319 lines
34 KiB
XML
319 lines
34 KiB
XML
<document id="50194B49049343B5D604FF6BEE9E59BC" ID-DOI="10.11646/zootaxa.4817.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954698" ID-ZooBank="7CBDB6A9-9AF9-495F-A55A-83BF36A4934D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595340362484" checkinUser="plazi" docAuthor="Isler, Morton L., Chesser, Terry, Robbins, Mark B., Cuervo, Andrés M., Cadena, Carlos Daniel & Hosner, Peter A." docDate="2020" docId="465F87DEFFB3745DFF07FBB2FEB6FC37" docLanguage="en" docName="zootaxa.4817.1.1.pdf" docOrigin="Zootaxa 4817 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Grallaria saturata Domaniewski & Stolzmann 1918" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="BA66FFA6FFBE7453FF90FFAFFFB2FFFC" masterDocTitle="Taxonomic evaluation of the Grallaria rufula (Rufous Antpitta) complex (Aves: Passeriformes: Grallariidae) distinguishes sixteen species" masterLastPageNumber="74" masterPageNumber="1" pageNumber="14" updateTime="1698783870222" updateUser="plazi">
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<mods:title id="19F9D74B873EF06023B9E15A30777039">Taxonomic evaluation of the Grallaria rufula (Rufous Antpitta) complex (Aves: Passeriformes: Grallariidae) distinguishes sixteen species</mods:title>
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<mods:namePart id="1D9DF37118B3DE2D0780FDBCE9C3AB37">Isler, Morton L.</mods:namePart>
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<mods:namePart id="E4B971485AB9EDE99F919470971991CF">Chesser, Terry</mods:namePart>
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<mods:namePart id="EF7D6ED31403A934168EDDE4D5F456A5">Robbins, Mark B.</mods:namePart>
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<mods:namePart id="CE0642025648279E104421061D738797">Cuervo, Andrés M.</mods:namePart>
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<mods:namePart id="2077DFBD8CEF090751653971CDA50C47">Cadena, Carlos Daniel</mods:namePart>
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<mods:namePart id="AC1633FB28BA4902CC3F1EA3ABCE8D93">Hosner, Peter A.</mods:namePart>
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<mods:title id="0562EAD5686B6B39927BEC2D4BB8C621">Zootaxa</mods:title>
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<mods:date id="4644979488E93715CDAA093DDFAD1C3F">2020</mods:date>
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<mods:number id="4F302A91BE7380CB2BD3EAE8E5F93BDA">2020-07-21</mods:number>
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<mods:number id="A5A98F5C1CFD65B6F74D22A9F39BD7BB">4817</mods:number>
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<mods:classification id="14EC343626AD9D221076BFD4248D26DA">journal article</mods:classification>
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<mods:identifier id="504E2BD596BE357B3AE6A1CB5417B0BE" type="DOI">10.11646/zootaxa.4817.1.1</mods:identifier>
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<treatment id="465F87DEFFB3745DFF07FBB2FEB6FC37" LSID="urn:lsid:plazi:treatment:465F87DEFFB3745DFF07FBB2FEB6FC37" httpUri="http://treatment.plazi.org/id/465F87DEFFB3745DFF07FBB2FEB6FC37" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<subSubSection id="86EC6543FFB3745EFF07FBB2FEDFFBA7" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="CE4936C8FFB3745EFF07FBB2FAD1FBCB" blockId="13.[151,1379,1053,1115]" box="[151,1379,1053,1080]" pageId="13" pageNumber="14">
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<heading id="950181A4FFB3745EFF07FBB2FAD1FBCB" bold="true" box="[151,1379,1053,1080]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="FC82EADAFFB3745EFF07FBB2FAD1FBCB" bold="true" box="[151,1379,1053,1080]" pageId="13" pageNumber="14">
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<taxonomicName id="09F64D4BFFB3745EFF07FBB2FCA9FBCB" authority="Domaniewski & Stolzmann, 1918" authorityName="Domaniewski & Stolzmann" authorityYear="1918" box="[151,795,1053,1080]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="saturata">
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<emphasis id="FC82EADAFFB3745EFF07FBB2FEC0FBCB" bold="true" box="[151,370,1053,1080]" italics="true" pageId="13" pageNumber="14">Grallaria saturata</emphasis>
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<bibRefCitation id="AA674B39FFB3745EFEEAFBB2FCA9FBCB" author="Domaniewski, J. & Stolzmann, J." box="[378,795,1053,1079]" pageId="13" pageNumber="14" pagination="479 - 484" refId="ref21909" refString="Domaniewski, J. & Stolzmann, J. (1918) Contribution a la connaissance des formes du genre Grallaria Vieillot. In: Comptes rendus des seances: Classe 4, Sciences biologiques. Societe scientifique de Varsowie, Varsowie, pp. 479 - 484." type="book chapter" year="1918">Domaniewski & Stolzmann, 1918</bibRefCitation>
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</taxonomicName>
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, subspecies resurrected and elevated to species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="CE4936C8FFB3745EFF07FBEEFEDFFBA7" blockId="13.[151,1379,1053,1115]" box="[151,365,1089,1115]" pageId="13" pageNumber="14">Equatorial Antpitta</paragraph>
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</subSubSection>
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<subSubSection id="86EC6543FFB3745EFF07FB26FB27FB5F" box="[151,1173,1161,1188]" pageId="13" pageNumber="14" type="description">
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<paragraph id="CE4936C8FFB3745EFF07FB26FB27FB5F" blockId="13.[151,1437,1161,1476]" box="[151,1173,1161,1188]" pageId="13" pageNumber="14">
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Includes populations designated
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<emphasis id="FC82EADAFFB3745EFD95FB26FDDFFB5F" box="[517,621,1161,1187]" italics="true" pageId="13" pageNumber="14">
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<taxonomicName id="09F64D4BFFB3745EFD95FB26FDFBFB5F" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[517,585,1161,1187]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
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2a
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</emphasis>
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,
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<emphasis id="FC82EADAFFB3745EFDEBFB26FD50FB5F" box="[635,738,1161,1187]" italics="true" pageId="13" pageNumber="14">
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<taxonomicName id="09F64D4BFFB3745EFDEBFB26FD0DFB5F" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[635,703,1161,1187]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
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2b
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</emphasis>
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,
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<emphasis id="FC82EADAFFB3745EFD60FB26FCE4FB5F" box="[752,854,1161,1187]" italics="true" pageId="13" pageNumber="14">
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<taxonomicName id="09F64D4BFFB3745EFD60FB26FC86FB5F" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[752,820,1161,1187]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
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2c
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</emphasis>
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and
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<emphasis id="FC82EADAFFB3745EFC1DFB26FC54FB5F" box="[909,998,1161,1187]" italics="true" pageId="13" pageNumber="14">
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<taxonomicName id="09F64D4BFFB3745EFC1DFB26FC63FB5F" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[909,977,1161,1187]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
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<quantity id="090E9B2DFFB3745EFC48FB26FC54FB5F" box="[984,998,1161,1187]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="13" pageNumber="14" unit="in" value="5.0">5</quantity>
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</emphasis>
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in the analysis.
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</paragraph>
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<subSubSection id="86EC6543FFB3745EFF57FB02FE03FA38" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="CE4936C8FFB3745EFF57FB02FE03FA38" blockId="13.[151,1437,1161,1476]" pageId="13" pageNumber="14">
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<emphasis id="FC82EADAFFB3745EFF57FB02FEF0FB3B" bold="true" box="[199,322,1197,1223]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
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Overall plumage typically reddish-brown, underparts paler, but overall intensity variable. Vocalizations of
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<taxonomicName id="09F64D4BFFB3745EFF66FB7EFEC5FB17" authorityName="Domaniewski & Stolzmann" authorityYear="1918" box="[246,375,1233,1259]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="saturata">
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<emphasis id="FC82EADAFFB3745EFF66FB7EFEC5FB17" box="[246,375,1233,1259]" italics="true" pageId="13" pageNumber="14">G. saturata</emphasis>
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</taxonomicName>
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(
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<figureCitation id="56CD2A4DFFB3745EFE19FB7EFE7CFB10" box="[393,462,1233,1260]" captionStart="FIGURE 7" captionStartId="13.[151,250,1876,1900]" captionTargetBox="[158,1433,1501,1843]" captionTargetId="figure@13.[151,1435,1497,1850]" captionTargetPageId="13" captionText="FIGURE 7. Songs of Grallaria saturata: (A) long song, Cerro Mongus, Carchi, Ecuador (Krabbe XC248386); (B) short song, Cajanuma, Loja, Ecuador (Krabbe XC250311)." figureDoi="http://doi.org/10.5281/zenodo.3954713" httpUri="https://zenodo.org/record/3954713/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
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) are distinguished from all other taxa in the
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<taxonomicName id="09F64D4BFFB3745EFC5EFB7EFB84FB17" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[974,1078,1233,1259]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rufula">
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<emphasis id="FC82EADAFFB3745EFC5EFB7EFB84FB17" box="[974,1078,1233,1259]" italics="true" pageId="13" pageNumber="14">G. rufula</emphasis>
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</taxonomicName>
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complex except
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<taxonomicName id="09F64D4BFFB3745EFB6AFB7EFAC5FB17" authorityName="Cuervo, Cadena, Isler, & Chesser" authorityYear="2020" box="[1274,1399,1233,1259]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="alvarezi">
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<emphasis id="FC82EADAFFB3745EFB6AFB7EFAC5FB17" box="[1274,1399,1233,1259]" italics="true" pageId="13" pageNumber="14">G. alvarezi</emphasis>
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</taxonomicName>
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by a short song that consists of a single note, followed by a longer interval and ending with an even longer terminal phrase whose structure appears to vary geographically (discussed in App. 1). Short songs of
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<taxonomicName id="09F64D4BFFB3745EFB09FAB6FAAAFACF" authorityName="Domaniewski & Stolzmann" authorityYear="1918" box="[1177,1304,1305,1331]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="saturata">
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<emphasis id="FC82EADAFFB3745EFB09FAB6FAAAFACF" box="[1177,1304,1305,1331]" italics="true" pageId="13" pageNumber="14">G. saturata</emphasis>
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</taxonomicName>
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differ diagnosably from those of
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<taxonomicName id="09F64D4BFFB3745EFE00FA92FDB8FAAB" authorityName="Cuervo, Cadena, Isler, & Chesser" authorityYear="2020" box="[400,522,1341,1367]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="alvarezi">
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<emphasis id="FC82EADAFFB3745EFE00FA92FDB8FAAB" box="[400,522,1341,1367]" italics="true" pageId="13" pageNumber="14">G. alvarezi</emphasis>
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</taxonomicName>
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by flat or downslurred rather than rounded notes. Long songs of
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<taxonomicName id="09F64D4BFFB3745EFB4AFA92FAEAFAAB" authorityName="Domaniewski & Stolzmann" authorityYear="1918" box="[1242,1368,1341,1367]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="saturata">
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<emphasis id="FC82EADAFFB3745EFB4AFA92FAEAFAAB" box="[1242,1368,1341,1367]" italics="true" pageId="13" pageNumber="14">G. saturata</emphasis>
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</taxonomicName>
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differ from those of
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<taxonomicName id="09F64D4BFFB3745EFEA1FACEFE19FA87" authorityName="Cuervo, Cadena, Isler, & Chesser" authorityYear="2020" box="[305,427,1377,1403]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="alvarezi">
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<emphasis id="FC82EADAFFB3745EFEA1FACEFE19FA87" box="[305,427,1377,1403]" italics="true" pageId="13" pageNumber="14">G. alvarezi</emphasis>
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</taxonomicName>
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in having note peaks at the same frequency or nearly so, whereas peaks of
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<taxonomicName id="09F64D4BFFB3745EFB71FACEFAE9FA87" authorityName="Cuervo, Cadena, Isler, & Chesser" authorityYear="2020" box="[1249,1371,1377,1403]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="alvarezi">
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<emphasis id="FC82EADAFFB3745EFB71FACEFAE9FA87" box="[1249,1371,1377,1403]" italics="true" pageId="13" pageNumber="14">G. alvarezi</emphasis>
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</taxonomicName>
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notes decline substantially in frequency. Long song note shapes are typically more downslurred and wider in band width than notes of
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<taxonomicName id="09F64D4BFFB3745EFEBFFA06FE18FA3F" authorityName="Cuervo, Cadena, Isler, & Chesser" authorityYear="2020" box="[303,426,1449,1475]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="alvarezi">
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<emphasis id="FC82EADAFFB3745EFEBFFA06FE18FA3F" box="[303,426,1449,1475]" italics="true" pageId="13" pageNumber="14">G. alvarezi</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<caption id="9A896640FFB3745EFF07F8FBFDCAF86C" ID-DOI="http://doi.org/10.5281/zenodo.3954713" ID-Zenodo-Dep="3954713" httpUri="https://zenodo.org/record/3954713/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1876,1900]" targetBox="[158,1433,1501,1843]" targetPageId="13">
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<paragraph id="CE4936C8FFB3745EFF07F8FBFDCAF86C" blockId="13.[151,1436,1876,1936]" pageId="13" pageNumber="14">
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<emphasis id="FC82EADAFFB3745EFF07F8FBFEBFF890" bold="true" box="[151,269,1876,1900]" pageId="13" pageNumber="14">FIGURE 7</emphasis>
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. Songs of
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<taxonomicName id="09F64D4BFFB3745EFEE9F8FBFD87F890" authorityName="Domaniewski & Stolzmann" authorityYear="1918" box="[377,565,1876,1900]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="saturata">
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<emphasis id="FC82EADAFFB3745EFEE9F8FBFD87F890" box="[377,565,1876,1900]" italics="true" pageId="13" pageNumber="14">Grallaria saturata</emphasis>
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</taxonomicName>
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: (A) long song, Cerro Mongus, Carchi, Ecuador (Krabbe XC248386); (B) short song, Cajanuma, Loja, Ecuador (Krabbe XC250311).
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</paragraph>
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</caption>
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<subSubSection id="86EC6543FFB3745DFF57F811FBC7FE0B" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="CE4936C8FFB3745DFF57F811FD25FF24" blockId="13.[151,1437,1982,2045]" lastBlockId="14.[151,1437,153,971]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<emphasis id="FC82EADAFFB3745EFF57F811FED3F824" bold="true" box="[199,353,1982,2008]" pageId="13" pageNumber="14">Distribution.</emphasis>
|
||
Iguaque Massif in the west slope of Eastern Andes in
|
||
<collectingRegion id="0C32F82AFFB3745EFC26F811FBB8F824" box="[950,1034,1982,2008]" country="Colombia" name="Boyaca" pageId="13" pageNumber="14">Boyacá</collectingRegion>
|
||
and extreme southwestern Santand- er,
|
||
<collectingCountry id="B6E17658FFB3745EFF29F84DFE99F800" box="[185,299,2018,2044]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
|
||
; Central Andes from southern
|
||
<collectingRegion id="0C32F82AFFB3745EFDEEF84DFD5CF800" box="[638,750,2018,2044]" country="Colombia" name="Antioquia" pageId="13" pageNumber="14">Antioquia</collectingRegion>
|
||
to
|
||
<collectingRegion id="0C32F82AFFB3745EFC80F84DFCEDF800" box="[784,863,2018,2044]" country="Colombia" name="Tolima" pageId="13" pageNumber="14">Tolima</collectingRegion>
|
||
and in
|
||
<collectingRegion id="0C32F82AFFB3745EFC20F84DFC4BF800" box="[944,1017,2018,2044]" country="Colombia" name="Cauca" pageId="13" pageNumber="14">Cauca</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB3745EFB94F84DFBC4F800" box="[1028,1142,2018,2044]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
|
||
; and the Andes from Nari- ño and western
|
||
<collectingRegion id="0C32F82AFFB0745DFED9FF35FE0FFF48" box="[329,445,154,180]" country="Colombia" name="Putumayo" pageId="14" pageNumber="15">Putumayo</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB0745DFE59FF36FD88FF4F" box="[457,570,153,179]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
|
||
, south through
|
||
<collectingCountry id="B6E17658FFB0745DFD79FF36FCF4FF4F" box="[745,838,153,179]" name="Ecuador" pageId="14" pageNumber="15">Ecuador</collectingCountry>
|
||
to
|
||
<collectingRegion id="0C32F82AFFB0745DFCFBFF36FC14FF4F" box="[875,934,153,179]" country="Peru" name="Piura" pageId="14" pageNumber="15">Piura</collectingRegion>
|
||
and
|
||
<collectingRegion id="0C32F82AFFB0745DFC4EFF36FBEBFF4F" box="[990,1113,153,179]" country="Peru" name="Cajamarca" pageId="14" pageNumber="15">Cajamarca</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB0745DFBF4FF35FB28FF48" box="[1124,1178,154,180]" name="Peru" pageId="14" pageNumber="15">Peru</collectingCountry>
|
||
, north of Río Marañón and east of Río Huancabamba,
|
||
<quantity id="090E9B2DFFB0745DFE65FF12FD21FF24" box="[501,659,189,216]" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.65" metricValueMin="2.55" pageId="14" pageNumber="15" unit="m" value="3100.0" valueMax="3650.0" valueMin="2550.0">2550–3650 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CE4936C8FFB0745DFF57FF4EFBC7FE0B" blockId="14.[151,1437,153,971]" pageId="14" pageNumber="15">
|
||
<emphasis id="FC82EADAFFB0745DFF57FF4EFE85FF07" bold="true" box="[199,311,225,251]" pageId="14" pageNumber="15">Plumage.</emphasis>
|
||
Overall plumage of upperparts, including rectrices, remiges (primaries edged paler), crown and auriculars, typically reddish-brown (5YR 4/4) but variable; specimens from the southern end of the range of
|
||
<emphasis id="FC82EADAFFB0745DFAD2FEAAFA2EFEE3" box="[1346,1436,261,287]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFAD2FEAAFA34FEE3" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[1346,1414,261,287]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5
|
||
</emphasis>
|
||
(
|
||
<emphasis id="FC82EADAFFB0745DFF31FE86FEB5FEBF" box="[161,263,297,323]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFF31FE86FF57FEBF" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[161,229,297,323]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5b
|
||
</emphasis>
|
||
) in
|
||
<collectingRegion id="0C32F82AFFB0745DFEA1FE86FDBAFEBF" box="[305,520,297,323]" country="Ecuador" name="Zamora-Chinchipe" pageId="14" pageNumber="15">Zamora-Chinchipe</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB0745DFD81FE86FDC2FEBF" box="[529,624,297,323]" name="Ecuador" pageId="14" pageNumber="15">Ecuador</collectingCountry>
|
||
, and from the northern end of the range of
|
||
<emphasis id="FC82EADAFFB0745DFBD2FE86FB28FEBF" box="[1090,1178,297,323]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFBD2FE86FB34FEBF" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[1090,1158,297,323]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
2
|
||
</emphasis>
|
||
(
|
||
<emphasis id="FC82EADAFFB0745DFB39FE86FABDFEBF" box="[1193,1295,297,323]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFB39FE86FB5FFEBF" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[1193,1261,297,323]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
2b
|
||
</emphasis>
|
||
) in
|
||
<collectingRegion id="0C32F82AFFB0745DFAA9FE86FA2AFEBF" box="[1337,1432,297,323]" country="Colombia" name="Quindio" pageId="14" pageNumber="15">Quindío</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB0745DFF07FEE2FEBAFE9B" box="[151,264,333,359]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
|
||
, are lighter and browner than those from intervening populations at the northern end of the range of
|
||
<emphasis id="FC82EADAFFB0745DFAC8FEE2FF17FE77" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFAC8FEE2FA2EFE9B" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[1368,1436,333,359]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5
|
||
</emphasis>
|
||
(
|
||
<emphasis id="FC82EADAFFB0745DFF25FEDEFEAFFE77" box="[181,285,369,395]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFF25FEDEFF4BFE77" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[181,249,369,395]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5a
|
||
</emphasis>
|
||
) and from
|
||
<collectingRegion id="0C32F82AFFB0745DFE09FEDEFE50FE77" box="[409,482,369,395]" country="Colombia" name="Cauca" pageId="14" pageNumber="15">Cauca</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB0745DFE7CFEDEFDE8FE77" box="[492,602,369,395]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
|
||
(
|
||
<emphasis id="FC82EADAFFB0745DFDFBFEDEFD68FE77" box="[619,730,369,395]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFDFBFEDEFD1DFE77" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[619,687,369,395]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
2c)
|
||
</emphasis>
|
||
; coloration shades to color of underparts in malar area. Overall plumage of underparts typically reddish yellow-brown (7.5YR 4/8) but lighter or darker consistent with upperparts; center of belly and undertail coverts slightly or substantially paler and yellower (7.5YR 6/6–8/2); flank coloration like upperparts or breast (specimen photographs App. 6,
|
||
<figureCitation id="56CD2A4DFFB0745DFC82FE72FC33FE0B" box="[786,897,477,503]" captionStart="FIGURE 20" captionStartId="27.[151,250,1835,1859]" captionTargetBox="[268,1323,190,1798]" captionTargetId="figure@27.[255,1332,181,1809]" captionTargetPageId="27" captionText="FIGURE 20. Map showing the taxonomy and distribution of the Grallaria rufula complex as revised herein, Part 2: G. rufocinerea and populations formerly ascribed to G. blakei. Solid circles represent sectors (Isler 1997) that contain locations of genetic samples used in a companion paper (Chesser et al. 2020) and stars are sectors containing locations where specimens have been collected or vocalizations recorded. Sectors may contain more than one genetic or vocal sample or specimen. Lines encompass sectors attributed to the same taxon and are not projections of geographic range which can be found in the companion paper (Chesser et al. 2020)." figureDoi="http://doi.org/10.5281/zenodo.3954739" httpUri="https://zenodo.org/record/3954739/files/figure.png" pageId="14" pageNumber="15">Figs. A20</figureCitation>
|
||
, A21, A33, and A34).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="86EC6543FFB0745DFF57FDAEFF44FDBC" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="CE4936C8FFB0745DFF57FDAEFF44FDBC" blockId="14.[151,1437,153,971]" pageId="14" pageNumber="15">
|
||
<emphasis id="FC82EADAFFB0745DFF57FDAEFEFCFDE7" bold="true" box="[199,334,513,539]" pageId="14" pageNumber="15">Etymology.</emphasis>
|
||
The English name reflects the geographic location of the range of this species, which straddles the Equator.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="86EC6543FFB0745DFF57FDE6FEB6FC37" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="CE4936C8FFB0745DFF57FDE6FEB6FC37" blockId="14.[151,1437,153,971]" pageId="14" pageNumber="15">
|
||
<emphasis id="FC82EADAFFB0745DFF57FDE6FE89FD9F" bold="true" box="[199,315,585,611]" pageId="14" pageNumber="15">Remarks.</emphasis>
|
||
See remarks under
|
||
<taxonomicName id="09F64D4BFFB0745DFDB6FDE6FD17FD9F" authorityName="Cuervo, Cadena, Isler, & Chesser" authorityYear="2020" box="[550,677,585,611]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alvarezi">
|
||
<emphasis id="FC82EADAFFB0745DFDB6FDE6FD17FD9F" box="[550,677,585,611]" italics="true" pageId="14" pageNumber="15">G. alvarezi</emphasis>
|
||
</taxonomicName>
|
||
for possible additional vocal differences from that species. Vocal analysis in this clade was made difficult by geographically unbalanced availability of recordings and specimens. Although recordings of 112 individuals were available for
|
||
<emphasis id="FC82EADAFFB0745DFCD0FD3EFC2FFD57" box="[832,925,657,683]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFCD0FD3EFC36FD57" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[832,900,657,683]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5
|
||
</emphasis>
|
||
, only a handful of recordings was available for the three study populations comprising
|
||
<emphasis id="FC82EADAFFB0745DFDEEFD1AFD6AFD33" box="[638,728,693,719]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFDEEFD1AFD70FD33" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[638,706,693,719]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
2
|
||
</emphasis>
|
||
, and nearly all were of short songs. Short songs varied but not sufficiently to distinguish populations with certainty. Notably, genetic distance was ~4% between
|
||
<emphasis id="FC82EADAFFB0745DFB78FD76FAF0FD0F" box="[1256,1346,729,755]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFB78FD76FA9EFD0F" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[1256,1324,729,755]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5
|
||
</emphasis>
|
||
and the
|
||
<emphasis id="FC82EADAFFB0745DFF07FD52FF41FCEB" box="[151,243,765,791]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFF07FD52FF69FCEB" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[151,219,765,791]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
2
|
||
</emphasis>
|
||
populations. Additional field work in
|
||
<collectingCountry id="B6E17658FFB0745DFD21FD52FCADFCEB" box="[689,799,765,791]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
|
||
is needed to provide recordings, specimens, and tissue samples and to determine whether populations are continuous, parapatric or allopatric.
|
||
<typeStatus id="114D886AFFB0745DFBE7FC8DFB1DFCC0" box="[1143,1199,802,828]" pageId="14" pageNumber="15">Type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="09F64D4BFFB0745DFAA4FC8EFDCEFCA3" authority="Domaniewski & Stolzmann, 1918" authorityName="Domaniewski & Stolzmann" authorityYear="1918" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="rufula" subSpecies="saturata">
|
||
<emphasis id="FC82EADAFFB0745DFAA4FC8EFF46FCA3" italics="true" pageId="14" pageNumber="15">G. rufula saturata</emphasis>
|
||
<bibRefCitation id="AA674B39FFB0745DFF6CFCEAFDCEFCA3" author="Domaniewski, J. & Stolzmann, J." box="[252,636,837,863]" pageId="14" pageNumber="15" pagination="479 - 484" refId="ref21909" refString="Domaniewski, J. & Stolzmann, J. (1918) Contribution a la connaissance des formes du genre Grallaria Vieillot. In: Comptes rendus des seances: Classe 4, Sciences biologiques. Societe scientifique de Varsowie, Varsowie, pp. 479 - 484." type="book chapter" year="1918">Domaniewski & Stolzmann, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
is San Rafael,
|
||
<collectingRegion id="0C32F82AFFB0745DFCB7FCEAFC02FCA3" box="[807,944,837,863]" country="Ecuador" name="Tungurahua" pageId="14" pageNumber="15">Tungurahua</collectingRegion>
|
||
,
|
||
<collectingCountry id="B6E17658FFB0745DFC2CFCEAFBA9FCA3" box="[956,1051,837,863]" name="Ecuador" pageId="14" pageNumber="15">Ecuador</collectingCountry>
|
||
, in the geographic range of
|
||
<emphasis id="FC82EADAFFB0745DFAC8FCEAFF17FC7F" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFAC8FCEAFA2EFCA3" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[1368,1436,837,863]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufula">rufula</taxonomicName>
|
||
5
|
||
</emphasis>
|
||
. The surprising occurrence of
|
||
<taxonomicName id="09F64D4BFFB0745DFE6CFCC6FDC9FC7F" authorityName="Domaniewski & Stolzmann" authorityYear="1918" box="[508,635,873,899]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="saturata">
|
||
<emphasis id="FC82EADAFFB0745DFE6CFCC6FDC9FC7F" box="[508,635,873,899]" italics="true" pageId="14" pageNumber="15">G. saturata</emphasis>
|
||
</taxonomicName>
|
||
in a small region on the west slope of the Eastern Andes surrounded by populations of
|
||
<emphasis id="FC82EADAFFB0745DFED0FC22FD8EFC5B" box="[320,572,909,935]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="09F64D4BFFB0745DFED0FC22FE15FC5B" baseAuthorityName="Lafresnaye" baseAuthorityYear="1843" box="[320,423,909,935]" class="Aves" family="Grallariidae" genus="Grallaria" kingdom="Animalia" order="Passeriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" sensu="stricto" species="rufula">G. rufula</taxonomicName>
|
||
<taxonomicNameLabel id="E7B157A1FFB0745DFE3FFC21FD8EFC5B" box="[431,572,910,935]" pageId="14" pageNumber="15" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
</emphasis>
|
||
to the north, east, and south is discussed in the companion paper (
|
||
<bibRefCitation id="AA674B39FFB0745DFAB6FC22FF4AFC37" author="Chesser, R. T. & Isler, M. L. & Cuervo, A. M. & Cadena, C. D. & Galen, S. C. & Bergner, L. M. & Fleischer, R. C. & Bravo, G. A. & Lane, D. F. & Hosner, P. A." pageId="14" pageNumber="15" pagination="1 - 25" refId="ref21648" refString="Chesser, R. T., Isler, M. L., Cuervo, A. M., Cadena, C. D., Galen, S. C., Bergner, L. M., Fleischer, R. C., Bravo, G. A., Lane, D. F. & Hosner, P. A. (2020) Conservative plumage masks extraordinary genetic diversity in the Grallaria rufula (Rufous Antpitta) complex of the humid Andes. The Auk: Ornithological Advances, 137, 1 - 25. https: // doi. org / 10.1093 / auk / ukaa 009" type="journal article" year="2020">
|
||
Chesser
|
||
<emphasis id="FC82EADAFFB0745DFA18FC21FF06FC37" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |