAE96FB7C" bold="true" box="[1180,1269,1231,1253]" italics="true" pageId="10" pageNumber="438">T. similis</emphasis>
|
||
(Endrödi, 1968)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8046A1B0242CFF97FF4FC6BCAFFFFA86" pageId="10" pageNumber="438">
|
||
<paragraph id="3B0DB915242CFF97FF4FC6BCAFFFFA98" blockId="10.[151,1437,761,1693]" box="[151,1436,1260,1282]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC6BCAFFFFA98" box="[151,1436,1260,1282]" pageId="10" pageNumber="438">7. Frontal tubercles triangular to conical..................................................................... 8</keyLead>
|
||
</paragraph>
|
||
<paragraph id="3B0DB915242CFF97FF4FC75BAFFFFA86" blockId="10.[151,1437,761,1693]" box="[151,1436,1289,1311]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC75BAFFFFA86" box="[151,1436,1289,1311]" pageId="10" pageNumber="438">- Frontal tubercles transverse, difficult to see in lateral view.................................................... 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8046A1B0242CFF97FF4FC776AFFFFAEE" pageId="10" pageNumber="438">
|
||
<paragraph id="3B0DB915242CFF97FF4FC776AFFFFAA5" blockId="10.[151,1437,761,1693]" box="[151,1436,1319,1341]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC776AFFFFAA5" box="[151,1436,1319,1341]" pageId="10" pageNumber="438">8. Pronotal fovea transversely oval to rounded and deep. Pronotal tubercle acute in lateral view..........................9</keyLead>
|
||
</paragraph>
|
||
<paragraph id="3B0DB915242CFF97FF4FC714AFFFFAEE" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC714AFFFFAEE" pageId="10" pageNumber="438">
|
||
- Pronotal fovea narrow and shallow, as an elongate striate area. Pronotal tubercle rounded in lateral view...........................................................................................
|
||
<taxonomicName id="FCB2C296242CFF97FBBEC733AF2BFAEF" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[1126,1352,1377,1399]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="09C66507242CFF97FBBEC733AF2BFAEF" bold="true" box="[1126,1352,1377,1399]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FBBEC733AEC9FAEE" bold="true" box="[1126,1194,1378,1399]" italics="true" pageId="10" pageNumber="438">T. gyas</emphasis>
|
||
<bibRefCitation id="5F23C4E4242CFF97FB68C730AF2BFAEF" author="Erichson, W. F." box="[1200,1352,1377,1399]" pageId="10" pageNumber="438" pagination="553 - 617" refId="ref6626" refString="Erichson, W. F. (1848) Die Insekten. In: Schomburg, R. (Ed.), Reisen in British-Guiana in den Jahren 1840 - 1844. Vol. 3. J. J. Weber, Leipzig, pp. 553 - 617." type="book chapter" year="1848">Erichson, 1848</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8046A1B0242CFF97FF4FC72EAFFFFA28" pageId="10" pageNumber="438">
|
||
<paragraph id="3B0DB915242CFF97FF4FC72EAFFEFA0C" blockId="10.[151,1437,761,1693]" box="[151,1437,1407,1429]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC72EAFFEFA0C" box="[151,1437,1407,1429]" pageId="10" pageNumber="438">
|
||
9. Pronotal fovea transversely oval, as wide as interocular distance............ ..
|
||
<taxonomicName id="FCB2C296242CFF97FC07C72EAFFEFA0C" authority="(Palisot de Beauvois, 1811)" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1811" box="[991,1437,1407,1429]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="bituberculatus">
|
||
<emphasis id="09C66507242CFF97FC07C72EAFFEFA0C" bold="true" box="[991,1437,1407,1429]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FC07C72EAEE4FA0C" bold="true" box="[991,1159,1407,1429]" italics="true" pageId="10" pageNumber="438">T. bituberculatus</emphasis>
|
||
(
|
||
<bibRefCitation id="5F23C4E4242CFF97FB4BC72EAFF5FA0C" author="Palisot de Beauvois, A. M. F. J." box="[1171,1430,1407,1429]" pageId="10" pageNumber="438" pagination="101 - 120" refId="ref7053" refString="Palisot de Beauvois, A. M. F. J. (1811) Insectes Recueillis en Afrique et en Amerique, dans les Royaumes d'Oware et de Benin, a Saint-Dominigue et dans les Etats-Unis, pendant les Annees 1786 - 1797. Livraison 7. Levrault, Schoell, et Cie, Paris, 20 pp. [pp. 101 - 120]" type="book chapter" year="1811">Palisot de Beauvois, 1811</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="3B0DB915242CFF97FF4FC7CCAFFFFA28" blockId="10.[151,1437,761,1693]" box="[151,1436,1436,1458]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC7CCAFFFFA28" box="[151,1436,1436,1458]" pageId="10" pageNumber="438">- Pronotal fovea rounded, narrower (about 1/3–2/3 times interocular distance)..................................... 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8046A1B0242CFF97FF4FC7EBAFFFF9BE" pageId="10" pageNumber="438">
|
||
<paragraph id="3B0DB915242CFF97FF4FC7EBAFFFFA75" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC7EBAFFFFA75" pageId="10" pageNumber="438">
|
||
10. Pronotal fovea width 1/3 times interocular distance. Protibia tridentate without basal tooth. Apex of metatibia with 16–20 spi- nules. Parameres with a short ventral tooth each side (
|
||
<figureCitation id="A389A590242CFF97FD30C786A94EFA74" box="[744,813,1495,1517]" captionStart="FIGURES 13–15" captionStartId="4.[151,264,1854,1879]" captionTargetBox="[166,1432,1063,1755]" captionTargetId="figure@4.[153,1438,1082,1764]" captionTargetPageId="4" captionText="FIGURES 13–15. Tomarus maimon: 13, habitus; 14, parameres in frontal view; 15, locality records in Peru. Habitus photograph by Yony Callohuari." httpUri="https://zenodo.org/record/2839279/files/figure.png" pageId="10" pageNumber="438">Fig. 14</figureCitation>
|
||
)............................ ..
|
||
<taxonomicName id="FCB2C296242CFF97FB4CC786AFFFFA75" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" box="[1172,1436,1495,1517]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="maimon">
|
||
<emphasis id="09C66507242CFF97FB4CC786AFFFFA75" bold="true" box="[1172,1436,1495,1517]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FB4CC786AE9DFA75" bold="true" box="[1172,1278,1495,1516]" italics="true" pageId="10" pageNumber="438">T. maimon</emphasis>
|
||
<bibRefCitation id="5F23C4E4242CFF97FADCC786AFFFFA75" author="Erichson, W. F." box="[1284,1436,1495,1517]" pageId="10" pageNumber="438" pagination="67 - 185" refId="ref6597" refString="Erichson, W. F. (1847) Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte, 13, 67 - 185." type="journal article" year="1847">Erichson, 1847</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="3B0DB915242CFF97FF4FC7A4AFFFF9BE" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC7A4AFFFF9BE" pageId="10" pageNumber="438">
|
||
- Pronotal fovea width 2/3 times interocular distance. Protibia tridentate and with a fourth, basal, small tooth. Apex of metatibia with 25 spinules. Parameres with a long dorsal tooth on each side (
|
||
<figureCitation id="A389A590242CFF97FCBBC440A9BBF9B1" box="[867,984,1553,1576]" captionStart="FIGURES 9–12" captionStartId="3.[151,264,1828,1853]" captionTargetBox="[157,1429,1032,1770]" captionTargetId="figure@3.[153,1437,1052,1776]" captionTargetPageId="3" captionText="FIGURES 9–12. Tomarus gyas: 9, habitus; 10, parameres in frontal view; 11, parameres in lateral view; 12, locality records in Peru. Habitus photograph by Yony Callohuari." httpUri="https://zenodo.org/record/2839275/files/figure.png" pageId="10" pageNumber="438">Figs. 10–11</figureCitation>
|
||
)...........
|
||
<taxonomicName id="FCB2C296242CFF97FBBEC443AF2BF9BF" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[1126,1352,1553,1575]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="09C66507242CFF97FBBEC443AF2BF9BF" bold="true" box="[1126,1352,1553,1575]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FBBEC443AEC9F9BE" bold="true" box="[1126,1194,1554,1575]" italics="true" pageId="10" pageNumber="438">T. gyas</emphasis>
|
||
<bibRefCitation id="5F23C4E4242CFF97FB68C440AF2BF9BF" author="Erichson, W. F." box="[1200,1352,1553,1575]" pageId="10" pageNumber="438" pagination="553 - 617" refId="ref6626" refString="Erichson, W. F. (1848) Die Insekten. In: Schomburg, R. (Ed.), Reisen in British-Guiana in den Jahren 1840 - 1844. Vol. 3. J. J. Weber, Leipzig, pp. 553 - 617." type="book chapter" year="1848">Erichson, 1848</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8046A1B0242CFF97FF4FC47EAFFFF905" pageId="10" pageNumber="438">
|
||
<paragraph id="3B0DB915242CFF97FF4FC47EAFFEF9FB" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC47EAFFEF9FB" pageId="10" pageNumber="438">
|
||
11. Apex of metatibia with 14–17 spinules. Pronotum with sparse, small punctures. Pygidium of males rugopunctate on basal third.............................................................................
|
||
<taxonomicName id="FCB2C296242CFF97FB7CC41CAFFEF9FB" authority="(Prell, 1937)" baseAuthorityName="Prell" baseAuthorityYear="1937" box="[1188,1437,1612,1634]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="maternus">
|
||
<emphasis id="09C66507242CFF97FB7CC41CAFFEF9FB" bold="true" box="[1188,1437,1612,1634]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FB7CC41CAF7AF9FB" bold="true" box="[1188,1305,1613,1634]" italics="true" pageId="10" pageNumber="438">T. maternus</emphasis>
|
||
(
|
||
<bibRefCitation id="5F23C4E4242CFF97FAFEC41DAFF6F9FB" author="Prell, H." box="[1318,1429,1612,1634]" pageId="10" pageNumber="438" pagination="89 - 90" refId="ref7181" refString="Prell, H. (1937) Beitrage zur Kenntnis der Dynastiden. XV. Neue Arten und Rassen. Entomologische Zeitschrift, 51, 89 - 90." type="journal article" year="1937">Prell, 1937</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="3B0DB915242CFF97FF4FC43BAFFFF905" blockId="10.[151,1437,761,1693]" pageId="10" pageNumber="438">
|
||
<keyLead id="80431A20242CFF97FF4FC43BAFFFF905" pageId="10" pageNumber="438">
|
||
- Apex of metatibia with 33 spinules. Pronotum with dense, large punctures. Pygidium of males completely ru- gose…………...…...............................................
|
||
<taxonomicName id="FCB2C296242CFF97FC1AC4D6AFFFF905" authority="Lopez-Garcia & Deloya, 2019" authorityName="Lopez-Garcia & Deloya" authorityYear="2019" box="[962,1436,1670,1693]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="pilcopataensis">
|
||
<emphasis id="09C66507242CFF97FC1AC4D6AFFFF905" bold="true" box="[962,1436,1670,1693]" pageId="10" pageNumber="438">
|
||
<emphasis id="09C66507242CFF97FC1AC4D6AE04F904" bold="true" box="[962,1127,1671,1693]" italics="true" pageId="10" pageNumber="438">T. pilcopataensis</emphasis>
|
||
<bibRefCitation id="5F23C4E4242CFF97FBB5C4D7AFFFF905" author="Lopez-Garcia, M. M. & Deloya, C." box="[1133,1436,1670,1693]" pageId="10" pageNumber="438" pagination="127 - 141" refId="ref6849" refString="Lopez-Garcia, M. M. & Deloya, C. (2019) Five new species of the dynastine genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an illustrated key to species. The Coleopterists Bulletin, 73, 127 - 141. https: // doi. org / 10.1649 / 0010 - 065 X- 73.1.127" type="journal article" year="2019">López-García & Deloya, 2019</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |