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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.65767" ID-GBIF-Dataset="cbd813c3-228e-4bce-b30a-1a150d778c5b" ID-PMC="PMC8096803" ID-Pensoft-Pub="1313-2970-1035-145" ID-Pensoft-UUID="55CAE096B4A451D88EA433175454DA3F" ID-PubMed="33958933" ID-ZooBank="ADD73A162DB34873890B1E490C2C8841" ModsDocID="1313-2970-1035-145" checkinTime="1619617199447" checkinUser="pensoft" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2021" docId="46CFFF9DFBBB518D9F9802AC3F372F3E" docLanguage="en" docName="ZooKeys 1035: 145-193" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.65767" docTitle="Acanthinozodium elburzicum Zamani &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="61538C61-2E8A-4F21-8289-B9129E5E4965" docUuidSource="ZooBank" docVersion="4" id="55CAE096B4A451D88EA433175454DA3F" lastPageNumber="145" masterDocId="55CAE096B4A451D88EA433175454DA3F" masterDocTitle="Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species" masterLastPageNumber="193" masterPageNumber="145" pageNumber="145" updateTime="1668150222111" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, Alireza</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation>Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1035</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>193</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1035.65767</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.65767</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1035-145</mods:identifier>
<mods:identifier type="ZooBank">ADD73A162DB34873890B1E490C2C8841</mods:identifier>
<mods:identifier type="Pensoft-UUID">55CAE096B4A451D88EA433175454DA3F</mods:identifier>
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<treatment ID-GBIF-Taxon="181753563" LSID="urn:lsid:zoobank.org:act:61538C61-2E8A-4F21-8289-B9129E5E4965" httpUri="http://treatment.plazi.org/id/46CFFF9DFBBB518D9F9802AC3F372F3E" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/61538C61-2E8A-4F21-8289-B9129E5E4965" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum" status="sp. nov.">Acanthinozodium elburzicum</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">Figs 18E-J</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">, 21D-F</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">, 23A-C</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">, 26E</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">, 29A</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">, 30A-E</figureCitation>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">, 33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="type material">
<paragraph pageId="0" pageNumber="145">Type material.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Holotype</emphasis>
♂ (MHNG), Iran:
<emphasis italics="true" pageId="0" pageNumber="145">Tehran Province</emphasis>
: Jamshidieh Mts.,
<geoCoordinate degrees="35" direction="north" minutes="49" orientation="latitude" precision="925" value="35.816666">35°49'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="27" orientation="longitude" precision="925" value="51.45">51°27'E</geoCoordinate>
, 05.2015 (A. Zamani).
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Paratypes</emphasis>
: 1♀ (MHNG), Pardisan Park,
<geoCoordinate degrees="35" direction="north" minutes="44" orientation="latitude" precision="925" value="35.733334">35°44'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="21" orientation="longitude" precision="925" value="51.35">51°21'E</geoCoordinate>
, 05.2015 (A. Zamani); 5♂2♀1j. (MMUE), Latian Dam,
<geoCoordinate degrees="35" direction="north" minutes="48" orientation="latitude" precision="925" value="35.8">35°48'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="08" orientation="longitude" precision="925" value="51.133335">51°08'E</geoCoordinate>
, 06-19.06.2000 (Y.M. Marusik); 3♂1♀ (MMUE), Plant Protection Institute,
<geoCoordinate degrees="35" direction="north" minutes="40" orientation="latitude" precision="925" value="35.666668">35°40'N</geoCoordinate>
,
<geoCoordinate degrees="51" direction="east" minutes="24" orientation="longitude" precision="925" value="51.4">51°24'E</geoCoordinate>
, 07-22.06.2000 (Y.M. Marusik).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">The specific epithet refers to the Elburz Mountain Range, in which the type locality of the new species is situated.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
The male of the new species is similar to
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. by the similar shape of the RTA and they both have a small retrolateral indentation apically on the conductor, but differs by the shape of the median apophysis, which is with a distinctly larger prolateral outgrowth. The female of the new species is most similar to
<taxonomicName lsidName="A. dorsa" pageId="0" pageNumber="145" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">A. dorsa</emphasis>
</taxonomicName>
sp. nov. but differs by the epigynal hood being wider than long (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
longer than wide) and receptacles separated by about 4 diameters (
<emphasis italics="true" pageId="0" pageNumber="145">vs.</emphasis>
ca. 2.5).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male</emphasis>
(holotype). Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18F-H</figureCitation>
(holotype: 18F). Total length 4.30. Carapace 2.08 long, 0.97 wide at pars cephalica, 1.49 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.14, ALE: 0.12, PME: 0.08, PLE: 0.09, PME-PME: 0.15. Carapace dark brown, lighter in pars cephalica, with irregular dark patches and lines. Sternum, labium and maxillae light brown. Chelicera (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Males of Parazodarion raddei (A-C) and Acanthinozodium elburzicum sp. nov. (E) and female of Trygetus susianus sp. nov. (D) A, D habitus, dorsal B, C palp, ventral and retrolateral E chelicerae, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure26" httpUri="https://binary.pensoft.net/fig/536903" pageId="0" pageNumber="145">26E</figureCitation>
) dark brown, with one retromarginal tooth. Legs slightly lighter than carapace, without annulations. Abdomen dorsally black and with large scutum; ventrally grayish, without any pattern. Spinnerets pale, uniform in color. Measurements of legs: I: 7.52 (1.79, 0.71, 1.75, 2.03, 1.24), II: 6.79 (1.87, 0.69, 1.40, 1.83, 1.00), III: 6.90 (1.78, 0.74, 1.40, 2.06, 0.92), IV: 9.19 (2.48, 0.83, 2.15, 2.84, 0.89).
</paragraph>
<paragraph pageId="0" pageNumber="145">
Palp as in Figs
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male palps of Acanthinozodium dorsa sp. nov. (A-C) and A. elburzicum sp. nov. (D-F) A, D ventral B, E retrolateral C, F dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145">21D-F</figureCitation>
,
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A-C) and A. diara sp. nov. (D-F) A, D ventral B, F retrolateral C, E dorsoretrolateral. Abbreviations: Cg - cymbial groove, Em - embolus, Ma - median apophysis, Ra - retrolateral tibial apophysis, Va - ventral apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145">23A-C</figureCitation>
,
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. SEM images of the male palps of Acanthinozodium elburzicum sp. nov. (A) and Z. talyschicum (B, C) A, B ventral C ventroretrolateral. Abbreviations: Tm - median tegular process, Tp - prolateral tegular process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1035.65767.figure29" httpUri="https://binary.pensoft.net/fig/536906" pageId="0" pageNumber="145">29A</figureCitation>
. RTA (
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
) long and conical, with a small projection apically; ventral apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
) small and conical; tegulum with posterior process; median apophysis (
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
) wider than long, with outgrowths on all four sides, the triangular prolateral one the largest; embolus (
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
) originating at about the 7
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
position; embolus proper thin and steadily curving.
</paragraph>
<caption doi="10.3897/zookeys.1035.65767.figure21" httpUri="https://binary.pensoft.net/fig/536898" pageId="0" pageNumber="145" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 21.</emphasis>
Male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium dorsa" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="dorsa">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium dorsa</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. elburzicum" pageId="0" pageNumber="145" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">A. elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, E</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, F</emphasis>
dorsal, with arrow pointing to cymbial groove. Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure22" httpUri="https://binary.pensoft.net/fig/536899" pageId="0" pageNumber="145" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 22.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium armita" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="armita">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium armita</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C, F</emphasis>
) and
<taxonomicName lsidName="A. ovtchinnikovi" pageId="0" pageNumber="145" rank="species" species="ovtchinnikovi">
<emphasis italics="true" pageId="0" pageNumber="145">A. ovtchinnikovi</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D, E, G-I</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, H</emphasis>
retroventral and retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">B, C, G</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
proventral, apicoproventral and apicoventral
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
retrolateral tibial and cymbial groove. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Co</emphasis>
- conductor,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Er</emphasis>
-embolar ridge,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis. Scale bars: 0.1 mm, unless stated otherwise.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1035.65767.figure23" httpUri="https://binary.pensoft.net/fig/536900" pageId="0" pageNumber="145" start="Figure 23" startId="F23">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 23.</emphasis>
SEM images of the male palps of
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthinozodium elburzicum" order="Araneae" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="elburzicum">
<emphasis italics="true" pageId="0" pageNumber="145">Acanthinozodium elburzicum</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
) and
<taxonomicName lsidName="A. diara" pageId="0" pageNumber="145" rank="species" species="diara">
<emphasis italics="true" pageId="0" pageNumber="145">A. diara</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="145">A, D</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="145">B, F</emphasis>
retrolateral
<emphasis bold="true" pageId="0" pageNumber="145">C, E</emphasis>
dorsoretrolateral. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="145">Cg</emphasis>
- cymbial groove,
<emphasis italics="true" pageId="0" pageNumber="145">Em</emphasis>
- embolus,
<emphasis italics="true" pageId="0" pageNumber="145">Ma</emphasis>
- median apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Ra</emphasis>
- retrolateral tibial apophysis,
<emphasis italics="true" pageId="0" pageNumber="145">Va</emphasis>
- ventral apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Habitus of Acanthinozodium armita sp. nov. (A), A. diara sp. nov. (B), A. dorsa sp. nov. (C, D), A. elburzicum sp. nov. (E-J), A. kiana sp. nov. (K) and A. masa sp. nov. (L) A-C, F, G, K, L males, dorsal H male, ventral D, E, I females, dorsal J female, ventral. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.1035.65767.figure18" httpUri="https://binary.pensoft.net/fig/536895" pageId="0" pageNumber="145">18E, I, J</figureCitation>
. Total length 4.95. Carapace 1.95 long, 0.96 wide at pars cephalica, 1.40 wide at pars thoracica. Eye sizes and interdistance of PMEs: AME: 0.15, ALE: 0.12, PME: 0.08, PLE: 0.11, PME-PME: 0.15. Coloration as in male, with paler abdomen lacking a scutum. Measurements of legs: I: 6.54 (1.50, 0.70, 1.44, 1.76, 1.14), II: 6.27 (1.59, 0.67, 1.33, 1.67, 1.01), III: 6.23 (1.60, 0.71, 1.21, 1.74, 0.97), IV: 8.11 (2.04, 0.78, 1.81, 2.46, 1.02).
</paragraph>
<paragraph pageId="0" pageNumber="145">
Epigyne as in Fig.
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Epigynes of Acanthinozodium elburzicum sp. nov. (A-E) and A. dorsa sp. nov. (F-I) A, F intact, ventral B macerated, ventral C-E, G-I macerated, dorsal. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1035.65767.figure30" httpUri="https://binary.pensoft.net/fig/536907" pageId="0" pageNumber="145">30A-E</figureCitation>
. Epigynal plater over 3 times wider than long, lacking fovea; anterior hood wider than long; receptacles with rounded posterior parts, separated by about 4 diameters.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
Known from the listed localities in Tehran Province, northern Iran (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Distribution records of Acanthinozodium spp. (partim 1 - 3, 5, 6, 8, 9), Zodariellum spp. (10, 11) and Zodarion spp. (4, 7, 12) in Iran and Turkmenistan: 1 A. armita sp. nov. 2 A. diara sp. nov. 3 A. elburzicum sp. nov. 4 Z. buettikeri 5 A. dorsa sp. nov. 6 A. kiana sp. nov. 7 Z. lutipes 8 A. masa sp. nov. 9 A. parmida sp. nov. 10 Z. proszynskii 11 Z. sytchevskajae 12 Z. talyschicum." figureDoi="10.3897/zookeys.1035.65767.figure33" httpUri="https://binary.pensoft.net/fig/536910" pageId="0" pageNumber="145">33</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>