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<document id="63B4B5A4CE55B63DFA479BAB798D7C2F" ID-DOI="10.11646/phytotaxa.601.3.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="8141856" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1689234448989" checkinUser="plazi" docAuthor="Trujillo, William, Jaramillo, M. Alejandra, Toro, Diego &amp; Balslev, Henrik" docDate="2023" docId="4B0087ACFFB14260FF28FE99FBFEDDEE" docLanguage="en" docName="phytotaxa.601.3.4.pdf" docOrigin="Phytotaxa 601 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.601.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Piper mituense Trel. &amp; Yunck." docType="treatment" docVersion="3" lastPageNumber="280" masterDocId="B739FFD4FFA04271FFA0FF93FFD6D946" masterDocTitle="Over- and under-described: new species, new synonyms, and a new name in the megadiverse genus Piper (Piperaceae) on the east Andean slopes" masterLastPageNumber="284" masterPageNumber="263" pageNumber="280" updateTime="1689274511015" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="403B6EEA796ACE04B5EAA1CD7A2F100B">Over- and under-described: new species, new synonyms, and a new name in the megadiverse genus Piper (Piperaceae) on the east Andean slopes</mods:title>
</mods:titleInfo>
<mods:name id="396AA53E6908BBAEB7B523D6CF2F14F9" type="personal">
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<mods:namePart id="C76EDADECE8AFD585EAA4A0D946FA803">Trujillo, William</mods:namePart>
<mods:nameIdentifier id="B8F8F0D321537AE87AAE2E3D0A3CEDC8" type="ORCID">0000-0002-0170-016X</mods:nameIdentifier>
<mods:affiliation id="AEE5328A486C5343D6D387F530C916D2">Grupo Investigaciones territoriales para el uso y conservación de la biodiversidad. Fundación La Palmita, Centro de Investigación. Cra 4 # 58 - 59 piso 2, Bogotá, Colombia. &amp; williamtrujilloca @ gmail. com; https: // orcid. org / 0000 - 0002 - 0170 - 016 X</mods:affiliation>
<mods:nameIdentifier id="C57F9336F7DCE08120C7780887278988" type="email">williamtrujilloca@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="87F7BF87A923304FF6A5B8118EA3C99A" type="personal">
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</mods:role>
<mods:namePart id="27C3A5EC0A259B7BAA3D972FD4300FAF">Jaramillo, M. Alejandra</mods:namePart>
<mods:nameIdentifier id="5211AD1E3D98859DB855C0682D6B960F" type="ORCID">0000-0002-6539-4149</mods:nameIdentifier>
<mods:affiliation id="6D41E1E06FDFFEE7A283EEC637EB49FC">Grupo DIVERSITAS, Facultad de Ciencias Básicas y Aplicadas, Universidad Militar Nueva Granada, km 2 Vía Cajicá-Zipaquirá, Cajicá, Colombia. &amp; maria. jaramillo @ unimilitar. edu. co; https: // orcid. org / 0000 - 0002 - 6539 - 4149</mods:affiliation>
<mods:nameIdentifier id="014167553F2D9E19EE23388ACDD80599" type="email">maria.jaramillo@unimilitar.edu.co</mods:nameIdentifier>
</mods:name>
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<mods:role id="946A0BA0B717A44A58C0DBEE456316FB">
<mods:roleTerm id="C4E0B7A074BEF8FA6F610E9DEB7B340E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7482B09629E86BADF0E3BE031C53F94">Toro, Diego</mods:namePart>
<mods:nameIdentifier id="FE0A3F75877246DEABBECA4E2FE1566B" type="ORCID">0009-0007-5970-8598</mods:nameIdentifier>
<mods:affiliation id="8CFEC39CC4F92DD52BED4BBF752E7091">Centro de Investigaciones Amazónicas CIMAZ-MACAGUAL, Laboratorio de Agrobiotecnología, Universidad de la Amazonia, Florencia 180002, Colombia. &amp; diegoalextoro @ gmail. com; https: // orcid. org / 0009 - 0007 - 5970 - 8598</mods:affiliation>
<mods:nameIdentifier id="DB553B0F78C66735203EE37F7BE73384" type="email">diegoalextoro@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2C05E71E1B93E9A36A2334686B4D1679" type="personal">
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<mods:roleTerm id="EEECB8052538E9510E5D1E28AC855CE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="601C35F21BCA3CACB9BEF1D86E3916E4">Balslev, Henrik</mods:namePart>
<mods:nameIdentifier id="E6EED5B76147FDAF5C3F9A325462074F" type="ORCID">0000-0002-7101-7120</mods:nameIdentifier>
<mods:affiliation id="28F1BEBFD6AA13FFB09A0142397254A6">Department of Biology, Ecoinformatics and Biodiversity, Aarhus University, DK- 8000 Aarhus C, Denmark. &amp; henrik. balslev @ bio. au. dk; https: // orcid. org / 0000 - 0002 - 7101 - 7120</mods:affiliation>
<mods:nameIdentifier id="7D589107ABCD2789938A949800377020" type="email">henrik.balslev@bio.au.dk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="D2AE9D9FC4058F1D28DE10DEB0A3EE4F">text</mods:typeOfResource>
<mods:relatedItem id="50C881FA5B92B2E5F254F7CBFAF87EBD" type="host">
<mods:titleInfo id="D570EE56E4C5406F42466C085CDF5838">
<mods:title id="1B3E57D44723E025F95A6A974B3FA5D7">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F39E171405C576516BB3DDBE92C63EFD">
<mods:date id="7DFFFC5F55CA5C149F5FD1AE988E3ACE">2023</mods:date>
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<mods:number id="D050637D817C07C960A07DB01CC3B73D">2023-07-11</mods:number>
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<mods:detail id="CAB4EC042810447F9F24AFD4AAFE1FC4" type="volume">
<mods:number id="4E009FFF6EB9D17C57642EB616B9B5AE">601</mods:number>
</mods:detail>
<mods:detail id="CAA0023AEC7C1AE2C2A4B64AF5691BC6" type="issue">
<mods:number id="42CC16BCFF5F9F36CA9256697326BA6A">3</mods:number>
</mods:detail>
<mods:extent id="C202F0EE465497AA719E8DD828AABE93" unit="page">
<mods:start id="0F92E8EC857B8C6ADF684191A0AA1971">263</mods:start>
<mods:end id="D97E7347A29C1D878A23AEC1EF187147">284</mods:end>
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<mods:location id="265BEECC7AFEFB6F5B421233C40E8A9C">
<mods:url id="5604DC2B8F0CC40B7AD8A4111379CEFD">http://dx.doi.org/10.11646/phytotaxa.601.3.4</mods:url>
</mods:location>
<mods:classification id="5A36DB80E3723BDFA26DB32FC1BE3CD3">journal article</mods:classification>
<mods:identifier id="19797C22029724E3753C1F5DC4F33A05" type="DOI">10.11646/phytotaxa.601.3.4</mods:identifier>
<mods:identifier id="1FE9633BBCA1C6DA611C2E5ED16F797B" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="DAB48E021F6E1769F882BB89F7B261B4" type="Zenodo-Dep">8141856</mods:identifier>
</mods:mods>
<treatment id="4B0087ACFFB14260FF28FE99FBFEDDEE" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE" httpUri="http://treatment.plazi.org/id/4B0087ACFFB14260FF28FE99FBFEDDEE" lastPageNumber="280" pageId="17" pageNumber="280">
<subSubSection id="8BB36531FFB14260FF28FE99FBB5D863" box="[136,1123,266,293]" pageId="17" pageNumber="280" type="nomenclature">
<paragraph id="C31636BAFFB14260FF28FE99FBB5D863" blockId="17.[136,1123,266,293]" box="[136,1123,266,293]" pageId="17" pageNumber="280">
<heading id="985E81D6FFB14260FF28FE99FBB5D863" box="[136,1123,266,293]" fontSize="11" level="2" pageId="17" pageNumber="280" reason="3">
<taxonomicName id="04A94D39FFB14260FF28FE99FE35D862" ID-CoL="4J4VL" authority="Trel. &amp; Yunck." authorityName="Trel. &amp; Yunck." box="[136,483,266,292]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="species" species="mituense">
<emphasis id="F1DDEAA8FFB14260FF28FE99FEE3D862" bold="true" box="[136,309,266,292]" italics="true" pageId="17" pageNumber="280">Piper mituense</emphasis>
Trel. &amp; Yunck.
</taxonomicName>
The
<treatmentCitation id="420810ABFFB14260FDBCFE99FB88D863" author="" box="[540,1118,266,293]" journal="The Piperaceae of northern South America" page="382" pageId="17" pageNumber="280" score="3.018367056741883" volume="1" year="1950">Piperaceae of northern South America 1: 382. 1950</treatmentCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BB36531FFB14260FF28FEC5FAE1D82D" box="[136,1335,342,364]" pageId="17" pageNumber="280" type="materials_examined">
<paragraph id="C31636BAFFB14260FF28FEC5FAE1D82D" blockId="17.[136,1452,342,436]" box="[136,1335,342,364]" pageId="17" pageNumber="280">
<materialsCitation id="73C13CE7FFB14260FF28FEC5FE7DD82D" box="[136,427,342,363]" country="Colombia" location="Vaupes" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Vaupes">
Type:—
<collectingCountry id="BBBE762AFFB14260FF76FEC5FE8CD82D" box="[214,346,342,363]" name="Colombia" pageId="17" pageNumber="280">COLOMBIA</collectingCountry>
:
<collectingRegion id="016DF858FFB14260FEC3FEC5FE7DD82D" box="[355,427,342,363]" country="Colombia" name="Vaupes" pageId="17" pageNumber="280">Vaupés</collectingRegion>
</materialsCitation>
,
<materialsCitation id="73C13CE7FFB14260FE14FEC5FAE5D82D" box="[436,1331,342,364]" collectingDate="1939-09-14" collectionCode="E, J, A" country="United States of America" elevation="200" location="Cuatrecasas" municipality="Perez Arbelaez" pageId="17" pageNumber="280" specimenCount="1" typeStatus="holotype">
Mitú,
<quantity id="04519B5FFFB14260FE4FFEC5FDF8D82D" box="[495,558,342,363]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="280" unit="m" value="200.0">
<elevation id="4884D189FFB14260FE4FFEC5FDF8D82D" box="[495,558,342,363]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="280" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="B717107AFFB14260FD97FEC5FD22D82D" box="[567,756,342,364]" pageId="17" pageNumber="280" value="1939-09-14">
<collectingDate id="A753E992FFB14260FD97FEC5FD22D82D" box="[567,756,342,364]" pageId="17" pageNumber="280" value="1939-09-14">14 September 1939</collectingDate>
</date>
,
<emphasis id="F1DDEAA8FFB14260FD5EFEC4FB4CD82D" box="[766,1178,342,364]" italics="true" pageId="17" pageNumber="280">
<collectionCode id="A5B8AE7FFFB14260FD5EFEC4FCC6D82A" box="[766,784,343,364]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="17" pageNumber="280" type="Herbarium">E</collectionCode>
.
<collectingMunicipality id="2372ACC0FFB14260FCB9FEC5FC7FD82D" box="[793,937,342,363]" pageId="17" pageNumber="280">Pérez Arbeláez</collectingMunicipality>
&amp;
<collectionCode id="A5B8AE7FFFB14260FC68FEC4FC00D82A" box="[968,982,343,364]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="17" pageNumber="280" type="Herbarium">J</collectionCode>
.
<location id="C6766061FFB14260FC7FFEC5FB80D82D" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FC7FFEC5FB80D82D" box="[991,1110,342,363]" country="United States of America" municipality="Perez Arbelaez" name="Cuatrecasas" pageId="17" pageNumber="280">Cuatrecasas</location>
6810
<collectionCode id="A5B8AE7FFFB14260FB2BFEC5FB4CD82D" box="[1163,1178,342,363]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="17" pageNumber="280" type="Herbarium">A</collectionCode>
</emphasis>
(
<typeStatus id="1C128818FFB14260FB06FEC5FB2AD82A" box="[1190,1276,342,364]" pageId="17" pageNumber="280">holotype</typeStatus>
<collectingCountry id="BBBE762AFFB14260FAA2FEC5FAFED82D" box="[1282,1320,342,363]" name="United States of America" pageId="17" pageNumber="280">US</collectingCountry>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BB36531FFB14260FF28FEE8FA80D8F2" pageId="17" pageNumber="280" type="reference_group">
<paragraph id="C31636BAFFB14260FF28FEE8FA80D8F2" blockId="17.[136,1452,342,436]" pageId="17" pageNumber="280">
<emphasis id="F1DDEAA8FFB14260FF28FEE8FF4ED8D6" box="[136,152,379,400]" italics="true" pageId="17" pageNumber="280">=</emphasis>
<taxonomicName id="04A94D39FFB14260FF3EFEE9FDA3D8D6" authority="Trel. &amp; Yunck." authorityName="Trel. &amp; Yunck." box="[158,629,378,400]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="variety" species="metanum" variety="villibracteum">
<emphasis id="F1DDEAA8FFB14260FF3EFEE9FEE7D8D6" box="[158,305,378,400]" italics="true" pageId="17" pageNumber="280">Piper metanum</emphasis>
<materialsCitation id="73C13CE7FFB14260FE96FEE9FDA3D8D6" box="[310,629,378,400]" collectingDate="1939-09-14" collectionCode="E, J" country="Colombia" county="The" elevation="200" location="Type" municipality="northern South America" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Meta" typeStatus="holotype">
var.
<emphasis id="F1DDEAA8FFB14260FEC1FEE9FE37D8C9" box="[353,481,378,399]" italics="true" pageId="17" pageNumber="280">villibracteum</emphasis>
Trel. &amp; Yunck.
</materialsCitation>
</taxonomicName>
<collectingCounty id="2A774E36FFB14260FDD9FEE9FD48D8D6" box="[633,670,378,400]" pageId="17" pageNumber="280">The</collectingCounty>
<treatmentCitation id="420810ABFFB14260FD04FEE9FB47D8D6" author="" box="[676,1169,378,400]" journal="The Piperaceae of northern South America" page="343" pageId="17" pageNumber="280" score="3.018367056741883" volume="1" year="1950">
<taxonomicName id="04A94D39FFB14260FD04FEE9FCDAD8D6" ID-CoL="625NL" box="[676,780,378,400]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
of
<collectingMunicipality id="2372ACC0FFB14260FC8BFEE9FBC7D8D6" box="[811,1041,378,400]" pageId="17" pageNumber="280">northern South America</collectingMunicipality>
1: 343. 1950
</treatmentCitation>
.
<location id="C6766061FFB14260FB39FEE9FB1DD8C9" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FB39FEE9FB1DD8C9" box="[1177,1227,378,399]" country="Colombia" county="The" municipality="northern South America" name="Type" pageId="17" pageNumber="280" stateProvince="Meta">Type</location>
:—
<collectingCountry id="BBBE762AFFB14260FB47FEE9FABDD8C9" box="[1255,1387,378,399]" name="Colombia" pageId="17" pageNumber="280">COLOMBIA</collectingCountry>
:
<collectingRegion id="016DF858FFB14260FAD4FEE9FA7ED8C9" box="[1396,1448,378,399]" country="Colombia" name="Meta" pageId="17" pageNumber="280">Meta</collectingRegion>
,
<location id="C6766061FFB14260FF1DFE0DFF38D8F2" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FF1DFE0DFF38D8F2" box="[189,238,414,436]" country="Colombia" county="The" municipality="northern South America" name="Mitu" pageId="17" pageNumber="280" stateProvince="Meta">Mitú</location>
, at
<collectingRegion id="016DF858FFB14260FEAFFE0DFE82D8F5" box="[271,340,414,435]" country="Colombia" name="Vaupes" pageId="17" pageNumber="280">Vaupés</collectingRegion>
boundary,
<quantity id="04519B5FFFB14260FE1FFE0DFE28D8F5" box="[447,510,414,435]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="280" unit="m" value="200.0">
<elevation id="4884D189FFB14260FE1FFE0DFE28D8F5" box="[447,510,414,435]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="280" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="B717107AFFB14260FDA8FE0DFD13D8F5" box="[520,709,414,436]" pageId="17" pageNumber="280" value="1939-09-14">
<collectingDate id="A753E992FFB14260FDA8FE0DFD13D8F5" box="[520,709,414,436]" pageId="17" pageNumber="280" value="1939-09-14">14 September 1939</collectingDate>
</date>
,
<emphasis id="F1DDEAA8FFB14260FD6FFE0CFB89D8F5" box="[719,1119,414,436]" italics="true" pageId="17" pageNumber="280">
<collectionCode id="A5B8AE7FFFB14260FD6FFE0CFD37D8F2" box="[719,737,415,436]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="17" pageNumber="280" type="Herbarium">E</collectionCode>
. Pérez-Arbeláez &amp;
<collectionCode id="A5B8AE7FFFB14260FC3BFE0CFC7FD8F2" box="[923,937,415,436]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="17" pageNumber="280" type="Herbarium">J</collectionCode>
. Cuatrecasas 6810
</emphasis>
(
<typeStatus id="1C128818FFB14260FBCBFE0DFB17D8F2" box="[1131,1217,414,436]" pageId="17" pageNumber="280">holotype</typeStatus>
US!).
<emphasis id="F1DDEAA8FFB14260FAA1FE0CFA80D8F2" box="[1281,1366,415,436]" italics="true" pageId="17" pageNumber="280">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="8BB36531FFB14260FF28FE71FAC5DA23" pageId="17" pageNumber="280" type="discussion">
<paragraph id="C31636BAFFB14260FF28FE71FAC5DA23" blockId="17.[136,1452,482,1193]" pageId="17" pageNumber="280">
<emphasis id="F1DDEAA8FFB14260FF28FE71FEDFD8BA" bold="true" box="[136,265,482,508]" pageId="17" pageNumber="280">Comments</emphasis>
:—
<taxonomicName id="04A94D39FFB14260FE8DFE70FCD8D8BA" authority="Trel. &amp; Yunck. (1950: 382)" authorityName="Trel. &amp; Yunck." authorityPageNumber="382" authorityYear="1950" box="[301,782,482,509]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="species" species="mituense">
<emphasis id="F1DDEAA8FFB14260FE8DFE70FE00D8BA" box="[301,470,483,508]" italics="true" pageId="17" pageNumber="280">Piper mituense</emphasis>
Trel. &amp; Yunck. (1950: 382)
</taxonomicName>
was described from the collection
<emphasis id="F1DDEAA8FFB14260FB37FE70FE87DB66" italics="true" pageId="17" pageNumber="280">E. Pérez Arbelaez and J. Cuatrecasas 6810</emphasis>
and
<taxonomicName id="04A94D39FFB14260FE24FD94FC28DB66" authority="Trel. &amp; Yunck. (1950: 343)" authorityName="Trel. &amp; Yunck." authorityPageNumber="343" authorityYear="1950" box="[388,1022,518,545]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="variety" species="metanum" variety="villibracteum">
<emphasis id="F1DDEAA8FFB14260FE24FD94FDD3DB66" box="[388,517,519,544]" italics="true" pageId="17" pageNumber="280">P. metanum</emphasis>
var.
<emphasis id="F1DDEAA8FFB14260FD99FD95FD18DB66" box="[569,718,518,544]" italics="true" pageId="17" pageNumber="280">villibracteum</emphasis>
Trel. &amp; Yunck. (1950: 343)
</taxonomicName>
was described from
<emphasis id="F1DDEAA8FFB14260FB42FD94FE7ADB02" italics="true" pageId="17" pageNumber="280">E. Pérez-Arbelaez and J. Cuatrecasas 6810A</emphasis>
. The two collections were made at the same locality and on the same day. The protologue of
<taxonomicName id="04A94D39FFB14260FF28FDDCFEDEDB2E" box="[136,264,591,616]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="species" species="mituense">
<emphasis id="F1DDEAA8FFB14260FF28FDDCFEDEDB2E" box="[136,264,591,616]" italics="true" pageId="17" pageNumber="280">P. mituense</emphasis>
</taxonomicName>
highlighted: “the villous twigs, pinnately nerved, oblong-elliptic leaves, and short spikes with substylose ovaries characterize this species”, while for
<taxonomicName id="04A94D39FFB14260FDD7FDE0FC11DBCA" box="[631,967,626,653]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="variety" species="metanum" variety="villibracteum">
<emphasis id="F1DDEAA8FFB14260FDD7FDE0FD2CDBCA" box="[631,762,627,652]" italics="true" pageId="17" pageNumber="280">P. metanum</emphasis>
var.
<emphasis id="F1DDEAA8FFB14260FC92FDE1FC11DBCA" box="[818,967,626,652]" italics="true" pageId="17" pageNumber="280">villibracteum</emphasis>
</taxonomicName>
the authors note: “Differs from the species in having somewhat less pubescent twigs, slightly thicker leaves, and with the floral bracts rounded and densely long-fringed”. In addition,
<taxonomicName id="04A94D39FFB14260FE16FD28FDEFDB92" box="[438,569,699,724]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="species" species="metanum">
<emphasis id="F1DDEAA8FFB14260FE16FD28FDEFDB92" box="[438,569,699,724]" italics="true" pageId="17" pageNumber="280">P. metanum</emphasis>
</taxonomicName>
is remarkably different from the
<taxonomicName id="04A94D39FFB14260FC0CFD29FB8CDB92" box="[940,1114,698,724]" pageId="17" pageNumber="280" rank="variety" variety="because">variety because</taxonomicName>
it has spikes
<quantity id="04519B5FFFB14260FB50FD29FA95DB93" box="[1264,1347,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="280" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
long and erect (vs. spikes
<quantity id="04519B5FFFB14260FEE3FD4DFE7DDBBF" box="[323,427,734,761]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="17" pageNumber="280" unit="cm" value="1.4" valueMax="1.8" valueMin="1.0">1–1.8 cm</quantity>
long and pendulous). The protologues do not recognize the similarity between the two taxa discussed here. After a detailed examination, we conclude that there are no notable morphological differences that allow considering the
<specimenCount id="D5AFFD33FFB14260FEDDFCB4FDF4DA06" box="[381,546,806,833]" pageId="17" pageNumber="280" type="generic">two specimens</specimenCount>
as belonging to different species, for which reason we propose to synonymize the variety
<taxonomicName id="04A94D39FFB14260FF7DFCD8FDFDDA22" box="[221,555,842,869]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="variety" species="metanum" variety="villibracteum">
<emphasis id="F1DDEAA8FFB14260FF7DFCD8FE89DA22" box="[221,351,843,868]" italics="true" pageId="17" pageNumber="280">P. metanum</emphasis>
var.
<emphasis id="F1DDEAA8FFB14260FE36FCD9FDFDDA22" box="[406,555,842,868]" italics="true" pageId="17" pageNumber="280">villibracteum</emphasis>
</taxonomicName>
under the species
<taxonomicName id="04A94D39FFB14260FD5AFCD8FCACDA22" box="[762,890,843,868]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="species" species="mituense">
<emphasis id="F1DDEAA8FFB14260FD5AFCD8FCACDA22" box="[762,890,843,868]" italics="true" pageId="17" pageNumber="280">P. mituense</emphasis>
</taxonomicName>
, giving priority to the species name.
</paragraph>
</subSubSection>
<subSubSection id="8BB36531FFB14260FF1DFCFDFC53DAEA" pageId="17" pageNumber="280" type="distribution">
<paragraph id="C31636BAFFB14260FF1DFCFDFC53DAEA" blockId="17.[136,1452,482,1193]" pageId="17" pageNumber="280">
<emphasis id="F1DDEAA8FFB14260FF1DFCFDFE36DACE" bold="true" box="[189,480,878,904]" pageId="17" pageNumber="280">Distribution and habitat</emphasis>
:—
<taxonomicName id="04A94D39FFB14260FDA3FCFCFD7ADACE" box="[515,684,879,904]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="17" pageNumber="280" phylum="Tracheophyta" rank="species" species="mituense">
<emphasis id="F1DDEAA8FFB14260FDA3FCFCFD7ADACE" box="[515,684,879,904]" italics="true" pageId="17" pageNumber="280">Piper mituense</emphasis>
</taxonomicName>
is distributed in the Amazon of
<collectingCountry id="BBBE762AFFB14260FBB1FCFDFBA9DACE" box="[1041,1151,878,904]" name="Colombia" pageId="17" pageNumber="280">Colombia</collectingCountry>
and
<collectingCountry id="BBBE762AFFB14260FB15FCFDFAC2DACE" box="[1205,1300,878,904]" name="Ecuador" pageId="17" pageNumber="280">Ecuador</collectingCountry>
, at elevations below
<specimenCount id="D5AFFD33FFB14260FF73FC01FEF4DAEB" box="[211,290,914,941]" pageId="17" pageNumber="280" type="generic">
<quantity id="04519B5FFFB14260FF73FC01FECBDAEB" box="[211,285,914,941]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="280" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
It is a shade-loving species growing in the understory.
</paragraph>
</subSubSection>
<subSubSection id="8BB36531FFB14260FF1DFC25FBFEDDEE" pageId="17" pageNumber="280" type="materials_examined">
<paragraph id="C31636BAFFB14260FF1DFC25FBFEDDEE" blockId="17.[136,1452,482,1193]" pageId="17" pageNumber="280">
<emphasis id="F1DDEAA8FFB14260FF1DFC25FDE5DA96" bold="true" box="[189,563,950,976]" pageId="17" pageNumber="280">Additional specimens examined</emphasis>
:—
<materialsCitation id="73C13CE7FFB14260FDF4FC25FDE4DAB2" collectingDate="2010-03-13" collectionCode="W" collectorName="Florencia &amp; Sebastopol" country="Colombia" county="Albania" elevation="246" latitude="1.4998056" location="El Porvenir" longLatPrecision="1" longitude="-75.65608" municipality="Florida" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Caqueta">
<collectingCountry id="BBBE762AFFB14260FDF4FC25FD26DA96" box="[596,752,950,976]" name="Colombia" pageId="17" pageNumber="280">COLOMBIA</collectingCountry>
:
<collectingRegion id="016DF858FFB14260FD58FC25FC81DA96" box="[760,855,950,976]" country="Colombia" name="Caqueta" pageId="17" pageNumber="280">Caquetá</collectingRegion>
,
<collectingCounty id="2A774E36FFB14260FCFEFC25FC6CDA96" box="[862,954,950,976]" pageId="17" pageNumber="280">Albania</collectingCounty>
, trail to
<collectingMunicipality id="2372ACC0FFB14260FBAFFC25FBB6DA96" box="[1039,1120,950,976]" pageId="17" pageNumber="280">Florida</collectingMunicipality>
1,
<location id="C6766061FFB14260FBDEFC25FB2ADA96" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FBDEFC25FB2ADA96" box="[1150,1276,950,976]" country="Colombia" county="Albania" municipality="Florida" name="El Porvenir" pageId="17" pageNumber="280" stateProvince="Caqueta">El Porvenir</location>
farm,
<date id="B717107AFFB14260FAE2FC25FF15DAB2" pageId="17" pageNumber="280" value="2010-03-13">
<collectingDate id="A753E992FFB14260FAE2FC25FF15DAB2" pageId="17" pageNumber="280" value="2010-03-13">13 March 2010</collectingDate>
</date>
,
<collectionCode id="A5B8AE7FFFB14260FF6DFC48FF33DAB2" box="[205,229,987,1012]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="280" type="Herbarium">
<emphasis id="F1DDEAA8FFB14260FF6DFC48FF33DAB2" box="[205,229,987,1012]" italics="true" pageId="17" pageNumber="280">W</emphasis>
</collectionCode>
<emphasis id="F1DDEAA8FFB14260FF45FC48FF3FDAB2" box="[229,233,987,1012]" italics="true" pageId="17" pageNumber="280">.</emphasis>
<emphasis id="F1DDEAA8FFB14260FF4EFC49FE6DDAB2" box="[238,443,986,1012]" italics="true" pageId="17" pageNumber="280">
<location id="C6766061FFB14260FF4EFC49FE94DAB2" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FF4EFC49FE94DAB2" box="[238,322,986,1012]" country="Colombia" name="Trujillo" pageId="17" pageNumber="280">Trujillo</location>
et al. 1358
</emphasis>
(
<collectionCode id="A5B8AE7FFFB14260FE69FC49FDF4DAB2" box="[457,546,986,1012]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="17" pageNumber="280" type="Herbarium">HUAZ</collectionCode>
!).
</materialsCitation>
<materialsCitation id="73C13CE7FFB14260FD97FC49FA8ADAB2" box="[567,1372,986,1013]" collectingDate="2010-03-13" collectionCode="W" collectorName="Florencia &amp; Sebastopol" country="Colombia" county="Albania" elevation="246" latitude="1.4998056" location="El Porvenir" longLatPrecision="1" longitude="-75.65608" municipality="Florida" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Caqueta">
Florencia, trail to Sebastopol,
<date id="B717107AFFB14260FC22FC49FB98DAB2" box="[898,1102,986,1013]" pageId="17" pageNumber="280" value="2010-09-05">
<collectingDate id="A753E992FFB14260FC22FC49FB98DAB2" box="[898,1102,986,1013]" pageId="17" pageNumber="280" value="2010-09-05">5 September 2010</collectingDate>
</date>
,
<collectionCode id="A5B8AE7FFFB14260FBF7FC48FBBCDAB2" box="[1111,1130,987,1012]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="17" pageNumber="280" type="Herbarium">
<emphasis id="F1DDEAA8FFB14260FBF7FC48FBBCDAB2" box="[1111,1130,987,1012]" italics="true" pageId="17" pageNumber="280">L</emphasis>
</collectionCode>
<emphasis id="F1DDEAA8FFB14260FBCAFC48FBB8DAB2" box="[1130,1134,987,1012]" italics="true" pageId="17" pageNumber="280">.</emphasis>
<emphasis id="F1DDEAA8FFB14260FBD3FC48FB32DAB2" box="[1139,1252,986,1013]" italics="true" pageId="17" pageNumber="280">
<location id="C6766061FFB14260FBD3FC48FB14DAB3" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FBD3FC48FB14DAB3" box="[1139,1218,987,1013]" country="Colombia" name="Zuniga" pageId="17" pageNumber="280">Zúñiga</location>
02
</emphasis>
(
<collectionCode id="A5B8AE7FFFB14260FB52FC49FA9DDAB2" box="[1266,1355,986,1012]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="17" pageNumber="280" type="Herbarium">HUAZ</collectionCode>
!);
</materialsCitation>
<materialsCitation id="73C13CE7FFB14260FAC1FC49FB98DD5E" collectingDate="2010-03-13" collectionCode="W" collectorName="Florencia &amp; Sebastopol" country="Colombia" county="Albania" elevation="246" latitude="1.4998056" location="El Porvenir" longLatPrecision="1" longitude="-75.65608" municipality="Florida" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Caqueta">
Center Macagual, Monitoreo research station,
<date id="B717107AFFB14260FD9FFC6DFD01DD5E" box="[575,727,1022,1049]" pageId="17" pageNumber="280" value="2008-06-30">
<collectingDate id="A753E992FFB14260FD9FFC6DFD01DD5E" box="[575,727,1022,1049]" pageId="17" pageNumber="280" value="2008-06-30">30 June 2008</collectingDate>
</date>
,
<collectionCode id="A5B8AE7FFFB14260FD42FC6CFD2BDD5E" box="[738,765,1023,1048]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="17" pageNumber="280" type="Herbarium">
<emphasis id="F1DDEAA8FFB14260FD42FC6CFD2BDD5E" box="[738,765,1023,1048]" italics="true" pageId="17" pageNumber="280">M</emphasis>
</collectionCode>
<emphasis id="F1DDEAA8FFB14260FD5DFC6CFCD7DD5E" box="[765,769,1023,1048]" italics="true" pageId="17" pageNumber="280">.</emphasis>
<emphasis id="F1DDEAA8FFB14260FCA7FC6CFC03DD5E" box="[775,981,1022,1049]" italics="true" pageId="17" pageNumber="280">
<collectingMunicipality id="2372ACC0FFB14260FCA7FC6CFC81DD5F" box="[775,855,1023,1049]" pageId="17" pageNumber="280">Correa</collectingMunicipality>
et al. 6648
</emphasis>
(
<collectionCode id="A5B8AE7FFFB14260FC45FC6DFBE8DD5E" box="[997,1086,1022,1048]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="17" pageNumber="280" type="Herbarium">HUAZ</collectionCode>
!);
</materialsCitation>
<materialsCitation id="73C13CE7FFB14260FBF5FC6DFE63DD26" collectingDate="2010-03-13" collectionCode="W" collectorName="Florencia &amp; Sebastopol" country="Colombia" county="Albania" elevation="246" latitude="1.4998056" location="El Porvenir" longLatPrecision="1" longitude="-75.65608" municipality="Florida" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Caqueta">
San Martin, trail to El Venado,
<location id="C6766061FFB14260FF28FBB1FCC0DD7A" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FF28FBB1FCC0DD7A" box="[136,790,1058,1084]" country="Colombia" latitude="1.4998056" longLatPrecision="1" longitude="-75.65608" municipality="Correa" name="Macagual Research Center of Universidad de la Amazonia" pageId="17" pageNumber="280">Macagual Research Center of Universidad de la Amazonia</location>
,
<quantity id="04519B5FFFB14260FC80FBB1FCBEDD7B" box="[800,872,1058,1085]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="17" pageNumber="280" unit="m" value="246.0">
<elevation id="4884D189FFB14260FC80FBB1FCBEDD7B" box="[800,872,1058,1085]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="17" pageNumber="280" unit="m" value="246.0">246 m</elevation>
</quantity>
,
<geoCoordinate id="A69D507DFFB14260FCD3FBB1FBC2DD7B" box="[883,1044,1058,1085]" degrees="01" direction="north" minutes="29" orientation="latitude" pageId="17" pageNumber="280" precision="1" seconds="59.3" value="1.4998056">01°29’59.3”N</geoCoordinate>
,
<geoCoordinate id="A69D507DFFB14260FBBEFBB1FB15DD7B" box="[1054,1219,1058,1085]" degrees="75" direction="west" minutes="39" orientation="longitude" pageId="17" pageNumber="280" precision="1" seconds="21.9" value="-75.65608">75°39’21.9”W</geoCoordinate>
,
<date id="B717107AFFB14260FB6DFBB1FA71DD7A" box="[1229,1447,1058,1084]" pageId="17" pageNumber="280" value="2011-09-30">
<collectingDate id="A753E992FFB14260FB6DFBB1FA71DD7A" box="[1229,1447,1058,1084]" pageId="17" pageNumber="280" value="2011-09-30">30 September 2011</collectingDate>
</date>
,
<collectionCode id="A5B8AE7FFFB14260FF28FBD4FF75DD26" box="[136,163,1095,1120]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="17" pageNumber="280" type="Herbarium">
<emphasis id="F1DDEAA8FFB14260FF28FBD4FF75DD26" box="[136,163,1095,1120]" italics="true" pageId="17" pageNumber="280">M</emphasis>
</collectionCode>
<emphasis id="F1DDEAA8FFB14260FF03FBD4FF71DD26" box="[163,167,1095,1120]" italics="true" pageId="17" pageNumber="280">.</emphasis>
<emphasis id="F1DDEAA8FFB14260FF0EFBD4FEEBDD26" box="[174,317,1094,1121]" italics="true" pageId="17" pageNumber="280">
<location id="C6766061FFB14260FF0EFBD4FF28DD27" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FF0EFBD4FF28DD27" box="[174,254,1095,1121]" country="Colombia" latitude="1.2319167" longLatPrecision="1" longitude="-75.491745" name="Correa" pageId="17" pageNumber="280">Correa</location>
7120
</emphasis>
(
<collectionCode id="A5B8AE7FFFB14260FEEDFBD5FE70DD26" box="[333,422,1094,1120]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="17" pageNumber="280" type="Herbarium">HUAZ</collectionCode>
!).
</materialsCitation>
<materialsCitation id="73C13CE7FFB14260FE1CFBD5FD52DDC2" collectingDate="2010-03-13" collectionCode="W" collectorName="Florencia &amp; Sebastopol" country="Colombia" county="Albania" elevation="246" latitude="1.4998056" location="El Porvenir" longLatPrecision="1" longitude="-75.65608" municipality="Florida" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Caqueta">
Milán: Bombay, La Juansoco canyon,
<quantity id="04519B5FFFB14260FCCCFBD5FC63DD27" box="[876,949,1094,1121]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="280" unit="m" value="230.0">
<elevation id="4884D189FFB14260FCCCFBD5FC63DD27" box="[876,949,1094,1121]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="280" unit="m" value="230.0">230 m</elevation>
</quantity>
,
<geoCoordinate id="A69D507DFFB14260FC62FBD5FB82DD27" box="[962,1108,1094,1121]" degrees="1" direction="north" minutes="13" orientation="latitude" pageId="17" pageNumber="280" precision="1" seconds="54.9" value="1.2319167">1°13’54.9”N</geoCoordinate>
,
<geoCoordinate id="A69D507DFFB14260FBC0FBD5FAD2DD27" box="[1120,1284,1094,1121]" degrees="75" direction="west" minutes="29" orientation="longitude" pageId="17" pageNumber="280" precision="1" seconds="30.3" value="-75.491745">75°29’30.3”W</geoCoordinate>
,
<date id="B717107AFFB14260FAB0FBD5FF15DDC2" pageId="17" pageNumber="280" value="2007-09-18">
<collectingDate id="A753E992FFB14260FAB0FBD5FF15DDC2" pageId="17" pageNumber="280" value="2007-09-18">18 September 2007</collectingDate>
</date>
,
<collectionCode id="A5B8AE7FFFB14260FF6CFBF8FF32DDC2" box="[204,228,1131,1156]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="17" pageNumber="280" type="Herbarium">
<emphasis id="F1DDEAA8FFB14260FF6CFBF8FF32DDC2" box="[204,228,1131,1156]" italics="true" pageId="17" pageNumber="280">W</emphasis>
</collectionCode>
<emphasis id="F1DDEAA8FFB14260FF45FBF8FF3FDDC2" box="[229,233,1131,1156]" italics="true" pageId="17" pageNumber="280">.</emphasis>
<emphasis id="F1DDEAA8FFB14260FF4DFBF9FE7BDDC2" box="[237,429,1130,1156]" italics="true" pageId="17" pageNumber="280">
<location id="C6766061FFB14260FF4DFBF9FE97DDC2" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FF4DFBF9FE97DDC2" box="[237,321,1130,1156]" country="Colombia" name="Trujillo" pageId="17" pageNumber="280">Trujillo</location>
et al. 713
</emphasis>
(
<collectionCode id="A5B8AE7FFFB14260FE1AFBF9FDC7DDC2" box="[442,529,1130,1156]" country="Colombia" httpUri="http://grbio.org/cool/2v9m-7g3r" name="Universidad de la Amazonia" pageId="17" pageNumber="280" type="Herbarium">HUAZ</collectionCode>
!,
<collectionCode id="A5B8AE7FFFB14260FDBFFBF9FDAFDDC2" box="[543,633,1130,1156]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15419" lsid="urn:lsid:biocol.org:col:15419" name="Instituto Amazónico de Investigaciones Científicas SINCHI" pageId="17" pageNumber="280" type="Herbarium">COAH</collectionCode>
!)
</materialsCitation>
.
<materialsCitation id="73C13CE7FFB14260FD2DFBF9FBF3DDEE" collectingDate="1985-07-04" collectionCode="H" collectorName="Anangu &amp; Rio" country="Ecuador" elevation="246" latitude="-0.53333336" location="Napo" longLatPrecision="1308" longitude="-76.38333" pageId="17" pageNumber="280" specimenCount="1" stateProvince="Napo">
<collectingCountry id="BBBE762AFFB14260FD2DFBF9FCCFDDC2" box="[653,793,1130,1156]" name="Ecuador" pageId="17" pageNumber="280">ECUADOR</collectingCountry>
:
<collectingRegion id="016DF858FFB14260FC81FBF8FCB6DDC3" box="[801,864,1131,1157]" country="Ecuador" name="Napo" pageId="17" pageNumber="280">Napo</collectingRegion>
, Añangu, south bank of Río Napo
<location id="C6766061FFB14260FB78FBF9FED7DDEE" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FB78FBF9FED7DDEE" country="Ecuador" latitude="-0.53333336" longLatPrecision="1308" longitude="-76.38333" name="95 km downstream from Coca" pageId="17" pageNumber="280" stateProvince="Napo">
<quantity id="04519B5FFFB14260FB78FBF9FACBDDC2" box="[1240,1309,1130,1157]" metricMagnitude="4" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="280" unit="km" value="95.0">95 km</quantity>
downstream from Coca
</location>
,
<quantity id="04519B5FFFB14260FEADFB1DFE80DDEF" box="[269,342,1166,1193]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="17" pageNumber="280" unit="m" value="246.0">
<elevation id="4884D189FFB14260FEADFB1DFE80DDEF" box="[269,342,1166,1193]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="17" pageNumber="280" unit="m" value="246.0">246 m</elevation>
</quantity>
,
<geoCoordinate id="A69D507DFFB14260FEC2FB1DFE64DDEE" box="[354,434,1166,1192]" degrees="0" direction="south" minutes="32" orientation="latitude" pageId="17" pageNumber="280" precision="925" value="-0.53333336">0°32’S</geoCoordinate>
,
<geoCoordinate id="A69D507DFFB14260FE1EFB1DFDF3DDEE" box="[446,549,1166,1192]" degrees="76" direction="west" minutes="23" orientation="longitude" pageId="17" pageNumber="280" precision="925" value="-76.38333">76°23’W</geoCoordinate>
,
<date id="B717107AFFB14260FD90FB1DFD60DDEF" box="[560,694,1166,1193]" pageId="17" pageNumber="280" value="1985-07-04">
<collectingDate id="A753E992FFB14260FD90FB1DFD60DDEF" box="[560,694,1166,1193]" pageId="17" pageNumber="280" value="1985-07-04">4 July 1985</collectingDate>
</date>
,
<collectionCode id="A5B8AE7FFFB14260FD61FB1CFD0FDDEE" box="[705,729,1167,1192]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="17" pageNumber="280" type="Herbarium">
<emphasis id="F1DDEAA8FFB14260FD61FB1CFD0FDDEE" box="[705,729,1167,1192]" italics="true" pageId="17" pageNumber="280">H</emphasis>
</collectionCode>
<emphasis id="F1DDEAA8FFB14260FD79FB1CFD0BDDEE" box="[729,733,1167,1192]" italics="true" pageId="17" pageNumber="280">.</emphasis>
<emphasis id="F1DDEAA8FFB14260FD43FB1DFC14DDEE" box="[739,962,1166,1192]" italics="true" pageId="17" pageNumber="280">
<location id="C6766061FFB14260FD43FB1DFCE3DDEE" LSID="urn:lsid:plazi:treatment:4B0087ACFFB14260FF28FE99FBFEDDEE:C6766061FFB14260FD43FB1DFCE3DDEE" box="[739,821,1166,1192]" country="Ecuador" name="Balslev" pageId="17" pageNumber="280">Balslev</location>
et al. 60620
</emphasis>
(
<collectionCode id="A5B8AE7FFFB14260FC71FB1DFBCEDDEE" box="[977,1048,1166,1192]" country="Ethiopia" httpUri="http://grbio.org/cool/5dxx-xet2" name="Addis Ababa University, Department of Biology" pageId="17" pageNumber="280">AAU</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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